Agrilus of East Asia

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Digitally signed by Eduard Jendek DN: cn=Eduard Jendek, o=CFIA, ou, email=jendeke@g mail.com, c=CA Date: 2013.12.31 17:43:12 -05'00'

Eduard Jendek Vasily Grebennikov

Eduard Jendek Vasily Grebennikov

AGRILUS of East Asia

AGRILUS (Coleoptera, Buprestidae) of East Asia

Nakladatelsví Jan Farkač

Eduard Jendek Vasily Grebennikov

AGRILUS (Coleoptera, Buprestidae)

of East Asia

2 011 Nakladatelství Jan Farkač

Recommended format for citation: Jendek, E. & Grebennikov, V., 2011: Agrilus (Coleoptera, Buprestidae) of East Asia. Prague, Jan Farkač, 362 pages.

Date issued: September 30, 2011

Author‘s address: Ottawa Plant Laboratory, Canadian Food Inspection Agency, K.W. Neatby Bldg., 960 Carling Avenue, Ottawa, Ontario, K1A 0C6, Canada

EDUARD JENDEK & VASILY GREBENNIKOV AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Cover design: © Martin Česanek, Bratislava, Slovakia Typeset by: Jakub Rolčík - CLAIRON Production, Prague, Czech Republic Published by: Nakladatelství Jan Farkač, Prague, Czech Republic Print by: 3P, spol. s r.o., Prague, Czech Republic © Eduard Jendek © Nakladatelství Jan Farkač © Jakub Rolčík - CLAIRON Production ISBN: 978-80-903590-6-2

This book is dedicated to amateur entomologists in appreciation of their immense contributions to the exploration of the World’s insect biodiversity. Their field work and specimens they have collected allowed this book to be completed.

CONTENTS

5 INTRODUCTION ...................................................................................................................................... 7 ACKNOWLEDGEMENTS ........................................................................................................................ 9 ABBREVIATIONS AND CODENS .......................................................................................................... 10 MATERIAL AND METHODS .................................................................................................................... 11 GENERAL CONVENTIONS .............................................................................................................. 11 GEOGRAPHICAL COVERAGE ....................................................................................................................... 11 INFORMATION RESOURCES ....................................................................................................................... 12 DATA GATHERING AND PROCESSING ......................................................................................................... 12 FORMAT OF TAXONOMIC DATA ................................................................................................................. 13 MORPHOLOGY ...................................................................................................................................... 15 SUBGENERIC CLASSIFICATION .......................................................................................................... 19 SPECIES GROUPS ................................................................................................................................ 22 TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA .......................................................... 35 AGRILUS TAXA OUTSIDE THE TARGET AREA ...................................................................................... 235 MISIDENTIFICATIONS, NOMINA NUDA, AND OTHER UNAVAILABLE NAMES ....................................................... 236 ANALYSIS ............................................................................................................................................... 237 TAXONOMIC-NOMENCLATURAL ANALYSIS .......................................................................................... 237 EXAMINED MATERIAL ............................................................................................................................... 237 HOST PLANTS ........................................................................................................................................ 238 TAXA DIVERSITY ..................................................................................................................................... 238 DISTRIBUTION ........................................................................................................................................ 239 BIOGEOGRAPHIC ANALYSIS ...................................................................................................................... 240 SIMILARITY OF SUB-REGIONAL FAUNAS ..................................................................................................... 242 CONCLUSIONS ....................................................................................................................................... 242 LITERATURE CITED .............................................................................................................................. 244 APPENDICES ......................................................................................................................................... 267 APPENDIX I. NEW TAXONOMIC AND NOMENCLATURAL ACTS ................................................................. 267 APPENDIX II. SYNOPTIC CATALOG OF EAST ASIAN AGRILUS ..................................................................... 270 APPENDIX III. CHECKLIST OF SPECIES-GROUP NAMES .............................................................................. 276 APPENDIX IV. REGIONAL DISTRIBUTION OF AGRILUS OF EAST ASIA ......................................................... 285 APPENDIX V. MORPHOLOGICAL CHARACTERISTIC OF AGRILUS SPECIES-GROUPS OF EAST ASIA .................. 292 PLATES ................................................................................................................................................... 295

ABSTRACT ..............................................................................................................................................

5

ABSTRACT This is the first comprehensive revision of the east Asian species of the hyperdiverse beetle genus Agrilus (Coleoptera, Buprestidae). The target area includes continental China, Korean peninsula, Mongolia, eastern most Russia and the main Japanese islands. All 278 Agrilus taxa known from the target area of East Asia are treated, among them 74 (26.6%) are described as new. Eight taxa are not confirmed for the region but cited as possible; three species outside the target area are also included to address their taxonomic and nomenclatural issues. All 641 species group names of east Asian Agrilus, both available and unavailable, are cited and annotated. Misidentifications, nomina nuda and other unavailable species group names are briefly mentioned to explain their taxonomic history and status. A total of 459 primary types (89.3% of all relevant) and an additional 9,112 Agrilus specimens originating from 2,627 sampling records have been examined. Comprehensive nomenclatural, taxonomic, typological, bibliographic, biological and biogeographic information is provided for each Agrilus taxon. Exhaustive bibliography comprises 748 publications linked to the 5220 taxon level references. Data from specimens locality labels are used to compose distributional, seasonal and altitudinal data for each species. The core of the book consists of three chapters: Morphology, Taxonomy and Analysis, illustrated with 68 full page plates of 1,053 individual color images. The Morphology chapter presents an overview of morphological characters with their character states illustrated with 173 images on plates 1 7. The Taxonomy chapter is the most extensive part of the book providing A) an overview of all 36 subgenera proposed within the genus Agrilus with critical re examination of their type species; B) the definition and diagnosis of 34 informal Agrilus species groups employed throughout the book with a list of included taxa, host plants and link to related Agrilus species groups; and C) a comprehensive taxonomic inventory of all Agrilus species recorded from East Asia. For each Agrilus taxon the following information is provided: a) adult habitus, aedeagus and type images, whenever possible; b) detailed bibliography with annotated taxonomic and nomenclatural acts and the type of published information; c) complete list of type material with information on the original spelling, type locality, type status, type fixation, locality label data of primary type and reference to publication fixing the primary type; d) a brief diagnosis, assignment to a particular species group and the name of the most closely related species; e) overview on the specimens examined grouped by the country of origin with information on number, sex, verbatim locality and deposition data and specimen depository; f) seasonal and altitudinal adult occurrences compiled from the original locality labels; g) critically re evaluated larval host plant data from published sources complemented with original data; and h) global and regional distribution. The taxonomy chapter is illustrated with 264 habitus images, 248 aedeagus images and 325 type images thus covering the vast majority of treated taxa and types. The Analysis chapter provides statistical and biogeographical analyses. Statistical analyses highlight various aspects of the names, type specimens, examined material, host plants, altitudinal distribution, regional diversity and the origin of East Asia Agrilus taxa illustrated with 14 graphs. Biogeographical analyses provide insight into differences and similarities between various regional Agrilus faunas illustrated with 29 color maps. The New taxonomic and nomenclatural act, proposals and changes: Generic Level: Teres Harris, 1829 is considered an available junior synonym of Agrilus Curtis, 1825; Buprestis granulata Say, 1823 is designated as the type species of Teres

Harris, 1829 (fixation by subsequent designation); type species of Paradomorphus is P. albicollis Waterhouse, 1887 and not P. frontalis Gory & Laporte, 1839; type species of Engyaulus is Agrilus pulchellus Bland, 1865 and not Engyaulus rubrovittatus Waterhouse 1889. Species Level: New Agrilus taxa: 73 new species and 1 new subspecies are described and are attributed to the senior author: A. aureofasciatus, A. aurigaster, A. aurinotus, A. aurithorax, A. bai, A. bicuspidatus, A. bocaki, A. brevinotus, A. businskorum, A. caudicula, A. chaeto coxalis, A. ciping, A. compacticornis, A. conspersus, A. cordicollis, A. daur, A. diao, A. diversornatus, A. dureli, A. eunuchus, A. fora menifer, A. fornicatus, A. fusciapex, A. fusiformis, A. han, A. hani, A. hmong, A. hui, A. improcerus, A. jindrai, A. kucerai, A. lacrima, A. lahu, A. lei, A. lisu, A. lobatus, A. lugu, A. magnus, A. manchu, A. mandala, A. miao, A. mio, A. muongoides, A. naxi, A. ornatoides, A. paralleloides, A. pleuralis, A. pleurostigmus, A. presbys, A. pro latus, A. prosternalis, A. pumi, A. qiang, A. qinlingcola, A. recticollis, A. sagittifer, A. schillhammeri, A. sentis, A. shan, A. she, A. sichuanus, A. siskai, A. smaragdinus habros, A. smetanai, A. spathulifer, A. spiculum, A. stigma, A. tai, A. terracotta, A. tuber, A. ulmicola, A. vigil, A. zhuang, A. zhui. New revalidated names: 2 names are revalidated: olivaceidorsis Obenberger, 1917 is resurrected from the synonymy of A. auriventris Saunders, 1873 as a valid specific name; sapporoensis Obenberger, 1935 is resurrected from the synonymy of A. komareki Obenberger, 1926 as a valid name of the subspecies A. komareki sapporoensis Obenberger, 1935. Change in rank: 1 name is downgraded to subspecies as A. sinuatus sachalinensis Obenberger, 1935. Transfers in synonymy: 4 names are tranferred: cojbalsanensis Cobos, 1968 from synonymy of A. gebleri Obenberger, 1924 to synonymy of A. viridis (Linné, 1758); kiricenkoi Obenberger, 1935 from synonymy of A. attila Obenberger, 1930 to synonymy of A. viridis (Linné, 1758); scutellatus Abeille de Perrin, 1897 from synonymy of A. ecarinatus Marseul, 1866 to synonymy of A. zigzag Marseul, 1866; sieversi Abeille de Perrin, 1897 from synonymy of A. zigzag Marseul, 1866 to synonymy of A. albogularis albogularis Gory, 1841. New synonyms: 69 new synonyms are proposed: alazon Lewis, 1893 synonym of A. decoloratus Kerremans, 1892; aridosus Obenberger, 1936 synonym of A. macrotatopus Obenberger, 1936; aritai Tôyama, 1985 synonym of A. alesi Obenberger, 1935; assamensis Obenberger, 1936 synonym of A. caligans Bourgoin, 1925; attila Obenberger, 1930 synonym of A. tschitscherini Semenov, 1895; brevitarsis Lewis, 1893 synonym of A. viridis (Linné, 1758); brydli Obenberger, 1924 synonym of A. viridis (Linné, 1758); candidiventris Obenberger, 1936 synonym of A. nivosus Abeille de Perrin, 1900; cardaces Obenberger, 1936 synonym of A. viduus Kerremans, 1914; chinganicus Obenberger, 1922 synonym of A. viduus Kerremans, 1914; chopardi Descarpentries & Villiers, 1963 synonym of A. auropictus Kerremans, 1912; daghestanicus Obenberger, 1930 synonym of A. sericans Kiesenwetter, 1857; diolaus Obenberger, 1958 synonym of A. malloti Théry, 1930; djukini Obenberger, 1935 synonym of A. pratensis (Ratzeburg, 1837); duporti Bourgoin, 1922 synonym of A. olivaceidorsis Obenberger, 1917; emisi Ohmomo, 2005 synonym of A. komareki sapporoensis Obenberger, 1935; fairmairei Kerremans, 1892 synonym of A. sinensis sinensis Thomson, 1879; filigranus Obenberger, 1916 synonym of A. salicis Frivaldszky, 1877; fleutiauxi Bourgoin, 1922 synonym of A. auriventris Saunders, 1873; gebleri Obenberger, 1924 synonym of A. viridis (Linné, 1758); gerschuni Stepanov, 1958 synonym of A. tschitscherini Semenov, 1895; graptelytrus Obenberger, 1914 synonym of A. auriventris Saunders, 1873; hoscheki Obenberger, 1916 synonym

6 of A. viduus Kerremans, 1914; imbellis Kerremans, 1892 synonym of A. viridis (Linné, 1758); importunus Kerremans, 1903 synonym of A. viridis (Linné, 1758); kanohi Kurosawa, 1954 synonym of A. auropictus Kerremans, 1912; kanssuanus Obenberger, 1930 synonym of A. smaragdifrons Ganglbauer, 1889; kuldjensis Obenberger, 1936 synonym of A. nivosus Abeille de Perrin, 1900; kusakievici Obenberger, 1936 synonym of A. zigzag Marseul, 1866; lubischevi Stepanov, 1958 synonym of A. tschitscherini Semenov, 1895; lugovoi Stepanov, 1958 synonym of A. salicis Frivaldszky, 1877; machnovskii Stepanov, 1958 synonym of A. tschitscherini Semenov, 1895; malyi Obenberger, 1936 synonym of A. subrobustus Saunders, 1873; melanolucidus Ohmomo, 2005 synonym of A. nipponigena Obenberger, 1935; mentor Kerremans, 1912 synonym of A. plagiatus Ganglbauer, 1889; milletae Baudon, 1968 synonym of A. coomani Bourgoin, 1925; mushanus Kurosawa, 1976 synonym of A. madeci Baudon, 1968; nagaoi Nakane, 1983 synonym of A. maculifer Saunders, 1873; nalajchanus Cobos, 1968 synonym of A. friebi Obenberger, 1922; nipponicola Kurosawa, 1963 synonym of A. fleischeri Obenberger, 1925; omissulus Obenberger, 1935 synonym of A. tyndarus Kerremans, 1913; ornamentifer Obenberger, 1940 synonym of A. olivaceidorsis Obenberger, 1917; panhensis Baudon, 1968 synonym of A. adelphinus Kerremans, 1895; pidjinus Obenberger, 1924 synonym of A. auriventris Saunders, 1873; pistaciophagus Alexeev & Kulinitsh, 1962 synonym of A. chlorophyllus Abeille de Perrin, 1904; poppiusi Obenberger, 1924 synonym of A. viridis (Linné, 1758); praeclarus Krogerus, 1925 synonym of A. pratensis (Ratzeburg, 1837); pseudograminis Jendek, 2007 synonym of A. andrusi Baudon, 1968; pseudoussuricola Alexeev, 1979 synonym of A. kaluganus Obenberger, 1940; richteri Alexeev, 1975 synonym of A. morawitzi Obenberger, 1936; rokuyai Kurosawa, 1976 synonym of A. madeci Baudon, 1968; ryukyuensis Tôyama, 1985 synonym of A. auropictus Kerremans, 1912; sacchari Obenberger, 1940 synonym of A. alesi Obenberger, 1935; salicivola Kurosawa, 1963 synonym of A. viridis (Linné, 1758); semirjeciae Obenberger, 1936 synonym of A. zigzag Marseul, 1866; sericarius Abeille de Perrin, 1897 synonym of A. sericans Kiesenwetter, 1857; sibiricola Obenberger, 1924 synonym of A. laticornis (Illiger, 1803); simbirensis Obenberger, 1925 synonym of A. zigzag Marseul, 1866; spesivcevi Obenberger, 1927 synonym of A. pilosovittatus Saunders, 1873; subviduus Kurosawa, 1957 synonym of A. viduus Kerremans, 1914; suensoni Obenberger, 1940 synonym of A. insulicola Kerremans, 1912; therondi Descarpentries & Villiers, 1963 synonym of A. tyndarus Kerremans, 1913; towadensis Miwa & Chûjô, 1940 synonym of A. viridis (Linné, 1758); toyoshimai Tôyama, 1988 synonym of A. esakii Kurosawa, 1964; transbaicalensis Obenberger, 1935 synonym of A. viridis (Linné, 1758); uzbekistanus Stepanov, 1958 synonym of A. tschitscherini Semenov, 1895; wilsoni Baudon, 1968 synonym of A. quentini

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Descarpentries & Villiers, 1963; yokoyamai Iga, 1955 synonym of A. sinuatus sachalinensis Obenberger, 1935; and zhelochovtsevi Alexeev, 1979 synonym of A. sinuatus sachalinensis Obenberger, 1935. Lectotype designations: 72 lectotype designations are proposed (taxa are cited in the original combination and rank): Agrilus aerosus Ganglbauer, 1889; Agrilus alesi Obenberger, 1935; Agrilus aridosus Obenberger, 1936; Agrilus assamensis Obenberger, 1936; Agrilus attila Obenberger, 1930; Agrilus auriventris Saunders, 1873; Agrilus brodjaga Obenberger, 1914; Agrilus brydli Obenberger, 1924; Agrilus caligans Bourgoin, 1925; Agrilus candidiventris Obenberger, 1936; Agrilus cardaces Obenberger, 1936; Agrilus chinganicus Obenberger, 1922; Agrilus chlorophyllus Abeille de Perrin, 1904; Agrilus daghestanicus Obenberger, 1930; Agrilus dentiapex Obenberger, 1940; Agrilus discalis Saunders, 1873; Agrilus duporti Bourgoin, 1922; Agrilus egorovi Alexeev, 1989; Agrilus emmerichi Obenberger, 1935; Agrilus fleutiauxi Bourgoin, 1922; Agrilus fortunatus Lewis, 1893; Agrilus gebleri Obenberger, 1924; Agrilus graptelytrus Obenberger, 1914; Agrilus hoscheki Obenberger, 1916; Agrilus imbellis Kerremans, 1892; Agrilus imitans Lewis, 1893; Agrilus insulicola Kerremans, 1912; Agrilus jenissejensis Obenberger, 1924; Agrilus kanssuanus Obenberger, 1930; Agrilus kuldjensis Obenberger, 1936; Agrilus kusakievici Obenberger, 1936; Agrilus lugovoi Stepanov, 1958; Agrilus machnovskii Stepanov, 1958; Agrilus macrotatopus Obenberger, 1930; Agrilus malloti Théry, 1930; Agrilus malyi Obenberger, 1936; Agrilus mentor Kerremans, 1912; Agrilus mongolorum Obenberger, 1936; Agrilus morawitzi Obenberger, 1936; Agrilus ningpoensis Obenberger, 1927; Agrilus obscurosericeus Obenberger, 1940; Agrilus olivaceidorsis Obenberger, 1917; Agrilus ornamentifer Obenberger, 1940; Agrilus peregrinus prasinulus Obenberger, 1924; Agrilus pidjinus Obenberger, 1924; Agrilus pliculipennis Obenberger, 1940; Agrilus priamus Kerremans, 1912; Agrilus przewalskii Obenberger, 1936; Agrilus pusillesculptus Obenberger, 1940; Agrilus robertii Chevrolat, 1840; Agrilus rossicus Obenberger, 1916; Agrilus sacchari Obenberger, 1940; Agrilus scutellatus Abeille de Perrin, 1897; Agrilus semirjeciae Obenberger, 1936; Agrilus sieversi Abeille de Perrin, 1897; Agrilus sikkimensis Obenberger, 1928; Agrilus smaragdifrons Ganglbauer, 1889; Agrilus sospes Lewis, 1893; Agrilus spinipennis Lewis, 1893; Agrilus suensoni Obenberger, 1940; Agrilus transbaicalensis Obenberger, 1935; Agrilus trinotatus Saunders, 1873; Agrilus tyndarus Kerremans, 1913; Agrilus unguiculosus Obenberger, 1936; Agrilus uzbekistanus Stepanov, 1958; Agrilus viridicaerulans esfandiarinus Davatchi, 1958; Agrilus viridis cernyi Obenberger, 1925; Agrilus viridis krogerusi Obenberger, 1924; Agrilus viridis poppiusi Obenberger, 1924; Agrilus yunnanus Obenberger, 1927; Agrilus zigzag Marseul, 1866; Buprestis (Agrilus) capreae Chevrolat, 1838.

7

INTRODUCTION The beetle genus Agrilus is by far the largest genus of the family Buprestidae with 2,784 species cited by BELLAMY (2008) [#8576]. In fact, as presently known, Agrilus appears to be the most speciose conventionally accepted genus in the entire Animal Kingdom. Adult Agrilus beetles can be distinguished from other related genera of the jewel-beetle subtribe Agrilina mainly by the presence of a doubled lateral pronotal margin consisting of marginal and submarginal carinae (Fig. 1C); by the presence of a prosternal lobe (Fig. 1B), by the shape of the scutellum (Fig. 1A), and, in most cases, by the presence of a pronotal carina (prehumerus) (Fig. 1C) and by the relatively long metatarsal segment (Figs. 7O, 7P). Agrilus larvae develop inside dicotyledonous plants, adults are also known to feed to attain sexual maturity. Larval and adult host plants can be occasionally different. Gymnosperm plants, except for a single plausible record by SCHWERDTFEGER (1953 [#9021]), are not known to be attacked by Agrilus (FISHER, 1928 [#980]). Many Agrilus species are forest or agricultural pests causing a whole plant or its parts to die prematurely, acting either as primary agents by predominantly larval feeding activity, or serving as vectors of various plant diseases, especially tracheomycoses (KRYUKOVA & TEREKHOVA, 1974 [#8643]). The demand for the present work was primarily triggered by recent North American outbreak of the highly invasive A. planipennis Fairmaire, 1888, colloquially known as Emerald Ash Borer and best known for its ability to attack and kill healthy ash trees. This east-Asian species had been first recorded in North America less than a decade ago (HAACK et al., 2002 [#5632]). Within a very few years the beetle became widespread in the northeastern part of the continent and this expansion is continuing. Economic losses caused by A. planipennis in North America drew much attention to the genus. It became apparent that the continent is highly vulnerable to colonisation by alien Agrilus, since at least eight other exotic species have already arrived to North America across the vastness of the surrounding oceans: A. cuprescens (Ménétriés, 1832); A. cyanescens (Ratzeburg, 1837); A. derasofasciatus Lacordaire, 1835; intentionally introduced A. hyperici (Creutzer, 1799); A. pilosovittatus Saunders, 1873; A. sinuatus (Olivier, 1790); A. subrobustus Saunders, 1873; and A. sulcicollis Lacordaire, 1835. In all cases of unwitting introduction, the original Agrilus stock was embedded in freshly cut wood dunnage or other wood products brought by maritime traffic. The vast majority of the 38 Agrilus border interceptions reported by HAACK et al. (2002) [#5632] for 1985-2000 in the United States were indeed of maritime origin. The recent outbreak of A. planipennis was remarkable by taking the North American taxonomic community entirely by surprise. Initial attempts to identify the first detected specimens of what was a very large and obviously exotic Agrilus were unsuccessful. Any other result could hardly be expected since this species, despite its natural distribution in six countries from Russia to Laos and being among the

largest and most colorful within the genus, also happened to be among the most obscure. Zoological Record, the main reference source in animal taxonomy, has only five pre-2002 secondary references to the species under three different names. The present day synonymy and lectotype of A. planipennis (and two among its three synonyms) have been designated only more than hundred years after the species was first described (JENDEK, 1994 [#1355]). It is plausible to assume that prior to its arrival, not a single specimen of A. planipennis existed in North American collections, either as an identified voucher or among unsorted material. Positive identification of this species was finally made by the first author, then in Bratislava, after receiving a shipment of North American specimens. The unassuming story of this A. planipennis identification revealed an acute lack of basic taxonomic expertise pertaining to this, and potentially other insect taxa, including those of high economic significance. Difficulties in Agrilus identification, although seemingly surprising, can hardly be avoided. Even though the rate of new Agrilus species descriptions rose markedly during the past decades (Fig. 55C), the overall taxonomic understanding of the genus, particularly when seen in a global prospective, remains highly unsatisfactory. Hundreds of new species remain to be discovered and described, while the current subgeneric classification is highly confusing if not outright misleading. The taxonomic concepts of the vast majority among the described species have never been critically reexamined, type specimens of older names are scattered in various collections and primary types are often not designated, while apparently hundreds of synonyms remain undetected. These underlying taxonomic hurdles prevent further progress and, owning to them, hardly any world region has its Agrilus fauna adequately revised. Overall taxonomy of the jewelbeetle genus Agrilus is just another sobering reminder to the all-mighty humans about how little we actually know about the world around us. The current project was envisioned, undertaken and completed after having realized the current unsatisfactory state of East Asian Agrilus taxonomy and the risk it poses to the plant resources in North America. The target region of East Asia (Fig. 58A), as defined for the purpose of our work, is a vast land with highly diverse types of ecological biomes (Fig. 58A) varying from tropical and subtropical forests, through broadleaf and coniferous forests, to grasslands, tundra and deserts. This area forms the eastern-most part of the Palaearctic zoogeographical region bordering with the Oriental region and therefore has a mix of species from both, along with a number of endemics. Different authors attempted to summarize Agrilus data at the national level for China (PENG ZHONGLIANG, 1987 [#2429]; HUA LI ZHONG, 2002 [#6696]), Japan (e.g., MIWA & CHUJO, 1936 [#1809]; IGA, 1955 [#3828] and 1962 [#3351]; KUROSAWA, 1963 [#1618]; TÔYAMA, 1985 [#3678] and AKIYAMA & OHMOMO, 1997 [#4091]), Mongolia (ALEXEEV & VOLKOVITSH, 1989 [#3241]), and Russia (ALEXEEV, 1989 [#64]), while the

8 comprehensive Agrilus revision for the entire region has never been attempted. The present book is aimed to bring all available information on east-Asian Agrilus beetles, either new or previously published, under a single cover. More specifically, it has the following main goals: - Present thoroughly revised up-to-date taxonomic concepts for every Agrilus species native to East Asia known to us, either new or previously named; - Promote a maximally optimized approach to methodology and terminology of Agrilus species description by providing a set of abundantly illustrated standardized morphological characters and their states that can be utilized consistently; - Facilitate species identification by providing every treated species, whenever possible, with a detailed differential diagnosis supplemented with habitus and male genitalia images; - Assign each east-Asian Agrilus species to previously suggested or newly established informal species-groups; - Complete a directory of all east-Asian Agrilus speciesgroup names, both available and unavailable, with indication of their original and current rank, availability and validity by accessing, verifying and summarizing all relevant publications; - Bring together all available information about all namebearing type specimens of east-Asian Agrilus including their original number, status and type locality together with verified data on their present depository, numbers and status, as well as to identify primary types by providing verbatim label data and specimen images; - Compile data on global and regional distribution for each east-Asian Agrilus taxa based on critically evaluated published sources and newly examined specimens; - Present seasonal and altitudinal occurrence of each eastAsian Agrilus species-group taxon based on statistic assessment of examined specimens and published faunal records; - Provide new as well as critically re-evaluated published information on larval Agrilus host plants; - Undertake spatial analyses of east-Asian Agrilus distributional data to examine regional diversity, faunal similarity and endemism rate among various sub-regions. Summing up, this publication should offer the first comprehensive overview on the east-Asian Agrilus fauna. The book is expected to provide a much needed taxonomic and intellectual framework for all subsequent studies, such as ecological, phylogenetic, or conservational, where precise Agrilus species-level identification matters. We hope our work will also enhance knowledge in such areas as biodiversity and biogeography, and will find its applications into the problem

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA of the invasive alien species. Besides such pragmatic goals, we also hope to make the extraordinarily diverse genus Agrilus somewhat more familiar to us humans, and not necessarily as feared and despised “pests”, but as esthetically beautiful and fascinating biological phenomenon. Having indicated what this book is expected to be, we feel compelled to say what it is not. First of all, there is no classical dichotomous species identification key. To do so for 270+ species, some of them inadequately known, is not only too much of a challenge for the authors, but also likely of little use to the reader. To identify an Agrilus species the reader should instead go through the habitus illustrations to find the species looking most alike, then see what species-group it belongs to, then look at the species diagnostic characters of the group itself and the species it includes trying to find the best match, and finally attempt using male genital characters. An alternative strategy would be to send east-Asian Agrilus specimens to us for identification, preferably unmounted in ethanol, and let us keep them. We always welcome an influx of new material, since hardly any sub-region of East Asia has been adequately sampled. Despite the considerably increased collecting efforts made in East Asia in the last two decades (Fig. 56B), vast areas have no or very few Agrilus records (Fig. 58B, 59A). The shortage of recently collected specimens was considered the main limitation when completing this work. Even with the presently recorded 278 Agrilus taxa, we firmly believe that East Asia still harbours hundreds of undetected species (Fig. 67B). Other areas this book does not intend to cover are immature stages, behaviour, parasites, dispersal and other aspects too numerous to mention. Interested readers are referred to the rather extensive body of literature accumulated over the last few years by mainly North American authors on A. planipennis and then use it as a starting point by cautiously applying these data to other congeners. Last, but not the least, we did not attempt to answer the seemingly main Agrilus-question: why are there so many of them? Indeed, on the one hand, what were those natural forces to diversify the Agrilus lineage into countless species making it the largest genus of the entire Animalia, and, on the other hand, to keep all of them so alike preventing overzealous entomologists from splitting it into a multitude of smaller units? Were these forces represented by the “higher” plants rapidly appearing everywhere on the planet to feed hungry Agrilus larvae? Could they work through some peculiarities of adult locomotion letting beetles fly, run, mate and oviposit in a way more efficiently than anything else? Was it a still unknown usage of the larval terminal hooks unique for Agrilini or, could it be their most peculiar bent pre-pupal posture? What is the likeliest of all is that we are simply yet too ignorant even to imagine the true causes. In private we seek comfort in hoping to live long enough to witness some well-grounded hypothesis attempting to answer this main Agrilus question.

9

ACKNOWLEDGEMENTS Numerous people kindly, and in a variety of ways, facilitated our work on this project. All individuals mentioned in the list of accessed collections (see “Material and Methods”), either the owners or curators, are sincerely acknowledged for their help in accessing the material under their care. Among them Vít Kubáň and Svatopluk Bílý (both Prague, Czech Republic), Jean J. Menier, Thierry Deuve and Isabelle Bruneau de Miré (all Paris, France), Malcolm Kerley and Maxwell Barclay (both London, UK), Peggy Albert (Beziers, France) and Philippe Rilliard (Narbonne, France) went well beyond their duty in facilitating our work. Numerous specimen collectors, whose names are mentioned when citing examined Agrilus material for every species, are acknowledged for their contribution to increasing Agrilus holdings and knowledge. Aleš Smetana (Ottawa, Canada) helped with everyday discussions on various aspects constantly arising in the course of the work and critically read early versions of the manuscript. Yves Bousquet (Ottawa, Canada) commented on old taxonomic literature, drew our attention to the date problem of the Laporte & Gory publications, and reviewed an early version of the manuscript. Sadahiro Ohmomo (Inashiki, Japan) provided specimens of Japanese Agrilus and commented on the Agrilus fauna of Japan.

The Ernst Mayr Fellowship (Museum of Comparative Zoology, Harvard University, Cambridge, USA) supported the first author’s 2009 travel to Prague, Czech Republic and St. Petersburg, Russia, to re-examine Agrilus type specimens. Karen McLachlan-Hamilton, Bruce Gill and Doug Parker (all Ottawa, Canada) critically read the MS prior to its submission. Analytical and assembling work on this project has been performed during the years 2009-2011 at the Canadian National Collection of Insects, Arachnids and Nematodes in Ottawa, Canada. Funding for this work came through the Visiting Fellowship Program to Canadian Governmental Laboratories awarded to the first author and administrated through the Entomology Research Unit, Ottawa Plant Laboratory, led by Linda DeVerno, itself a part of the Canadian Food Inspection Agency (CFIA). This publication forms a part of a broader CFIA research project “Phytophagous beetles from the Asia Pacific region posing a risk to Canadian Plant resources” administrated by the second author. Bruce Gill and Linda DeVerno envisioned completion of this project by facilitating both authors joining the Agency and engaging in Agrilus research. Last but not least, our wives, Janka and Tatiana, respectively, who gave lasting patience, understanding and support in completing the book.

10

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

ABBREVIATIONS

AND CODENS

Codens for collections used in the text follow those listed in ARNETT et al., 1993 [#6167] and in: http://hbs.bishopmuseum. org/codens/codens-r-us.html with some addition and modifications. Names of the main contact person(s) when accessing Agrilus holdings in respective public collections are given in brackets (where applicable). Abbreviations for collections AAOR AGCM APCE BMNH CBCS DEI DGCC EIHU EJCB EKCS FACF GNCW HHCC HHCR HMMG HNHM IMCM ISCK ISNB IZAS JLCS JMCZ JVCO MCCD MCSN MDCM MHNB MHNG MKCY MNCA MNHE MNHN MNMS MSCI NHMB NMBS NMPC NMW NSCF NSMT

Alexeev, A. V., Orekhovo Zuyevo, Russia Gorodinski, A., Moscow, Russia Pütz, A., Eisenhüttenstadt, Germany The Natural History Museum, London, United Kingdom (Kerley, M.; Barclay, M.) Bellamy, C., California, Sacramento, USA Deutsches Entomologisches Institut, Eberswalde Finow, Germany [currently moved to Müncheberg] (Zerche, L.) Gianasso, D., Castelnuovo Don Bosco, Italy Hokkaido University, Sapporo, Japan Jendek, E., Bratislava, Slovak Republic [presently in Ottawa, Canada] Kučera, E., Soběslav, Czech Republic Angelini, F., Francavilla Fontana, Italy Novak, G., Wien, Austria Collection of H. A. Hespenheide, University of California, Los Angeles, USA Hebauer, H., Rain, Germany Mühle, H., München, Germany Hungarian Natural History Museum, Budapest, Hungary (Merkl, O.) Melnik, I. V., Moscow, Russia Smatana, I., Košice, Slovakia Institut Royal des Sciences naturelles de Belgique, Brussels, Belgium Institute of Zoology, Academia Sinica, Beijing, China Lee, Jun gu, Seoul, South Korea Marek, J., Zbecno, Czech Republic Vávra, J., Ostrava, Czech Republic Curikov, M. N., Donskoe, Russia Museo Civico di Storia Naturale “Giacomo Doria”, Genova, Italy (Poggi, R.) Danilevski, M., Moscow, Russia Muséum d’histoire naturelle, Béziers, France (Albert, P.) Muséum d’Histoire Naturelle, Geneva, Switzerland (Löbl, I., Cuccudoro, G.) Kalashian, M., Yerevan, Armenia Niehuis, M., Albersweiler, Germany Museum of Natural History, Tallinn, Tallinn, Estonia Muséum National d’Histoire Naturelle, Paris, France (Menier, J.J; Deuve, T.; Bruneau de Miré, I) Museo Nacional de Ciencias Naturales, Madrid, Spain Smirnov, M. E., Ivanovo, Russia Naturhistorisches Museum, Basel, Switzerland (Sprecher, E.) Naturhistorisches Museum, Bern, Switzerland National Museum (Natural History), Prague, Czech Republic (Bílý, S.; Kubáň V.) Naturhistorisches Museum Wien, Wien, Austria (Schillhammer, H.) Sugiura, N., Fukushima, Japan National Science Museum (Natural History), Tokyo, Japan (Nomura, S.)

PMCL Moravec, P., Litoměřice, Czech Republic RKCK Królik, R., Kluczbork, Poland RMNH Nationaal Natuurhistorische Museum (“Naturalis”), Leiden, Netherlands SGCB Gotwald, S., Berlin, Germany SMNS Staatliches Museum für Naturkunde, Stuttgart, Germany SOCI Ohmomo, S., Ibaraki ken, Japan SZMN Institute of Animal Systematics and Ecology, Siberian Zoological Museum, Novosibirsk, Russia TARI Taiwan Agricultural Research Institute, Taichung, Taiwan TSCB Sainval Neef de, Thierry, Brussels, Belgium TSCO Sitek, T., Ostrava, Czech Republic USNM National Museum of Natural History, Washington D.C., USA (Furth, D.G.; Lingafelter, S.W.) VNCO Nemkov, V. A., Orenburg, Russia VSCS Sakalian, V., Sofia, Bulgaria WBCW Barries, W., Wien, Austria ZIN Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia (Volkovitsh, M.) ZMAN nstituut voor Taxonomische Zoologie, Zoologisch Museum, Universiteit van Amsterdam, Netherlands (Lackner, T.) ZMHB Museum für Naturkunde der Humboldt Universität, Berlin, Germany (Hieke, F.) ZMUM Zoological Museum, Moscow State University, Moscow, Russia (Nikitsky, N.B.) ZSCB Šustek, Z., Bratislava, Slovakia ZSMC Zoologische Staatssammlung, München, Germany (Baehr, M.) Other abbreviations and symbols [] … AT Code ed(s) [h] HT HTM ICZN LT MS nom. rev. [p] pers. com PLT PT sp. inqurenda sp. nov. ssp. nov. ST stat. nov. syn. nov.

annotation or comment piece of irrelevant text or illegible handwriting allotype International Code of Zoological Nomenclature editor(s) handwritten holotype holotype by monotypy International Commission on Zoological Nomenclature lectotype handwriting, manuscript revalidated name printed personal communication paralectotype(s) paratype(s) species inqurenda new species new subspecies syntype(s) new status new synonym

11

MATERIAL

AND

METHODS

GENERAL CONVENTIONS Adherence to the Code. Nomenclatural issues were resolved by following the fourth edition of the International Code of Zoological Nomenclature (ICZN, 1999 [#4793]). Articles or Recommendations of the Code cited in the references are simply cited as “Article” or “Recommendation” with the relevant index. Citations of names. Taxonomic taxa (taxonomic concepts) are currently recognized species (or subspecies) and as such are cited as a binomina (or trinomina) either in the full form or with the generic name abbreviated. Species synonyms (valid or invalid) are cited as uninomina. Their original generic (subgeneric) combination, along with their original rank reflecting their use in the former taxonomic concepts, is cited in the references enclosed in parenthesis after the author and year. Because the original and current generic assignments are explicitly specified, the author’s name is never enclosed in brackets. The names are listed with all mandatory changes in spelling consequent upon changes in gender of various generic assignments. Types in the Chapter “Taxonomic inventory” are cited as binomina in the original generic combination and rank with the author(s) and year. This convention best conforms to the practice of type specimen labeling and record keeping in collections. Images of the primary types are displayed in alphabetical order based on the respective species-group names. Throughout the text, all genus- and species-group names are cited in italic except for verbatim transcription of locality labels and when part of a title in the chapter “Literature cited”.

New taxonomic and nomenclatural acts. All new taxonomic and nomenclatural acts are attributed to the senior author. Newly described taxa, type fixations, changes in binominal combination, rank, and validity are explicitly indicated and summarized in Appendix I. Species inquirenda and nomina dubia. Some deficiently described Agrilus species with untraceable or lost types have unknown identity. Names for these species inquirenda are called nomina dubia (see Glossary of the Code). These unidentifiable species-concepts are listed among other taxa and flagged with the phrase “species inquirenda”. Because these names are available, some of them can be synonyms of currently recognized species. If they are senior synonyms, then ICZN, Article 23.9 can be later applied to protect the younger names. Unavailable names. Unavailable names are excluded from the synonymy of species in the chapter “Taxonomic Inventory”, but they are cited in the Synoptic Catalog of East Asian Agrilus (Appendix II) and Checklist of species-group names (Appendix III) to avoid possible confusion or questions about their status. Transliteration. Titles of articles, type localities or locality labels cited in Cyrillic are transliterated to Latin using the British standard. The original language is mentioned in square brackets. In very few cases, the locality labels are cited in original Cyrillic. English titles of articles translated from Chinese or Japanese are given in square brackets.

GEOGRAPHICAL COVERAGE As defined for the purpose of our work, the entire region termed “East Asia” includes in full or in part the territories of the following six countries (Figs. 57B, 58A, 58B): Russian Federation (Russia), Mongolia, People’s Republic of China (China), Democratic People’s Republic of Korea (Korea North), Republic of Korea (Korea South) and Japan. Mongolia, Korea North and Korea South are included entirely. Of Russia only the following eastern-most administrative units are included: Amurskaya oblast’; Buryatiya Respublika; Chukotskiy avtonomnyy okrug; Irkutskaya oblast’; Kamchatskiy kray; Khabarovskiy kray; Magadanskaya oblast’; Primorskiy kray; Sakha Respublika; Sakhalinskaya oblast’; Tyva Respublika; Yevreyskaya avtonomnaya oblast’; and Zabaykalskiy kray. Except for the southern islands of Hainan and Taiwan, the whole continental China is included with the following administrative units: Anhui; Beijing; Fujian; Gansu; Guangdong; Guangxi; Guizhou; Hebei; Heilongjiang; Henan; Hong Kong; Hubei; Hunan; Jiangxi; Jiangsu; Jilin; Liaoning; Macau; Nei Mongol; Ningxia; Qinghai; Shaanxi; Shandong; Shanghai; Shanxi;

Sichuan; Tianjin; Xinjiang; Xizang; Yunnan; and Zhejiang. The territory of Japan is subdivided geographically after the main islands (Hokkaido, Honshu, Kyushu, Shikoku, and Tsushima), rather than following administrative subunits, and the predominantly Oriental archipelagos Bonin and Ryukyu are excluded. The species distribution in this book is defined up to the first administration level for China and Russia, and by the main islands for Japan. The species distribution for Mongolia and both Koreas is given on the country level without further subdivision. These geographical units are consistently termed “sub-regions”, while the term “region” normally refers to the entire target area of East Asia. The political subdivision and general natural conditions of the target area of East Asia are shown in Figs. 57B, 58A, and 58B. The map of terrestrial biomes (Fig. 58A) was adopted from the vector digital map: “Terrestrial Ecoregions of the World”, Version 2.0, 2004, provided by the World Wildlife Fund – US, and based on Olson et al., 2001 [#8903].

12

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

INFORMATION RESOURCES Examined specimens. The following institutions or private collections have been personally visited to study type or nontype Agrilus specimens: AAOR, BMNH, CBCS, DEI, HNHM, MCSN, MDCM, MHNB, MHNG, MNCA, MNHN, NHMB, NMPC, NMW, VSCS, WBCW, ZIN, ZMHB, ZMUM, ZSCB. The rest of the material was made available by post either via institutional loans (mainly EIHU, NSMT, SOCI, USNM, ZSMC) or through personal arrangements. Type specimens. Among the total of 513 Agrilus primary type specimens pertaining to East Asia, 459 (89.5%) were personally examined; for more details refer to the “Examined material” in the chapter “Analysis”. All relevant type specimen data, such as their number, sex, status, current depository and label data of at least the primary type specimen, were recorded. Characters of ventral side and male genitalia of the majority of the primary types have been examined by dissection and remounting using a novel style of L-shaped mounting cards allowing specimen observation from all sides. The subsequently designated type specimens were provided with a red printed label bearing the specific name in original rank and generic combination, name of designator, and year of labeling. The habitus of each examined primary type was photographed (see below), often along with the aedeagus and important body parts. During the process of type examination, every effort was made to find a conspecific specimen in the collection of the first author. If such a specimen was found, it has been labeled by the determination label with the inscription “Cum typo comparavit” to highlight its conspecifity with the name bearing type. If a conspecific specimen was not found, the most similar specimen has been labeled with the tag inscribed “Near to”, to indicate its similarity with the examined type specimen. As a result of this multi-year routine, the collection currently comprises of the most reliably determined species to make it, along with the database (see below), the most complete and reliable source of information for Agrilus of the eastern Palaearctic and Oriental regions. Other specimens. Altogether 2,627 Agrilus faunal records and 9,112 specimens (not counting primary types) have been examined in the course of the present work. The vast majority of these specimens form part of the first author’s collection assembled over the last 30 years (Fig. 56B). The overview on various numerical aspects for studied specimens, such as their numbers, geographic origin, altitudinal distribution and

current depository, is given in the chapter “Analysis”. All cited specimens, except for very few annotated exceptions, were personally determined and their verbatim label data was entered in the database (see below). Each faunal record was also entered in the database to contain, if known, information on the number of specimens, their sex, collection date, altitude, collector’s name, country and county of origin, geographic coordinates, current depository, and the identifier’s name. The Bibliography at the end of this book includes 748 of the most relevant titles which are linked to more than 5,220 commented taxon-level references. Each reference was examined for the type of information included. For taxa occurring exclusively in East Asia, all available articles are reported. For widely distributed Eurosiberian taxa, most of the European publications of local importance were omitted. Emphasis was placed on monographs, revisions, catalogs, keys and publications providing new nomenclatural or taxonomic acts or new information on biology, host plants, distribution, and pest management, etc. All relevant publications prior to 2010 are included, while some later titles are also referred to. Publication of Laporte and Gory. NELSON & BELLAMY, 1993 [#3167] published a critical article on the authorship and publication dates of Histoire naturelle et iconographie des insectes coléoptères by Laporte and Gory. Their list with publication dates of various “livraisons” arose, without further scrutiny, “as they appear in the Inter Documentation Company micro-edition”. Yves Bousquet (Ottawa, Canada) informed us about the incorrect date of publications mentioned by NELSON & BELLAMY, 1993 [#3167], at least for the “livraison” where Agrilus names were mentioned. He drew our attention to the article of ERICHSON, 1840 [#8752] announcing the commented overview of entomological publications from the year 1839. On page 240, Erichson comments about the continuation of Laporte and Gory’s work and cites genera included with Agrilus. It becomes evident that the Agrilus “livraisons” was available to Erichson already in 1839, and therefore 1839 is the earliest proven date of publication of Laporte and Gory’s Agrilus taxa. There are also several further corroborating announcements in the Archiv für Naturgeschichte published by various authors on other “livraisons” of Laporte and Gory’s work and matching the citations of SHERBORN, 1922 [#4372] who also used them to resolve publication dates.

DATA GATHERING AND PROCESSING Database. The taxonomic parts and bibliography were outputted from the taxonomic relational database MS Access 2003 to the text format RTF (Rich Text Format). The text was further formatted to the final layout in MS Office Word 2003.

The first, very simple version of the database used for the present work was created by the first author in 1992 using Access Version 1.0. In subsequent years the database has gradually grown to its current state in both complexity

MATERIAL AND METHODS and data volume. The current version is believed to be the most complete and updated data source on Agrilus of the Palaearctic and Oriental regions storing all relevant published or unpublished information. The information is kept in a modular and consistent way through various interrelated tables. Data in the database can be easily and efficiently entered, edited, filtered, analyzed, figured and outputted. Some types of data, such as images and PDF files, are stored outside the database and linked to the relevant records via hyperlink. Illustrations. The quality of the images of the primary types in Figs. 42-54 varies widely, depending on the circumstances of when the image was taken. Many of the earlier images were of the type specimens taken on-site using simple equipment. The first digital camera used in 1999 was a Nikon Coolpix 950, which was later replaced by a Nikon Coolpix 4500. Both were

13 armed with close-up macro adapters, fixed to a provisional stand and illuminated by standard compact fluorescent bulbs. All other beetle images are recent and were taken with a Leica DC500 digital camera mounted on a Leica Z16 APO stereoscope with a Planapo 1.0x/WD 97mm lens. The open source CombineZM image processing software package developed by Alan Hadley (http://www.hadleyweb.pwp. blueyonder.co.uk) was used to assemble multiple images from successive focal planes into a single deep-focus image. All photos and drawings were processed and assembled into plates using Adobe Photoshop CS3. A few line drawings were made using vector-based CorelDraw 12 (Corel Corporation). Graphs on Figs. 55-57 were produced using the graphic tools available in MS Access 2003. Geographic analysis and maps on Figs. 57-68 were generated by exporting data from the taxonomic database format in Access 2003 to Esri ArcMap 9.2.

FORMAT OF TAXONOMIC DATA General structure. Species and subspecies are arranged in alphabetical order. The layout for taxa consists of: Synonymy, References, Type material, Diagnosis, Species group, Related species, Specimens examined, Adult occurrence, Altitude range, Host plants, Distribution and Remarks. Newly described taxa have a layout similar except for the parts Description and Etymology. Synonymy. The valid synonym is cited first, followed by the chronological list of invalid ones (if any). Each invalid synonym is preceded by the sign “=”. Preoccupied names are annotated by the phrase (preoccupied) and replacement names by the phrase (replacement name for) following by the preoccupied name and by both, the junior and the senior author(s), separated by the phrase “not”. Only available synonyms are cited in the chapter “Taxonomic Inventory”, while a complete list of all names is given in full in Appendices II “Synoptic catalog of East Asian Agrilus” and III “Checklist of species-group names of Agrilus of East Asia”. References are grouped by specific names. The original publication establishing the name is cited first with the annotation “description” or “replacement name proposal” enclosed in round brackets “( )”. Subsequent references follow chronologically, separated by a dash “–”. Each reference includes information about author, year, page(s) and numeric identifier (see cross-indexing below). The reference author, unlike the taxon author, is highlighted using SMALL CAPITALS. All references are briefly annotated in round brackets with information, whenever applicable, on the generic assignment (if other than Agrilus), subgeneric placement, original rank (if other than species), brief indication of information, as well as country or region from where the taxon is reported. In addition to the annotations, remarks or explanations concerning problems or inconsistencies are cited in square brackets “[ ]”. Cross-indexing between references and “Literature cited”. References are cross-indexed with the “Literature

cited” through a unique number beginning with “#” and enclosed in square brackets. This method was favored over the classical way of indexing by unique combination of Author, year, and letter; mostly due to the database origin of the data. The numeric identifier is cited beyond each reference providing thus an unambiguous link to the related publication listed in the bibliography. Type material. Type specimen data are cited for each nominal taxon in the same sequence as they appear in the synonymy. The term Type locality refers to the originally published type locality. The original language, if other than those using the Latin alphabet, is mentioned in the brackets. Locatities published in Cyrillic are transliterated into the Latin alphabet using the British Standard. If no type locality is cited, the phrase “not given” is used, while in the case of indirect indication of the locality in the original publication, this fact is implicitly mentioned. Comments and additions are given in square brackets. Since the type locality of a nominal species-group taxon is the place of origin of the namebearing type (ICZN, Article 76.1), the originally published type locality can be altered or corrected based on the other related sources of information (ICZN, Recommendation 76A), such as the locality label of the primary type. When ascertaining and clarifying the type locality (ICZN, Article 76A.1), both the originally published type locality and the primary type locality label should be considered. Type specimens. The primary type is cited separately in the following sequence: Status (holotype, holotype by monotypy, lectotype), sex, reference to subsequent designation (if applicable), depository, and verbatim locality data in quotation marks. The secondary type citations, whenever applicable, have the following format: number of specimens, sex, status and depository. If the primary type has not been fixed, subdivision to primary and secondary types is disregarded. The last sentence specifies the number of originally published specimens in the type series. Unless stated otherwise, all data on the type specimens were

14 personally verified. Untraced types are indicated as “not found”. Paratypes of newly described taxa are grouped by country and, when applicable, by county. Descriptions. Descriptions for newly described taxa were generated using the diagnostic software Lucid 3, Version 3.4 Builder (Centre for Biological Information Technology, Queensland University, Australia). Descriptions are based on the multi-character states matrix which can be transformed to “natural” descriptions using preformatted templates. All character states used in the descriptions are explained and illustrated in the chapter “Morphology”. Diagnosis. This section provides the most important and readily useful diagnostic characters to distinguish a taxon from the most similar taxon or taxa listed in “relevant species”. Each species is assigned to a species group and one or more closest or most similar species are cited in the section “relevant species”. Specimens examined. Except for a very few annotated examples, all specimens listed in this section were personally determined and recorded. Data for each collecting record, whenever applicable, are given in the following sequence: administrative region, number of specimens, sex, current specimen depository given in brackets, followed by a colon and verbatim quote of locality label data. If specimens have been previously reported, a reference is provided. Records are grouped by countries and first-level subordinate administration units, as stated in the Geographic Coverage. Adult occurrence and Altitude range represent the months when adult beetles were collected and the altitude in meters, respectively. Both are derived from the locality labels of examined specimens. The number of records (n) used for each calculation is given in round brackets. The number of records for “Adult occurrence” is a number of unique combinations of three parameters: locality, date and collector. The number of records for “Altitude range” is counted equally, but only for the localities with coordinates. In a few cases, additional faunal records published by other

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA authors are included in calculations and a reference to such a source is always given. Host plants. Data on host plants are extracted from published sources or from the locality labels of examined specimens. If a host plant was published under a synonymic name of that taxon, this fact is stated in parentheses. The Latin names of host plants are cited in the original, unaltered form, without regard to its current availability or validity. Vernacular names of plants are translated to the genus level. Names are arranged alphabetically followed by the colon and references in chronological order separated from each other by a dash. Each reference includes the author, year and RefID (see crossindexing). The list of references is not comprehensive and does not include every published mention for each host plant. The host plant of Agrilus beetles is the plant where the larval stage develops. Reliable host plant records should be ideally based on adults reared from reliably determined plants. In most publications, however, host plants are cited without reference to the source of information. The situation becomes more complicated by misidentifications of Agrilus species, by frequent taxonomic changes in both beetles and plants, and by unreliable host plant records (e.g., plant on which adult was collected). Unverified or likely erroneous host plant records are grouped at the end of the list with comments given in parentheses following the host plant name. Distribution. Distributional data are provided as an alphabetical list of countries. First level administrative units are given for China and Russia. Japan is subdivided geographically. For some widely distributed species, local and global distribution data are separated into two paragraphs with the global distribution being further separated for records from Europe, Asia and North America. Russia and Turkey, two countries extending over Europe and Asia, are mentioned independently for each continent. Countries where a species has been introduced by humans, either consciously or not, are noted as such.

15

MORPHOLOGY Definitions of adult Agrilus morphological structures given in this chapter explain and illustrate characters and character states consistently used throughout this work, particularly when generating verbal descriptions of new taxa from a numerical matrix and when defining Agrilus species-groups. The majority of Agrilus specimens possess a characteristically prolonged and elevated keel-like structure on each side of the pronotal disk, frequently called pronotal carina. Since the expression of this structure varies markedly from being sharply carinal, through filamentary and tubercular, to obsolete and absent, we use a more neutral term prehumerus regularly found in French literature and indicating its close position to elytral humerus. Body Size - small, ≤ 5 mm; medium, > 5 mm and ≤ 10 mm; large > 10 mm. Shape - subparallel (Fig. 2A), body subparallel, apex of elytra broad; cuneiform (Fig. 2B), apical third of elytra markedly converging, elytral apex narrow; fusiform (Fig. 2C), body narrowed on both ends, head small, pronotum widest at basal half. Build - slender (Fig. 2D); robust (Fig. 2E). Color (dorsally) - unicolored (Fig. 2F); bicolored (Fig. 2G). Color (dominant) - blue; green; brown; red. Dominant color of dorsal side of body. Head Shape - markedly convex (Fig. 2H); obviously flat (Fig. 2I). Medial impression - absent (Fig. 2J); present (Fig. 2K). Medial impression (depth) - shallow; deep (Fig. 2K). Medial impression (extent) - vertex; frons; vertex and frons. Epistoma - in plane with frons; with raised upper margin (Fig. 2CC). Frons Shape - flat (Fig. 2L); moderately convex (Fig. 2M); markedly convex (Fig. 2N). Outline - protruding from head outline (Fig. 2O); not protruding from head outline (Fig. 2P). Microsculpture - present (Fig. 2R), sculpture of frons is complemented with fine but distinct microsculpture. Vertex Shape - markedly convex, in postero-dorsal view. Outline - protruding from head outline (Fig. 2Q); not protruding from head outline, in postero-dorsal view. Sculpture (predominant) - punctures (Figs. 2S, 2T); rugae (Figs. 2U, 2V, 2W, 2Y, 2Z). Sculpture (aspect) - irregular (Fig. 2S); regular (Fig. 2T); semispherical (Fig. 2V, 2X); subparallel (Figs. 2Y, 2Z). Sculpture (density) - sparse (Figs. 2S, 2Z); dense (Figs. 2U, 2W, 2Y). Sculpture (intensity) - superficial (Figs. 2S, 2Z); rough (Fig. 2U). Eyes Size - small (Fig. 2I), width of eye is smaller than half width of vertex; moderate (Figs. 2N, 2O), width of eye is subequal to half width of vertex; large (Figs. 2H, 2J, 2P), width of eye is larger than half width of vertex (dorsal view). Shape - not protruding from head outline (Fig. 2P); protruding from head outline (Fig. 2O). Lower margin - above antennal socket (Fig. 2AA); in line with antennal socket; below antennal socket (Fig. 2BB). Antennae Length - short (Figs. 15B, 28B), antennae reach to about anterior pronotal angles; moderate (Fig. 8A), antennae reach to middle of pronotal length; long (Fig. 29B), antennae reach to or beyond posterior pronotal angles. Shape - slender (Fig. 2DD); stout (Fig. 2EE), with antennomeres 5-10 obviously short and wide. Sexual modification in male - not apparent; antennae longer; antennomeres modified (Figs. 2FF, 23K, 24F). Pronotum Shape - visually elongate (Figs. 10C, 25B); visually square (Fig. 12K); transverse (Fig. 12G). Remark: Truly elongate or square pronotum is rarely found, but some transverse shapes may visually appear as elongate, mostly in slender species with large head. Sides - straight (Fig. 3T); arcuate (Fig. 3U); markedly arcuate (Fig. 3V); subangulate (Fig. 3W). Maximal width - at anterior margin (Fig. 3K); at middle (Fig. 3L); at posterior margin (Fig. 3M). Anterior margin - wider than posterior (Fig. 3N); subequal to posterior (Fig. 3O); narrower than posterior (Fig. 3P).

16

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Anterior lobe Presence - absent (Fig. 3B); vague (Fig. 3C); moderate (Fig. 3D); obvious (Fig. 3E). Shape - arcuate (Fig. 3F); subangulate (Fig. 3G). Position to anterior pronotal angles - not reaching level of anterior pronotal angles (Fig. 3H); at level with anterior pronotal angles (Fig. 3I); projecting beyond anterior pronotal angles (Fig. 3J). Posterior angles Apex - sharp (Fig. 3CC); blunt (Fig. 3DD). Shape - acute (Fig. 3HH); rectangular (Fig. 3II); obtuse (Fig. 3JJ). Disk Convexity - flat (Fig. 11E); moderately convex (Fig. 12C); strongly convex (Fig. 15D). Disk modifications - with obvious tomentose spots (Fig. 10F); with two black longitudinal stripes (Fig. 16A). Disk impressions Presence - absent; medial only; medial and lateral (Figs. 3Q, 3R); lateral only; posterolateral (Fig. 3S). Medial impression (shape) - entire (Fig. 3Q); anteromedial and posteromedial (Fig. 3S); posteromedial only. Medial impression (deepness) - shallow; deep. Anteromedial and posteromedial impressions - smaller anteromedial and larger posteromedial; subequal in size. Lateral impressions (deepness) - shallow; deep. Lateral impressions (width) - narrow; wide. Prehumerus Development - absent (Fig. 4A); obsolete, vestigial and only feebly indicated in pronotal sculpture (Fig. 4B); tubercular (Fig. 4C); filamentary, flat and indicated by smooth, polished sculpture (Fig. 4D); carinal, elevated and sharp (Fig. 4E). Shape - straight (Fig. 4B); arcuate (Fig. 3A); bisinuate (Figs. 3BB, 4E). Extent - close to posterior pronotal angle (Fig. 3X); to third of pronotal length (Fig. 3Y); to half of pronotal length (Fig. 3Z); beyond half of pronotal length (Fig. 3AA); to anterior pronotal angle (Figs. 3BB, 4J). Modifications - with rudiment at anterior pronotal angle; obsolete in anterior part (Fig. 4K). Posterior end - joining posterior pronotal angle (Fig. 3EE); joining posterior pronotal margin (Fig. 3FF); distant from pronotal angle or margin (Fig. 3GG). Anterior end - distant from pronotal marginal carina (Figs. 4F, 4G); close to pronotal marginal carina (Fig. 4I); joining pronotal marginal carina (Fig. 4H); joining anterior pronotal angle (Fig. 4J). Arc - obvious (Fig. 4G); moderate (Fig. 3Z). Marginal and submarginal carinae Interspace - narrow (Fig. 4L); moderate (Fig. 4N); wide (Fig. 4M). Convergence - subparallel (Fig. 4N); moderately convergent (Fig. 4L); strongly convergent (Fig. 4M). Junction - absent (Figs. 4L, 4N); present (Fig. 4M). Narrowest point - at posterior angles; at posterior 1/5 1/4 of pronotal length (Fig. 4L); at posterior third of pronotal length (Fig. 4M). Modifications - submarginal carina posteriorly obliterate; submarginal carina divergent at posterior angles (Fig. 4L). Scutellum Size - moderate (Fig. 4O); rudimentary (Fig. 4P); large (Fig. 4Q). Disk - not impressed (Figs. 4O, 4Q); impressed (Fig. 4P). Scutellar carina - present (Figs. 4O, 4Q); obsolete (Fig. 4P).

Elytra Color - unicolored (Fig. 5N); unicolored with darker hue in apical half (Fig. 5O); bicolored (Fig. 5P); bicolored complementary with pubescence (Fig. 5AA). Alternative color (location) - perisutural drop (Fig. 5Q); perisutural stripe (Fig. 5P); pleural portions; transverse band before apex (Fig. 5R); apical portion (Fig. 5S). Humeral carina - absent; obsolete; present (Fig. 1C). Elytra (apical part) - markedly attenuate (Fig. 10K); feebly attenuate (Fig. 14K); spatulate (Fig. 14A). Apices Arrangement - separate (Fig. 5B); conjoint (Fig. 5A). Width - narrow (Fig. 9A); wide (Fig. 9D). Shape - arcuate (Fig. 5B); subangulate (Fig. 5C); subtruncate (Fig. 5D); cuspidate (Fig. 5E); spinose (Fig. 5F). Truncation - transverse (Fig. 5H); oblique with projecting sutural angle (Fig. 5J); oblique with projecting pleural angle (Fig. 5I).

MORPHOLOGY

17

Position of dominant cusp or spine - sutural margin (Fig. 5K); medial (Fig. 5L); pleural margin (Fig. 5M). Modifications - sutural rim spinuliform (Fig. 5G); margin obviously denticulate (Fig. 13L). Elytral pubescence Presence - absent (Fig. 5N); entire (Fig. 5T); entire except epipleura; entire except glabrous part at apical half (Fig. 5O); sutural wedge (Fig. 5U); perisutural stripes (Fig. 5V); proximal and distal (Figs. 5Y, 5Z); proximal only (Fig. 5W); distal only (Fig. 5X); ornamental (Figs. 5AA, 5BB). Proximal - humeral; quadrangular (Fig. 5W); triangular; like V or upturned T (Figs. 5Y, 5Z); perisutural stripes. Distal - apical (Fig. 5Y); preapical (Fig. 5Z); perisutural stripe (Fig. 5X). Color - unicolored; bicolored (Fig. 5BB). Character - homogenous (Figs. 5T, 5Y); with patches of denser pubescence (Fig. 5Z); with spots of denser pubescence (Fig. 5O). Elytral tomentum Presence - absent; stripes (Fig. 5DD); spots (Fig. 5CC). Spots location - humeral, located in humeral part; premedial, located before middle of elytra; postmedial, located beyond middle of elytra. Sternum Prosternum (sexual modification in male) - without obvious sexual modifications; with longer and denser pubescence (Fig. 6P); with medial row of setae (Fig. 6O). Prosternal lobe Size - narrow (Fig. 6A); moderate (Fig. 6B); large (Fig. 6C). Anterior margin - arcuate (Fig. 6A); subtruncate (Fig. 6B); arcuately emarginate (Fig. 6F); angulately emarginate (Fig. 6G). Emargination (deepness) - shallow (Fig. 6H); moderate (Fig. 6I); deep (Fig. 6J). Emargination (width) - narrow (Fig. 6K); moderate (Fig. 6F); wide (Fig. 6J). Prosternal process Size - narrow (Figs. 6N, 6X), widest part of prosternal process is subequal or narrower than width of coxa; moderate (Fig. 6L), widest part of prosternal process is slightly wider than width of coxa; wide (Fig. 6AA), widest part of prosternal process is distinctly wider than width of coxa. Shape - narrowed (Fig. 6Q); subparallel (Fig. 6R); dilated (Fig. 6S). Remark: Shape of prosternal process is defined by width ratio of prosternal process immediately before and behind procoxae. If width before and behind coxae are subequal, then the shape is subparallel, if width before coxae is wider then the shape is narrowed, if it is narrower then the shape is dilated. Sides - straight (Fig. 6V); arcuate (Fig. 6W). Angles - absent (Fig. 6X); acute (Fig. 6AA); rectangular (Fig. 6Z); obtuse (Fig. 6Y). This character is defined by the shape of angle delimited by arms regardless the shape of the tip. Angles (tips) - sharp (Fig. 6T); blunt (Fig. 6U). Disk - flat; convex; impressed; impressed, with sides protruding. Modification - sides crenulate; disk in male with two smooth glabrous protuberances; disk in male with transverse rugae. Metasternal projection - flat (Fig. 6L); impressed (Fig. 6M); with pit (Fig. 6N). Abdomen Male sexual modification - not apparent; impression at intercoxal projection; medial sulcus on ventrite 1; single medial tubercle on ventrite 1; pair of medial tubercles on ventrite 1; pair of medial tubercles delimited with sulcus on ventrite 1; transverse protrusion on ventrite 1; medial strip of erect hairs on two apical ventrites; ventrite 1 attenuate; last ventrite impressed on apex. Sternal groove Shape on apex of last ventrite - arcuate (Fig. 7A); subangulate (Fig. 7B); subtruncate (Fig. 7C); arcuately sinuate (Fig. 7G); angulately sinuate (Fig. 7H). Sinuosity (deepness) - shallow (Fig. 7D); markedly deep (Fig. 7G). Sinuosity (width) - narrow (Fig. 7F); markedly wide (Fig. 7E). Pygidium (apical margin) - arcuate (Fig. 7I); extended into short protrusion (Fig. 7J); extended into long spine (Fig. 7K). Last ventrite (apical margin) - arcuate (Fig. 7H); subtruncate (Fig. 7F); laterally emarginate (Fig. 7A). Legs Male sexual modifications - medial posterior angle of metacoxa protruding; metacoxa medially with cluster of setae; posterior margin of metacoxa with dent; lower margin of metafemur with row of hairs; metafemur markedly incrassate (Figs. 7L, 7M).

18

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Metatarsus - about as long as mesotarsus (Fig. 19A); somewhat longer than mesotarsus (Fig. 15E); distinctly longer than mesotarsus (Fig. 8H). Tarsomere 1 (relative size) - shorter or subequal to 2 3 combined (Fig. 7N); longer than 2 3 combined but shorter than 2 4 combined (Fig. 7O); subequal or longer than 2 4 combined (Fig. 7P).

Genitalia Aedeagus Symmetry - symmetrical (Fig. 7S); feebly asymmetrical (Fig. 7T); distinctly asymmetrical (Fig. 7U). Modifications - weakly sclerotised (Fig. 7V); markedly bent dorso ventrally (Fig. 7W); parameres with obvious membranous lobes (Fig. 7X); medial lobe distinctly wider than parameres (Fig. 7Y); apex of medial lobe markedly obtuse (Fig. 7Z); apex of medial lobe sharply pointed (Fig. 7AA). Ovipositor - uritiform (Fig. 7R); elongate (Fig. 7Q).

19

SUBGENERIC

CLASSIFICATION

Thirty six valid Agrilus subgenera have been proposed so far. Nineteen of them appear to pertain to the east-Asian fauna, namely: Agrilosambus Tôyama, 1987; Agrilus Curtis, 1825; Anambus Thomson, 1864; Aridagrilus Alexeev, 1998; Austragrilus Alexeev, 1998; Convexagrilus Alexeev, 1998; Dentagrilus Alexeev, 1998; Igagrilus Kurosawa, 1981; Micragrilus Alexeev, 1998; Orientagrilus Alexeev, 1998; Pseudoquercagrilus Alexeev, 1998; Quercuagrilus Alexeev, 1998; Robertius Théry, 1947; Rosagrilus Alexeev, 1998; Sinagrilus Alexeev, 1998; Sinuatiagrilus Alexeev, 1998; Spiragrilus Alexeev, 1998; Uragrilus Semenov, 1935 and Xeragrilus Alexeev, 1998. The majority of these names were coined by ALEXEEV, 1998 [#4322] based on a limited number of analyzed species, with a few exceptions, within the boundaries of the former Soviet Union. Alexeev’s subgenera were introduced without diagnostic characters through mere redescription of the type species. Any attempt to follow Alexeev’s subgeneric system is bound to leave the vast majority of known Palaearctic and Oriental Agrilus species unassigned, let alone more “exotic” faunas of other zoogeographical regions. At present we consider the existing Agrilus subgeneric system highly unsatisfactory. We firmly believe that any attempt to propose a satisfactory subgeneric classification for the hyperdiverse genus Agrilus should be based on sufficient knowledge of the world fauna, a task seemingly well outside of the present-day reach. For these reasons, the Agrilus subgeneric division is not employed in our work. Instead, informal species groups were used to cluster apparently most similar or most closely related species and, simultaneously, to avoid burdening nomenclature with seemingly meaningless subgeneric names. More on the species-groups approach might be seen below. Although arguing against using Agrilus subgeneric classification, we are compelled to presented it below in order to correct and clarify some nomenclatorial ambiguities (see Remarks) cited by BELLAMY, 2008 [#8576] in his jewel-beetle World Catalog.

Agrilus Curtis, 1825 CURTIS, 1825: No 67 [#3915] (species included: viridis, chryseis). TYPE SPECIES: Buprestis viridis Linné, 1758 fixed by original designation. Teres Harris, 1829 HARRIS, 1829: 2 3 [#1166] (subgenus of Buprestis; species included granulata, ignara, ruficollis, innuba). TYPE SPECIES: Buprestis granulata Say, 1823 fixed by present designation. Remarks: BELLAMY, 2003: 88 [#5879] and 2008: 1939 [#8576] cited Teres as an unavailable name, but Harris originally included in it four species that are presently assigned to three genera: Agrilus, Eupristocereus and Taphrocerus. The subsequent designation of type species was not found in the subsequent references of Teres. This situation might threaten the status of two valid genera Eupristocereus Deyrolle, 1864 and Taphrocerus Solier, 1833 that are younger than Teres. To avoid this situation Buprestis granulata Say, 1823 is designated herein as the type species of Teres. Paradomorphus Waterhouse, 1887 WATERHOUSE, 1887: 455 [#3053] (described as genus; species included frontalis, albicollis). TYPE SPECIES: Paradomorphus albicollis Waterhouse, 1887 fixed by subsequent designation by HESPENHEIDE, 1974: 49 [#1221]. Remarks: BELLAMY, 2008: 1939 [#8576] erroneously cited Paradomorphus frontalis as the type species of Paradomorphus. The statement of Waterhouse: “I propose this name for Agrilus frontalis, L. & G., and some allied species, which differ from Agrilus in having the basal joint of the posterior tarsi comparatively short” can not be considered as a fixation by orginal designation (Article 67.5). Samboides Kerremans, 1900 KERREMANS, 1900: 4, 16 [#1435] (described as genus; species included: viridana). TYPE SPECIES: Samboides viridana Kerremans, 1900 fixed by monotypy. Callichitones Obenberger, 1931 OBENBERGER, 1931: 181 182 [#2097] (described as genus; species included: semenovi). TYPE SPECIES: Callichitones semenovi Obenberger, 1931 fixed by original designation and monotypy. Therysambus Descarpentries & Villiers, 1967 DESCARPENTRIES & VILLIERS, 1967: 1007 1008 [#712] (described as genus; species included: apicalis). TYPE SPECIES: Sambus apicalis Bourgoin, 1923 fixed by original designation and monotypy. Wallaceilus Holynski, 2003 HOLYNSKI, 2003: 16 17 [#7129] (subgenus of Agrilus; species included: scutellaris, papua). TYPE SPECIES: Agrilus scutellaris Deyrolle, 1864 fixed by original designation.

subgenus Agrartus Curletti, 2001 CURLETTI, 2001: 3 4 [#5025] (described as subgenus of Agrilus; species included: deyrollei, rectus; characters in key). TYPE SPECIES: Agrilus deyrollei Kerremans, 1892 a replacement name for A. elongatus Deyrolle, 1864 fixed by original designation.

subgenus Agrilomorpha Théry, 1909 THÉRY, 1909: 442 443 [#2708] (described as genus; species included: rothschildi). TYPE SPECIES: Agrilomorpha rothschildi Théry, 1909 fixed by monotypy. Agrilophotus Bellamy & Westcott, 1992 BELLAMY & WESTCOTT, 1992: 126 128 [#190] (described as genus; species included: acutipennis). TYPE SPECIES: Amorphosoma acutipennis Boheman, 1851 fixed by original designation and monotypy.

subgenus Agrilosambus Tôyama, 1987 TÔYAMA, 1987: 298 299 [#2931] (described as genus; species included: taoi). TYPE SPECIES: Agrilosambus taoi Tôyama, 1987 fixed by original designation and monotypy.

subgenus Agrilus Curtis, 1825 CURTIS, 1825: No 67 [#3915] (species included: viridis, chryseis). TYPE SPECIES: Buprestis viridis Linné, 1758 fixed by original designation.

subgenus Agriphylus Curletti, 1998 CURLETTI, 1998: 95 96 [#4514] (described as subgenus of Agrilus; species included: maddalenae, schoutedeni, gestroi, limnophilus, rhopalocerus, rossi, trico). TYPE SPECIES: Agrilus (Agriphylus) maddalenae Curletti, 1998 fixed by original designation.

subgenus Anambus Thomson, 1864 THOMSON, 1864: 38 [#2898] (described as genus; species included: biguttatus). TYPE SPECIES: Buprestis biguttata Fabricius, 1776 fixed by monotypy.

subgenus Aridagrilus Alexeev, 1998 ALEXEEV, 1998: 382 383 [#4322] (described as subgenus of Agrilus; species included: niveosignatus, suturisignatus, transversesulcatus, validiusculus). TYPE SPECIES: Agrilus transversesulcatus Reitter, 1890 fixed by original designation.

subgenus Austragrilus Alexeev, 1998 ALEXEEV, 1998: 374 [#4322] (described as subgenus of Agrilus; species included: kurumi, malotiellus). TYPE SPECIES: Agrilus kurumi Kurosawa,

20

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

1957 fixed by original designation.

subgenus Nigritius Curletti, 1998

subgenus Bubagrilus Curletti, 1993

CURLETTI, 1998: 127, 131, 131 132, 133 [#4357] (described as subgenus of Agrilus; 40 species included). TYPE SPECIES: Agrilus graueri Kerremans, 1914 fixed by original designation.

CURLETTI, 1993: 437, 438 [#657] (described as subgenus of Agrilus; species included: embrikstrandellus, apamaeus, praemulus, decellei). TYPE SPECIES: Agrilus embrikstrandellus Obenberger, 1935 fixed by original designation.

subgenus Convexagrilus Alexeev, 1998 ALEXEEV, 1998: 377 [#4322] (described as subgenus of Agrilus; species included: beauprei, convexicollis). TYPE SPECIES: Agrilus convexicollis Redtenbacher, 1849 fixed by original designation.

subgenus Coroebilus Curletti, 1993 CURLETTI, 1993: 436, 437 [#657] (described as subgenus of Agrilus; species included: grandis, congolanus, ogoouensis, pangalae, pretiosissimus, colmanti). TYPE SPECIES: Agrilus grandis Gory & Laporte, 1839 fixed by original designation.

subgenus Dentagrilus Alexeev, 1998 ALEXEEV, 1998: 377 378 [#4322] (described as subgenus of Agrilus; species included: bidentulus, coeruleus [sic!], gussakovskii, pooli, rudicolllis, stepanovi). TYPE SPECIES: Buprestis caerulea Rossi, 1792 fixed by original designation.

subgenus Diplolophotus Abeille de Perrin, 1897 ABEILLE DE PERRIN, 1897: 1 2 [#19] (described as genus; species included: martini, desertus). TYPE SPECIES: Buprestis deserta Klug, 1829 fixed by subsequent designation by Bellamy, 1998: 13 [#4510]. Remarks: Subsequent designation of Bellamy from May 25, 1998 preceeded that of CURLETTI, 1998 [#4357] from August 31, 1998.

subgenus Orientagrilus Alexeev, 1998 ALEXEEV, 1998: 379 380 [#4322] (described as subgenus of Agrilus; species included: tempestivus). TYPE SPECIES: Agrilus tempestivus Lewis, 1893 fixed by original designation and monotypy.

subgenus Pantherina Curletti, 1998 CURLETTI, 1998: 127, 128, 132, 133 [#4357] (described as subgenus of Agrilus; species included: benningseni, hastulatus, nero, portonovensis, woodi). TYPE SPECIES: Agrilus benningseni 1899 Kerremans, fixed by original designation.

subgenus Paralophotus Curletti, 1998 CURLETTI, 1998: 127, 128, 130, 132 133 [#4357] (described as subgenus of Agrilus; species included: baringanus, arenatus, desertus, elgonicus, eupractus, holubiellus, olifantinus, quaestionis, sordidans, sperator). TYPE SPECIES: Agrilus baringanus Obenberger, 1931 fixed by original designation.

subgenus Personatus Curletti, 1993 CURLETTI, 1993: 436, 438 [#657] (subgenus of Agrilus; species included: sexguttatus, discolor, discoloriformis, andresi, dewetti). TYPE SPECIES: Buprestis sexguttata Thunberg, 1789 fixed by original designation.

subgenus Pinarinus Curletti, 2001 CURLETTI, 2001: 3, 35 39 [#5025] (described as subgenus of Agrilus; species included: bispinosus, woodlarkianus, fidjiensis, thylacinus, dingo). TYPE SPECIES: Agrilus bispinosus Carter, 1924 fixed by original designation.

subgenus Duttus Curletti, 1993 CURLETTI, 1993: 436 437, 438 [#657] (described as subgenus of Agrilus; species included: purpuratus, rhabdiscus, braunsi, delagoanus, polyctor, colligatus, atricapillus, euglenes, arabeon, barkeri, lepinaeyi, breyeri, lukuledianus, kituinus, raffray, elongatulus, baxii, mirei). TYPE SPECIES: Agrilus purpuratus Klug, 1829 fixed by original designation.

subgenus Pinarius Curletti, 1996 CURLETTI, 1996: 123 124 [#4144] (described as subgenus of Agrilus; species included: sellatus, coraebiformis, subgriseus, salmo, sainvali). TYPE SPECIES: Agrilus sellatus Kerremans, 1898 fixed by original designation.

subgenus Pseudoquercagrilus Alexeev, 1998 subgenus Engyaulus Waterhouse, 1889 WATERHOUSE, 1889: 50 [#3054] (described as genus; species included: pulchellus, rubrovittatus). TYPE SPECIES: Agrilus pulchellus Bland, 1865 fixed by subsequent designation by CHAMBERLIN, 1926: 200 201 [#468]. Remarks: BELLAMY, 2003 [#5879] and 2008 [#8576] erroneously cited Engyaulus rubrovittatus WATERHOUSE, 1889 as the type species of Engyaulus fixed by monotypy.

ALEXEEV, 1998: 378 379 [#4322] (described as subgenus of Agrilus; species included: asiaticus, sauteri). TYPE SPECIES: Agrilus asiaticus Kerremans, 1898 fixed by original designation.

subgenus Quercuagrilus Alexeev, 1998 ALEXEEV, 1998: 372 374 [#4322] (described as subgenus of Agrilus; 29 species included). TYPE SPECIES: Buprestis angustula Illiger, 1803 fixed by original designation.

subgenus Igagrilus Kurosawa, 1981 KUROSAWA, 1981: 5 6 [#1637] (described as subgenus of Agrilus; species included: elongatissimus). TYPE SPECIES: Agrilus (Igagrilus) elongatissimus Kurosawa, 1981 fixed by original designation and monotypy.

subgenus Liliput Curletti, 1998 CURLETTI, 1998: 90 91 [#4514] (described as subgenus of Agrilus; species included: paracuspidatus, pseudolituratus, canus, sangadzinus, minimus, forschhammeri; characters in key). TYPE SPECIES: Agrilus paracuspidatus Obenberger, 1939 fixed by original designation.

subgenus Melagrilus Curletti, 1993 CURLETTI, 1993: 437, 438 [#657] (described as subgenus of Agrilus; species included: escalerai, obdurescens, vincentae, notatus, schultzei, pujoli). TYPE SPECIES: Agrilus escalerai Obenberger, 1921 fixed by original designation.

subgenus Micragrilus Alexeev, 1998 ALEXEEV, 1998: 374 375 [#4322] (described as subgenus of Agrilus; species included: ganglbaueri, lituratus). TYPE SPECIES: Agrilus ganglbaueri Semenov, 1891 fixed by original designation.

subgenus Robertius Théry, 1947 THÉRY, 1947: 670 671 [#2891] (described as subgenus of Agrilus; species included: robertii, taylori). TYPE SPECIES: Agrilus robertii Chevrolat, 1840 fixed by original designation. Arquagrilus Alexeev, 1998 ALEXEEV, 1998: 369 370 [#4322] (described as subgenus of Agrilus; 31 species included). TYPE SPECIES: Buprestis pratensis Ratzeburg, 1837 fixed by original designation.

subgenus Rosagrilus Alexeev, 1998 ALEXEEV, 1998: 375 377 [#4322] (described as subgenus of Agrilus; species included: convexifrons, marozzinii, pistaciophagus, roscidus, viridicoerulans). TYPE SPECIES: Agrilus roscidus Kiesenwetter, 1857 fixed by original designation.

subgenus Sinagrilus Alexeev, 1998 ALEXEEV, 1998: 379 [#4322] (described as subgenus of Agrilus; species included: coreanus, sinensis). TYPE SPECIES: Agrilus sinensis Thomson, 1879 fixed by original designation.

subgenus Sinuatiagrilus Alexeev, 1998 ALEXEEV, 1998: 372 [#4322] (described as subgenus of Agrilus; species

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SUBGENERIC CLASSIFICATION included: mali, mendax, sachalinensis, sinuatus, zhelochovtsevi). TYPE SPECIES: Buprestis sinuata Olivier, 1790 fixed by original designation.

subgenus Spiragrilus Alexeev, 1998 ALEXEEV, 1998: 381 382 [#4322] (described as subgenus of Agrilus; species included: antiquus, cinctus, constantini, hyperici, hypericicola, lopatini, macroderus, pseudolimoniastri, solieri, sulcifer, unguiculosus, istjurti, vaginalis). TYPE SPECIES: Agrilus antiquus Mulsant & Rey, 1863 fixed by original designation.

subgenus Uragrilus Semenov, 1935 SEMENOV, 1935: 276 277 [#2623] (described as subgenus of Agrilus; species included: guerini). TYPE SPECIES: Agrilus guerini Lacordaire, 1835 fixed by original designation and monotypy.

subgenus Xenagrilus Obenberger, 1924 OBENBERGER, 1924: 35 36 [#2035] (described as genus; species included: binderi). TYPE SPECIES: Xenagrilus binderi Obenberger, 1924 fixed by original designation and monotypy. Agrilacanthus Théry, 1940 THÉRY, 1940: 160 162 [#2865] (described as genus; species included: lloydi). TYPE SPECIES: Agrilacanthus lloydi Théry, 1940 fixed by original designation and monotypy.

subgenus Xeragrilus Alexeev, 1998 ALEXEEV, 1998: 375 [#4322] (described as subgenus of Agrilus; species included: aerosus, albogularis, aridosus, ecarinatus, mongolorum, plavilstshikovi, sericans, zigzag). TYPE SPECIES: Agrilus sericans Kiesenwetter, 1857 fixed by original designation.

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SPECIES

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

GROUPS

The enormous Agrilus species diversity plainly suggests the high plasticity and variability among morphological characters and, therefore, only few morphological features are discrete and distinctive enough to be used for species identification. Furthermore, it is plausible to assume that similar features might often develop independently in unrelated Agrilus species resulting in relative uniformity and sameness among distantly related species. Due to these factors Agrilus species-level identification is a highly challenging task, requiring sufficient experience and availability of adequately identified voucher specimens. Originally we intended to provide a key to species, however when attempting to do so it was found that it will not be a proper and useful tool due to the difficulties mentioned above. Instead of driving a reader through hundreds of preset and difficult-to-decide choices, we offer an alternative Agrilus species identification strategy by means of informal species-groups supplemented with the character state matrix permitting usage of reader-chosen random combination of most noticeable characters (as opposed to a rigid key). The proposed 34 species-groups represent the first attempt to cluster east-Asian Agrilus diversity in smaller and manageable groups. Despite all our efforts to make these groupings as meaningful as we could, this attempt does not claim to be phylogenetic. Our main goal was purely practical, i.e. to allow species-level identification of east-Asian Agrilus. Some species groups were established through the presence of a highly distinctive combination of characters and therefore are likely non-monophyletic. Others might be natural, i.e. monophyletic, and may later serve as nuclei of future subgenera. We spend no time or effort in this work trying to argue the phylogenetic value of the employed characters, and this is because any phylogenetic attempt is far beyond the scope and potential of our present work. Our knowledge of Agrilus taxonomy, either of the target area of East Asia or on the global level, is still too unsatisfactory to permit such an attempt. Only within the realms of the Oriental region, hundreds of Agrilus await their discovery, while the majority of already described species are in acute need of revision. Any future serious attempt to phylogenetically define eastAsian Agrilus species-grouping has to be based on, at the very least, adequate alpha-taxonomic knowledge of both Palaearctic and Oriental regions, and, therefore, is presently postponed. For more on this subject see also discussion on the existing subgeneric Agrilus classification. Guide to the species determination. To arrive to the most plausible species identification for an Agrilus specimen originating from East Asia the following strategy should be followed. Reader should select the most distinct or striking feature on the specimen, or their combination, and refer to the table of species-group characters (Appendix V) to find a species-group satisfying chosen criteria. The process should be repeated until, optimally, a single species-group remains. Then reader should refer to the text with description and diagnosis for the chosen species-group to see whether it fits

the specimen at hand. Relevant (related or similar) speciesgroups are mentioned in the description and they should be referred to if in doubt. For each species-group, the included species and known host plants are given to facilitate further identification. Species of each species-group are illustrated together to simplify further taxa elimination. In case of uncertainty, the user should refer to the diagnostic characters provided for each species. If the species with the matching set of characters is not found, the unassigned taxa should be consulted. Although the majority of east-Asian Agrilus species are distinctive enough to be reliably identified by habitus and aedeagus, the existence of a few less straightforward taxa might make their determination more demanding. EastAsian species not found in this book should be assumed as either new for the region or new to science. Acutus species-group Description: BODY: Size: medium, Shape: cuneiform, Build: medium, Color (dorsally): unicolored. EYES: Size: moderate. ANTENNAE: Length: short or moderate, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: transverse, Maximal width: at middle or at posterior margin, Anterior lobe: obvious, Disk: moderately convex, Impressions: medial and lateral, Prehumerus: carinal, Prehumerus shape: bisinuate, Prehumerus extent: beyond half of pronotal length, Prehumerus - anterior end: joining pronotal marginal carina, rarely close to pronotal marginal carina, Marginal and submarginal carinae - interspace: moderate, Marginal and submarginal carinae - convergence: subparallel, Marginal and submarginal carinae - junction: absent, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: spinose, Pubescence: proximal and/or distal, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Elytral apex with spine on pleural side; pronotum with mediobasal impression; prehumerus carinal and long; elytra with proximal and distal pubescence; aedeagus strongly sclerotised. Relevant group(s): Plasoni. Taxa: A. dianthus Kerremans, 1892. Remark. This mostly Oriental species-group with only a single east-Asian species, has been recently revised by JENDEK, 2004 [#6675]. Host plants: Malvaceae: Abelmoschus; Corchorus; Hibiscus; Urena. See remarks under A. dianthus Kerremans, 1892. Albogularis species-group Description: BODY: Size: small or medium, Shape: subparallel, Build: slender, medium, robust, strongly convex, Color (dorsally): unicolored. EYES: Size: small. ANTENNAE: Length: short, moderate or long, Shape:

23

SPECIES GROUPS slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square or transverse, Maximal width: at anterior margin or at middle, Anterior lobe: absent or moderate, Disk: flat, moderately convex or strongly convex, Impressions: absent, medial only, lateral only or medial and lateral, Prehumerus: absent, obsolete or filamentary, Prehumerus shape: arcuate or bisinuate, Prehumerus extent: close to posterior pronotal angle, to half of pronotal length, beyond half of pronotal length or to anterior pronotal angle, Prehumerus - anterior end: close to pronotal marginal carina, joining pronotal marginal carina or joining anterior pronotal angle, Marginal and submarginal carinae - interspace: moderate or wide, Marginal and submarginal carinae convergence: subparallel, Marginal and submarginal carinae - junction: absent, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices arrangement: separate, Apices - shape: arcuate, Pubescence: entire or on most of elytra, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Aedeagus: rarely markedly dorsoventrally bent, Ovipositor: elongate. Diagnosis: Body strongly convex; eyes small, rarely medium; antennae slender; marginal and submarginal carinae subparallel; ovipositor elongate; elytral pubescence entire, often with denser perisutural stripes. Relevant group(s): Hyperici, Pekinensis. Taxa: A. aerosus Ganglbauer, 1889; A. daur sp. nov.; A. ecarinatus Marseul, 1866; A. macrotatopus Obenberger, 1930; A. mongolorum Obenberger, 1936; A. morawitzi Obenberger, 1936; A. nivosus Abeille de Perrin, 1900; A. plavilstshikovi Alexeev, 1989; A. pleuralis sp. nov.; A. przewalskii Obenberger, 1936; A. pseudosericans Obenberger, 1936 Host plants: Asteraceae: Ajania; Artemisia; Chenopodiaceae: Eurotia; Nitrariaceae: Nitraria. Angulatus species-group Description: BODY: Size: medium, Shape: subparallel, Build: slender, medium or strongly convex, Color (dorsally): unicolored. EYES: Size: small. ANTENNAE: Length: moderate or long, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square, Maximal width: at anterior margin or at middle, Anterior lobe: moderate or obvious, Disk: strongly convex, Impressions: lateral only, rarely medial and lateral, Prehumerus: absent, Marginal and submarginal carinae - interspace: moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: present, Marginal and submarginal carinae - narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate, Pubescence: ornamental, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: sinuate, Pygidium: arcuate. GENITALIA: Ovipositor: uritiform. Diagnosis: Medium or large species; elytra with ornamental

pubescence; elytral apices arcuate; prehumerus absent; pygidium arcuate. Relevant group(s): Occipitalis, Muscarius. Taxa: A. grisator Kerremans, 1893; A. olivaceidorsis Obenberger, 1917 Host plants: Rutaceae: Citrus. Barrati species-group Description: BODY: Size: small or medium, Shape: subparallel, Build: robust, Color (dorsally): unicolored. EYES: Size: small or moderate. ANTENNAE: Length: short or moderate, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: transverse, Maximal width: at anterior margin or at middle, Anterior lobe: obvious, Disk: strongly convex, Impressions: lateral only or posterolateral, Prehumerus: absent or carinal, Prehumerus shape: arcuate, Prehumerus extent: beyond half of pronotal length, Prehumerus - anterior end: distant from pronotal marginal carina, Marginal and submarginal carinae - interspace: moderate, Marginal and submarginal carinae - convergence: convergent or subparallel, Marginal and submarginal carinae - junction: absent, Marginal and submarginal carinae narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate, Pubescence: pairs of pubescent-tomentose spots, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Aedeagus: parameres rarely with obvious membranous lobes, Ovipositor: uritiform. Diagnosis: Small to medium sized, robust species; pronotal lobe prominent, extending conspicuously beyond anterior pronotal angles; pronotal disk without medial impression, strongly convex antero-medially. Relevant group(s): Cyaneovirens. Taxa: A. barrati Descarpentries & Villiers, 1963; A. holzschuhi Jendek, 1994. Host plants: Uknown. Betuleti species-group Description: BODY: Size: small or medium, Shape: subparallel or cuneiform, Build: slender, medium, robust, rarely strongly convex, Color (dorsally): unicolored, rarely bicolored. EYES: Size: small or moderate. ANTENNAE: Length: short or moderate, Shape: slender, rarely stout, Antennomeres in male: not modified. PRONOTUM: Shape: visually square or transverse, Maximal width: at anterior margin or at middle, Anterior lobe: moderate, Disk: flat or moderately convex, Impressions: medial and lateral, Prehumerus: carinal, rarely absent, obsolete or tubercular, Prehumerus shape: straight or arcuate, Prehumerus extent: close to posterior pronotal angle or to half of pronotal length, Prehumerus - anterior end: distant from pronotal marginal carina, Marginal and submarginal carinae - interspace: narrow or moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent, Marginal and submarginal carinae - narrowest point: distant from posterior angles, rarely at posterior angles,

24 Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, rarely conjoint, Apices - shape: arcuate or subtruncate, Pubescence: absent. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications or with sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Body glabrous; pronotum with deep lateral impressions; ovipositor elongate. Relevant group(s): Viridis, Smaragdinus. Taxa: A. alutaceicollis Obenberger, 1930; A. betuleti (Ratzeburg, 1837); A. cupes Lewis, 1893; A. delphinensis Abeille de Perrin, 1897; A. fareastensis Jendek, 1995; A. jindrai sp. nov.; A. mio sp. nov.; A. nicolanus Obenberger, 1924; A. ogatai Ohmomo, 2002; A. pratensis (Ratzeburg, 1837); A. sibiricus Obenberger, 1912; A. sibiricus fukushimenis Jendek, 1994; A. smetanai sp. nov.; A. soudeki Obenberger, 1926; A. voriseki Jendek, 1995. Host plants: Betulaceae: Betula; Fagaceae: Quercus; Salicaceae: Populus; Salix; Sapindaceae: Acer; Staphyleaceae: Staphylea. Biguttatus species-group Description: BODY: Size: large, Shape: subparallel or cuneiform, Build: robust, rarely strongly convex, Color (dorsally): unicolored. EYES: Size: small, moderate or large. ANTENNAE: Length: moderate, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square or transverse, Maximal width: at middle or at posterior margin, Anterior lobe: absent, moderate or obvious, Disk: moderately convex, Impressions: medial and lateral or lateral only, Prehumerus: absent, obsolete, tubercular or carinal, Prehumerus shape: straight or arcuate, Prehumerus extent: close to posterior pronotal angle, Prehumerus anterior end: distant from pronotal marginal carina, Marginal and submarginal carinae - interspace: moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent or present, Marginal and submarginal carinae - narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored or bicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate, Pubescence: absent or pairs of pubescent-tomentose spots, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate or sinuate, Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Large species; prehumerus absent, tubercular or short carinal; elytra glabrous or with pairs of pubescent spots; elytral apices separately arcuate. Relevant group(s): Spinipennis, Niveoguttatus, Quadrisignatus. Taxa: A. decoloratus Kerremans, 1892; A. magnus sp. nov.; A. proditor Obenberger, 1917; A. ventricosus Fairmaire, 1888. Host plants: Cannabaceae: Celtis; Fagaceae: Quercus.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Cyaneoniger species-group Description: BODY: Size: large, Shape: cuneiform, Build: medium, robust, rarely strongly convex, Color (dorsally): unicolored. EYES: Size: moderate or large. ANTENNAE: Length: moderate or long, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square or transverse, Maximal width: at anterior margin, at middle or at posterior margin, Anterior lobe: absent or moderate, Disk: flat or moderately convex, Impressions: medial and lateral, Prehumerus: absent, obsolete, tubercular or filamentary, Prehumerus shape: straight, Prehumerus extent: beyond half of pronotal length or to anterior pronotal angle, Prehumerus - anterior end: close to pronotal marginal carina, Marginal and submarginal carinae - interspace: narrow or moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent or present, Marginal and submarginal carinae - narrowest point: distant from posterior angles, Scutellum: rudimentary. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate, Pubescence: absent, rarely pairs of pubescent-tomentose spots, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications or with sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate or sinuate, Pygidium: arcuate, extended into short protrusion or extended into long spine. GENITALIA: Ovipositor: elongate. Diagnosis: Very large species; pronotum with deep medial pronotal impression; scutellum rudimentary; elytra glabrous, rarely with sutural apical or preapical spot; elytral apices separately arcuate. Relevant group(s): Sinensis. Taxa: A. agnatus Kerremans, 1892; A. auristernum Obenberger, 1924; A. cyaneoniger Saunders, 1873; A. lubopetri Jendek, 2000; A. planipennis Fairmaire, 1888; A. qinling Jendek, 2000. Host plants: Fagaceae: Quercus; Juglandaceae: Juglans; Pterocarya; Oleaceae: Fraxinus; Ulmaceae: Ulmus. Cyaneovirens species-group Description: BODY: Size: small or medium, Shape: subparallel or cuneiform, Build: slender, medium, robust, rarely strongly convex, Color (dorsally): unicolored, bicolored, rarely tricolored. EYES: Size: moderate or large. ANTENNAE: Length: moderate, long, rarely short, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square, transverse, rarely visually elongate, Maximal width: at middle, rarely at anterior margin, Anterior lobe: moderate or obvious, Disk: moderately convex, strongly convex, rarely flat, Impressions: medial and lateral, rarely lateral only or posterolateral only, Prehumerus: carinal, Prehumerus shape: arcuate or bisinuate, Prehumerus extent: to half of pronotal length, beyond half of pronotal length or to anterior pronotal angle, Prehumerus - anterior end: distant from pronotal marginal carina, close to pronotal marginal carina, joining pronotal marginal carina or joining anterior pronotal angle, Marginal and submarginal carinae - interspace: moderate, Marginal and submarginal carinae convergence: convergent, Marginal and submarginal carinae

25

SPECIES GROUPS - junction: absent, rarely present, Marginal and submarginal carinae - narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored or bicolored, Humeral carina: absent, Apices - arrangement: separate or conjoint, Apices - shape: arcuate or subtruncate, Pubescence: proximal and/or distal, rarely entire or on most of elytra, Pubescence - color: unicolored or bicolored. STERNUM: Metasternal projection: flat or impressed. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications or with sexual modifications, Groove on apex of last ventrite: sinuate, Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Small or medium species, eyes large or moderate; prehumerus carinal, extending at least to half of pronotal length; prosternal process dilated; groove on apex of last ventrite distinctly sinuate. Relevant group(s): Barrati. Taxa: A. bai sp. nov.; A. bonnottei Bourgoin, 1925; A. brevinotus sp. nov.; A. businskorum sp. nov.; A. cyaneovirens Bourgoin, 1922; A. euonymi Tôyama, 1985; A. gussakovskiji Alexeev, 1981; A. hani sp. nov.; A. hui sp. nov.; A. kucerai sp. nov.; A. lahu sp. nov.; A. manchu sp. nov.; A. naxi sp. nov.; A. pterostigma Obenberger, 1927; A. qinlingcola sp. nov.; A. rougeoti Descarpentries & Villiers, 1963; A. sagittifer sp. nov.; A. schillhammeri sp. nov.; A. subtilipilis Obenberger, 1940; A. terracotta sp. nov.; A. tsushimanus Kurosawa, 1963; A. yamabusi Miwa & Chûjô, 1940. Host plants: Celastraceae: Euonymus; Fagaceae: Quercus; Juglandaceae: Juglans; Rosaceae: Prunus; Rutaceae: Chloroxylon; Ulmaceae: Ulmus. Cyanescens species-group Description: BODY: Size: medium, Shape: subparallel, Build: robust, strongly convex, Color (dorsally): unicolored. EYES: Size: small. ANTENNAE: Length: short or moderate, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: transverse, Maximal width: at middle, rarely at anterior margin, Anterior lobe: absent or moderate, Disk: moderately convex, Impressions: lateral only, rarely medial and lateral, Prehumerus: absent or tubercular, Prehumerus extent: to half of pronotal length, Marginal and submarginal carinae - interspace: moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent or present, Marginal and submarginal carinae - narrowest point: at posterior angles or distant from posterior angles, Scutellum: well developed, rarely rudimentary. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate, Pubescence: absent, Pubescence color: unicolored. STERNUM: Metasternal projection: flat or impressed. ABDOMEN: Basal ventrite or metacoxae in male: with sexual modifications, Groove on apex of last ventrite: sinuate, Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Groove on apex of last ventrite deeply sinuate; body large and convex; elytra blue, green or bronze, glabrous; aedeagus slender and convex. Relevant group(s): Laetecyanescens. Taxa: A. arsenevi Jendek, 2009; A. asahinai Kurosawa, 1956; A. bidentulus

Ganglbauer, 1889; A. cyanescens (Ratzeburg, 1837); A. dali Jendek, 2009; A. thibetanus Obenberger, 1928; A. zhongdian Jendek, 2009 Host plants: Caprifoliaceae: Lonicera; Symphoricarpos; Rhamnaceae: Rhamnus. Densetomentosus species-group Description: BODY: Size: small, Shape: fusiform, Build: robust, strongly convex, Color (dorsally): unicolored. EYES: Size: moderate or large. ANTENNAE: Length: short, moderate, rarely long, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square or transverse, Maximal width: at posterior margin, rarely at middle, Anterior lobe: moderate or obvious, Disk: moderately convex or strongly convex, Impressions: medial only or medial and lateral, Prehumerus: carinal, Prehumerus shape: bisinuate, Prehumerus extent: beyond half of pronotal length or to anterior pronotal angle, Prehumerus - anterior end: joining pronotal marginal carina or joining anterior pronotal angle, Marginal and submarginal carinae - interspace: wide, rarely moderate, Marginal and submarginal carinae convergence: convergent, Marginal and submarginal carinae - junction: present, Marginal and submarginal carinae narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, obsolete or present, Apices - arrangement: conjoint, Apices - shape: arcuate, Pubescence: entire or on most of elytra or proximal and/or distal, Pubescence - color: unicolored. STERNUM: Metasternal projection: impressed or with pit. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications or with sexual modifications, Groove on apex of last ventrite: sinuate, Pygidium: arcuate. GENITALIA: Ovipositor: uritiform. Diagnosis: Body small, fusiform, robust; head strongly convex; pronotum widest at basal half; distinctly tapering anteriorly; prehumerus extends to anterior pronotal angles; mesosternal projection impressed or with pit. Relevant group(s): Viridiobscurus, Japanocarinatus. Taxa: A. densetomentosus Obenberger, 1940; A. fusiformis sp. nov.; A. uenoi Kurosawa, 1963. Host plants: Euphorbiaceae: Mallotus. Hyperici species-group Description: BODY: Size: small or medium, Shape: subparallel, Build: medium, robust, strongly convex, Color (dorsally): unicolored. EYES: Size: small. ANTENNAE: Length: short or moderate, Shape: stout, Antennomeres in male: not modified. PRONOTUM: Shape: visually square or transverse, Maximal width: at anterior margin or at middle, Anterior lobe: absent, rarely moderate, Disk: moderately convex or strongly convex, Impressions: medial and lateral, Prehumerus: obsolete, tubercular or carinal, Prehumerus shape: arcuate, Prehumerus extent: to half of pronotal length, Prehumerus - anterior end: distant from pronotal marginal carina or close to pronotal marginal carina, Marginal and submarginal carinae - interspace: moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent, rarely present, Marginal and submarginal carinae - narrowest point: distant

26 from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices arrangement: separate, Apices - shape: arcuate, Pubescence: entire or on most of elytra, sutural wedge or perisutural stripes, rarely absent, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Aedeagus: rarely weakly sclerotised or parameres with obvious membranous lobes, Ovipositor: uritiform. Diagnosis: Body strongly convex; eyes small, rarely medium; antennae stout and short; marginal and submarginal carinae convergent; ovipositor uritiform; elytral pubescence entire, often with denser perisutural stripes. Relevant group(s): Albogularis, Pekinensis. Taxa: A. lugu sp. nov.; A. myopic Jendek, 2000; A. pseudolimoniastri Cobos, 1968; A. unguiculosus Obenberger, 1936. Remark: Only some species of this Palaearctic species-group are occurring East Asia. Hosts plants: Fabaceae: Caragana; ?Rosaceae: Spirea. Imitans species-group Description: BODY: Size: large, rarely medium, Shape: cuneiform, Build: slender or medium, Color (dorsally): bicolored. EYES: Size: moderate, rarely small. ANTENNAE: Length: moderate or long, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square, transverse, rarely visually elongate, Maximal width: at anterior margin, Anterior lobe: moderate or obvious, Disk: moderately convex, Impressions: medial and lateral, rarely lateral only, Prehumerus: filamentary or carinal, Prehumerus shape: bisinuate, Prehumerus extent: to anterior pronotal angle, Prehumerus - anterior end: joining pronotal marginal carina or joining anterior pronotal angle, Marginal and submarginal carinae - interspace: narrow, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent or present, Marginal and submarginal carinae - narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate, rarely subangulate or subtruncate, Pubescence: entire or on most of elytra or sutural wedge or perisutural stripes, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications or with sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Body cuneiform, dorsally bicolored; pronotum widest at anterior margin, golden-yellow or golden-orange; prehumerus long, bisinuate, extending to anterior pronotal angles; marginal and submarginal carinae with narrow interspace; elytra blackish or greenish. Relevant group(s): Plagiatus; Sinuatus. Taxa: A. blatteicollis Bourgoin, 1922; A. gaoligong Jendek, 2000; A. imitans Lewis, 1893; A. nepalensis Tôyama, 1988; A. rivalieri Descarpentries & Villiers, 1963; A. sichuanus sp.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA nov.; A. sikkimensis Obenberger, 1928; A. smaragdifrons Ganglbauer, 1889. Host plants: Cannabaceae: Celtis; Ulmaceae: Ulmus; Zelkova. Japanocarinatus species-group Description: BODY: Size: small or medium, Shape: cuneiform or fusiform, Build: slender or medium, Color (dorsally): unicolored. EYES: Size: moderate. ANTENNAE: Length: moderate or long, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually elongate or visually square, Maximal width: at middle or at posterior margin, Anterior lobe: obvious, Disk: moderately convex, Impressions: medial and lateral or posterolateral, Prehumerus: carinal, Prehumerus shape: arcuate or bisinuate, Prehumerus extent: to half of pronotal length, beyond half of pronotal length or to anterior pronotal angle, Prehumerus - anterior end: distant from pronotal marginal carina or joining pronotal marginal carina, Marginal and submarginal carinae - interspace: narrow or moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: present, Marginal and submarginal carinae - narrowest point: at posterior angles or distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: present, Apices - arrangement: conjoint, Apices - shape: arcuate or subtruncate, Pubescence: entire or on most of elytra or proximal and/or distal, Pubescence - color: unicolored. STERNUM: Metasternal projection: impressed or with pit. ABDOMEN: Basal ventrite or metacoxae in male: with sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Elytra with humeral carina; mesosternal projection impressed or with pit; last ventrite arcuate. Relevant group(s): Densetomentosus, Viridiobscurus. Taxa: A. bocaki sp. nov.; A. japanocarinatus Ohmomo, 2002; A. lei sp. nov. Host plants: Oleaceae: Fraxinus. Laetecyanescens species-group Description: BODY: Size: medium, Shape: cuneiform, Build: medium, Color (dorsally): unicolored or bicolored. EYES: Size: small, moderate, rarely large. ANTENNAE: Length: moderate, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square or transverse, Maximal width: at middle, Anterior lobe: moderate or obvious, Disk: moderately convex or strongly convex, Impressions: medial and lateral or lateral only, Prehumerus: tubercular or carinal, Prehumerus shape: arcuate, Prehumerus extent: to half of pronotal length, Prehumerus anterior end: distant from pronotal marginal carina or close to pronotal marginal carina, Marginal and submarginal carinae - interspace: moderate, Marginal and submarginal carinae convergence: convergent, Marginal and submarginal carinae - junction: absent or present, Marginal and submarginal carinae - narrowest point: at posterior angles or distant from posterior angles, Scutellum: well developed. ELYTRA: Color:

27

SPECIES GROUPS unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate, rarely cuspidate or spinose, Pubescence: absent or proximal and/or distal, Pubescence color: unicolored. STERNUM: Metasternal projection: flat, rarely impressed. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: sinuate, Pygidium: arcuate or extended into long spine. GENITALIA: Ovipositor: elongate. Diagnosis: Groove on apex of last ventrite deeply sinuate; body large and convex; elytra blue, green or bronze, glabrous or with preapical perisutural stripe of white pubescence; aedeagus wide and flattened. Relevant group(s): Cyanescens. Taxa: A. aurithorax sp. nov.; A. laetecyanescens Obenberger, 1940; A. paralleloides sp. nov.; A. vigil sp. nov.; A. yunnanus Obenberger, 1927. Host plants: Unknown. Maculifer species-group Description: BODY: Size: small or medium, Shape: cuneiform, Build: medium, Color (dorsally): unicolored. EYES: Size: small or moderate. ANTENNAE: Length: short, moderate or long, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: transverse, Maximal width: at middle, Anterior lobe: moderate or obvious, Disk: flat or moderately convex, Impressions: medial and lateral or posterolateral, Prehumerus: filamentary, Prehumerus shape: bisinuate, Prehumerus extent: to anterior pronotal angle, Prehumerus - anterior end: joining pronotal marginal carina or joining anterior pronotal angle, Marginal and submarginal carinae - interspace: narrow, rarely moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent, rarely present, Marginal and submarginal carinae - narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices arrangement: separate, Apices - shape: arcuate or subtruncate, Pubescence: proximal and/or distal, ornamental or pairs of pubescent-tomentose spots, Pubescence - color: unicolored, rarely bicolored. STERNUM: Metasternal projection: flat, rarely impressed. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Medium or small species; eyes small or moderate; prehumerus filamentary, long, bisinuate, extending to anterior pronotal angles. Relevant group(s): Quadrisignatus. Taxa: A. daimio Obenberger, 1936; A. imasakai Tôyama, 1985; A. maculifer Saunders, 1873; A. nelsonodaimio Ohmomo, 2006; A. pleurostigmus sp. nov.; A. shashamboe Kurosawa, 1963; A. sudai Kurosawa, 1985. Host plants: Betulaceae: Alnus; Carpinus; Celastraceae: Euonymus; Ericaceae: Rhododendron; Symplocos; Vaccinium; Lauraceae: Parabenzoin; Rosaceae: Rhaphiolepis. Muscarius species-group Description: BODY: Size: small, Shape: fusiform, Build: medium or robust, Color (dorsally): unicolored or bicolored.

EYES: Size: small. ANTENNAE: Length: short, rarely moderate, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: transverse, Maximal width: at posterior margin, Anterior lobe: moderate or obvious, Disk: moderately convex or strongly convex, Impressions: absent or posterolateral, Prehumerus: carinal, Prehumerus shape: arcuate, rarely bisinuate, Prehumerus extent: to half of pronotal length or beyond half of pronotal length, Prehumerus - anterior end: distant from pronotal marginal carina, Marginal and submarginal carinae - interspace: moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: present, Marginal and submarginal carinae - narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored or bicolored, Humeral carina: absent, Apices - arrangement: separate, rarely conjoint, Apices - shape: arcuate, Pubescence: ornamental, Pubescence - color: unicolored or bicolored. STERNUM: Metasternal projection: flat, rarely impressed. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: sinuate, Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Small, fusiform species; elytra with ornamental pubescence. Relevant group(s): Angulatus, Occipitalis. Taxa: A. hunanus Jendek, 2009; A. kurumi Kurosawa, 1957; A. liscapia Jendek, 2003; A. mallotiellus Kurosawa, 1985; A. muscarius Kerremans, 1895. Remark: This Palaearctic and Oriental species-group was revised by JENDEK & GREBENNIKOV, 2009 [#8563]. Host plants: Euphorbiaceae: Mallotus; Fabaceae: Indigofera; Juglandaceae: Juglans. Niveoguttatus species-group Description: BODY: Size: medium, Shape: cuneiform, Build: slender or medium, Color (dorsally): unicolored. EYES: Size: large. ANTENNAE: Length: moderate or long, Shape: slender. PRONOTUM: Shape: visually elongate, visually square or transverse, Maximal width: at anterior margin, rarely at middle, Anterior lobe: moderate or obvious, Disk: flat or moderately convex, Impressions: medial and lateral, Prehumerus: absent, obsolete, rarely filamentary, Prehumerus shape: bisinuate, Prehumerus extent: beyond half of pronotal length or to anterior pronotal angle, Prehumerus - anterior end: joining pronotal marginal carina or joining anterior pronotal angle, Marginal and submarginal carinae - interspace: narrow or moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent, Marginal and submarginal carinae - narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: cuspidate or spinose, Pubescence: pairs of pubescent-tomentose spots. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: with sexual modifications, Groove on apex of last ventrite: sinuate, Pygidium: extended into long spine. GENITALIA: Aedeagus: rarely markedly dorsoventrally bent, Ovipositor: elongate.

28 Diagnosis: Medium sized species; pronotum enlarged anteriorly; prehumerus absent or obsoletely filamentary; elytra with 2-4 pair of tomentose spots; elytral apices cuspidate or spinose; pygidium with long spine; groove on apex of last ventrite sinuate. Relevant group(s): Spinipennis, Biguttatus, Quadrisignatus. Taxa: A. auropictus Kerremans, 1912; A. jaminae Baudon, 1968; A. madeci Baudon, 1968; A. niveoguttatus Kerremans, 1892; A. satoi Kurosawa, 1954. Host plants: Fagaceae: Quercus. Occipitalis species-group Description: BODY: Size: medium or large, Shape: cuneiform, Build: medium or robust, Color (dorsally): unicolored. EYES: Size: moderate, rarely small. ANTENNAE: Length: moderate or long, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: transverse, Maximal width: at middle, rarely at anterior margin, Anterior lobe: absent, moderate or obvious, Disk: moderately convex, rarely strongly convex, Impressions: medial and lateral, Prehumerus: carinal, rarely tubercular or filamentary, Prehumerus shape: arcuate, Prehumerus extent: to half of pronotal length, Prehumerus - anterior end: distant from pronotal marginal carina, rarely close to pronotal marginal carina or joining pronotal marginal carina, Marginal and submarginal carinae - interspace: moderate, rarely narrow, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent or present, Marginal and submarginal carinae narrowest point: at posterior angles or distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate, Pubescence: ornamental, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: extended into short protrusion or extended into long spine. GENITALIA: Ovipositor: elongate. Diagnosis: Medium or large species; elytra with ornamental pubescence; elytral apices arcuate; prehumerus present; pygidium with short protrusion or long spine Relevant group(s): Angulatus, Muscarius. Taxa: A. alesi Obenberger, 1935; A. auriventris Saunders, 1873; A. inamoenus Kerremans, 1892; A. perroti Descarpentries & Villiers, 1963; A. tonkineus Kerremans, 1895; A. yamawakii Kurosawa, 1957; A. zanthoxylumi Li Meng Lou, 1989. Remark: This species-group is distributed in Palaearctic and Oriental regions. Some species are known as serious pest of Citrus and Zanthoxylum. Host plants: Aurantioideae: Citrus; Rutaceae: Fagara; Zanthoxylum. Pekinensis species-group Description: BODY: Size: medium, Shape: subparallel or cuneiform, Build: medium, robust, often strongly convex, Color (dorsally): unicolored. EYES: Size: small or moderate. ANTENNAE: Length: moderate or long, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape:

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA visually square or transverse, Maximal width: at anterior margin, at middle, rarely at posterior margin, Anterior lobe: moderate, rarely absent or obvious, Disk: flat, moderately convex or strongly convex, Impressions: medial and lateral or lateral only, Prehumerus: absent, filamentary or carinal, Prehumerus shape: arcuate or bisinuate, Prehumerus extent: to half of pronotal length, beyond half of pronotal length or to anterior pronotal angle, Prehumerus - anterior end: distant from pronotal marginal carina, close to pronotal marginal carina or joining anterior pronotal angle, Marginal and submarginal carinae - interspace: narrow or moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent or present, Marginal and submarginal carinae - narrowest point: at posterior angles or distant from posterior angles, Scutellum: well developed, rarely rudimentary. ELYTRA: Color: unicolored, Humeral carina: absent, Apices arrangement: separate, Apices - shape: arcuate, Pubescence: absent, entire or on most of elytra, rarely proximal and/or distal, Pubescence - color: unicolored. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Ovipositor: uritiform or elongate. Diagnosis: Medium sized; eyes small or moderate; antennae slender; marginal and submarginal carinae convergent. Relevant group(s): Hyperici, Albogularis. Taxa: A. alashanensis Obenberger, 1936; A. mandala sp. nov.; A. pekinensis Obenberger, 1924; A. pekinensis accolus Jendek, 1995; A. souliei Jendek, 1994. Host plants: Fabaceae: Caragana; Lespedeza. Plagiatus species-group Description: BODY: Size: medium, Shape: cuneiform, Build: medium, Color (dorsally): unicolored or bicolored. EYES: Size: moderate, rarely small or large. ANTENNAE: Length: moderate or long, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square or transverse, Maximal width: at anterior margin or at middle, Anterior lobe: moderate, Disk: flat, moderately convex, rarely strongly convex, Impressions: medial and lateral, Prehumerus: filamentary, carinal, rarely obsolete, Prehumerus shape: bisinuate, rarely arcuate, Prehumerus extent: to half of pronotal length or to anterior pronotal angle, Prehumerus - anterior end: joining pronotal marginal carina or joining anterior pronotal angle, Marginal and submarginal carinae - interspace: narrow or moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent or present, Marginal and submarginal carinae - narrowest point: at posterior angles or distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored or bicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate or subangulate, Pubescence: entire or on most of elytra, proximal and/or distal or ornamental, Pubescence - color: unicolored or bicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate,

29

SPECIES GROUPS Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Elytra distinctly attenuate in apical third, prehumerus long, bisinuate, rarely arcuate. Relevant group(s): Imitans; Sinuatus. Taxa: A. aureofasciatus sp. nov.; A. conspersus sp. nov.; A. discalis Saunders, 1873; A. kostali Jendek, 2007; A. majzlani Jendek, 2007; A. miao sp. nov.; A. plagiatus Ganglbauer, 1889; A. pumi sp. nov.; A. spathulifer sp. nov.; A. stigma sp. nov. Host plants: Cannabaceae: Celtis; Ulmaceae: Zelkova. Plasoni species-group Description: BODY: Size: medium, Shape: cuneiform, Build: slender or medium, Color (dorsally): unicolored, bicolored, rarely tricolored. EYES: Size: moderate, rarely small. ANTENNAE: Length: short, moderate or long, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: transverse, Maximal width: at middle, Anterior lobe: absent or moderate, Disk: moderately convex, Impressions: medial and lateral, Prehumerus: absent or obsolete, Prehumerus shape: straight or arcuate, Prehumerus extent: to half of pronotal length, Prehumerus - anterior end: distant from pronotal marginal carina, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent, Marginal and submarginal carinae - narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored or bicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: cuspidate or spinose, Pubescence: proximal and/or distal, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Aedeagus: weakly sclerotised, rarely parameres with obvious membranous lobes, Ovipositor: uritiform. Diagnosis: Elytral apex with cusp or spine on pleural side; prehumerus absent or obsolete and short; aedeagus weakly sclerotised, often with obvious membranous lobes. Relevant group(s): Acutus. Taxa: A. bicuspidatus sp. nov.; A. chujoi Kurosawa, 1985; A. diaolin Jendek, 2001; A. huashanus Jendek, 2001; A. plasoni Obenberger, 1917. Host plants: Fabaceae: Pueraria. Prolatus species-group Description: BODY: Size: small, Shape: cuneiform or fusiform, Build: slender, Color (dorsally): unicolored. EYES: Size: small. ANTENNAE: Length: short or moderate, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square, Maximal width: at middle or at posterior margin, Anterior lobe: moderate, Disk: moderately convex or strongly convex, Impressions: absent, medial and lateral or lateral only, Prehumerus: obsolete or carinal, Prehumerus shape: straight or arcuate, Prehumerus extent: to half of pronotal length, Prehumerus - anterior end: distant from pronotal marginal carina, rarely close to pronotal marginal carina, Marginal and submarginal carinae - interspace: narrow, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: present, Marginal and submarginal carinae - narrowest point:

at posterior angles or distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent or obsolete, Apices - arrangement: separate, Apices - shape: arcuate, rarely subtruncate, Pubescence: entire or on most of elytra, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications or with sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate or sinuate, Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Small and slender species; prehumerus obsolete or carinal; marginal and submarginal carinae with narrow interspace; elytra entirely pubescent. Relevant group(s): not asserted. Taxa: A. prolatus sp. nov.; A. qiang sp. nov.; A. recticollis sp. nov. Host plants: Unknown. Quadrisignatus species-group Description: BODY: Size: medium, Shape: cuneiform, Build: medium, Color (dorsally): unicolored. EYES: Size: moderate or large. ANTENNAE: Length: moderate, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square or transverse, Maximal width: at anterior margin or at middle, Anterior lobe: moderate or obvious, Disk: flat or moderately convex, Impressions: medial and lateral, rarely medial only, Prehumerus: absent, obsolete, tubercular or carinal, Prehumerus shape: arcuate, Prehumerus extent: close to posterior pronotal angle, Prehumerus - anterior end: distant from pronotal marginal carina, Marginal and submarginal carinae - interspace: narrow or moderate, Marginal and submarginal carinae convergence: convergent, Marginal and submarginal carinae - junction: absent or present, Marginal and submarginal carinae - narrowest point: distant from posterior angles, rarely at posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate, Pubescence: pairs of pubescent-tomentose spots, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: with sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Medium sized species; pronotum flat or moderately convex, with large lateral impressions; elytra with pairs of pubescent spots; elytral apices separately arcuate. Relevant group(s): Spinipennis, Niveoguttatus, Biguttatus, Maculifer. Taxa: A. emmerichi Obenberger, 1935; A. fortunatus Lewis, 1893; A. nipponigena Obenberger, 1935; A. ornatoides sp. nov.; A. prosternalis sp. nov.; A. quadrisignatus Marseul, 1866; A. sospes Lewis, 1893. Host plants: Ebenaceae: Diospyros; Fagaceae: Fagus; Quercus; Ulmaceae: Ulmus; Zelkova. Roscidus species-group Description: BODY: Size: small or medium, Shape: cuneiform, Build: medium, Color (dorsally): unicolored. EYES: Size: small, rarely moderate. ANTENNAE: Length: moderate, Shape: slender, Antennomeres in male: not

30 modified. PRONOTUM: Shape: visually square, Maximal width: at anterior margin, Anterior lobe: moderate, Disk: moderately convex, Impressions: medial and lateral, rarely lateral only, Prehumerus: obsolete, filamentary or carinal, Prehumerus shape: arcuate or bisinuate, Prehumerus extent: to half of pronotal length or to anterior pronotal angle, Prehumerus - anterior end: distant from pronotal marginal carina or joining anterior pronotal angle, Marginal and submarginal carinae - interspace: moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent, Marginal and submarginal carinae - narrowest point: at posterior angles or distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate, Pubescence: entire or on most of elytra, rarely absent, Pubescence color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications or with sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Aedeagus: rarely markedly dorsoventrally bent, Ovipositor: elongate. Diagnosis: Eyes small, rarely medium; pronotum enlarged anteriorly; pronotal sides bisinuate; groove on apex of last ventrite arcuate. Relevant group(s): not asserted. Taxa: A. chlorophyllus Abeille de Perrin, 1904; A. eunuchus sp. nov.; A. moerens Saunders, 1873; A. spiculum sp. nov.; A. ulmicola sp. nov. Remark: This is Palaearctic species-group with additional species occurring outside East Asia. Host plants: Anacardiaceae: Pistacia; Fagaceae: Quercus. Sinensis species-group Description: BODY: Size: large, Shape: subparallel or cuneiform, Build: robust, Color (dorsally): bicolored. EYES: Size: moderate. ANTENNAE: Length: moderate, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square or transverse, Maximal width: at middle or at posterior margin, Anterior lobe: moderate, obvious, rarely absent, Disk: moderately convex, Impressions: medial and lateral, Prehumerus: obsolete or filamentary, Prehumerus shape: bisinuate, Prehumerus extent: to anterior pronotal angle, Prehumerus - anterior end: joining pronotal marginal carina or joining anterior pronotal angle, Marginal and submarginal carinae - convergence: subparallel, Marginal and submarginal carinae - junction: absent, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate, Pubescence: absent. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: with sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Large, ventrally bicolored species; marginal and submarginal carinae subparallel; elytra widely arcuate; aedeagus robust. Relevant group(s): Cyaneoniger. Taxa: A. coreanus Obenberger, 1935; A. sinensis Thomson, 1879; A. sinensis splendidicollis Fairmaire, 1889.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Host plants: Unknown. Sinuatus species-group Description: BODY: Size: medium, rarely large, Shape: cuneiform, Build: medium or robust. EYES: Size: moderate. ANTENNAE: Length: moderate, rarely long, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Maximal width: at middle, rarely at anterior margin, Anterior lobe: moderate, rarely absent, Disk: moderately convex, Impressions: medial and lateral, Prehumerus: carinal, Prehumerus shape: arcuate, Prehumerus extent: to half of pronotal length, Prehumerus - anterior end: distant from pronotal marginal carina, Marginal and submarginal carinae - interspace: narrow, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent or present, Marginal and submarginal carinae narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate, Pubescence: absent or proximal and/ or distal, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Medium-sized, dorsally coppery or reddish coppery; prehumerus carinal. Relevant group(s): Plagiatus; Imitans. Taxa: A. mali Matsumura, 1924; A. sinuatus sachalinensis Obenberger, 1935. Remark: This is Palaearctic species-group with additional species occurring outside East Asia. Host plants: Rosaceae: Crataegus; Malus; Prunus; Pyrus; Sorbus. Smaragdinus species-group Description: BODY: Size: medium, Shape: cuneiform, Build: medium, rarely robust, strongly convex, Color (dorsally): bicolored or tricolored. EYES: Size: moderate, rarely small or large. ANTENNAE: Length: moderate or long, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square, Maximal width: at anterior margin or at middle, Anterior lobe: moderate or obvious, Disk: flat, moderately convex or strongly convex, Impressions: absent, medial and lateral or lateral only, Prehumerus: absent, obsolete or carinal, Prehumerus shape: arcuate, Prehumerus extent: close to posterior pronotal angle, Prehumerus - anterior end: distant from pronotal marginal carina, Marginal and submarginal carinae - interspace: narrow, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent or present, Marginal and submarginal carinae narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored or bicolored, Humeral carina: absent, Apices - arrangement: separate, rarely conjoint, Apices - shape: subangulate or subtruncate, Pubescence: absent or proximal and/or distal, Pubescence color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without

SPECIES GROUPS sexual modifications or with sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Ovipositor: elongate. Diagnosis: Medium-sized, body dorsally bicolored or tricolored; pronotum often with blackish medial stripe; elytra often with blackish perisutural part; prehumerus very short or absent. Relevant group(s): Viridis, Betuleti. Taxa: A. aurinotus sp. nov.; A. fusciapex sp. nov.; A. lacrima sp. nov.; A. loosdregti Baudon, 1965; A. notoclavus Jendek, 2000; A. odetteae Baudon, 1968; A. peregrinus Kiesenwetter, 1879; A. priamus Kerremans, 1912; A. smaragdinus habros ssp. nov.; A. smaragdinus smaragdinus Solsky, 1876. Host plants: Betulaceae: Alnus; Ulmaceae: Ulmus. Spinipennis species-group Description: BODY: Size: medium or large, Shape: subparallel or cuneiform, Build: medium, robust, rarely strongly convex, Color (dorsally): unicolored or bicolored. EYES: Size: large, rarely moderate. ANTENNAE: Length: moderate, Shape: slender, Antennomeres in male: not modified, rarely modified. PRONOTUM: Shape: visually square, transverse, rarely visually elongate, Maximal width: at middle, Anterior lobe: moderate or obvious, Disk: moderately convex, strongly convex, rarely flat, Impressions: medial and lateral, rarely lateral only, Prehumerus: carinal, Prehumerus shape: straight, arcuate or bisinuate, Prehumerus extent: close to posterior pronotal angle, to half of pronotal length, beyond half of pronotal length or to anterior pronotal angle, Prehumerus - anterior end: distant from pronotal marginal carina, close to pronotal marginal carina, joining pronotal marginal carina or joining anterior pronotal angle, Marginal and submarginal carinae - interspace: moderate, Marginal and submarginal carinae - convergence: convergent, rarely subparallel, Marginal and submarginal carinae - junction: absent or present, Marginal and submarginal carinae narrowest point: at posterior angles or distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: subangulate, cuspidate, spinose, rarely arcuate, Pubescence: pairs of pubescent-tomentose spots, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: with sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate or sinuate, Pygidium: arcuate, extended into short protrusion or extended into long spine. GENITALIA: Aedeagus: markedly dorsoventrally bent, Ovipositor: elongate. Diagnosis: Large or medium sized species; eyes large, rarely moderate; pronotum widest at middle; prehumerus carinal; elytra with 2-4 pair of tomentose spots; elytral apices subangulate, cuspidate or spinose, rarely arcuate. Relevant group(s): Niveoguttatus, Biguttatus, Quadrisignatus. Taxa: A. ashlocki Baudon, 1968; A. coomani Bourgoin, 1925; A. cordicollis sp. nov.; A. cyanipennis Gory & Laporte, 1839; A. dureli sp. nov.; A. erythrostictus Bourgoin, 1922; A. fleischeri Obenberger, 1925; A. fornicatus sp. nov.; A. hmong sp. nov.; A. insulicola Kerremans, 1912; A. kawarai Kurosawa, 1963; A. sachalinicola Obenberger,

31 1940; A. spinipennis Lewis, 1893; A. tokyoensis Kurosawa, 1985; A. zhui sp. nov. Host plants: Betulaceae: Alnus; Fabaceae: Dalbergia; Fagaceae: Quercus; Salicaceae: Populus; Salix; Ulmaceae: Ulmus; Zelkova. Subrobustus species-group Description: BODY: Size: small or medium, Shape: cuneiform, Build: slender, medium, robust, strongly convex, Color (dorsally): unicolored. EYES: Size: small, moderate or large. ANTENNAE: Length: short or moderate, Shape: slender or stout, Antennomeres in male: not modified. PRONOTUM: Shape: visually square or transverse, Maximal width: at middle, rarely at anterior margin or at posterior margin, Anterior lobe: absent, moderate or obvious, Disk: strongly convex, Impressions: absent or medial only, Prehumerus: absent or obsolete, Prehumerus shape: arcuate, Prehumerus extent: close to posterior pronotal angle, Prehumerus - anterior end: distant from pronotal marginal carina, Marginal and submarginal carinae - interspace: narrow or moderate, Marginal and submarginal carinae convergence: convergent, Marginal and submarginal carinae - junction: absent or present, Marginal and submarginal carinae - narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate or conjoint, Apices - shape: arcuate, Pubescence: absent, sutural wedge or perisutural stripes or proximal and/or distal, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat or impressed. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications or with sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate or sinuate, Pygidium: arcuate. GENITALIA: Aedeagus: rarely parameres with obvious membranous lobes, Ovipositor: uritiform or elongate. Diagnosis: Antennae short or medium, slender or stout; pronotum strongly convex, without lateral impressions, disk regularly transversely rugose; prehumerus absent, rarely obsolete and short. Relevant group(s): Viduus. Taxa: A. aurigaster sp. nov.; A. bonadonai Descarpentries & Villiers, 1963; A. caligans Bourgoin, 1925; A. ineptus Kerremans, 1892; A. ningpoensis Obenberger, 1927; A. presbys sp. nov.; A. quentini Descarpentries & Villiers, 1963; A. subrobustus Saunders, 1873; A. yunnanicola Obenberger, 1936. Host plants: Fabaceae: Albizzia. Sulcicollis species-group Description: BODY: Size: small or medium, Shape: subparallel or cuneiform, Build: slender or medium, Color (dorsally): unicolored. EYES: Size: large, rarely moderate. ANTENNAE: Length: moderate, rarely long, Shape: slender, Antennomeres in male: not modified or modified. PRONOTUM: Shape: visually elongate, visually square or transverse, Maximal width: at anterior margin, rarely at middle, Anterior lobe: moderate or obvious, Disk: flat or moderately convex, Impressions: medial and lateral, Prehumerus: filamentary or carinal, Prehumerus shape: straight or arcuate, Prehumerus extent: to half of pronotal

32 length, rarely beyond half of pronotal length, Prehumerus anterior end: distant from pronotal marginal carina, rarely close to pronotal marginal carina or joining pronotal marginal carina, Marginal and submarginal carinae - interspace: moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: present, rarely absent, Marginal and submarginal carinae narrowest point: at posterior angles or distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate or conjoint, Apices - shape: arcuate or subtruncate, Pubescence: absent or proximal and/or distal, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat, rarely impressed. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications or with sexual modifications, Groove on apex of last ventrite: sinuate, Pygidium: arcuate. GENITALIA: Aedeagus: often asymmetrical, Ovipositor: elongate. Diagnosis: Pronotum with narrow medial impression; groove on apex of last ventrite deeply sinuate; male often with sexual modifications on antennae and abdominal ventrites; aedeagus often asymmetrical. Relevant group(s): Tempestivus. Taxa: A. adelphinus Kerremans, 1895; A. andrusi Baudon, 1968; A. bituberculatus Jendek, 2007; A. chaetocoxalis sp. nov.; A. fissus Obenberger, 1917; A. foliatus Jendek, 2007; A. friebi Obenberger, 1922; A. kaluganus Obenberger, 1940; A. laticornis (Illiger, 1803); A. latipalpis Jendek, 2007; A. marginicollis Saunders, 1873; A. miwai Obenberger, 1936; A. obscuricollis Kiesenwetter, 1857; A. obscurosericeus Obenberger, 1940; A. pseudobscuricollis Jendek, 2007; A. reichardti Obenberger, 1935; A. ribbei Kiesenwetter, 1879; A. suginoi Tôyama, 1985; A. sulcinotus Jendek, 2007; A. suturicuspidatus Jendek, 2007; A. takahashii Tôyama, 1988; A. trinotatus Saunders, 1873; A. truncatus Jendek, 2007; A. ussuricola Obenberger, 1924; A. varius Kerremans, 1895; A. volkovitshi Jendek, 2007; A. yakushimensis Tôyama, 1985. Remark: This is Palaearctic species-group with additional species occurring outside East Asia. Host plants: Betulaceae: Betula; Carpinus; Corylus; Ostrya; Fagaceae: Castanea; Quercus; Malvaceae: Tilia; Moraceae: Ficus; Rhamnaceae: Rhamnus; Sapindaceae: Acer; Ulmaceae: Zelkova; Vitaceae: Vitis. Tempestivus species-group Description: BODY: Size: small or medium, Shape: cuneiform, Build: slender or medium, Color (dorsally): unicolored. EYES: Size: moderate or large. ANTENNAE: Length: moderate or long, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually elongate, visually square, rarely transverse, Maximal width: at anterior margin, rarely at middle, Anterior lobe: moderate or obvious, Disk: flat or moderately convex, Impressions: medial and lateral or posterolateral, Prehumerus: carinal, rarely obsolete or filamentary, Prehumerus shape: arcuate or bisinuate, Prehumerus extent: to half of pronotal length, beyond half of pronotal length or to anterior pronotal angle, Prehumerus - anterior end: distant from pronotal marginal carina, close to pronotal marginal carina or joining pronotal marginal

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA carina, Marginal and submarginal carinae - interspace: moderate, rarely narrow, Marginal and submarginal carinae convergence: convergent, Marginal and submarginal carinae - junction: absent or present, Marginal and submarginal carinae - narrowest point: distant from posterior angles, rarely at posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate, rarely subangulate or subtruncate, Pubescence: entire or on most of elytra, proximal and/or distal, rarely sutural wedge or perisutural stripes, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat, rarely impressed. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications or with sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, rarely sinuate, Pygidium: arcuate, extended into short protrusion or extended into long spine. GENITALIA: Ovipositor: elongate. Diagnosis: Pronotum enlarged anteriorly; disk with medial and lateral, or posterolateral impression; prehumerus long, carinal, extending at least to half of pronotum, rarely obsolete. Relevant group(s): Sulcicollis. Taxa: A. asiaticus Kerremans, 1898; A. chekiangensis Gebhardt, 1929; A. ciping sp. nov.; A. diao sp. nov.; A. esakii Kurosawa, 1964; A. fevreae Descarpentries & Villiers, 1963; A. han sp. nov.; A. malloti Théry, 1930; A. sentis sp. nov.; A. she sp. nov.; A. shibatai Kurosawa, 1964; A. tempestivus Lewis, 1893; A. tsutsumiuchii Ohmomo, 2006; A. zhuang sp. nov. Host plants: Betulaceae: Carpinus; Euphorbiaceae: Mallotus; Fagaceae: Castanopsis; Quercus; Lauraceae: Machilus; Ulmaceae: Zelkova. Viduus species-group Description: BODY: Size: medium, Shape: cuneiform, Build: medium or robust, Color (dorsally): unicolored. EYES: Size: moderate or large. ANTENNAE: Length: short, Shape: stout, Antennomeres in male: not modified. PRONOTUM: Shape: visually square or transverse, Maximal width: at anterior margin, Anterior lobe: absent or moderate, Disk: flat or moderately convex, Impressions: medial and lateral or lateral only, Prehumerus: tubercular, carinal, rarely obsolete, Prehumerus shape: arcuate, Prehumerus extent: to half of pronotal length, Prehumerus - anterior end: distant from pronotal marginal carina, Marginal and submarginal carinae - interspace: narrow or moderate, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: absent, Marginal and submarginal carinae - narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices shape: arcuate, Pubescence: entire or on most of elytra or sutural wedge or perisutural stripes, Pubescence - color: unicolored. STERNUM: Metasternal projection: impressed. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, rarely sinuate, Pygidium: arcuate. GENITALIA: Ovipositor: uritiform. Diagnosis: Frons longitudinally rugose; antennae short and stout; pronotum widest at anterior margin.

33

SPECIES GROUPS Relevant group(s): Subrobustus. Taxa: A. compacticornis sp. nov.; A. shan sp. nov.; A. viduus Kerremans, 1914. Host plants: Cannabaceae: Celtis; Rosaceae: Prunus; Ulmaceae: Ulmus; Zelkova. Viridiobscurus species-group Description: BODY: Size: small, rarely medium, Shape: cuneiform, Build: robust, strongly convex, Color (dorsally): unicolored. EYES: Size: moderate or large. ANTENNAE: Length: short, moderate or long, Shape: slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square or transverse, Maximal width: at middle, Anterior lobe: moderate or obvious, Disk: moderately convex or strongly convex, Impressions: medial and lateral, posterolateral, rarely medial only, Prehumerus: carinal, Prehumerus shape: bisinuate, Prehumerus extent: beyond half of pronotal length or to anterior pronotal angle, Prehumerus - anterior end: joining pronotal marginal carina or joining anterior pronotal angle, Marginal and submarginal carinae - interspace: narrow, moderate or wide, Marginal and submarginal carinae - convergence: convergent, Marginal and submarginal carinae - junction: present, Marginal and submarginal carinae - narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, rarely obsolete or present, Apices - arrangement: conjoint, rarely separate, Apices - shape: arcuate, rarely subtruncate, Pubescence: entire or on most of elytra, Pubescence - color: unicolored. STERNUM: Metasternal projection: impressed or with pit. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications or with sexual modifications, Groove on apex of last ventrite: sinuate, Pygidium: arcuate. GENITALIA: Ovipositor: uritiform. Diagnosis: Body small or medium, cuneiform; pronotum widest around middle; prehumerus extends to anterior pronotal angles; mesosternal projection impressed or with pit. Relevant group(s): Densetomentosus, Japanocarinatus. Taxa: A. caudicula sp. nov.; A. foramenifer sp. nov.; A. lobatus sp. nov.; A. pliculipennis Obenberger, 1940; A. pusillesculptus Obenberger, 1940; A. tai sp. nov.; A. tuber sp. nov.; A. viridiobscurus Saunders, 1873 Host plants: Betulaceae: Alnus; Carpinus; Hamamelidaceae: Hamamelis; Juglandaceae: Juglans; Lauraceae: Parabenzoin; Ulmaceae: Zelkova. Viridis species-group Description: BODY: Size: medium, Shape: cuneiform, Build: medium, robust, rarely strongly convex, Color (dorsally): unicolored, rarely bicolored. EYES: Size: moderate, rarely small or large. ANTENNAE: Length: short, rarely moderate, Shape: stout, rarely slender, Antennomeres in male: not modified. PRONOTUM: Shape: visually square or transverse, Maximal width: at anterior margin or at middle,

Anterior lobe: absent or moderate, Disk: flat, moderately convex, rarely strongly convex, Impressions: medial and lateral, Prehumerus: absent, obsolete, tubercular, rarely carinal, Prehumerus shape: straight or arcuate, Prehumerus extent: close to posterior pronotal angle, rarely to half of pronotal length, Prehumerus - anterior end: distant from pronotal marginal carina, Marginal and submarginal carinae - interspace: moderate, Marginal and submarginal carinae - convergence: convergent, rarely subparallel, Marginal and submarginal carinae - junction: absent, Marginal and submarginal carinae - narrowest point: distant from posterior angles, Scutellum: well developed. ELYTRA: Color: unicolored, Humeral carina: absent, Apices - arrangement: separate, Apices - shape: arcuate, Pubescence: absent, rarely proximal and/or distal, Pubescence - color: unicolored. STERNUM: Metasternal projection: flat. ABDOMEN: Basal ventrite or metacoxae in male: without sexual modifications, Groove on apex of last ventrite: arcuate, subangulate or subtruncate, Pygidium: arcuate. GENITALIA: Ovipositor: uritiform. Diagnosis: Prehumerus tubercular, obsolete or absent, rarely carinal; pronotum with medial and lateral impressions; elytra in apical third distinctly attenuate; elytral apices separately arcuate; ovipositor uritiform. Relevant group(s): Smaragdinus, Betuleti. Taxa: A. cuprescens (Ménétriés, 1832); A. pecirkai Obenberger, 1916; A. ribesi Schaefer, 1946; A. salicis Frivaldszky, 1877; A. suvorovi Obenberger, 1935; A. tschitscherini Semenov, 1895; A. viridis (Linné, 1758). Host plants: Betulaceae: Alnus; Betula; Carpinus; Corylus; Ostrya; Fagaceae: Castanea; Fagus; Grossulariaceae: Grossularia; Ribes; Malvaceae: Tilia; Myricaceae: Myrica; Rhamnaceae: Rhamnus; Rosaceae: Rosa; Sorbus; Salicaceae: Populus; Salix; Sapindaceae: Acer; Ulmaceae: Ulmus. Unassigned taxa Taxa: A. adspersus (Kerremans, 1890); A. baroni Gory & Laporte, 1839; A. charismaticus Jendek, 2000; A. confinis Faldermann, 1835; A. convictor Descarpentries & Villiers, 1963; A. cyaneomicans Nonfried, 1892; A. diversornatus sp. nov.; A. ganglbaueri Semenov, 1891; A. hattorii Nakane, 1983; A. improcerus sp. nov.; A. komareki Obenberger, 1926; A. komareki sapporoensis Obenberger, 1935; A. koyoi Ohmomo, 2002; A. kuangcenensis Obenberger, 1940; A. lisu sp. nov.; A. muongoides sp. nov.; A. pilosovittatus Saunders, 1873; A. pseudoconstantini Alexeev, 1981; A. rudicollis Alexeev, 1979; A. shaowensis Jendek, 1994; A. siskai sp. nov.; A. subauratus amurensis Obenberger, 1922; A. sugiurai Jendek, 2007; A. tazoei Kurosawa, 1985; A. tyndarus Kerremans, 1913; A. ulaangomiensis Cobos, 1972; A. violaceus Nonfried, 1892; A. yamajii Tôyama, 1985. Host plants: Cannabaceae: Celtis; Ericaceae: Symplocos; Fabaceae: Caragana; Wisteria; Fagaceae: Quercus; Moraceae: Morus; Salicaceae: Populus; Pueraria; Ulmaceae: Ulmus.

34

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

35

TAXONOMIC

INVENTORY OF

AGRILUS

Agrilus adelphinus Kerremans Fig. 22I (habitus); Fig. 37R (aedeagus); Figs. 42C, 45S, 49W, 50C, 50M (types) adelphinus Kerremans, 1895 (Agrilus) KERREMANS, 1895: 222 [#1416] (description). SOLSKY, 1871: 360 [#4756] ([Note: Misidentified as tenuis]; faunal records; East Siberia). KERREMANS, 1903: 278 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 939 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 354 [#2429] (checklist; China). JENDEK, 1994: 14 [#1355] (lectotype designation; synonyms; characters; Russian Far East; Korea; China). ALEXEEV, 1998: 373 [#4322] (subgenus Quercuagrilus; subgeneric classification). HUA LI ZHONG, 2002: 89 [#6696] (cited as adelphillus; checklist; China). OHMOMO, 2002: 163 [#5750] (characters; Japan). JENDEK, 2003: 186 [#5823] (characters). FUKUTOMI & HORI, 2004: 13 [#6851] (checklist; faunal records; Japan (Hokkaido)). OHMOMO, 2004: 140 141 [#6850] (notes). JENDEK, 2006: 391 [#7441] (subgenus Quercuagrilus; Palaearctic catalog). BELLAMY, 2008: 1950 [#8576] (subgenus Quercuagrilus; world catalog). VOLKOVITSH, 2009: 135 [#8850] (subgenus Quercuagrilus; faunal records; Russia: Primorskiy kray). nonfriedi Obenberger, 1914 (Agrilus; [preoccupied]) OBENBERGER, 1914: 44 (in Czech), 49 50 (in German) [#1965] (description). OBENBERGER, 1923: 64 [#2026] (synonym of nonfriedanus). OBENBERGER, 1936: 1016 [#2191] (synonym of nonfriedanus). JENDEK, 1994: 14 [#1355] (synonym of adelphinus; lectotype designation). JENDEK, 2006: 391 [#7441] (synonym of adelphinus; Palaearctic catalog). BELLAMY, 2008: 1950 [#8576] (synonym of adelphinus; world catalog).

= nonfriedanus Obenberger, 1923 (Agrilus; replacement name for nonfriedi Obenberger not Kerremans) OBENBERGER, 1923: 64 [#2026] (replacement name proposal). OBENBERGER, 1924: 38 [#2040] (note). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1016 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 356 357 [#2429] (checklist; China). JENDEK, 1994: 14 [#1355] (synonym of adelphinus). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Shandong). JENDEK, 2006: 391 [#7441] (synonym of adelphinus; Palaearctic catalog). BELLAMY, 2008: 1950 [#8576] (synonym of adelphinus; world catalog). nigrocoerulans Obenberger, 1924 (Agrilus) OBENBERGER, 1924: 39 [#2040] (description). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1016 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 356 [#2429] (cited as nigrocoerulaus; checklist; China). JENDEK, 1994: 14 [#1355] (synonym of adelphinus; lectotype designation). HUA LI ZHONG, 2002: 90 [#6696] (cited as nigrocaerulans; checklist; China: Shandong, Hubei, Guangdong). JENDEK, 2006: 391 [#7441] (synonym of adelphinus; Palaearctic catalog). BELLAMY, 2008: 1950 [#8576] (synonym of adelphinus; world catalog). panhensis Baudon, 1968 (Agrilus) syn. nov. BAUDON, 1968: 108, 117 118 [#162] (description). BELLAMY, 2008: 2224 [#8576] (subgenus Sinagrilus; world catalog). egorovi Alexeev, 1989 (Agrilus) JENDEK, 1994: 14 ALEXEEV, 1989: 480 [#64] (description). [#1355] (as unavailable synonym of adelphinus). JENDEK, 2006: 391 [#7441] (available synonym of adelphinus; Palaearctic

TAXA OF

EAST ASIA

catalog). BELLAMY, 2008: 1950 [#8576] (as unavailable synonym of adelphinus; world catalog).

Type material: Agrilus adelphinus Kerremans, 1895 Type locality: Chine: Ho-Chan. Type specimens: Primary: Lectotype (Fig. 42C) , ♂, designated by JENDEK (1994: 14) [#1355], (MNHN): “Chine Ho Chan R. P. Moulton [p] \ adelphinus Kerr. Type [h] \ Kerremans vidit 1895 [p]”. Secondary: 1 PLT (BMNH). Described from unknown number of syntypes. Agrilus nonfriedi Obenberger, 1914 Type locality: China: Kiautschou. Type specimens: Primary: Lectotype (Fig. 50C), ♂, designated by JENDEK (1994: 14) [#1355], (NMPC): “China ex Nonfried [h] \ Agrilus Nonfriedanus m. (= Nonfriedi m.) Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24887 [h] [orange label]”. Described from 2 syntypes. Agrilus nonfriedanus Obenberger, 1923 Type locality and specimens: See: Agrilus nonfriedi Obenberger, 1914. Agrilus nigrocoerulans Obenberger, 1924 Type locality: China: Kiau-Čeu. Type specimens: Primary: Lectotype (Fig. 49W), ♀, designated by JENDEK (1994: 14) [#1355], (NMPC): “Kiautschou China [p] \ typus [p] [red label] \ Agrilus nigrocoerulans m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24883 [h] [orange label]”. Described from unknown number of syntypes. Agrilus panhensis Baudon, 1968 Type locality: Laos: Muong Panh. Type specimens: Primary: Holotype (Fig. 50M), ♀, (MHNB): “Muong Panh 1.iv.64 [h] Laos (Baudon) [p] \ Type [p] [red label] \ Agrilus panhensis Type mihi [h] A. Baudon det. [p]”. Secondary: Two specimens from the same locality labeled as paratypes (MHNB). Described from 1 specimen (HT). Agrilus egorovi Alexeev, 1989 Type locality: YU Prim. [= south of Primorskiy kray] [in Russian]. Type specimens: Primary: Lectotype by present designation: Lectotype (Fig. 45S), ♂, (ZIN): “Primoree 15 [illegible handwriting] Kraskino 23.VI.1974 Egorov [h] \ Agrilus egorovi sp. n. Alexeev det [h] \ Holotypus [sic!] [p] [red label]”. Secondary: 2 PLT (ZIN). Described from unknown number of syntypes. Remark: The name egorovi was cited first time by ALEXEEV, 1989 [#64] in the key without the explicit intention to establish a new nominal taxon and fix name-bearing types. The ICZN, Article 73.1.3 declares that “The holotype of a new nominal species group taxon can only be fixed in the original publication and by the original author”. Diagnosis: 5.2-8.3 mm; pronotum visually elongate; medial pronotal impression deep; parameres of aedeagus with

36 obvious angular sides. Species group: Sulcicollis. Relevant species: A. ribbei Kiesenwetter, 1879.

Specimens examined: CHINA: 1 ♀ (MNHN): “Chasseurs Thibétains 1899”. Anhui: 1 ♂ (BMNH): “Chine, Ngan Hoei, R. P. Moulton [leg.]”. Hebei: 1 ♀ (EJCB): “China, 3 June 2000, Fengning env. 1200m, NW slope of Yunwu Shan, Zd. Jindra lgt.”; 1 ♀ (EJCB): “China, N Hebei, 3.vi.2000, Fengning env. NW slope of Yunwu Shan, 41.2, 116.6E, 1200m, Jaroslav Turna leg.”. Hubei: 2 ♂, 1 ♀ (EJCB): “China, central Hubei, Dahongshan, 31.5N 113.0E, 30.iv 1.v., Jaroslav Turna leg., 2005”. Shaanxi: 1 ♂ (EJCB): “China, S Shaanxi, Qinling Mts. S slope, Xunyangba S+W env., 33º28 37’ / 108º23 33’, 1400 2100m, 5 9.6.[19]95, L.+R. Businský lgt.”; 1 ♀ (EJCB): “China Shaanxi, Lüeang 33°07’N 106°05’E, 8.6. 14.6.1996, lgt. E. Kučera”; 1 ♀ (JMCZ): “China c., 17 18.V., Ganguyi 110,3’ 36,8’, 35 km NE of Yanan, leg. J. Halada 1996”; 2 ♂, 1 ♀ (EJCB): “China, 1000 1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v. 13.vi.1998, I. H. Marshal leg.”; 16 (EJCB): “China, 1150 1300 m, Shaanxi, Qinling mts., Foping (6 km N), 20 21.VI.1998, I. H. Marshal leg.”; 2 (EJCB): “China Shaanxi, Lüeang, 18.5. 21.5.2000, leg. E. Kučera”; 7 (EJCB): “C China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000 1300m, 23.V. 13.VI. leg. C. Holzschuh 2000”; 1 ♀ (EJCB): “China Shaanxi, Lueang env., 15 km NW, 26.5. 31.5.2001, lgt. E. Kučera”. Shanxi: 9 (EJCB): “China c. 29 30.v., Jingangling, 111.7’ 50km W Linfen, 36.2’, leg. J. Halada 1996”; 2 ♀ (EJCB): “China, Shanxi, 9.June.2000, Lüliang Shan, 1000m, road Fangshan Jiaocheng, Hengjian env., Zd. Jindra”; 1 ♂, 2 ♀ (EJCB): “China, W Shanxi, 9.VI.2000, 37.7N, 111.6E, Lüliang Shan, road Fangshan Jiaocheng, Hengjian env., ~1000m, Jaroslav Turna leg.”. Sichuan: 1 ♂ (EJCB): “China Sichuan, Pingwu, 32°15’N, 104°16’E, 3 9.6.1997, lgt. E. Kučera”. Yunnan: 1 (NMPC): “Yünnan” (JENDEK, 1994 [#1355]); 38 (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.” (JENDEK, 1994 [#1355]); 1 ♂ (EJCB): “Yunnan, 8 10. vii.1996, 28°06’N 98°53’E, 3300 m, Hengduan mts part Meili, Vít Kubáň leg.”. KOREA NORTH: 1 ♀ (EJCB): “D.P.R. Korea, Wonsan botanic garden, lgt. M. Slovák 28.V.1988” (JENDEK, 1994 [#1355]); 5 (EJCB): “N Korea 19.7.1990, Toxong env., Omdouri”; 6 (EJCB): “N. Korea, Hamgen South prov., Sinpkho 15.v.1990”. KOREA SOUTH: Ch’ungch’ong-namdo: 1 ♀ (JMCZ): “Korea: Chungnam Prov., Cheingyang gun, Mt. Chilgapsan, Naengcheun vallaey, 13 VII 2000, S. J. Park”. RUSSIA: Primorskiy kray: 1 ♀ (ZIN): “Primorsk. krai 15 17 km NNO st. Khasan Loginova 20.VI.[19]74 [in Russian]”; 1 ♀ (EJCB): “USSR, Ussuri prov. Arsenev env., VII.1990, leg. B. Šiška & R. Čermák”; 1 ♀ (EJCB): “Prim. Lazovsk, Kievka 18.VI.[19]91, Filimonov leg.” (JENDEK, 1994 [#1355]); 1 ♂ (EJCB): “USSR, Primorsk. kraj, Arsenev env., VI.1991, 44.11N, 133.19E, leg. O. Šauša & M. Štrba”; 1 ♂, 1 ♀ (EJCB): “USSR, Primorsk. kraj, Arsenev env., VI.1991, 44.11N, 133.19E, O. Šausa leg.” (JENDEK, 1994 [#1355]); 1 ♂, 1 ♀ (EJCB): “USSR, Primorskij kraj Arseneev env., VI.1991, 44.11 N, 133.19 E, M. Štrba leg.” (JENDEK, 1994 [#1355]); 1 ♂ (EJCB): “Prim. Khasansk. Rjazanovka, 5.VI.[19]91, Filimonov leg.” (JENDEK, 1994 [#1355]); 1 ♂, 1 ♀ (EJCB): “Russia, Primorsk. prov. Arsenev env., VII.1992 B. Šiska leg.” (JENDEK, 1994 [#1355]); 1 ♀ (JMCZ): “Ross. or, Prim. kr., Kraskino, Azajsan, 13 16.VII.1992, Snížek leg.”; 2 ♂ (EJCB): “Rossia, 2 10.VIII., Primorskij kraj, 1992, Ussurijsk, Kajmanovka, lgt. Smrž”; 1 ♂ (EJCB): “Rossia, Primorsk, Kraskino env. mer., 13 16.VII.1992, P. Smrž leg.” (JENDEK, 1994 [#1355]); 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Preobrazhenie env. 9 17.07.06 leg. М. & L. Smirnov’s”. Adult occurence: 5-6-7-8 (n=33). Altitude range: 10003300 m (n=9). Host plants: Quercus: ALEXEEV, 1989 [#64] (as egorovi).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Distribution: CHINA: Anhui, Guangdong, Hebei, Hubei, Shaanxi, Shandong, Shanxi, Sichuan, Xizang, Yunnan. JAPAN: Hokkaido, Honshu, Kyushu. LAOS: Xiangkhoang. KOREA NORTH. RUSSIA: Primorskiy kray.

Agrilus adspersus (Kerremans) Fig. 30A (habitus); Fig. 41D (aedeagus); Fig. 42D (type) adspersus Kerremans, 1890 (Cisseis) KERREMANS, 1890: CCIV CCV [#1389] (description). KERREMANS, 1892: 224 [#1397] (Cisseis; catalog). KERREMANS, 1903: 278 [#1441] (catalog). OBENBERGER, 1935: 842 [#2123] (Cisseis). OBENBERGER, 1936: 1071 [#2191] (world catalog). JENDEK, 2000: 502 [#4851] (lectotype designation; notes). JENDEK, 2006: 396 [#7441] (Palaearctic catalog). BELLAMY, 2008: 1951 [#8576] (world catalog).

Type material: Cisseis adspersa Kerremans, 1890 Type locality: Konbir. Type specimens: Primary: Lectotype (Fig. 42D), ♂, designated by JENDEK (2000: 502) [#4851], (MNHN): “Konbir [p] \ Cisseis adspersa Kerr. Type [h] [Kerremans’ MS] \ COLLECTIO Oberthur [p]”. Secondary: 2 PLT (BMNH). Described from 3 syntypes. Diagnosis: 6.6-7.9 mm; body with four pairs of large tomentose spots; head and pronotum with fine sculpure; prehumerus tubercular. Species group: Unassigned. Relevant species: Unknown. Specimens examined: BANGLADESH: Rajshahi: 1 (EJCB): “East Pakistan, Andharkola (Rajshahi Dt.) VII 1963, Mapelli”. INDIA: 2 ♀ (MNHN): “Chota Nagpore, Nowatoli, R. P. Cardon, V., VI, 1896”. Karnataka: 1 ♂, 1 ♀ (MNHN): “Samanahally, près Bangalore, R. P. Tabourel, IV, V, 1899”. Tamil Nadu: 1 ♀ (EJCB): “India S, Tamil Nadu, Nilgiris, 15 km SE of Kotagiri, Kunchappanai, 900 m, 11.22 N 76.56 E, 7 22.v.2000, leg. P. Pacholátko”. NEPAL: Kosi: 3 (EJCB): “E Nepal, Koshi, Dharan, 18.vi.1985, leg. Holzschuh”. Adult occurence: 4-5-6-7 (n=5). Altitude range: ~900 m (n=1). Host plants: Unknown. Distribution: INDIA. BANGLADESH. NEPAL. [Note: Not in Pakistan. Record of JENDEK, 2006 [#7441] is a confusion with Bangladesh, see East Pakistan in Specimens examined]. Remarks: Not recorded from East Asia but possible in south China.

Agrilus aerosus Ganglbauer Fig. 21G (habitus of LT); Fig. 37D (aedeagus) aerosus Ganglbauer, 1889 (Agrilus) GANGLBAUER, 1889: 22, 33 35 [#3427] (description). HEYDEN, 1893: 90 [#3450] (catalog). KERREMANS, 1903: 281 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 939 [#2191] (world catalog). ALEXEEV, 1975: 156 157 [#52] ([Note: Misidentification of przewalskii];

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA notes; faunal records; distributional overview; Mongolia). PENG ZHONGLIANG, 1987: 354 [#2429] (checklist; China). ALEXEEV & VOLKOVITSH, 1989: 315, 344, 356 [#3241] ([Note: Misidentification of przewalskii]; characters in key; faunal records; Mongolia). ALEXEEV, 1998: 375 [#4322] (subgenus Xeragrilus; [Note: Misidentification of przewalskii]; subgeneric classification). NONNAIZAB et al., 1999: 112 [#8247] (checklist; China: Inner Mongolia). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Gansu). JENDEK, 2006: 395 [#7441] (subgenus Xeragrilus; Palaearctic catalog). BELLAMY, 2008: 1954 [#8576] (subgenus Xeragrilus; world catalog).

Type material: Agrilus aerosus Ganglbauer, 1889 Type locality: Kan-ssu [Berg I-dschuschan, unwait der Stadt Nan-pin] [Note: Locality was complemented from the list of localities on pages 21-22]. Type specimens: Primary: Lectotype (Fig. 21G) by present designation, ♂, (NMW): “Kanssou 15/VII 1885 Potanin [h] \ Agr. aerosus Gglb. Typ [h] \ Typus [p] [red label]”. Secondary: 1 PLT (ZIN). Described from 3 syntypes. Note: The paralectotype in ZIN bears label “Holotype”. Diagnosis: (♂): 4.9-5.5 mm; body dark bronze, white pubescent, without tomentum; vertex with deep medial impression; pronotum widest in apical third; prehumerus present, long, extending from anterior to posterior angle. Species group: Albogularis. Relevant species: A. przewalskii Obenberger, 1936. Specimens examined: CHINA: Sichuan: 1 ♂ (EJCB): “China, NW Sichuan prov., Maowen, 1000 m, 31º30’N 103º50’E, 10 18. vii.1990, J. Kolibáč leg.”; 1 ♀ (EJCB): “China, N Sichuan, env. Nanping, 1800 m, 15.vii.2002”.

Adult occurence: 7 (n=2). Altitude range: 1000-1800 m (n=2). Host plants: Unknown. Host plants cited for A. aerosus by ALEXEEV, 1975 [#52] and ALEXEEV & VOLKOVITSH, 1989 [#3241] pertain to A. przewalskii. See also Remarks. Distribution: CHINA: Gansu, Sichuan. Remarks: All examined specimens determined by Alexeev as A. aerosus, were misidentifications of A. przewalskii. All distributional data published for A. aerosus, except the type locality, are dubious.

Agrilus agnatus Kerremans Fig. 8G (habitus); Fig. 32D (aedeagus); Fig. 42G (type) agnatus Kerremans, 1892 (Agrilus) KERREMANS, 1892: 820 821 [#1398] (description). KERREMANS, 1903: 281 [#1441] (catalog). OBENBERGER, 1936: 1071 [#2191] (world catalog). DESCARPENTRIES & VILLIERS, 1963: 8 [#701] (synonym of lafertei; characters; faunal records; Inde; Birmanie; Tonkin; Laos). TÔYAMA, 1988: 759 [#2932] ([Note: Specimens from Thailand are misidentified as lafertei according to JENDEK, 2000: 189 [#4852]]; faunal records; Thailand [pertains to agnatus]; Nepal [pertains to lafertei]). JENDEK, 2000: 189 190, 192 [#4852] (valid species; lectotype designation; notes; key; faunistic data). JENDEK, 2006: 396 [#7441] (Palaearctic catalog). BELLAMY, 2008: 1955 [#8576] (world catalog).

37 Type material: Agrilus agnatus Kerremans, 1892 Type locality: Carin Cheba, 900-1100 m. Type specimens: Primary: Lectotype (Fig. 42G), ♀, designated by JENDEK (2000: 189-190, 192) [#4852], (MCSN): “Carin Cheba 900-1100 m L. Fea V XII-[18]88 [p] \ Typus [p] [red ink, red border] \ agnathus [sic!] Kerr. [h] [black border] \ agnatus Kerr. Type [Kerremans’ MS] \ Agrilus agnathus [sic!] Kerr. typus! [h] \ Museo Civico di Genova [p]”. Secondary: 1 PLT (MNHN). Described from unknown number of syntypes. Diagnosis: 10-13 mm; vertex convex with distinct medial carina; eyes protruding beyond head outline; elytral apices narrowly arcuately-acuminate; pronotal sides with narrow patches of whitish pubescence. Species group: Cyaneoniger. Relevant species: A. lafertei Kerremans, 1892; A. qinling Jendek, 2000. Specimens examined: CHINA: Guizhou: 1 (NMW): “China: Guizhou, Leishan Co., SE Kaili, 15 km E Leishan, Leigong Shan, E slope, 26º23.39’N, 108º13.33’E / ca 2.5 km E of pass between Leishan and Fangxiang vill., ca 1600 m, 23/24.6.2001, leg. Schillhammer (17)”. Yunnan: 1 ♂ (EJCB): “Yun’an’ 765 km po dor[oge]. Kun’min Dalo, 1000 m, 26.IV.1957, Pu Fu di [in Russian]” (JENDEK, 2000 [#4852]); 1 ♂ (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.” (JENDEK, 2000 [#4852]); 1 ♀ (EJCB): “CHINA, Yunnan, 14 21.VI.1993, 100 km W of Baoshan, GAOLIGONGSHAN nat. res., E. Jendek & O. Šauša leg.” (JENDEK, 2000 [#4852]); 1 ♀ (EJCB): “Yunnan, 1500 2500 m, 25°22’N, 98°49’E, 17 24 / 5 Gaoligong mts, Vít Kubáň leg., 1995” (JENDEK, 2000 [#4852]); 3 (CBCS): “Yunnan, Jing Gu County, Mt. Yingpantou, 7 14.vi.2009 local collector”. LAOS: Houaphan: 8 (EJCB): “Laos NE, Hua Phan prov., Ban Saluei, Phu Phan Mt., 20°15’N, 104°02’E, 1500 2000m, D. Hauck leg., 26.iv. 11.v.2001”; 3 ♀ (EJCB): “Laos NE, Hua Phan prov., 20°15’N, 104°02’E, Phu Pan mt. (Ban Saluei), 1500 2000 m, 26.iv. 11.v.2001, J. Bezděk leg.”; 1 ♂, 1 ♀ (EJCB): “NE Laos, Hua Phan prov., Ban Saluei, 1300 2000m, 20°13’N, 103°59’E, 6 18.iv., Phu Phan Mt. env. 2004, F. & L. Kantner leg.”; 1 ♀ (EJCB): “LAOS, Hua Phan prov. 2004, Ban Saluei env., alt. 1500m, 30 km S of Xam Neua, Phu Phan Mt., 6 17.5.2004, 103°59’33’’E, 20°13’39’’N, P. Kresl leg.”; 7 (EJCB): “Lao NE, Hua Phan pr., ~20°12’N,104°01’E, Phu Phan Mt., 1500 1900m, 17.v. 3.vi.2007, Vít Kubáň leg.”; cca 30 (EJCB): “LAOS NE, Houa Phan prov., 20°12 13.5’N, 103°59’.5 104°01’E, Ban Saluei Phou Pane Mt., 1340 1870m, 15.iv. 15.v.2008, Lao collectors leg.”; cca 50 (NMPC): “LAOS NE, Houa Phan prov., 20°12 13.5’N, 103°59’.5 104°01’E, Ban Saluei Phou Pane Mt., 1340 1870m, 15.iv. 15.v.2008, Lao collectors leg.”. Louang Namtha: 1 ♀ (EJCB): “Laos north, 5 11.V.1997, 20 km NW Louang Namtha, N 21°09.2, E 101°18.7, alt. 900±100 m, E. Jendek & O. Šauša leg.” (JENDEK, 2000 [#4852]). Louangphrabang: 1 ♂, 3 ♀ (EJCB): “Laos N, 24.iv. 16.v.1999, Louang Phrabang prov. 20°33 4’ N 102°14’E, Ban Song Cha (5km W) ± 1200 m, Vít Kubáň leg.” (JENDEK, 2000 [#4852]); 1 ♂, 1 ♀ (EJCB): “Laos N, 23.iv.1999, Louangphrabang prov. 20°42’ N 102°54’E, 25 km E Muang Ngoy, 1000 m, Vít Kubáň leg.” (JENDEK, 2000 [#4852]). Phongsali: 1 ♀ (EJCB): “Lao N, Phongsaly prov., 21°41’ 2’N, 102°06’ 08’E, Phongsaly env., 28.v. 20.vi.2003, 1500m, P. Pacholátko leg.”. THAILAND: Chiang Mai: 1 ♀ (EJCB): “Phuping Palace, Ch eng Mai, N. Thailand, 25.V.1983, H. Akiyama leg.” (JENDEK, 2000 [#4852]); 1 ♀ (EJCB): “THAI, 17 24.V.1991, Chiang Dao, 1000 m, 19°25’N, 98°52’E, Vít Kubáň leg.” (JENDEK, 2000 [#4852]); 1 ♀

38 (EJCB): “THAI, 2 3/6.1995, 18°49’N, 98°54’E, DOI PUI, 1400 m, Vít Kubáň leg.” (JENDEK, 2000 [#4852]); 1 ♀ (EJCB): “Thai, Chiang Mai prov. 18°49’N, 98°54’E, 1600 m, Doi Pui mt., 2 6.V.1996, Vít Kubáň leg.” (JENDEK, 2000 [#4852]); 3 ♂ (EJCB): “Thailand, Chiang Mai prov., 19°19’N, 98°50’E, 1400 m, Sanpakia vill., 1 15.V.1998, Vít Kubáň leg.” (JENDEK, 2000 [#4852]). Mae Hong Son: 1 ♂ (EJCB): “NW Thailand, 19°19’N, 97°59’E, Mae Hong Son, 1991, Ban Huai Po, 1600 2000 m, 17 29.V., L. Dembický leg.” (JENDEK, 2000 [#4852]); 3 ♀ (EJCB): “Thai, Mae Hong Son pr., 19°27’N, 98°20’E, 1500 m, Soppong 7 12.V., Vít Kubáň leg., 1996” (JENDEK, 2000 [#4852]). VIETNAM: 1 ♀ (EJCB): “V’etnam gory V okr. SHA PA 1700 2000 m, 23.5.1963 g., Kabakov [leg.] [in Russian]” (JENDEK, 2000 [#4852]).

Adult occurence: 4-5-6 (n=27). Faunal records published by TÔYAMA, 1988 [#2932] are also included. Altitude range: 800-2500 m (n=22). Host plants: Unknown. Distribution: CHINA: Guizhou, Yunnan. LAOS. MYANMAR. THAILAND. VIETNAM. [Note: Not in Nepal. Record of JENDEK, 2006 [#7441] pertains to A. lafertei]

Agrilus alashanensis Obenberger Fig. 22G (habitus); Fig. 37P (aedeagus); Figs. 42I, 53H (types) alashanensis Obenberger, 1936 (Agrilus) OBENBERGER, OBENBERGER, 1936: 112 [#2205] (description). 1936: 940 [#2191] (world catalog). ALEXEEV, 1989: 485 [#64] (key to species; distributional summary; Chita; Buryatia; Tuva; China (Alashan); Mongolia). ALEXEEV & VOLKOVITSH, 1989: 319, 347, 356 [#3241] (characters in key; faunal records; lectotype designation; Mongolia). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). JENDEK, 2006: 393 [#7441] (subgenus Robertius; Palaearctic catalog). BELLAMY, 2008: 1955 [#8576] (subgenus Robertius; world catalog). rozhkovi Alexeev, 1964 (Agrilus) ALEXEEV, 1964: 13 14 [#3846] (description). ALEXEEV, 1968: 881 882 [#50] (redescription). ALEXEEV, 1975: 151 152 [#52] (notes; faunal records; distributional overview; Mongolia). ALEXEEV, 1989: 485 [#64] (synonym of alashanensis). ALEXEEV & VOLKOVITSH, 1989: 319 [#3241] (synonym of alashanensis; type examination). NONNAIZAB et al., 1999: 113 [#8247] (checklist; China: Inner Mongolia). JENDEK, 2006: 393 [#7441] (synonym of alashanensis; Palaearctic catalog). BELLAMY, 2008: 1955 [#8576] (synonym of alashanensis; world catalog). steppensis Cobos, 1968 (Agrilus; subgenus Agrilus) COBOS, 1968: 396 398 [#580] (description). COBOS, 1972: 51, 56 [#591] (subgenus Agrilus; faunal record; Mongolia). ALEXEEV, 1975: 151 [#52] (synonym of rozhkovi). ALEXEEV, 1989: 485 [#64] (synonym of alashanensis). NONNAIZAB et al., 1999: 113 [#8247] (cited as valid species and as steppesis in synonymy of rozhkovi; checklist; China: Inner Mongolia). JENDEK, 2006: 393 [#7441] (synonym of alashanensis; Palaearctic catalog). BELLAMY, 2008: 1955 [#8576] (synonym of alashanensis; world catalog).

Type material: Agrilus alashanensis Obenberger, 1936 Type locality: Asia centralis: Ala-Shan. Type specimens: Primary: Lectotype (Fig. 42I), ♂, designated by ALEXEEV & VOLKOVITSH (1989: 319, 347,

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA 356) [#3241], (NMPC): “Ala Shan [h]\ Typus [p] [red label] \ Agrilus alashanensis m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24321 [h] [orange label]”. Described from unknown number of syntypes. Agrilus rozhkovi Alexeev, 1964 Type locality: Zabaikal’e, ozero Gusinoe [in Russian]. Type specimens: Not located. Described from 2 specimens (HT, AT). Agrilus steppensis Cobos, 1968 Type locality: Central aimak, 26 km W von somon Lun, 1.180 m. alt. Type specimens: Primary: Holotype (Fig. 53H), ♂, (HNHM): “Mongolia, Central aimak, 26 km O [sic!] von somon Lun, 1.180 m. Exp. Dr. Z. Kaszab, 1964 [p] \ Nr. 262, 4. VII. 1964 [p] \ Holotypus A. Cobos [p] [red label] \ Agrilus steppensis nov. sp. Holotypus [h] A. Cobos det. 196 [p] 8 [p]”. Secondary: PT (MNMS); 2 PT (HNHM). Described from 5 specimens (HT, AT, PT). Diagnosis: 6.2-7.8 mm; marginal and submarginal carinae convergent and conjoined at posterior angles; prehumerus sharply carinal, arcuate, reaching almost to half of pronotal length. Species group: Pekinensis. Relevant species: A. pekinensis Obenberger, 1924. Specimens examined: MONGOLIA: 1 ♀ (ZIN): “Mongolia, Bayan Khongorskii aimak, 25 km Z [ West] Bumbègèra, G. Medvedev, 14.vi.[1]980 [in Russian]”. 1 ♂ (ZIN): “Mongolia, Vostochnyi aimak, oz. [ lake] Khukh Nur, M. Kozlov, 25.vi. [1]976 [in Russian]”. RUSSIA: Tyva Respublika: 2 ♀ (EJCB): “Tuvinskaja ASSR, 15 km W Kyzyl, 26.6., Z. Kletečka lg. [19]94”. Zabaykalskiy kray: 1 ♂, 1 ♀ (APCE): “Russia, Chitinskaja obl., Durbatschi, 23.VII.1984, leg. Makarin”; 1 ♂ (ZIN): “YuV Chitin, obl., r. Borzya, 26.vi.1989, Kadikov [in Russian]”; 1 ♂, 1 ♀ (ZIN): “YuV Chitin. obl., r. Borzya 26.vi.1989, Kabakov [in Russian]”; 2 ♂ (EJCB): “Chita Prov., Priargunsk, Distr. Duroy, 16 18.vi.2000, Kataev”.

Adult occurence: 6-7 (n=12). Faunal records published by ALEXEEV, 1964 [#3846] (as rozhkovi) and COBOS, 1972 [#591] (as steppensis) are also included. Altitude range: 1000-1630 m (n=5). Faunal records published by COBOS, 1972 [#591] (as steppensis) are also included. Host plants: Caragana: ALEXEEV, 1964 (as rozhkovi) [#3846]; ALEXEEV, 1968 (as rozhkovi) [#50]; ALEXEEV, 1975 (as rozhkovi) [#52]; ALEXEEV, 1989 [#64]; ALEXEEV & VOLKOVITSH, 1989 [#3241]. Distribution: CHINA: Nei Mongol. MONGOLIA. RUSSIA: Buryatia Respublika, Zabaykalskyi kray, Tyva Respublika.

Agrilus alesi Obenberger Fig. 19I (habitus); Fig. 36J (aedeagus); Figs. 42L, 42W, 52J (types) alesi Obenberger, 1935 (Agrilus) OBENBERGER, 1935: 164 [#2158] (description). MIWA & CHÛJÔ, 1936: 15 [#1809] (catalog; Japan). OBENBERGER, 1936: 940 [#2191] (world catalog). KUROSAWA, 1963: 152 [#1618] (subspecies of auriventris; characters; Japan). CHÛJÔ, 1970: 20 [#3350] (subspecies of auriventris; checklist; faunal records; Japan (Loo

39

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Choo Archipelago)). KUROSAWA, 1974: 2 3 [#4575] (characters; notes). TÔYAMA, 1985: 24 [#3678] (iconography; Japan). HIRASHIMA, 1989: 322 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 28 29 [#4091] (checklist; Japan: Ryukyus (Okinawa)). JENDEK, 2006: 396 [#7441] (Palaearctic catalog). BELLAMY, 2008: 1960 [#8576] (world catalog). sacchari Obenberger, 1940 (Agrilus) syn. nov. MIWA OBENBERGER, 1940: 175 176 [#2273] (description). & CHÛJÔ, 1940: 74 [#1810] (cited as sachari; faunal records; Okinawa: Loo Choo). KUROSAWA, 1974: 3 [#4575] (subspecies of alesi; characters; notes). HIRASHIMA, 1989: 322 [#8078]

(subspecies of alesi; checklist; Japan). – PENG ZHONGLIANG, 1994: 19 [#2432] (faunal records; China). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (subspecies of alesi; checklist; Japan). AKIYAMA & OHMOMO, 1997: 29 [#4091] (subspecies of alesi; checklist; Japan: Kyushu, Tanegashima I., Yakushima I., Ryukyus (Amami oshima I.)). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Sichuan). JENDEK, 2006: 396 [#7441] (subspecies of alesi; Palaearctic catalog). BELLAMY, 2008: 1960 [#8576] (subspecies of alesi; world catalog). aritai Tôyama, 1985 (Agrilus) syn. nov. TÔYAMA, 1985: 42 44 [#2924] (description). TÔYAMA, 1985: 25 [#3678] (iconography; Japan). HIRASHIMA, 1989: 322 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 29 [#4091] (checklist; Japan: Ryukyus (Ishigaki jima I.)). JENDEK, 2006: 396 [#7441] (Palaearctic catalog). BELLAMY, 2008: 1976 [#8576] (world catalog).

Type material: Agrilus alesi Obenberger, 1935 Type locality: Loo-choo: Okinawa. Type specimens: Primary: Lectotype (Fig. 42L) by present designation, ♀, (NMPC): “Okinawa Loo Choo 13-5, 1931 [h] \ Typus [p] [red label] \ Agrilus Aleši m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24282 [h] [orange label]”. Described from unknown number of syntypes. Agrilus sacchari Obenberger, 1940 Type locality: Insulae Loo-Choo; Okinawa. Type specimens: Primary: Lectotype (Fig. 52J) by present designation, ♀, (NMPC): “11/IX’19 Okinawa [h] Typus [p] [red label] \ Agrilus sacchari m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24295 [h] [orange label]”. Described from unknown number of syntypes. Agrilus aritai Tôyama, 1985 Type locality: Hirano, Ishigakijima Is. Type specimens: Primary: Holotype (Fig. 42W), ♀, (NSMT): “Hirano, Is. Ishigaki-jima August 30.1965, Y. Arita [h] \ Holotype [p] Agrilus aritai Tôyama, 1985 [h] [red label]”. Described from 1 specimen. Diagnosis: 7.2-8.5 mm; anterior pronotal lobe obvious; elytral pubescent striae narrow; pygidium extended into long spine. Species group: Occipitalis. Relevant species: A. auriventris Saunders, 1873. Specimens examined: JAPAN: Kyushu: 1 (EJCB): “Hiyamizu Kagoshima VI.5.1938 S.Hirayama”. Ryukyu isl. (Okinawa incl.): 1 ♀ (MNHN): “Archipel Liou Kiou, Ille d’Oshima, Ferrie 1895”; 1 (EJCB): “Shimama, Is.Tanegashima, Kagoshima pref., 1984.V.26, K.Mori leg.”; 1 (EJCB): “Izumi, Okinawa hontô Is., 13.VII.1984, T.Moriyama leg.”.

Adult occurence: 5-6-7-9 (n=6). Faunal records published by CHÛJÔ, 1970: [#3350] and MIWA & CHÛJÔ, 1940 [#1810] (as sachari [sic!]) are also included. Host plants: Citrus: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: JAPAN: Kyushu, Ryukyu Islands.

Agrilus alutaceicollis Obenberger Fig. 18A (habitus); Fig. 35Q (aedeagus); Fig. 42N (type) alutaceicollis Obenberger, 1930 (Agrilus) OBENBERGER, 1930: 110 111 [#2088] (description). ALEXEEV, 1979: 129 [#57] (synonym of soudeki). ALEXEEV, 1989: 484 [#64] (synonym of soudeki). JENDEK, 1994: 21 [#1355] (valid species; lectotype designation; characters; Russia: Primorsk region). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). JENDEK, 2006: 393 [#7441] (subgenus Robertius; Palaearctic catalog). BELLAMY, 2008: 1961 [#8576] (subgenus Robertius; world catalog). VOLKOVITSH, 2009: 135 [#8850] (subgenus Robertius; faunal records; Russia: Primorskiy kray).

Type material: Agrilus alutaceicollis Obenberger, 1930 Type locality: Siberia or.: Primorje. Type specimens: Primary: Lectotype (Fig. 42N), ♀, designated by JENDEK (1994: 21) [#1355], (NMPC): “Primorje [h] \ Typus [p] [red label] \ Agrilus alutaceicollis m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Diagnosis: 3.4-4.1 mm; pronotum with obvious microsculpture; elytral apices subtruncate, with spinuliform sutural margin. Species group: Betuleti. Relevant species: A. soudeki Obenberger, 1926. Specimens examined: CHINA: Beijing: 2 ♀ (EJCB): “China, Beijing distr., Yan Shan, Dongling Mts., 39º58’ N, E 115º26’ E, 1400 m, Xiaolongmen, 15 16.VI.2001”. Hebei: 1 ♂ (EJCB): “China, 1. 2. VI.2000, border Hebei Nei Mongol, road Chengde Chifeng, pass 1600 m, 41.6N, 118.2E, Jaroslav Turna leg.”. Shaanxi: 1 ♂, 1 ♀ (EJCB): “China, 1150 1300 m, Shaanxi, Qinling mts., Foping (6 km N), 20 21.VI.1998, I. H. Marshal leg.”; 1 ♀ (EJCB): “C China, Shaanxi, Qinling, Shan, 6 km N of Foping, 1150 1300m, 20 21.VI.2000, leg. C. Holzschuh”; 1 ♀ (EJCB): “China, Shaanxi, 11.VI.2000, pass 50 km S Xi’an, road Xi’an Ningshan, 2000 m, 33.8N, 108.8E, Qin Ling Shan, Jaroslav Turna leg.”; 1 ♂ (EKCS): “China Shaanxi, Lueyang, 29.5. 2.6.2005, lgt. E. Kučera”. Shanxi: 2 (EJCB): “China, Shanxi, 9.June.2000, Lüliang Shan, 1000m, road Fangshan Jiaocheng, Hengjian env., Zd. Jindra”. RUSSIA: Primorskiy kray: 1 ♀ (EJCB): “USSR, Primorskij kraj Arseneev env. 300 400 m 11. 19.VII.1990 B. Šiška & R. Čermák leg.”; 25 (EJCB): “USSR, Primorsk. kraj, Arsenev env., VI.1991, 44.11N, 133.19E, O. Šausa leg.” (JENDEK, 1994 [#1355]); 1 ♂ (EJCB): “USSR, Primorskij kraj Arseneev env., VI.1991, 44.11 N, 133.19 E, M. Štrba leg.” (JENDEK, 1994 [#1355]); 1 (EJCB): “Russia, Primorski reg., Khankayski distr., Novokachalinsk, 08.07.2003, Filimonov leg.”.

Adult occurence: 5-6-7 (n=11). Altitude range: 300-2000 m (n=7). Host plants: Unknown. Distribution: CHINA: Beijing, Hebei, Shaanxi, Shanxi. RUSSIA: Primorskiy kray.

40

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Agrilus andrusi Baudon

Adult occurence: 7 (n=1).

Fig. 23D (habitus of HT); Fig. 37X (aedeagus)

Host plants: Unknown.

andrusi Baudon, 1968 (Agrilus) BAUDON, 1968: 108, 118 120 [#162] (description). BELLAMY, 2008: 1964 [#8576] (subgenus Sinagrilus; world catalog). pseudograminis Jendek, 2007 (Agrilus) syn. nov. JENDEK, 2007: 34 [#8006] (description). BELLAMY, 2008: 2250 [#8576] (world catalog).

Distribution: RUSSIA: Primorskiy kray.

Type material: Agrilus andrusi Baudon, 1968 Type locality: Laos: Muong Panh, alt. circa 1100 m. Type specimens: Primary: Holotype (Fig. 23D), ♂, (MHNB): “Muong Panh 1.iv.64 [h] Laos (Baudon) [p] \ Type [p] [red label] \ Agrilus andrusi Type mihi [h] A, Baudon det. [p]”. Secondary: 1 PT (MHNB). Described from unknown number of specimens (HT, AT, PT). Agrilus pseudograminis Jendek, 2007 Type locality: China, Shaanxi, Qinling mts., 6 km E of Xunyangba, 1000-1300 m. Type specimens: Primary: Holotype, ♂, (EJCB): “China, 1000-1300m. Shaanxi, Qinling mts., Xunyangba (6km E), 23.v.-13.vi.1998, I. H. Marshal leg.”. Secondary: 7 PT (EJCB). Described from 8 specimens (HT, PT).

asahinai Kurosawa, 1956 (Agrilus) KUROSAWA, 1956: 38 40 [#1614] (description). KUROSAWA, 1963: 153 [#1618] (characters; Japan). KUROSAWA, 1974: 3 [#4575] (characters; notes). TÔYAMA, 1985: 30 [#3678] (iconography; Japan). LI JINGKE, 1992: 93 [#6755] (checklist; northeastern China). AKIYAMA & OHMOMO, 1997: 29 [#4091] (checklist; Russia: south Sakhalin). JENDEK, 2000: 503 [#4851] (synonymy). JENDEK, 2006: 390 [#7441] (subgenus Dentagrilus; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (checklist; Japan). BELLAMY, 2008: 1977 [#8576] (subgenus Dentagrilus; world catalog). JENDEK & GREBENNIKOV, 2009: 44, 53 54 [#8561] (characters in key; faunal records; synonymy; distributional and biological summary; China (Hebei, Heilongjian); Japan (Honshu); Korea North; Korea South; Russia (Primorsky kray, Sakhalin)). VOLKOVITSH, 2009: 134 [#8850] (subgenus Dentagrilus; checklist; Russia: Primorskiy kray). pseudorotundicollis Kurosawa, 1956 (Agrilus; subspecies of asahinai) KUROSAWA, 1956: 40 [#1614] (description). KUROSAWA, 1963: 153 [#1618] (subspecies of asahinai; characters; Japan). KUROSAWA, 1974: 3 [#4575] (subspecies of asahinai; characters; notes). TÔYAMA, 1985: 30 [#3678] (subspecies of asahinai; iconography; Japan). HIRASHIMA, 1989: 322 [#8078] (subspecies of asahinai; checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (subspecies of asahinai; checklist; Japan). AKIYAMA & OHMOMO, 1997: 29 [#4091] (subspecies of asahinai; checklist; Japan: Honshu). JENDEK, 2000: 503 [#4851] (synonym of asahinai; notes). JENDEK, 2006: 390 [#7441] (synonym of asahinai; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (synonym of asahinai). BELLAMY, 2008: 1977 [#8576] (synonym of asahinai; world catalog). JENDEK & GREBENNIKOV, 2009: 53 [#8561] (synonym of asahinai; HT examined). stepanovi Alexeev, 1979 (Agrilus; subgenus Dentagrilus) ALEXEEV, 1979: 134 136 [#57] (description). ALEXEEV, 1989: 483 [#64] (key to species; YU Primorsk; northeastern China). ALEXEEV, 1998: 378 [#4322] (subgenus Dentagrilus; subgeneric classification). JENDEK, 2000: 503 [#4851] (synonym of asahinai). JENDEK, 2006: 390 [#7441] (synonym of asahinai; Palaearctic catalog). BELLAMY, 2008: 1977 [#8576] (synonym of asahinai; world catalog). JENDEK & GREBENNIKOV, 2009: 53 54 [#8561] (synonym of asahinai; HT examined).

Diagnosis: 5.2-6.8 mm; elytral apices conjointly arcuate; prosternal process obviously enlarged. Species group: Sulcicollis. Relevant species: A. ribbei Kiesenwetter, 1879. Specimens examined: Known only from the type specimens.

Adult occurence: 4-5-6 (n=2). Altitude range: 1000-1300 m (n=2). Host plants: Unknown. Distribution: CHINA: Shaanxi. LAOS: Xiangkhoang.

Agrilus arsenevi Jendek Fig. 19E (habitus of HT); Fig. 36F (aedeagus) arsenevi Jendek, 2009 (Agrilus) JENDEK in JENDEK & GREBENNIKOV, 2009: 45, 55 56 [#8561] (description).

Type material: Agrilus arsenevi Jendek, 2009 Type locality: SU, Ussuri region, Jasnoe (= Yasnoe, 43°40’N, 134°00’E). Type specimens: Primary: Holotype (Fig. 19E), ♂, (EJCB): “SU, Ussuri reg., Jasnoe, 12–19.vii.1989, R. Červenka leg.”. Secondary: 1 PT (EJCB). Described from 2 specimens (HT, PT). Diagnosis: 6.6-6.9 mm; prosternal lobe deeply arcuately emarginate; median lobe of aedeagus very distinctive (Fig 36F). Species group: Cyanescens. Relevant species: A. asahinai Kurosawa, 1956; A. bidentulus Ganglbauer, 1889; A. cyanescens (Ratzeburg, 1837). Specimens examined: RUSSIA: Primorskiy kray: 1 ♂ PT (EJCB): “SU, Ussuri reg., Jasnoe, 500 m, 12 19.vii.1989, M. Nikodým leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]).

Agrilus asahinai Kurosawa Fig. 19F (habitus); Fig. 36G (aedeagus); Figs. 42X, 51K, 53G (types)

Type material: Agrilus asahinai Kurosawa, 1956 Type locality: Shirakaba, S. Saghalien. Type specimens: Primary: Holotype (Fig. 42X), ♀, (NSMT): “Konuma S-Saghalien VIII. 1932 S. Asahina [h] \ Holotype [p] Agrilus asahinai Y. Kurosawa, 1956 [h] [red label]”. Described from 2 specimens (HT, AT). Agrilus asahinai pseudorotundicollis Kurosawa, 1956 Type locality: Serio, Kyôto, Japan. Type specimens: Primary: Holotype (Fig. 51K), ♀, (NSMT): “SERIO KYOTO 9. VI. 1954 F. Takahashi lgt. [h] \ Holotype [p] Agrilus asahinai pseudorotundicollis Y. Kurosawa, 1956

41

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA [h] [red label]”. Described from 1 specimen (HT). Agrilus stepanovi Alexeev, 1979 Type locality: Primorskii kraj, Lazovskii r-n, pritok r. Kievka [in Russian]. Type specimens: Primary: Holotype (Fig. 53G), (ZIN). Secondary: 1 PT (ZIN); 4 PT (ZIN); 4 PT (AAOR). Described from 11 specimens (HT, PT). Diagnosis: 5.1-7.0 mm; sides of median lobe aedeagus denticulate (Fig. 36G); metacoxal plates sharply protruding. Species group: Cyanescens. Relevant species: A.

bidentulus Ganglbauer, 1889. Specimens examined: CHINA: Hebei: 1 ♂, 1 ♀ (EJCB): “China, Hebei (Nei Mongol) pass Chengde Chifeng 41.6N, 118.2E, 30 31.V., J.Turna leg. 2002” (JENDEK & GREBENNIKOV, 2009 [#8561]). KOREA NORTH: Hwanghae-namdo: 1 ♀ (EJCB): “Korea, Sujang san bei Hedzu, 4.6.1975, Josifov” (JENDEK & GREBENNIKOV, 2009 [#8561]). KOREA SOUTH: 2 (JLCS): “Aanmyeondo, Chungnam Prov., Korea, 23.v.2004, S. L. An leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]). RUSSIA: Primorskiy kray: 1 ♂, 1 ♀ (EJCB): “Primorskii kr. Lazovskii r n., d. Ta chin, 2 vi 1951, V. N. Stepanov” (JENDEK & GREBENNIKOV, 2009 [#8561]); 17 (EJCB): “Sib. or. Primorje, Komarovka flum., Kamenushka env. 300 m, Voříšek, J. leg. VI.1992” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 (MNCA): “Russia: Primorskiy Kray, Ussuriysky Zapovednik, 33 km SE Ussuriysk, 43.37N, 132.18E, 10/11.VI.1993, 300 m, leg. L. Zerche et al.” (JENDEK & GREBENNIKOV, 2009 [#8561]).

Adult occurence: 5-6-7-8 (n=8). Faunal records published by ALEXEEV, 1979 [#57] (as stepanovi) and KUROSAWA, 1956 [#1614] are also included. Altitude range: 300-1600 m (n=3). Host plants: Lonicera maackii: ALEXEEV, 1979 (as stepanovi) [#57]; ALEXEEV, 1989 (as stepanovi) [#64]. Distribution: CHINA: Hebei, Heilongjiang. JAPAN: Honshu. KOREA NORTH. KOREA SOUTH. RUSSIA: Primorskiy kray, Sakhalinskaya oblast’.

Agrilus ashlocki Baudon Fig. 10B (habitus); Fig. 32Q (aedeagus); Fig. 42Y (type) ashlocki Baudon, 1968 (Agrilus) BAUDON, 1968: 63, 67 68 [#162] (description). 1977 [#8576] (world catalog).

BELLAMY, 2008:

Type material: Agrilus ashlocki Baudon, 1968 Type locality: Laos: Paksong, alt. circa 1100 m. Type specimens: Primary: Holotype (Fig. 42Y), ♀, (MHNB): “Pakson VII.[19]65 [h] Laos (Baudon) \ Type [p] [red label] \ Agrilus ashlocki nov. [h] A. Baudon det. [p] [blue label]”. Described from: 1 specimen (HT). Diagnosis: 7.6-8.1 mm; head with deep medial impression; elytral apices with tomentose patches; aedeagus with large membranous lobes. Species group: Spinipennis. Relevant species: A. tokyoensis Kurosawa, 1985 Specimens examined: CHINA: Yunnan: 1 ♂ (EJCB): “Yunnan, Jing Gu County, Mt. Yingpantou, 7 14.vi.2009 local collector”. LAOS: Houaphan: 1 ♂ (EJCB): “Lao NE, Hua Phan pr.,

~20°12’N,104°01’E, Phu Phan Mt., 1500 1900m, 17.v. 3.vi.2007, Vít Kubáň leg.”.

Adult occurence: 5-6 (n=2). Altitude range: 1100-1900 m (n=2). Host plants: Unknown. Distribution: CHINA: Yunnan. LAOS.

Agrilus asiaticus Kerremans Fig. 25B (habitus); Fig. 38U (aedeagus); Figs. 42Z, 47B, 50W (types) asiaticus Kerremans, 1898 (Agrilus) KERREMANS, 1898: 178 [#1427] (description). HEYDEN, 1898: 21 [#5553] (catalog). KERREMANS, 1903: 276 [#1441] (catalog). JAKOBSON, 1913: 796 [#4386] (catalog; Russia and Europe). BODEMEYER, 1915: 447 [#8261] (faunal record; China: Heilongjiang). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 950 [#2191] (world catalog). KUROSAWA, 1974: 3 [#4575] (notes). KUROSAWA, 1976: 132 [#1627] (notes; type in BMNH examined). KUROSAWA, 1976: 132 133 [#1628] (note). ALEXEEV, 1979: 132 [#57] (subspecies of tibialis; notes). TÔYAMA, 1985: 29 [#3678] (iconography; Japan). ANONYMOUS, 1986: 177 [#8848] (insect pest; Korea). PENG ZHONGLIANG, 1987: 354 [#2429] (checklist; China). ALEXEEV, 1989: 480 [#64] ([Note: Misidentification of ribbei]; characters in key). LI JINGKE, 1992: 93 [#6755] (checklist; northeastern China). JENDEK, 1994: 14 15 [#1355] (synonyms; characters; notes; Russian Far East; northern China; Korea; Japan). AKIYAMA & OHMOMO, 1997: 29 [#4091] (checklist; Korea; northeastern China; Russia: East Siberia). ALEXEEV, 1998: 378, 379 [#4322] (subgenus Pseudoquercagrilus; genotype of the subgenus Pseudoquercagrilus). NONNAIZAB et al., 1999: 112 [#8247] (checklist; China: Inner Mongolia). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China). JENDEK, 2006: 391 [#7441] (subgenus Pseudoquercagrilus; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (checklist; Japan). BELLAMY, 2008: 1977 1978 [#8576] (subgenus Pseudoquercagrilus; world catalog). VOLKOVITSH, 2009: 135 [#8850] (subgenus Pseudoquercagrilus; faunal records; Russia: Primorskiy kray). hauserellus Obenberger, 1935 (Agrilus) OBENBERGER, 1935: 171 [#2158] (description). OBENBERGER, 1936: 997 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 355 [#2429] (checklist; China). ALEXEEV, 1989: 481 [#64] (key to species; Primorsk; China). JENDEK, 1994: 14, 15 [#1355] (synonym of asiaticus; lectotype designation). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Jiangxi). JENDEK, 2006: 391 [#7441] (synonym of asiaticus asiaticus; Palaearctic catalog). BELLAMY, 2008: 1977 [#8576] (synonym of asiaticus; world catalog). planefasciatus Obenberger, 1936 (Agrilus) OBENBERGER, 1936: 115 [#2205] (description). OBENBERGER, 1936: 1022 [#2191] (world catalog). KUROSAWA, 1949: 14, 28 [#5028] (characters). IGA, 1955: 79 [#3828] (iconography; Japan). IGA, 1962: 79 [#3351] (iconography; Japan). KUROSAWA, 1963: 153 [#1618] (characters; Japan). KUROSAWA, 1974: 3 [#4575] (notes). KUROSAWA, 1976: 132 [#1628] (synonym of asiaticus; type in NMPC examined). ALEXEEV, 1989: 481 [#64] (synonym of hauserellus). JENDEK, 1994: 14, 15 [#1355] (synonym of asiaticus; lectotype designation; notes). AKIYAMA & OHMOMO, 1997: 29 [#4091] (synonym of asiaticus). JENDEK, 2006: 391 [#7441] (synonym of asiaticus asiaticus; Palaearctic catalog). BELLAMY, 2008: 1977 [#8576] (synonym of asiaticus; world catalog).

42 panphilovi Alexeev, 1959 (Agrilus) ALEXEEV, 1959: 28 30 [#3231] (description). ALEXEEV, 1979: 137 138 [#57] (cited as species incertae sedis; redescription; notes). ALEXEEV, 1989: 481 [#64] (synonym of hauserellus). JENDEK, 1994: 14 [#1355] (synonym of asiaticus). JENDEK, 2006: 391 [#7441] (synonym of asiaticus asiaticus; Palaearctic catalog). BELLAMY, 2008: 1977 [#8576] (synonym of asiaticus; world catalog). igai Kurosawa, 1963 (Agrilus; subspecies of planefasciatus) KUROSAWA, 1963: 110 111 [#1617] (description). KUROSAWA, 1963: 153 [#1618] (subspecies of planefasciatus; characters; Japan). KUROSAWA, 1974: 3 [#4575] (subspecies of asiaticus; characters; notes). KUROSAWA, 1976: 133 [#1628] (subspecies of asiaticus). TÔYAMA, 1985: 29 [#3678] (subspecies of asiaticus; iconography; Japan). AKIYAMA, 1986: 25 [#3199] (subspecies of asiaticus; faunal records; Japan: Kanagawa prefecture). HIRASHIMA, 1989: 322 [#8078] (subspecies of asiaticus; checklist; Japan). JENDEK, 1994: 14, 15 [#1355] (synonym of asiaticus; notes). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (subspecies of asiaticus; checklist; Japan). AKIYAMA & AKIYAMA, 1996: 184 [#3962] (subspecies of asiaticus; faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 5, 29 [#4091] (subspecies of asiaticus; checklist; Japan: Honshu, Shikoku, Kyushu, Tsushima). AKIYAMA & OHMOMO, 2000: 87 [#6240] (subspecies of asiaticus; faunal records; Japan (Hiroshima)). JENDEK, 2006: 391 [#7441] (subgenus Pseudoquercagrilus; subspecies of asiaticus; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (synonym of asiaticus). BELLAMY, 2008: 1978 [#8576] (subspecies of asiaticus; world catalog).

Type material: Agrilus asiaticus Kerremans, 1898 Type locality: Mandchourie. Type specimens: Primary: Lectotype (Fig. 42Z), ♂, designated by KUROSAWA (1976: 132) [#1627], (BMNH): “Holo-type [p] [round label with red border] \ Mandchourie Boucard [h] \ asiaticus Kerr. Type [h] \ Kerremans 1903 59 [p]”. Described from unknown number of syntypes. Agrilus hauserellus Obenberger, 1935 Type locality: China: Kiang-Si. Chang-tsin-cheng. Type specimens: Primary: Lectotype (Fig. 47B), ♂, designated by JENDEK (1994: 14, 15) [#1355], (NMPC): “Kiang-Si Chang-tsin-cheng [p] \ Typus [p] [red label] \ Agrilus hauserellus m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24942 [h] [orange label]”. Described from unknown number of syntypes. Agrilus planefasciatus Obenberger, 1936 Type locality: Primorje: Trojckoje. Type specimens: Primary: Lectotype (Fig. 50W), ♀, designated by JENDEK (1994: 14, 15) [#1355], (NMPC): “Trojckoje Primorje [h] \ Typus [p] [red label] \ Agrilus planefasciatus m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24333 [h] [orange label]”. Described from unknown number of syntypes. Agrilus panphilovi Alexeev, 1959 Type locality: Sudzukhinskii zapovednik, bukhta Kit. [in Russian]. Type specimens: Not located. Described from 1 specimen (Holotype by monotypy). Agrilus planefasciatus igai Kurosawa, 1963 Type locality: Ôsaka Pref.: Myôken.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Type specimens: Not located. Described from unknown number of specimens (HT, AT, PT). Diagnosis: 5.7-8.9 mm; pronotum with medial and lateral impressions; medial impression disrupted in middle; margin of elytral apices with obvious denticulation; tubercles on basal abdominal ventrite transverse. Species group: Tempestivus. Relevant species: A. tempestivus Lewis, 1893 Specimens examined: CHINA: Beijing: 1 ♀ (EJCB): “China, Beijing distr., Yan Shan, Dongling Mts., 39º58’ N, E 115º26’ E, 1400 m, Xiaolongmen, 15 16.VI.2001”. Hebei: 1 ♀ (EJCB): “China, 3 June 2000, Fengning env. 1200m, NW slope of Yunwu Shan, Zd. Jindra lgt.”. Henan: 4 ♂, 3 ♀ (EJCB): “China, N. Henan, 11 12.VI., Jiuligou, 550 m, 35º12’N, 112º26’E, Jaroslav Turna leg. 2007”. Nei Mongol: 1 ♂ (EJCB): “Chahar Yangkiaping 15.VII.1931”. Shaanxi: 1 ♀ (EJCB): “China Shaanxi, Lüeyang, 8.6. 14.6.1996, lgt. E. Kučera”; 1 ♂, 2 ♀ (EJCB): “China, 1000 1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v. 13.vi.1998, I. H. Marshal leg.”; 1 ♂, 3 ♀ (EJCB): “China, 1150 1300 m, Shaanxi, Qinling mts., Foping (6 km N), 20 21.VI.1998, I. H. Marshal leg.”; 2 ♂, 3 ♀ (EJCB): “C China, Shaanxi, Qinling, Shan, 6 km N of Foping, 1150 1300m, 20 21.VI.2000, leg. C. Holzschuh”; 1 (EKCS): “China Shaanxi, Lueyang, 23 30.5.2009, lgt. E. Kučera”. Sichuan: 1 (GNCW): “China Pr. Sichuan nord, Jiuzhaigon [sic!], 4 7.Juli 1994, 80 km n Songpan, 3000 m, leg. Beneš”; 1 ♂, 1 ♀ (MKCY): “China N, Sechuan, Nanjiang, 21 23.5.2002, lgt. E. Kučera”. JAPAN: Honshu: 1 ♀ (EJCB): “1982.7.10, Mts. Abukuma, Fukushima Pref., Col. S. Ohmomo”; 1 ♂ (EJCB): “Japan, Osaka Pref. Iwakiyama, Amami 16.Jun 2.Jul 2002, 450 m, Bolm Leg.”; 1 ♂, 1 ♀ (EJCB): “Japan, Honshu, Osak fu, Kongosan, 300 m, 2 km E of Amami, 26. May 2002, Bolm lgt.”; 1 ♀ (EJCB): “Japan 16.6.2002 Amami, L. Bocák leg.”. Tsushima: 1 ♂, 1 ♀ (EJCB): “1975 VI., Mt. Ohboshi Tsushima Is. Nagasaki Pref. col. H. Makihara” (Jendek, 1994 [#1355]); 1 (NMPC): “Japan, Tsushima is., (Sasuna) 5 7.6.1981, A. Torikata leg.”; 1 ♀ (EJCB): “1986.7.19., Sasuna, Tsushima Is., Nagasaki Pref. Col. S. Urushibata” (Jendek, 1994 [#1355]). LAOS: Houaphan: 1 ♂, 3 ♀ (EJCB): “NE Laos, Hua Phan prov., Ban Saluei, 1300 2000m, 20°13’N, 103°59’E, 6 18.iv., Phu Phan Mt. env. 2004, F. & L. Kantner leg.”; 14 (EJCB): “Laos NE, Houa Phan prov., 20°12 13.5’N, 103°59’.5 104°01’E, Ban Saluei → Phou Pane Mt., 1340 1870m, 15.iv. 15.v.2008, Lao collectors leg.”. RUSSIA: Primorskiy kray: 1 ♂, 1 ♀ (EJCB): “Sib. or. m., Primorje Sichote Alin Mts. Sokolči 1. 15.7.1990 Kadlec + Voříšek lg.” (Jendek, 1994 [#1355]); 1 (EJCB): “Prim. Khasansk. Sukhanovskij per. 9.8.[19]90 Napolov” (Jendek, 1994 [#1355]); cca 50 (EJCB): “Russia or, Primorskiy Kray, 44°10’00’’N, 133°15’00’’E, ARSEN’YEV env., June 1991, O. Šauša & M. Štrba leg.”; 1 ♂ (EJCB): “USSR, Primorsk. kraj, Arsenev env., VI.1991, 44.11N, 133.19E, O. Šausa leg.” (Jendek, 1994 [#1355]); 1 (EJCB): “Rossia, Primorsk, Kraskino env. mer., 13 16.VII.1992, P. Smrž leg.” (Jendek, 1994 [#1355]); 1 (EJCB): “Rossia, Primorskij k., Rjazanovka 4 8.vii.1993, Z. Kletečka leg.”; 1 ♀ (JMCZ): “Russia, Ussuri, Kajmanovka, VIII.1994, Snížek”; 4 (FACF): “Russia Far East, S Primorie, Hanka lake, 30.vii.2002”; 1 (MKCY): “Russia Far East, Khanka Lake NW 0 3 km S Turiy Rog, 8.VII.2003, Lg. A. Napolov”; 1 (MKCY): “Russia Far East, 20 km SE Ussuriysk, Gornotaezhnoje vill. env., 27.06.2003, Lg. A. Napolov”; 2 (EJCB): “Russia Far East, Khanka Lake NW Novokachalinsk vill. env., 7.7.2003, Lg. A. Napolov”; 1 (MKCY): “Russia Far East, 20 km SE Ussuriysk, Gorno Tajozhnoje vill. env., 27.VI.2003, Lg. A. Napolov”; 4 (MKCY): “Russia, Primirski reg. Khankayski distr., Novokachalinsk, 07.07.2003, Filimonov leg.”; 1 (MKCY): “Russia, Primorski reg. Khankayski distr., Novokachalinsk, 05.07.2003, Filimonov leg.”; 1 (MKCY): “Russia Far East, Khanka Lake NW Novokachalinsk vill. env., 4.7.2003, Lg. A. Napolov”; 2 (EJCB):

43

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA “Russia, Far East, Lazovkiy reserve, 17 26.06.2007, V. Shokhrin”. VIETNAM: 1 ♀ (EJCB): “N. Vietnam, 14.5., Shon La prov., Moe Tjau, 1991”.

Adult occurence: 4-5-6-7-8 (n=37). Altitude range: 3003000 m (n=11). Host plants: Quercus: ALEXEEV, 1979 (as panphilovi) [#57]. − Q. acutissima: KUROSAWA, 1963 (as igai) [#1617]; AKIYAMA & OHMOMO, 1997 (as igai) [#4091]. − Q. mongolica: ALEXEEV, 1989 (as hauserellus) [#64]. − Q. variabilis: KUROSAWA, 1976 (as igai) [#1628]; AKIYAMA & OHMOMO, 1997 (as igai) [#4091]. Distribution: CHINA: Beijing, Hebei, Heilongjiang, Henan, Jiangxi, Nei Mongol, Shaanxi, Sichuan. JAPAN: Honshu, Kyushu, Shikoku, Tsushima. KOREA NORTH. KOREA SOUTH. RUSSIA: Primorskiy kray. LAOS. VIETNAM.

Agrilus aureofasciatus Jendek, sp. nov. Fig. 13L (habitus of HT); Fig. 34F (aedeagus)

Description: 5.6-9.3 mm (Holotype 8.1 mm). BODY: Shape: cuneiform, Color (dorsally): bicolored, Color (dominant): brown. HEAD: Medial impression: present, Impression (extent): vertex, Epistoma: in plane with frons. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): subparallel, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: moderate, Shape: protruding from head outline, Lower margin: in line with antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: vague, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: blunt. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): entire or anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow. Prehumerus: filamentary or carinal, Shape: bisinuate, Extent: to anterior pronotal angle, Posterior end: distant from pronotal angle or margin, Anterior end: joining anterior pronotal angle, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: absent, rarely present, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina divergent at posterior angles. Scutellum: Size: moderate or large, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: bicolored, complementary with pubescence, Humeral carina: absent, Apical part: sometimes spatulate. Apices: Arrangement: separate, Shape: subtruncate or cuspidate, Truncation: oblique with projecting pleural angle, Position of dominant cusp (spine): pleural margin, Modifications: margin obviously denticulate. Elytral pubescence: ornamental, Color: unicolored, Character: homogenous. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal

lobe: moderate, Anterior margin: arcuate. Prosternal process: moderate, Shape: subparallel, Sides: straight or arcuate, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: elongate. Diagnosis: glabrous portions of elytra black with blue tinge; elytral apices more elongate; aedeagus slender, slightly dilated apically. Species group: Plagiatus. Relevant species: A. radiolus Kerremans, 1913. Type locality: North Laos, 20 km northwest of Louang Namtha, N21°09.2, E101°18.7, altitude 900±100 m. Type specimens: Primary: Holotype (Fig. 13L), ♂, (EJCB): “Laos north, 5-11.V.1997, 20 km NW Louang Namtha, N 21°09.2, E 101°18.7, alt. 900±100 m, E. Jendek & O. Šauša leg.”. Secondary: CHINA: Guizhou: 1 PT (MNHN): “Kouy-Tcheou, Reg. de Pin Fa”; Yunnan: 1 ♀ PT (NMPC): “China: Yunnan province, Tengchong, 9-10.vi.2007, Laifeng Shan nat. Forest Park, 25°01.1’N, 098°29.1’E, 1700 m, J. Hájek & J. Růžička leg. (CH16).”; INDIA: West Bengal: 1 PT (MNHN): “British Bootang, Maria Basti, 1899”; LAOS: Borikhamxai: 3 ♂ PT (EJCB): “Laos centr., Bolikhamsai prov., Ban Nape - Kaew Nua Pass, 18.4.-1.5.1998, alt. 600±100 m, N 18°22.3, E 105°09.1 (GPS), E. Jendek & O. Šauša leg.”; Louang Namtha: 15 PT (EJCB): “Laos north, 5-11.V.1997, 20 km NW Louang Namtha, N 21°09.2, E 101°18.7, alt. 900±100 m, E. Jendek & O. Šauša leg.”; 1 PT (EJCB): “Laos north, 13-24.V.1997, 15 km NW Louang Namtha, N 21°07.5, E 101°21.0, alt. 750±100 m, E. Jendek & O. Šauša leg.”; 19 PT (EJCB): “Laos, Louang Namtha pr., 21°09’N, 101°19’E, Namtha - Muang Sing, 5-31.v.1997, 9001200 m, Vit Kubáň leg.”; Louangphrabang: 1 ♂ PT (EJCB): “Laos-N, 24.iv.- 16.v.1999, Louang Phrabang prov. 20°33-4’ N 102°14’E, Ban Song Cha (5km W) ± 1200 m, Vít Kubáň leg.”; Oudomxai: 1 ♀ PT (EJCB): “Laos-N (Oudomxai), 1-9.v.2002, ~1100m, 20°45’N, 102°09’E, Oudom Xai (17 km NEE), Vít Kubáň leg.”; Phongsali: 1 ♀ PT (EJCB): “Lao, Phongsaly prov., 21°21’N, 102°03’E, Ban Sano Mai, 19-26.v.2004, 1150 m, P. Pacholátko leg.”; NEPAL: Kosi: 1 PT (EJCB): “E-Nepal, Dhankuta, Arun-Valley, Lamobagar Gola, 27.5.-3.6.1980, 1000-1400 m, leg. C. Holzschuh”; 3 ♀ PT (EJCB): “E-Nepal, Koshi, Tamur River, Phulvari-Waku, 1200-1600 m, 9.VI., leg. Holzschuh, 1985”. Adult occurence: 4-5-6 (n=10). Altitude range: 500-1700 m (n=10). Host plants: Unknown. Distribution: CHINA: Guizhou, Yunnan. INDIA: West Bengal. LAOS: Borikhamxai, Louang Namtha, Louangphrabang, Oudomxai, Phongsali. NEPAL: Kosi. Etymology: The specific name is a combination of the Latin adjectives aureus (golden) and fasciatus (striped). It refers to the elytral pubescence of this species.

44

Agrilus aurigaster Jendek, sp. nov. Fig. 14I (habitus of HT); Fig. 34M (aedeagus)

Description: 5.4-7.5 mm (Holotype 5.4 mm). BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): blue. HEAD: Shape: markedly convex, Medial impression: absent or present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): subparallel, Sculpture (intensity): superficial. Eyes: Size: large, Shape: protruding from head outline, Lower margin: in line with antennal socket. Antennae: Length: short, Shape: stout, Sexual modification in male: not apparent. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, rarely at anterior margin, Anterior margin: subequal to posterior. Anterior lobe: absent or vague, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: strongly convex, Disk impressions: absent. Prehumerus: absent. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior third of pronotal length, Modifications: submarginal carina posteriorly obliterate or submarginal carina divergent at posterior angles. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: markedly attenuate. Apices: Arrangement: separate, Shape: arcuate, Modifications: margin obviously denticulate. Elytral pubescence: absent. Tomentum: absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: large, Anterior margin: arcuate, rarely arcuately emarginate, Emargination (deepness): shallow, Emargination (width): narrow. Prosternal process: moderate, Shape: narrowed, Sides: straight, Angles: absent, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): weakly sclerotised and parameres with obvious membranous lobes, Ovipositor: uritiform. Diagnosis: body dorsally dark blue; elytra glabrous; ventral side golden-cupreous. Species group: Subrobustus. Relevant species: A. yunnanicola Obenberger, 1936. Type locality: China, Yunnan province, Gaoligong mts., 25°22’N, 98°49’E, altitude 1500-2500 m. Type specimens: Primary: Holotype (Fig. 14I), ♂, (EJCB): “Yunnan, 1500-2500 m, 25°22’N, 98°49’E, 17-24 / 5 Gaoligong mts, Vít Kubáň leg., 1995”. Secondary: CHINA: Yunnan: 1 ♀ PT (EJCB): “Yunnan, 1500-2500 m, 25°22’N, 98°49’E, 17-24 / 5 Gaoligong mts, Vít Kubáň leg., 1995”;

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA MYANMAR: 1 ♀ PT (EJCB): “Burma, SW Shan State, Taunggyi, 1-18.vi.1997, J. Kaláb”. Adult occurence: 5-6 (n=2). Altitude range: 1500-2500 m (n=1). Host plants: Unknown. Distribution: CHINA: Yunnan. MYANMAR. Etymology: The specific name is a combination of the Latin adjective aureus (golden) and the Greek noun gaster (belly). It refers to the golden color of the abdominal ventrites of the species.

Agrilus aurinotus Jendek, sp. nov. Fig. 15K (habitus of HT); Fig. 34Y (aedeagus)

Description: Holotype 8.8 mm. BODY: Color (dorsally): bicolored, Color (dominant): black. HEAD: Medial impression: present, Impression (extent): vertex, Epistoma: in plane with frons. Frons: Shape: flat, Outline: not protruding from head outline. Vertex: Outline: protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): subparallel, Sculpture (density): sparse, Sculpture (intensity): superficial. Eyes: Size: small, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: long. PRONOTUM: Shape: visually elongate, Sides: arcuate, Maximal width: at middle, Anterior margin: wider than posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: flat, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial, Lateral impressions (deepness): deep. Prehumerus: obsolete or carinal, Shape: arcuate, Extent: to third of pronotal length, Posterior end: joining posterior pronotal margin, Anterior end: distant from pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: narrow or moderate, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Shape: arcuate. Tomentum: spots, Spots (locations): premedial or postmedial. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): moderate. Prosternal process: moderate, Shape: narrowed, Sides: arcuate, Angles: absent, Disk: flat, Modification: disk in male with transverse rugae. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus

45

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed. Diagnosis: dorsal side black except for golden-green pronotal sides; pronotum visually elongate with distinct anteromedial and posteromedial impressions. Species group: Smaragdinus. Relevant species: A. odetteae Baudon, 1968. Type locality: China, Guizhou province, Leishan Co., southestern Kaili, 15 km eastern Leishan, Leigong Shan, eastern slope, 26°23.39’N, 108°13.33’E, about 2.5 km eastern of pass between Leishan and Fangxiang village, altitude about 1600 m. Type specimens: Holotype (Fig. 15K), ♂, (NMW): “China: Guizhou, Leishan Co., SE Kaili, 15 km E Leishan, Leigong Shan, E-slope, 26°23.39’N, 108°13.33’E / ca 2.5 km E of pass between Leishan and Fangxiang vill., ca 1600 m, 23/24.6.2001, leg. Schillhammer (17)”. Adult occurence: 6 (n=1). Altitude range: 1600 m (n=1). Host plants: Unknown. Distribution: CHINA: Guizhou. Etymology: The specific name is a combination of the Latin adjective aureus (golden) and the Greek noun notos (back). It refers to the golden color of the pronotum of this species.

Agrilus auristernum Obenberger Fig. 8C (habitus); Fig. 43E (type) auristernum Obenberger, 1924 (Agrilus) OBENBERGER, 1924: 34 35, 37 [#2040] (description). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 958 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 354 [#2429] (cited as auristermum; checklist; China). JENDEK, 2000: 188, 190 [#4852] (lectotype designation; faunal records; key). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China). JENDEK, 2006: 396 [#7441] (Palaearctic catalog). BELLAMY, 2008: 1984 [#8576] (world catalog).

Type material: Agrilus auristernum Obenberger, 1924 Type locality: Yünan (China). Type specimens: Primary: Lectotype (Fig. 43E), sex not examined, designated by JENDEK (2000: 188, 190) [#4852], (NMPC): “Pe-yen-tsing [h] Yunnan \ Typus [p] [red label] \ Agrilus auristernum m. Type [Obenberger’s MS] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24789 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 13-15 mm; body markedly elongate; dorsally blue or green; pronotum enlarged anteriorly; elytra with pair of adsutural tomentose spots in apical half. Species group: Cyaneoniger. Relevant species: A. cyaneoniger Saunders, 1873. Specimens examined: CHINA: Yunnan: 1 ♀ (EJCB): “Djoukoula Yunnan” (JENDEK, 2000 [#4852]); 1 ♀ (EJCB): “Yunnanfou” (JENDEK, 2000 [#4852]); 1 ♀ (EJCB): “China, Yunnan, 6 10.VII, Ninglang env. 27.19N, 100.55E, E. Jendek leg., 1992” (JENDEK, 2000 [#4852]).

Adult occurence: 7 (n=1). Host plants: Quercus: JENDEK, 2000 [#4852]. Distribution: CHINA: Yunnan.

Agrilus aurithorax Jendek, sp. nov. Fig. 18L (habitus of HT); Fig. 36A (aedeagus)

Description: 6.1-6.6 mm (Holotype 6.6 mm). BODY: Shape: subparallel, Color (dorsally): bicolored, Color (dominant): black or blue. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: in plane with frons. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Shape: markedly convex, Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): semispherical, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: moderate, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: visually square, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate or obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow. Prehumerus: obsolete or carinal, Shape: arcuate, Extent: to third of pronotal length, Posterior end: distant from pronotal angle or margin, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate, Width: wide, Shape: arcuate. Elytral pubescence: absent or distal only, Distal: perisutural stripe, Color: unicolored. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: large, Anterior margin: arcuate, subtruncate or arcuately emarginate, Emargination (deepness): shallow, Emargination (width): wide. Prosternal process: moderate, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: medial strip of erect hairs on apical 2 ventrites, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed. Diagnosis: body bicolored, head and pronotum goldenorange, elytra blue; prehumerus obtusely carinal or obsolete,

46 feebly arcuate; aedeagus dilated apically. Species group: Laetecyanescens. Relevant species: A. laetecyanescens Obenberger, 1940. Type locality: China, Fujian province, Wuyi Shan Nat. Res., Sangan env., altitude 900 m. Type specimens: Primary: Holotype (Fig. 18L), ♂, (EJCB): “China, Fujian prov., Wuyi Shan Nat. Res., Sangan env. (900 m), 30.v.-12.vi.2001”. Secondary: CHINA: Beijing: 1 ♂ PT (EJCB): “China, Beijing distr., Yan Shan, Dongling Mts. (1400 m), Xiaolongmen, 15-16.vi.2001”; Guangxi: 1 ♂ PT (EJCB): “Guangxi, Lingchuan, 1984.vi.7”; LAOS: Houaphan: 1 ♂ PT (EJCB): “Laos-NE, Houa Phan prov., 20°12-13.5’N, 103°59’.5-104°01’E, Ban Saluei → Phou Pane Mt., 1340-1870m, 15.iv.-15.v.2008, Lao collectors leg.”. Adult occurence: 4-5-6 (n=4). Altitude range: 900-1870 m (n=3). Host plants: Unknown. Distribution: CHINA: Beijing, Fujian, Guangxi. LAOS: Houaphan. Etymology: The specific name is a combination of the Latin adjective aureus (golden) and the Latin noun thorax. It refers to the golden color of the thoracal portions of this species.

Agrilus auriventris Saunders Fig. 19J (habitus); Fig. 36K (aedeagus); Figs. 43F, 46J, 46U, 50R (types) auriventris Saunders, 1873 (Agrilus) SAUNDERS, 1873: 517 [#2554] (description). LEWIS, 1879: 15 [#3345] (catalog; Japan). KERREMANS, 1885: 152 [#1381] (catalog). SCHÖNFELDT, 1887: 113 [#8901] (Anambus; catalog; Japan). KERREMANS, 1892: 248 [#1397] (catalog). KERREMANS, 1903: 282 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA, 1931: 126 [#6101] (catalog; Formosa). OGLOBLIN & REICHARDT, 1932: 277 [#8654] (pest). YUASA, 1932: 653 [#3369] (characters; Japan). YUASA, 1933: 281 [#3132] (larva). MIWA & CHÛJÔ, 1936: 15 [#1809] (catalog; Japan). OBENBERGER, 1936: 958 [#2191] (world catalog). CHÛJÔ & MATUDA, 1940: 65 [#500] (checklist; faunal records; Japan (Kyushu)). MIWA, 1940: 72 [#4586] (checklist; Japan). YUASA, 1949: 128 [#8108] (characters; Japan). KUROSAWA, 1950: 1115 [#3274] (note). MÜHLMANN, 1954: 83 [#8152] (pest in Asia; biology). IGA, 1955: 78 [#3828] (iconography; Japan). TER MINASYAN, 1955: 450 [#7893] (notes). KUROSAWA, 1956: 1115 [#8107] (characters; Japan). BALACHOWSKI et al., 1962: 286 [#96] (economic importance). IGA, 1962: 78 [#3351] (iconography; Japan). ISRIGAYA, 1963: 351 355 [#7016] (biology; larva; economic importance; treatment; Japan (Wakayama)). OHGUSHI, 1963: 92 96 [#5415] (pest; ovary and pre oviposition period; Japan). WOO SHEH MING, 1964: 61 71 [#7020] (larval biology; protection; China: Chekiang). OHGUSHI, 1966: 55 63 [#8729] (biology). GURYEVA, 1974: 98 [#5428] (pest of agriculture cultures; references). KUROSAWA, 1974: 2, 3 [#4575] (characters; notes). OHGUSHI, 1978: 62 73 [#7015] (population ecology; Japan: Nagasaki prefecture). TÔYAMA, 1985: 24 [#3678] (iconography; Japan). PENG ZHONGLIANG, 1987: 354 [#2429] (checklist; China). HIRASHIMA, 1989: 322 [#8078] (checklist;

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Japan). PENG ZHONGLIANG, 1992: 398 399 [#2431] (characters; notes; Hunan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 184 [#3962] (faunal records; Japan: Honshu). NAKA & OHASHI, 1996: 43 44 [#4720] (occurence; chemical control; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 29 30 [#4091] (checklist; Japan: Honshu, Kyushu; China). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Hubei, Jiangxi, Zhejiang, Fujian, Taiwan, Guandong, Hunan, Guangxi, Sichuan). LIN GUI RUI, 2002: 233 [#7926] (checklist; pest). PENG ZHONGLIANG, 2002: 268 [#6628] (characters; Fujian). JENDEK, 2006: 396 [#7441] (Palaearctic catalog). WEI SHU JUN et al., 2006: 302 308 [#7653] (larval instars and characters; China: Zheijang). ZHENG HONG HAI et al., 2006: 806 [#7883] (spatial distribution pattern; attacking strategy; China). HUANGFU et al., 2007: 682 688 [#8169] (ovarian development; ovariole; oviposition duration; fecundity). WEI SHU JUN et al., 2007: 79 84 [#8166] (immature stages; chorion; hatching; pupation). BELLAMY, 2008: 1985 [#8576] (world catalog). graptelytrus Obenberger, 1914 (Agrilus) syn. nov. OBENBERGER, 1914: 43 (in Czech), 48 49 (in German) [#1965] (description). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1084 [#2191] (world catalog). DESCARPENTRIES & VILLIERS, 1963: 104, 118 [#700] (species incertae sedis). BELLAMY, 2008: 2116 [#8576] (world catalog). fleutiauxi Bourgoin, 1922 (Agrilus) syn. nov. BOURGOIN, 1922: 23 [#365] (description). THÉRY, 1935: 132 [#2818] (faunal record; China: Kiang Si Wei Yang). OBENBERGER, 1936: 1083 [#2191] (world catalog). DESCARPENTRIES & VILLIERS, 1963: 108, 118 [#700] ([Note: An inference of holotype: “Tonkin: Cho Ganh (Duport), holotype au Muséum de Paris” is not valid lectotype designation because body size interval in original description indicates more syntypes (ICZN, Article 74.5)]; characters; faunal records; Tonkin; Formose; Chine (Hong Kong, Kiangsi); Birmanie). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2096 [#8576] (world catalog). pidjinus Obenberger, 1924 (Agrilus) syn. nov. OBENBERGER, 1924: 53 54, 58 [#2040] (description; cited as podjinus on page 53 and as pidjinus on page 58; [Note: Multiple original spelling. Precedence of the name pidjinus has been fixed by the original author OBENBERGER, 1926: 654 [#2057] as the first reviser (Article 24.2.4). The name podjinus is incorect original spelling and unavailable name (Article 32.4)])). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1022 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 357 [#2429] (checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Hongkong). JENDEK, 2006: 400 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2235 [#8576] (world catalog).

Type material: Agrilus auriventris Saunders, 1873 Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 43F) by present designation, ♂, (BMNH): “Japan G. Lewis. 1910 - 320 [p] \ Agrilus auriventris E. S. Type [h] \”. Described from unknown number of syntypes. Agrilus graptelytrus Obenberger, 1914 Type locality: China; Tonkin: Phu-long-thuan. Type specimens: Primary: Lectotype (Fig. 46U) by present designation, ♂, (NMPC): “Phu-long-thuan, Tonkin Lamey [h] Typus [p] [red label] \ Agrilus graptelytrus m. Type [h] Det. Dr Obenberger [p]”. Secondary: 1 PLT (NMPC). Described from unknown number of syntypes.

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Agrilus fleutiauxi Bourgoin, 1922 Type locality: not given [Indo-Chine française is cited in the title of the publication]. Type specimens: Primary: Lectotype (Fig. 46J) by present designation, ♂, (MNHN): “Tonkin Cho Ganh L. Duport [p] \ Agrilus Fleutiauxi Bourgoin [h] [black ink] Type [h] [red ink] \ Muséum Paris 1935 coll. A. Théry [p]”. Secondary: 2 PLT (MCSN); 3 PLT (NMPC); 11 PLT (MNHN); 1 PLT (BMNH). Described from unknown number of syntypes. Agrilus pidjinus Obenberger, 1924 Type locality: China: Hong-Kong. Type specimens: Primary: Lectotype (Fig. 50R) by present designation, ♀, (NMPC): “Hong Kong [h] Typus [p] [red label] \ Agrilus pidjinus m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Diagnosis: 5.0-8.8 mm; anterior pronotal lobe moderate; elytral pubescent striae wider; pygidium extended into short protrusion. Species group: Occipitalis. Relevant species: A. alesi Obenberger, 1935. Specimens examined: CHINA: Zhejiang: 8 (EJCB): “CH Zhejiang prov., Xiangshan, October, 2004, Wei Shujun leg.”; 16 (EJCB): “CH Zhejiang prov., Zheng Honghai, May 2004, Wei Shujun leg.”; 8 (EJCB): “CH Zhejiang prov., Zheng Honghai, August 8th, 2004, Wei Shujun leg.”. JAPAN: Kyushu: 2 ♂ (EJCB): “Taku City, Saga pref. 14.6.1992, K. Adachi leg. On Citrus sp.”. LAOS: Borikhamxai: 1 ♀ (EJCB): “Laos centr., Bolikhamsai prov., Ban Nape Kaew Nua Pass, 18.4. 1.5.1998, alt. 600±100 m, N 18º22.3, E 105º09.1 (GPS), E. Jendek & O. Šauša leg.”. VIETNAM: 1 ♂ (EJCB): “Vietnam N, Tonkin, Cuc Phuong nat. park, 2 12.V.1991, E. Jendek leg.”. 3 ♂ (EJCB): “Vietnam N, 26 29.IV.1991, 60 km SW Hanoi, Chua Huong Huong Son, E.Jendek leg.”; 7 (EJCB): “N Vietnam, 21°35N, 106°30E, 52 km SW of Lang Son, 27.iv. 6.v.1996, 370 m, Dembický & Pacholátko leg.”.

Adult occurence: 4-5-6-8-10 (n=9). Faunal records published by CHÛJÔ & MATUDA, 1940 [#500] are also included. Altitude range: 370-700 m (n=2). Host plants: Citrus: OGLOBLIN & REICHARDT, 1932 [#8654]; YUASA, 1933 [#3132]; QUAYLE, 1938 (as citri) [#7340]; MÜHLMANN, 1954 [#8152]; BALACHOWSKY et al., 1962 [#96]; ISRIGAYA, 1963 [#7016]; OHGUSHI, 1963 [#5415]; WOO SHEH MING, 1964 [#7020]; OHGUSHI, 1966 [#8729]; GURYEVA, 1974 [#5428]; OHGUSHI, 1978 [#7015]; NAKA & OHASHI, 1996 [#4720]; AKIYAMA & OHMOMO, 1997 [#4091]; HUA LI ZHONG, 2002 [#6696]; LIN GUI RUI, 2002 [#7926]; WEI SHU JUN et al., 2006 [#7653]; ZHENG HONG HAI et al., 2006 [#7883]; WEI SHU JUN et al., 2007 [#8166]. Distribution: CHINA: Fujian, Guangdong, Guangxi, Hong Kong, Hubei, Hunan, Jiangxi, Sichuan, Taiwan, Zhejiang. JAPAN: Honshu, Kyushu. LAOS. MYANMAR. VIETNAM.

Agrilus auropictus Kerremans Fig. 10H (habitus); Fig. 32W (aedeagus); Figs. 43G, 44L, 52I (types) auropictus Kerremans, 1912 (Agrilus) KERREMANS, 1912: 208 [#1377] (description). MIWA, 1931: 124 [#6101] (catalog; Formosa). MIWA & CHÛJÔ, 1936: 15 [#1809]

47 (catalog; Japan). OBENBERGER, 1936: 1074 [#2191] (world catalog). KUROSAWA, 1963: 152 [#1618] (characters; Japan). KUROSAWA, 1974: 4 [#4575] (characters; notes). TÔYAMA, 1985: 25 [#3678] (iconography; Japan). PENG ZHONGLIANG, 1987: 354 [#2429] (checklist; China). AKIYAMA & OHMOMO, 1997: 30 [#4091] (checklist; Taiwan). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Taiwan). MÜHLE, 2003: 46 [#6155] (checklist; Taiwan). JENDEK, 2005: 4 [#7217] (lectotype designation). JENDEK, 2006: 396 [#7441] (Palaearctic catalog). BELLAMY, 2008: 1986 [#8576] (world catalog). kanohi Kurosawa, 1954 (Agrilus; subspecies of auropictus) syn. nov. KUROSAWA, 1954: 90 [#1613] (description). IGA, 1955: 78 [#3828] (subspecies of auropictus; iconography; Japan). IGA, 1962: 78 [#3351] (subspecies of auropictus; iconography; Japan). KUROSAWA, 1963: 152 [#1618] (subspecies of auropictus; characters; Japan). CHÛJÔ, 1970: 20 [#3350] (subspecies of auropictus; checklist; faunal records; Japan (Loo Choo Archipelago)). KUROSAWA, 1974: 4 [#4575] (subspecies of auropictus; characters; notes). TÔYAMA, 1985: 25 [#3678] (subspecies of auropictus; iconography; Japan). HIRASHIMA, 1989: 322 [#8078] (subspecies of auropictus; checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (subspecies of auropictus; checklist; Japan). AKIYAMA & OHMOMO, 1997: 30 [#4091] (subspecies of auropictus; checklist; Japan: Honshu, Kyushu, Tanegashima I., Yakushima I., Ryukyus (Amami oshima I., Okinawa I.)). JENDEK, 2006: 396 [#7441] (subspecies of auropictus; Palaearctic catalog). BELLAMY, 2008: 1986 [#8576] (subspecies of auropictus; world catalog). chopardi Descarpentries & Villiers, 1963 (Agrilus) syn. nov. DESCARPENTRIES & VILLIERS, 1963: 52 53 [#699] (description). DESCARPENTRIES & VILLIERS, 1967: 144 [#710] (diagnosed with chapaensis). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2025 [#8576] (world catalog). ryukyuensis Tôyama, 1985 (Agrilus; subspecies of auropictus) syn. nov. TÔYAMA, 1985: 47 [#2924] (description). TÔYAMA, 1985: 25 [#3678] (subspecies of auropictus; iconography; Japan). HIRASHIMA, 1989: 322 [#8078] (subspecies of auropictus; checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (subspecies of auropictus; checklist; Japan). AKIYAMA & OHMOMO, 1997: 30 [#4091] (subspecies of auropictus; checklist; Japan: Ryukyus (Ishigaki jima I., Iriomote jima I.)). JENDEK, 2006: 396 [#7441] (subspecies of auropictus; Palaearctic catalog). BELLAMY, 2008: 1986 [#8576] (subspecies of auropictus; world catalog).

Type material: Agrilus auropictus Kerremans, 1912 Type locality: Taihorin. Type specimens: Primary: Lectotype (Fig. 43G), ♀, designated by JENDEK (2005: 4) [#7217], (MNHN): “Taihorin Formosa H. Sauter 1911 [p] \ auropictus Kerr. Type [h] \ Museum Paris Coll. Ch. Kerremans 1923 [p] [yellow label]”. Secondary: 2 PLT (DEI). Described from unknown number of syntypes. Agrilus auropictus kanohi Kurosawa, 1954 Type locality: Ohdomari, Sata, Kagoshima Pref. Type specimens: Not located. Described from 1 specimen (HT). Agrilus chopardi Descarpentries & Villiers, 1963 Type locality: Tonkin: Chapa. Type specimens: Primary: Holotype (Fig. 44L), ♂, (MNHN): “TONKIN [p] Chapa 25. IV. [h] 191 [p] 8 [h]

48

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Specimens examined: CHINA: Fujian: 1 ♂ (EJCB): “China, Fujian prov. Shaowu env., 5 10.VII.1991”. Taiwan: 2 ♂ (EJCB): “Baibara Formosa, 8.VI.1978, K. Kusama”; 1 ♀ (EJCB): “Mt. Holuan, Nantou Hsien, Taiwan, 26.V.1982, T. Wakejima leg.”. INDIA: Assam: 1 ♂ (EJCB): “NE India, Assam, 27º08’N, 94º00’E, Pan Bari reserv. forest, 4 7.07.1997, leg. V. Siniaev”. JAPAN: Ryukyu isl. (Okinawa incl.): 1 ♂ (EJCB): “Nishinakama, Amami, Ryukyu, 1 7.VI.1970, H. Makihara leg.”; 1 (EJCB): “Chūō Rindō Amami oshima Is., 1.VII.1989, M. Itō”. VIETNAM: 1 ♂ (EJCB): “Vietnam, Tam Dao, 12.5.1984, M. Král lgt.”; 3 ♂ (EJCB): “N. Vietnam (Tonkin), Tamdao, 12 24.5.1989, A. Olexa leg.”; 1 ♀ (EJCB): “N Vietnam, 21º27N, 105º39E, 70 km NW of Hanoi, Tam Dao, 1 8.vi.1996, 900 1200 m, Dembický & Pacholátko leg.”; 2 ♂, 1 ♀ (EJCB): “Vietnam N, 55 km NNW Hanoi env., Tam Dao vill. 800 900 m, 11 14.08.1998, Napolov leg.”.

and posteromedial, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial. Prehumerus: carinal, Shape: arcuate, Extent: beyond half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina, Arc: obvious. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior third of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate or conjoint, Width: wide, Shape: arcuate. Elytral pubescence: proximal and distal or proximal only, Proximal: quadrangular, Distal: apical, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: obtuse, Angles (tips): sharp or blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed, Ovipositor: elongate.

Adult occurence: 4-5-6-7-8-11 (n=14). Faunal records published by TÔYAMA, 1985 [#2924] (as auropictus ryukyuensis) are also included. Altitude range: 800-1200 m (n=2).

Diagnosis: body bicolored ventrally, head golden-red, elytra black; prosternal process wide, strongly enlarged between coxae. Species group: Cyaneovirens. Relevant species: A. lahu sp. nov.

Host plants: Unknown.

Type locality: China, Yunnan province, Lijiang.

Distribution: CHINA: Fujian, Taiwan. INDIA. JAPAN: Honshu, Kyushu, Ryukyu. VIETNAM.

Type specimens: Primary: Holotype (Fig. 27J), ♂, (EJCB): “China, Yunnan, Lijiang, 8-24.VI.1994, Šiška & Spevár”. Secondary: CHINA: Yunnan: 1 PT (EJCB): “China, Yunnan 1994 Zhongdian 16-20.vi. leg. Šiška T. Spevár”; 1 PT (EJCB): “China, Yunnan, 25-27.VI.1994, Daju, B. Šiška & T. Spevár leg.”; 9 PT (EJCB): “China, Yunnan, Lijiang, 8-24.VI.1994, Šiška & Spevár”; 4 PT (EJCB): “ChinaYunnan, Lijiang, 11.6.-15.6.1995, lgt. E. Kučera”; 2 ♂ PT, 1 ♀ PT (EJCB): “China, Yunnan, Lijiang, 19.VI.1995, Šiška & Spevár & Pekarovič”.

Jeanvoine [p] \ Agrilus chopardi n. sp. Holotype nob[is]. A. Descarpentries et [h] A. Villiers det. 195 [p]”. Described from 1 specimen (HT). Agrilus auropictus ryukyuensis Tôyama, 1985 Type locality: Shirahama, Iriomote Is. Type specimens: Primary: Holotype (Fig. 52I), ♀, (NSMT): “Shirahama, Is. Iriomote, 29. III. 1965, Coll. H. Yokoyama [h] \ Holotype [p] Agrilus auropictus ryukyuensis Tôyama, 1985 [h] [red label]”. Described from 5 specimens (HT, PT). Diagnosis: 8.0-9.7 mm; elytra evenly attenuate apically, without expanded apices. Species group: Niveoguttatus. Relevant species: A. alpenae Baudon, 1968.

Agrilus bai Jendek, sp. nov. Fig. 27J (habitus of HT); Fig. 40C (aedeagus)

Description: 4.2-5.5 mm (Holotype 4.6 mm). BODY: Shape: subparallel, Color (dorsally): bicolored, Color (dominant): black. HEAD: Medial impression: absent, Epistoma: with raised upper margin. Frons: Shape: flat or moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture (density): dense. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: in line with antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square, Sides: straight or arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate or obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles, rarely not reaching level of anterior pronotal angles. Posterior angles: Shape: rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: posterolateral, rarely medial and lateral, Medial impression (shape): anteromedial

Adult occurence: 6 (n=5). Host plants: Unknown. Distribution: CHINA: Yunnan. Etymology: Noun in apposition; the specific name refers to the Chinese ethnic group Bai living mostly in the provinces of Yunnan (Dali area).

Agrilus baroni Gory & Laporte Fig. 30B (habitus); Fig. 41E (aedeagus); Figs. 43J, 49P (types) baroni Gory & Laporte, 1839 (Agrilus) GORY & LAPORTE, 1839: 23 [#3738] (description). GEMMINGER & HAROLD, 1869: 1437 [#1057] (catalog). SAUNDERS, 1871: 114

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TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA [#2552] (cited as baronii; catalog). WATERHOUSE, 1889: 123 [#3054] (uncertain in Mexico). KERREMANS, 1892: 248 [#1397] (catalog). KERREMANS, 1903: 282 [#1441] (catalog). OBENBERGER, 1960: 151 [#2394] (lectotype designation). DESCARPENTRIES & VILLIERS, 1963: 1, 6 [#701] (type locality “Mexique” is confused; characters; faunal record; Chine; Tonkin). NELSON & BELLAMY, 1993: 304 [#3167] (authorship and publication date). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). BELLAMY, 2008: 1990 [#8576] (subgenus Sinagrilus; world catalog). mopsus Théry, 1930 (Agrilus) THÉRY, 1930: 303 304 [#2777] (description). OBENBERGER, 1936: 1015 [#2191] (world catalog). OBENBERGER, 1960: 151 [#2394] (Agrilus cited as Agrolus; synonym of baroni). DESCARPENTRIES & VILLIERS, 1963: 6 [#701] (synonym of baroni). PENG ZHONGLIANG, 1987: 356 [#2429] (checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Hongkong). JENDEK, 2006: 397 [#7441] (synonym of baroni; Palaearctic catalog). BELLAMY, 2008: 1990 [#8576] (synonym of baroni; world catalog).

Type material: Agrilus baroni Gory & Laporte, 1839 Type locality: Mexique [Note: Locality confused. See DESCARPENTRIES & VILLIERS, 1963 [#701]]. Type specimens: 1 ST (Fig. 43J), (MNHN). Described from unknown number of syntypes. Agrilus mopsus Théry, 1930 Type locality: Hong-Kong, Chine. Type specimens: Holotype by monotypy, ♀ (Fig. 49P) (MNHN). Described from 1 ♀ specimen. Diagnosis: 15.7-18 mm; dorsally glabrous; pronotum without medial impression; prehumerus tubercular; scutellum large, flat, without transverse carina. Species group: Unassigned. Relevant species: Unknown. Specimens examined: VIETNAM: Hoa Binh: 1 ♂ (MNHN): “Hoa Binh, Tonkin R. P. de Cooman”.

Host plants: Unknown. Distribution: CHINA: Hong Kong. VIETNAM.

Agrilus barrati Descarpentries & Villiers Fig. 28A (habitus); Fig. 40F (aedeagus); Fig. 43K (type) barrati Descarpentries & Villiers, 1963 (Agrilus) DESCARPENTRIES & VILLIERS, 1963: 55, 57 58 [#699] (description). DESCARPENTRIES & VILLIERS, 1963: 106 [#700] (notes; characters). BELLAMY, 2008: 1990 [#8576] (world catalog).

Type material: Agrilus barrati Descarpentries & Villiers, 1963 Type locality: Tonkin: Hoa-Binh. Type specimens: Primary: Holotype (Fig. 43K), ♂, (MNHN): “Hoa Binh Tonkin de Cooman [h] \ Agrilus barrati n. sp. Holotype ♂ nob[is]. A. Descarpentries et [h] A. Villiers det. 19 [p] 62 [h]”. Secondary: 6 PT (MNHN). Described from unknown number of specimens (HT, AT, PT). Diagnosis: 3.9-5.5 mm, pronotum very distinctive, with obvious pronotal lobe and central gibbosity on disk. Dorsal color varies from deep blue to golden-green. Species group: Barrati. Relevant species: A. ctesias Kerremans, 1913.

Specimens examined: CHINA: Fujian: 1 (EJCB): “China, Fujian prov., Shaowu env., 13 16.6.1991”. Hubei: 1 (EJCB): “China, SE Hubei, 7.+17.vi, 20 km NW Tongshan, 29.7N 114.3E, Jaroslav Turna leg. 2004”. Hunan: 1 ♂, 2 ♀ (EJCB): “China, Hunan SE, 1994, Guidong env. 26 31.V., 26.04N 113.56E, E. Jendek & O. Šauša leg.”; 1 ♂ (EJCB): “China, N Hunan, 4 7.vii., Wuling Shan, 29.4N, 110.4E, Zhangjiajie, 700 m, Jaroslav Turna leg. 2003”. LAOS: Borikhamxai: 1 ♂ (EJCB): “Laos centr., Bolikhamsai prov., Ban Nape Kaew Nua Pass, 18.4. 1.5.1998, alt. 600±100 m, N 18º22.3, E 105º09.1 (GPS), E. Jendek & O. Šauša leg.”; 5 (EJCB): “Laos CE., 1 18.v.2001, Boli Kham Xai prov., 18º21’N, 105º08’E, Ban Nape (8 km NE), ~600m, Kubáň leg.”; 1 ♀ (EJCB): “Laos CE., 1 18.v.2001, Boli Kham Xai prov., 18º21’N, 105º08’E, Ban Nape (8 km NE), ~600m, Dembický leg.”; 2 ♀ (EJCB): “Laos C., Bolikhamsai pr., Ban Nape env., 7 16.V.2004, alt., 400±100 m, 18º20’N, 105º08’E, E. Jendek & O. Šauša leg.”. Louang Namtha: 1 ♀ (EJCB): “N Laos, 14 22.vi.1996, Louang Namtha pr. 10 30 km NW Louang Namtha, C. Holzschuh leg. 800 m”; cca 30 (EJCB): “Laos, Louang Namtha pr., 21°09’N, 101°19’E, Namtha Muang Sing, 5 31.v.1997, 900 1200 m, Vit Kubáň leg.”; cca 440 (EJCB): “Laos north, 5 11.V.1997, 20 km NW Louang Namtha, N 21°09.2, E 101°18.7, alt. 900±100 m, E. Jendek & O. Šauša leg.”. Phongsali: 1 ♂ (EJCB): “Lao N, Phongsaly prov., 21°41’ 2’N, 102°06’ 08’E, 28.v. 20.vi.2003 Phongsaly env., ~1500m, Vít Kubáň leg.”; 1 ♂ (EJCB): “Lao N, Phongsaly prov., 21°41’ 2’N, 102°06’ 08’E, Phongsaly env., 28.v. 20.vi.2003, 1500m, P. Pacholátko leg.”; 1 ♂ (EJCB): “Lao, Phongsaly prov. 21°41’N, 102°06’E Phongsaly env. 6 17.v.2004, 1500 m, P. Pacholátko leg.”. THAILAND: Chiang Mai: 1 (EJCB): “Chiang Dao Valley, North Thailand, 2.V.1980, Y. Komiya lgt”; 1 (EJCB): “Thailand, Chiang Mai prov., 19°19’N, 98°50’E, 1400 m, Sanpakia vill., 1 15.V.1998, Ivo Martinů leg.”. VIETNAM: 1 ♂ (EJCB): “Tonkin, Hoa Binh”; 1 ♀ (MNHN): “Tonkin, III, Hoa Binh, 1937, A. de Cooman”; 1 ♂ (EJCB): “Vietnam N, Tonkin, Cuc Phuong nat. park, 2 12.V.1991, E. Jendek leg.”; 1 ♂ (EJCB): “N Vietnam, 20°15N, 105°42E, 75 km SW of Hanoi, 21 22.v.1996, Cuc Phuong, Dembický & Pacholátko leg.”; 3 ♂, 3 ♀ (EJCB): “N Viet Nam (Tonkin), pr. Houang Lien Son, Sa Pa, 11 15.V.1990, Vít Kubáň leg.”; 1 ♀ (EJCB): “SaPa 11 18.VI., N. Vietnam, A. Olexa [leg.] 1990”; 1 (EJCB): “Vietnam, TamDao 900 m, 16 23.V.1991, Strnad, J. lgt.”; 1 ♂ (EJCB): “N Vietnam, 21°35N, 106°30E, 52 km SW of Lang Son, 27.iv. 6.v.1996, 370 m, Dembický & Pacholátko leg.”.

Adult occurence: 3-4-5-6-7 (n=23). Altitude range: 3001500 m (n=14). Host plants: Unknown. Distribution: CHINA: Fujian, Hubei, Hunan. LAOS. THAILAND. VIETNAM.

Agrilus betuleti (Ratzeburg) Fig. 17E (habitus); Fig. 35J (aedeagus); Figs. 47H, 52L (types) betuleti Ratzeburg, 1837 (Buprestis) RATZEBURG, 1837: 57 [#2498] (description). KIESENWETTER, 1857: 126, 153 154 [#3015] (characters in key, redescription; notes; Germany). MARSEUL, 1857: 92 [#4371] (checklist; Europe). MARSEUL, 1866: 436, 455 [#676] (characters in key; redescription; references; distribution overview). GEMMINGER & HAROLD, 1869: 1437 [#1057] (catalog). SAUNDERS, 1871: 119 [#2552] (catalog). HOCHHUTH, 1873: 158 [#3453] (faunal records; Gouvernements Kiew und Volhynien). KEPPEN, 1882: 232 [#8636] (notes; pest). MARSEUL, 1887: 251 [#4707] (catalog). JUDEICH & NITSCHE, 1889: 319, 322 [#5686] (characters; biology). KERREMANS, 1892:

50 248 [#1397] (catalog). RUPERTSBERGER, 1894: 161, 287 [#4465] (larval bibliography; catalog). ABEILLE DE PERRIN, 1897: 4 [#19] (characters in key). CSIKI, 1901: 89 [#505] (faunal records; Siberia: Baltim). KERREMANS, 1903: 275 [#1441] (catalog). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). BAU, 1914: 257 [#8668] (characters in key). BEDEL, 1921: 202, 210 [#170] (characters in key; faunal records; France). ESCHERICH, 1923: 148, 150 [#6717] (characters in key; notes). OBENBERGER, 1926: 656 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 959 961 [#2191] (world catalog). RIKHTER, 1945: 166 [#3507] (characters in key; European part of USSR except northernmost and south regions, eastward to Primorsk). PALMÉN, 1946: 120 [#7730] (faunal records; Russia (Karelia)). KANGAS, 1947: 110 [#3622] (notes on larval characters; Finnland). ILINSKIJ, 1948: 296 [#5670] (larval characters in key; notes). RIKHTER, 1948: 394 [#3879] (characters in key). STEPANOV, 1948: 243, 244 [#3535] (checklist; faunal record; Chelyabinskaya oblast’). SCHAEFER, 1949: 355, 387 389, 490 [#2565] (characters; references; faunal records; biology; notes; ditributional overview; notes). RIKHTER, 1950: 158, 165 [#3881] (adult and larval characters in key). MÜHLMANN, 1954: 79 [#8152] (pest; note). HORION, 1955: 92 93 [#1285] (distribution overview; faunal records; biology). STARK, 1955: 775 [#3533] (faunal record; USSR: Saratov). TER MINASYAN, 1955: 450 [#7893] (host plants; distribution; references). ALEXEEV, 1957: 153 [#3226] (characters in key). PLAVILSHCHIKOV, 1957: 210 [#8254] (characters; European part of Soviet Union). ALEXEEV, 1959: 9 [#3229] (characters in key; eastern to Primorsk). ALEXEEV, 1960: 1506 [#45] (larval characters in key). ZAGAIKEVICH, 1960: 257 [#5615] (checklist; Ukraine). ALEXEEV, 1961: 5 [#3243] (larval characters in key). BALACHOWSKI et al., 1962: 286 [#96] (notes). ZAGAIKEVICH, 1962: 60 [#5616] (checklist; Ukraine). ZAGAIKEVICH, 1962: 103 [#3249] (faunal records; biology; Ukraine; Krym). KHRAMCOV & PADIJ, 1965: 83 84 [#8649] (characters; larva; biology; harms; eradication; USSR). RIKHTER & ALEXEEV, 1965: 302 [#2522] (characters in key; European part of USSR). COBOS, 1968: 387 388 [#580] (subgenus Agrilus; characters; faunal records; Mongolia). ALEXEEV, 1971: 225 [#73] (phylogenetic origin). COBOS, 1972: 56 [#591] (subgenus Agrilus; checklist; Mongolia). KOSTIN, 1973: 110 [#3693] (characters in key; Kazakhstan). ALEXEEV, 1975: 152 [#52] (notes; Mongolia). LOMPE, 1979: 239 [#3628] (characters in key; Central Europe). ALEXEEV, 1981: 70 [#61] (larval characters in key). BÍLÝ, 1982: 64, 66, 71 72 [#295] (characters in key; distribution overview; biology). ALEXEEV, 1989: 483 484 [#64] (key to species; Khabarovsk; Amursk; Primorsk; Siberia; European USSR; central Mongolia). ALEXEEV & VOLKOVITSH, 1989: 318, 346, 356 [#3241] (characters in key; faunal records; Mongolia). GUTOWSKI, 1993: 301 [#3638] (characters). BÍLÝ, 1994: 109 [#8200] (larval characters in key). CURLETTI, 1994: 22, 164 164 [#658] (subgenus Agrilus; catalog; faunal records). JENDEK, 1994: 10 [#1355] (synonymy). LAGUNOV & NOVOZHENOV, 1996: 46 [#7461] (checklist; Russia (Chelyabinsk)). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). TLEPPAEVA, 1999: 184 [#6551] (notes; Kazakhstan). BÍLÝ, 2002: 13 [#5877] (bionomy; host plants; distribution). BRECHTEL & KOSTENBADER, 2002: 225 227 [#5766] (biology; ecology; distribution; forestry). JENDEK, 2006: 393 [#7441] (subgenus Robertius; Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 1995 1996 [#8576] (subgenus Robertius; world catalog). VOLKOVITSH, 2009: 135 [#8850] (subgenus Robertius; faunal records; Russia: Primorskiy kray). impressicollis Marseul, 1866 (Agrilus; [preoccupied]) MARSEUL, 1866: 436, 454 455 [#676] (description). POPOFF, 1853: 102 [#7853] (Mannerheim is cited as the author; faunal record; notes; no characters cited; Buryatia). GEMMINGER & HAROLD, 1869: 1440 [#1057] (synonym of foveicollis).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA MARSEUL, 1869: 122 [#674] (synonym of foveicollis). SAUNDERS, 1871: 119 [#2552] (synonym of foveicollis). HEYDEN, 1881: 118 [#5548] (synonym of foveicollis). MARSEUL, 1887: 251 [#4707] (synonym of foveicollis). KERREMANS, 1892: 257 [#1397] (synonym of foveicollis; catalog). ABEILLE DE PERRIN, 1897: 4 [#19] (Mannerheim is cited as the author; characters in key). KERREMANS, 1903: 285 [#1441] (synonym of foveicollis). JAKOBSON, 1913: 797 [#4386] (synonym of foveicollis; catalog). OBENBERGER, 1936: 990 [#2191] (synonym of foveicollis). ALEXEEV, 1959: 20 [#3229] (synonym of foveicollis). JENDEK, 1994: 10 [#1355] (synonym of betuleti). JENDEK, 1998: 323 324 [#4481] (lectotype designation). JENDEK, 2006: 393 [#7441] (synonym of betuleti; Palaearctic catalog). BELLAMY, 2008: 1995 [#8576] (synonym of betuleti; world catalog). foveicollis Marseul, 1869 (Agrilus; replacement name for impressicollis Marseul not Gory) MARSEUL, 1869: 122 [#674] (replacement name proposal). GEMMINGER & HAROLD, 1869: 1440 [#1057] (catalog). SAUNDERS, 1871: 119 [#2552] (catalog). HEYDEN, 1881: 118 [#5548] (catalog). MARSEUL, 1887: 251 [#4707] (catalog). KERREMANS, 1892: 257 [#1397] (catalog). KERREMANS, 1903: 285 [#1441] (catalog). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). OBENBERGER, 1924: 42 [#2040] (note). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 990 [#2191] (world catalog). RIKHTER, 1945: 168 [#3507] (characters in key; north of European part of USSR; Sibir). RIKHTER, 1948: 395 [#3879] (characters in key). TER MINASYAN, 1955: 451 [#7893] (host plants; distribution; references). ALEXEEV, 1959: 20 [#3229] (characters in key; north of European USSR). LUNDBERG, 1980: 92 [#7208] (notes; not in Sweden). LUNDBERG, 1980: 96 [#6031] (notes; not in Sweden). BÍLÝ, 1982: 71, 72 [#295] (synonym of betuleti). ALEXEEV, 1989: 487 [#64] (key to species; Khabarovsk; Amursk; Primorsk; Chita; Buryatia; Siberia; European USSR; northeastern Europe). JENDEK, 1994: 10 [#1355] (synonym of betuleti). JENDEK, 2006: 393 [#7441] (synonym of betuleti; Palaearctic catalog). BELLAMY, 2008: 1995 [#8576] (synonym of betuleti; world catalog). saliceti Obenberger, 1924 (Agrilus; variety of betuleti) OBENBERGER, OBENBERGER, 1924: 42 [#2040] (description). 1926: 656 [#2057] (variety of betuleti; Palaearctic catalog). OBENBERGER, 1936: 961 [#2191] (subspecies of betuleti; world catalog). THÉRY, 1942: 156 [#2873] (subgenus Agrilus; subspecies of betuleti). SCHAEFER, 1949: 388 [#2565] (subspecies of betuleti; notes). COBOS, 1972: 56 [#591] (uncertain synonym of betuleti). JENDEK, 1994: 10 [#1355] (synonym of betuleti; lectotype designation). JENDEK, 2006: 393 [#7441] (synonym of betuleti; Palaearctic catalog). BELLAMY, 2008: 1996 [#8576] (synonym of betuleti; world catalog).

Type material: Buprestis betuleti Ratzeburg, 1837 Type locality: in meiner Gegend [Note: Presumably “Germany, Eberswalde”, the residence of Ratzeburg]. Type specimens: Not located. Described from unknown number of syntypes. Agrilus impressicollis Marseul, 1866 Type locality: Sibérie. Type specimens: Primary: Lectotype (Fig. 47H), sex not examined, designated by JENDEK (1998: 323-324) [#4481], (MNHN): “Transcriptio [p] Impressicollis (Manh.) Mars. type Sibérie [h] \ Agrilus foveicollis Mars. 1869 = impressicollis Mars. 1865 nom. praeoc. syntype [h] A. Descarpentries det. [p] \ Agrilus [p] betuleti Ratz. [h] Sv. Bílý det. [p] 1981 [h] \ Muséum Paris 1952 Coll. R. Oberthur [p] [yellow label]”.

51

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Secondary: 1 PLT (MNHN); 1 PLT (ISNB). Described from unknown number of syntypes. Agrilus foveicollis Marseul, 1869 Type locality and specimens: See: Agrilus impressicollis Marseul, 1866. Agrilus betuleti variety saliceti Obenberger, 1924 Type locality: Semirječia: Kamenaja Ret. Type specimens: Primary: Lectotype (Fig. 52L), sex not examined, designated by JENDEK (1994: 10) [#1355], (NMPC): “Kamenaja - ret. Semirjetschensk coll. Winkler [p] \ Typus [p] [red label] \ Mus. Nat. Pragae Inv. [p] 24255 [h] [orange label] \ Agrilus betuleti var. saliceti m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Diagnosis: 4.2-6.7 mm; body black, silky lustrous; pronotum widest in anterior half; flat with large and deep lateral impressions; pronotal disk with very fine transverse sculpture; elytral apices widely, separarely arcuate. Species group: Betuleti. Relevant species: A. delphinensis Abeille de Perrin, 1897; A. fareastensis Jendek, 1995. Specimens examined: KAZAKHSTAN: 1 ♀ (EJCB): “W Kazakhstan, Aktyubinsk, Mugodzhany, Alinskol, 6.6.1995, leg. Dolin”. MONGOLIA: 1 (EJCB): “Mongolia, 5 9 July 2003, Cagan nuga env. 1500 m, Segneger riv. vall. / 35 km NE of Mandal ( Batsumber), cca 70 km N of Ulaanbaatar, Zd. Jindra lgt.”. RUSSIA: Amurskaya oblast’: 1 (EJCB): “USSR, 4.6.1975, Amurskaja obl., Umg. Svobodnij, leg. Kuznetsov”; 1 (ZIN): “Amurskaya obl., Svobodnensk. r n., s. Gaicenka, 8.6.[19]76 g, V. Kuznecov [in Russian]”. Chelyabinskaya oblast’: 1 (ZIN): “pos. Sokolovskii, Chelyab., u. Orenb. g. Infant’ev 19.v.1912 [in Russian]”. Khabarovskiy kray: 1 (ZIN): “Niz. Amura, Nizhne Tambovskoe, 10.6.1955 Kabakov [in Russian]”; 1 (MSCI): “Khabarovsky reg., Komsomolsky zapov., okr. kor. Kamennaya pad’, 28.VI.1991 leg. Smirnov”. Lipetskaya oblast’: 1 (MCCD): “1ex, 10км Ю г. Липецк, Троицкий Лес, 22.5.2002 Цуриков”; 1 (MCCD): “30км В г. Елец, окр. с. Донское, 3.7.2004 Артеменко”. Primorskiy kray: 1 ♀ (EJCB): “SU Ussuri reg., Novovarvarovka, 6 10.7.1989, M. Nikodým lgt.”; 2 (MNHE): “Russia, 27 29.06.1989, Primorye Distr., Buyanki, leg. U. Roosileht”; 1 ♂, 1 ♀ (EJCB): “SU Ussuri reg. Jasnoe 400 m 12 19.7.1989 M. Nikodým lgt.”; 1 ♂ (EJCB): “Sib. or. m., Primorje, Sichote Alin Mts., Sokolči, 1 15.7.1990, Kadlec + Voříšek lg.”; 1 ♀ (EJCB): “USSR, Primorskij kraj Arseneev env., VI.1991, 44.11 N, 133.19 E, M. Štrba leg.”; 2 ♀ (EJCB): “USSR, Primorsk. kraj, Arsenev env., VI.1991, 44.11N, 133.19E, O. Šausa leg.”; 1 ♂ (EJCB): “Russia Primorskij Kraj, Gribnoye env., Sinnij chrebet Mts., 17 19.VI.1997, leg. L. Roller & L. Vidlička”; 1 ♂, 1 ♀ (IMCM): “Yu Primor’e, Lazovskii raion, p. Preobrazhenie na usokhshikh Quercus mongolica, 9 14.vii.2006, leg. M. L. Smirnovy [in Russian]”; 2 (MSCI): “Primorskii krai, Lazovskii r n, okr. pos. Lazo, 7.07.06, leg. M. & L. Smirnov [in Russian]”; 1 (EJCB): “Russia, Primorsky Kray, Kaymanovka vil., N43º37’, E132º14’, 7 11.vii.2007, 100m, V. Grebennikov”. Sakhalinskaya oblast’: 1 (EJCB): “Sakhalin, SO Novoaleksandrovsk, 29.6.[19]91 Filimonov [in Russian]”. Saratovskaya oblast’: 2 (MSCI): “Saratovskaya obl. Novoburasskii r n, okr. st. Burasy, 52°12’N, 46°08’E, 21.06.2008, M. Smirnov”. Yevreyskaya avtonomnaya oblast’: 4 (EJCB): “Russia, ES Siberia, Khabarovsk., Oblutchie distr., S. Chingan Mts., 500 800 m, 15.VI. 17.VII.2001, R. Misustin & S. Duz leg.”. Zabaykalskiy kray: 1 (ZIN): “YuV Chitinsk. obl. Sherlovaya gora, 15.VI.1989 Kabakov [in Russian]”.

Adult occurence: 5-6-7 (n=25). Altitude range: 100-1500 m (n=4).

Host plants: Betula: RATZEBURG, 1837 [#2498]; MARSEUL, 1866 [#676]; KALTENBACH, 1874 [#8154]; JUDEICH & NITSCHE, 1889 [#5686]; ESCHERICH, 1923 [#6717]; SAINTE CLAIRE, 1935 [#8551]; RIKHTER, 1945 [#3507]; ILINSKIJ, 1948 [#5670]; TER MINASYAN, 1955 [#7893]; ALEXEEV, 1959 [#3229]; ALEXEEV, 1961 [#3243]; ZAGAIKEVICH, 1962 [#3249]; KHRAMCOV & PADIJ, 1965 [#8649]; RIKHTER & ALEXEEV, 1965 [#2522]; ALEXEEV, 1975 [#52]; KUBÁŇ, 1978 [#3474]; ALEXEEV, 1981 [#61]; ALEXEEV, 1989 [#64]; ALEXEEV & VOLKOVITSH, 1989 [#3241]; CURLETTI, 1994 [#658]; BRECHTEL & KOSTENBADER, 2002 [#5766]. − B. alba: BEDEL, 1921 [#170]; SCHAEFER, 1949 [#2565]; HORION, 1955 [#1285]; NIEHUIS, 1988 [#1912]. − B. nana: SCHAEFER, 1949 [#2565]; BÍLÝ, 2002 [#5877]; HASTIR & GASPAR, 2002 [#6211]; − B. odorata: KANGAS, 1947 [#3622]. − B. pendula: BÍLÝ, 2002 [#5877]. − B. pubescens: HORION, 1955 [#1285]; BÍLÝ, 2002 [#5877]; HASTIR & GASPAR, 2002 [#6211]. −B. tianshanica: TLEPPAEVA, 1999 [#6551]. − B. verrucosa: KANGAS, 1947 [#3622]; HORION, 1955 [#1285]; HASTIR & GASPAR, 2002 [#6211]. Erroneous: Quercus: ESCHERICH, 1923 [#6717]; TER MINASYAN, 1955 [#7893]; ALEXEEV, 1959 (as foveicollis) [#3229]; ALEXEEV, 1989 (as foveicollis) [#64]. Distribution: Local: MONGOLIA. RUSSIA: Amurskaya oblast’, Buryatiya Respublika, Irkutskaya oblast’, Khabarovskiy kray, Primorskiy kray, Sakha Respublika, Sakhalinskaya oblast’, Tyva Respublika, Yevreyskaya avtonomnaya oblast’, Zabaykalskiy kray. Global: ASIA: KAZAKHSTAN. RUSSIA. EUROPE: AUSTRIA. BELARUS. BELGIUM. BOSNIA & HERZEGOVINA. CROATIA. CZECH REPUBLIC. DENMARK. ESTONIA. FINLAND. FRANCE. GERMANY. HUNGARY. ITALY. LATVIA. LIECH-TENSTEIN. LITHUANIA. NETHERLANDS. NOR-WAY. POLAND. ROMANIA. RUSSIA. SERBIA & MONTENEGRO. SLOVAKIA. SLOVENIA. SWEDEN. SWITZERLAND. UKRAINE.

Agrilus bicuspidatus Jendek, sp. nov. Fig. 12G (habitus of HT)

Description: Holotype 7.7 mm. BODY: Shape: cuneiform, Color (dorsally): bicolored, Color (dominant): green. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): irregular, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: flat or moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial,

52 Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial. Prehumerus: obsolete or filamentary, Shape: arcuate or bisinuate, Extent: to third of pronotal length, Posterior end: distant from pronotal angle or margin, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: bicolored, Alternative color (location): perisutural stripe, Humeral carina: absent. Apices: Arrangement: conjoint, Shape: cuspidate, Position of dominant cusp (spine): pleural margin. Elytral pubescence: ornamental, Color: unicolored, Character: homogenous. Tomentum: absent. Prosternal lobe: moderate, Anterior margin: subtruncate. Prosternal process: moderate, Shape: narrowed, Sides: straight, Angles: absent, Disk: flat. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. GENITALIA: Ovipositor: uritiform or elongate. Diagnosis: mazarine spots on each elytron elongate in form of perisutural stripe; elytral apex with sharp cusp on pleural margin. Species group: Plasoni. Relevant species: A. chujoi Kurosawa, 1985, A. huashanus Jendek, 2001. Type locality: China, Kouy-Tschéou [= Guizhou province]. Type specimens: Primary: Holotype (Fig. 12G), ♀, (EJCB): “Kouy-Tschéou, R. Marchand, 1904”. Secondary: CHINA: Yunnan: 1 ♀ PT (ZIN): “China, Yunnan, Loc. 29, 17-20 km road Xiaguan-Dacang, 25°28’53.3’’N, 100°15’42.3’’E, 1981 m, 08.06.2002, Konstantinov & Volkovitsh leg.”. Adult occurence: 6 (n=1). Altitude range: 1981 m (n=1). Host plants: Unknown. Distribution: CHINA: Guizhou, Yunnan. Etymology: The specific name was derived from the Latin prefix bi- (two) and the noun cuspis (pointed end). It refers to the two dents at pleural margin of the elytral apex of the species.

Agrilus bidentulus Ganglbauer Fig. 19C (habitus); Fig. 36D (aedeagus); Fig. 43N (type) bidentulus Ganglbauer, 1889 (Agrilus) GANGLBAUER, 1889: 22, 31 [#3427] (description). HEYDEN, 1893: 90 [#3450] (catalog). KERREMANS, 1903: 282 [#1441] (catalog). JAKOBSON, 1913: 796 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 961 [#2191] (world catalog). ALEXEEV, 1959: 26 [#3230] (synonym of caeruleus). ALEXEEV, 1959: 7 [#3229] (synonym of caeruleus). PENG ZHONGLIANG, 1987: 354 [#2429] (checklist; China). ALEXEEV, 1998: 378 [#4322] (subgenus Dentagrilus; subgeneric classification). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Gansu). JENDEK, 2002: 3 [#5709] (valid species; taxonomic notes). JENDEK, 2006: 390 [#7441] (subgenus Dentagrilus; Palaearctic catalog). BELLAMY, 2008: 1996 1997 [#8576] (subgenus Dentagrilus; world catalog). JENDEK & GREBENNIKOV, 2009: 44, 53 [#8561]

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA (characters in key; type examined; faunal records; notes; China (Gansu, Sichuan)).

Type material: Agrilus bidentulus Ganglbauer, 1889 Type locality: Kan-ssu [Dorf Ter-ga] [Note: Locality is complemented from the list of localities on page 21] Type specimens: Holotype by monotypy (Fig. 43N), ♂, (ZIN): “Agr. bidentulus Ggl. Typ. [h] \ Coll. Semenov-TianShansky [p] \ kop. app. vzyat dlya izgot. preparata Alekseev [h] [in Russian] \”. Described from 1 specimen. Diagnosis: 6.1-7.5 mm; metacoxal plates in males flat, not protruding outward; median lobe of aedeagus smooth, without denticulate sides (Fig. 36D). Species group: Cyanescens. Relevant species: A. asahinai Kurosawa, 1956; A. cyanescens (Ratzeburg, 1837). Specimens examined: CHINA: Gansu: 1 ♀ (EJCB): “China 17.viii.1918 Kansou Hasitai” (JENDEK & GREBENNIKOV, 2009 [#8561]); 4 ♂ (EJCB): “China Kansu Ponggartang 30.Jun 1992 M. Bok lgt.” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 ♂, 1 ♀ (EJCB): “China Gansu prov. 120 km SW Lanzhou Ponggartang 30.VI. 2.VII.1992 Jaroslav Turna leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 ♂ (EJCB): “China, Gansu 20 km W Wu du, 1800 2600, 1 6.6.97 A. Shamajev leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]). Sichuan: 2 ♂ (EJCB): “China Gansu Juizhaigon [sic!] 12 17.6.2000 leg. E. Kučera” (JENDEK & GREBENNIKOV, 2009 [#8561]).

Adult occurence: 6-7-8 (n=5). Altitude range: 1800-2600 m (n=1). Host plants: Unknown. Distribution: CHINA: Gansu, Sichuan.

Agrilus bituberculatus Jendek Fig. 24E (habitus of HT); Fig. 38L (aedeagus) bituberculatus Jendek, 2007 (Agrilus) JENDEK, 2007: 39 [#8006] (description). [#8576] (world catalog).

BELLAMY, 2008: 2002

Type material: Agrilus bituberculatus Jendek, 2007 Type locality: China S, Shaanxi, Changan Co., Qinling Mts. N. slope, 33.56-59’ - 108.50’, 800-1200 m. Type specimens: Primary: Holotype (Fig. 24E), ♂, (EJCB): “China S, Shaanxi, 14-16.6.1995, Qinling Mts. N. slope, 33.56-59’ / 108.50’ 800-1200 m, Changan Co.”. Secondary: 1 ♀ PT (CBCS); 1 PT (SOCI); 25 PT (EJCB). Described from 28 specimens (HT, PT). Diagnosis: 3.5-5.0 mm; lateral pronotal impressions very weak; pronotal disk with very fine sculpture. Species group: Sulcicollis. Relevant species: A. truncatus Jendek, 2007. Specimens examined: CHINA: Beijing: 1 ♂ PT (EJCB): “China, Beijing Shi, 4.vi., ~ 30km N Huairou, 2000, Yunmeng Shan, Forest Park, 40.6N, 116.7E, Jaroslav Turna leg.” (JENDEK, 2007 [#8006]); 2 ♂ PT, 3 ♀ PT (EJCB): “China, Beijing distr., Yan Shan, Dongling Mts., 39º58’ N, E 115º26’ E, 1400 m, Xiaolongmen, 15 16. VI.2001” (JENDEK, 2007 [#8006]). Fujian: 1 ♀ PT (EJCB): “China, Fujian prov. bor. occ., Shaowu env. 25 km road, Shaowu Taining

53

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA 6 10.7., R. Červenka lgt. 1991” (JENDEK, 2007 [#8006]); 1 ♂ PT (EJCB): “China, Fujian prov., Wuyi Shan Nat. Par., Sangan env. (900 m), 30.v. 12.vi.2001, Hlaváč, P. & Cooter, J. lgt.” (JENDEK, 2007 [#8006]). Hebei: 1 ♂ PT (EJCB): “China, Hebei/Nei Mongol, pass Chengde Chifeng, 41.6N, 118.2E, 14 16.VI., Jaroslav Turna leg. 2001” (JENDEK, 2007 [#8006]). Henan: 17 (EJCB): “China, N. Henan, 11 12.VI., Jiuligou, 550 m, 35º12’N, 112º26’E, Jaroslav Turna leg. 2007”. Shaanxi: 5 ♂ PT, 8 ♀ PT (EJCB): “China S, Shaanxi, 14 16.6.1995, Qinling Mts. N. slope, 33.56 59’/108.50’ 800 1200 m, Changan Co., L.+R. Businský lgt.” (JENDEK, 2007 [#8006]); 2 ♀ PT (EJCB): “China Yunnan [sic!][should be Shaanxi], Lüyeang, 25.7. 30.7.1996, E. Kučera leg.” (JENDEK, 2007 [#8006]); 1 ♂ PT, 1 ♀ PT (EJCB): “China, Shaanxi prov., 21 23. June 1998 \ Qing Ling Shan mts, 1000 m road Baoji Taibai vill., pass 40 km S Baoji, Zd. Jindra lgt.” (JENDEK, 2007 [#8006]); 1 ♀ PT (EJCB): “China Shaanxi, Lüeang, 22.5. 25.5.2000, leg. E. Kučera” (JENDEK, 2007 [#8006]); 1 ♂ (EKCS): “China Shaanxi, Lueyang, 29.5. 2.6.2005, lgt. E. Kučera”.

Adult occurence: 5-6-7 (n=11). Altitude range: 550-1400 m (n=5). Host plants: Unknown. Distribution: CHINA: Beijing, Fujian, Hebei, Nei Mongol, Shaanxi.

Agrilus blatteicollis Bourgoin Fig. 12L (habitus); Fig. 33U (aedeagus); Fig. 43P (type) blatteicollis Bourgoin, 1922 (Agrilus) BOURGOIN, 1922: 168 [#366] (description). OBENBERGER, 1936: 1076 [#2191] (cited as blattaeicollis; world catalog). BAUDON, 1960: 220 [#157] (note). BAUDON, 1961: 73 [#143] (faunal records; Laos). DESCARPENTRIES & VILLIERS, 1963: 4, 14 [#701] ([Note: The original description neither implies not requires that there were syntypes, then Descarpentries & Villiers by the mention about “the holotype” validly fixed the lectotype (ICZN, Article 74.6.)]; lectotype designation; type locality determined by lectotype: “Hoa Binh (R. P. De Cooman), holotype female au Muséum de Paris”; characters; faunal records; Tonkin; Ile de Hainan; Laos; Bhoutan). BAUDON, 1968: 135, 168 [#162] (note). TÔYAMA, 1988: 759 [#2932] (faunal records; Thailand). JENDEK, 2003: 181 [#5823] (remark on lectotype designation). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2003 [#8576] (world catalog).

Type material: Agrilus blatteicollis Bourgoin, 1922 Type locality: not given [Note: Indo-Chine française is cited in the title of the publication]. Type specimens: Primary: Lectotype (Fig. 43P), ♀, designated by DESCARPENTRIES & VILLIERS (1963: 4, 14) [#701], (MNHN): “Hoo-Binh [sic!] Tonkin [p] \ Agrilus blatteicollis Bourgoin [h] [black ink] Type [h] [red ink] \ Muséum Paris 1935 Coll. A. Théry [p]”. Described from unknown number of syntypes. Diagnosis: 8.5-12.8 mm, body slender; elytra silky black; male with cluster of short erect setae in middle of prosternum, prosternal process without erect pubescence or setae. Species group: Imitans. Relevant species: A. nepalensis Tôyama, 1988; A. sichuanus sp. nov. Specimens examined: CHINA: Guizhou: 1 ♂ (EJCB): “China Guizhou 21 26.V., 60 km N of Kaili, 1995, Shibing Yuntai Shan,

E. Jendek & O. Šauša leg.”. Yunnan: 1 ♂ (EJCB): “Yunnan, Jing Gu County, Mt. Yingpantou, 7 14.vi.2009 local collector”. INDIA: Assam: 2 ♀ (EJCB): “NE India, Assam, 1999, 5 km N of Umrongso 700m, 25°27’N, 92°43’E, 17. 25.v., J. Rolčík leg.”. LAOS: Borikhamxai: 1 ♂, 3 ♀ (EJCB): “Laos centr., Bolikhamsai prov., Ban Nape Kaew Nua Pass, 18.4. 1.5.1998, alt. 600±100 m, N 18º22.3, E 105º09.1 (GPS), E. Jendek & O. Šauša leg.”. Louangphrabang: 19 (EJCB): “Laos N, 24.iv. 16.v.1999, Louang Phrabang prov. 20°33 4’ N 102°14’E, Ban Song Cha (5km W) ± 1200 m, Vít Kubáň leg.”. Oudomxai: 1 ♂, 5 ♀ (EJCB): “Laos N (Oudomxai), 1 9.v.2002, ~1100m, 20º45’N, 102º09’E, Oudom Xai (17 km NEE), Vít Kubáň leg.”. Phongsali: 4 (EJCB): “Lao, Phongsaly prov., 21°21’N, 102°03’E, Ban Sano Mai, 19 26.v.2004, 1150 m, P. Pacholátko leg.”. THAILAND: Chiang Mai: 1 (EJCB): “Doi Suthep near Chiang Mai, Thailand, 2.VI.1978, lgt. M. Akiyama”. VIETNAM: 1 (EJCB): “S Vietnam Gia Lai Kon Tum, 60 km N Ankhé, 900 m, Con Hanung 19 27.vi., Kannack, 1983”; 2 ♂ (EJCB): “S Vietnam, 14.10N 108.30E, 40 km NW Ankhe, Buon Luoi, 620 750 m, 28.3. 12.4.1995, Pacholátko & Dembický leg.”.

Adult occurence: 3-4-5-6 (n=12). Faunal records published by BAUDON, 1960 [#157] and TÔYAMA, 1988 [#2932] are also included. Altitude range: 500-1200 m (n=7). Host plants: Unknown. Distribution: CHINA: Guizhou, Hainan,Yunnan. INDIA: Assam, West Bengal. LAOS. THAILAND. VIETNAM. Remarks: Record from “Bhoutan - Maria-Basti” cited by DESCARPENTRIES & VILLIERS, 1963 [#701] pertains to A. nepalensis Tôyama, 1988

Agrilus bocaki Jendek, sp. nov. Fig. 29C (habitus of HT); Fig. 40S (aedeagus)

Description: Holotype 4.6 mm. BODY: Shape: subparallel, Build: slender, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Medial impression: present, Impression (extent): vertex. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: punctures, Sculpture (aspect): regular, Sculpture (density): sparse, Sculpture (intensity): superficial. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate or long. PRONOTUM: Shape: visually square, Sides: arcuate, Maximal width: at middle, Anterior margin: narrower than posterior. Anterior lobe: obvious, Shape: subangulate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: posterolateral. Prehumerus: carinal, Shape: bisinuate, Extent: to anterior pronotal angle, Posterior end: joining posterior pronotal angle, Anterior end: close to pronotal marginal carina or joining pronotal marginal carina, Arc: obvious. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior angles. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: present, Apical part: feebly attenuate. Apices: Arrangement: conjoint, Width: wide, Shape: subtruncate, Truncation:

54 transverse. Elytral pubescence: entire, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: narrow, Anterior margin: arcuate. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: rectangular, Angles (tips): sharp, Disk: impressed. ABDOMEN: Sexual modifications in male: medial sulcus on ventrite 1, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe markedly obtuse. Diagnosis: antennae short, reaching to half of pronotal length; elytral apices transversely subtruncate; basal ventrite with deep medial sulcus across whole length. Species group: Japanocarinatus. Relevant species: A. japanocarinatus Ohmomo, 2002. Type locality: China, Guizhou province, Leigongshan, Xijiang, altitude 1200 -1900 m. Type specimens: Holotype (Fig. 29C), ♂, (EJCB): “China, Guizhou prov., Leigongshan, Xijiang, 29 May - 2 June 1997, 1200 -1900 m, Bolm lgt.”. Adult occurence: 5-6 (n=1). Altitude range: 1200-1900 m (n=1). Host plants: Unknown. Distribution: CHINA: Guizhou. Etymology: Patronymic; the species was named in honor of the Czech entomologist Ladislav Bocák, the eminent specialist in Lycidae (Coleoptera).

Agrilus bonadonai Descarpentries & Villiers Fig. 15B (habitus); Fig. 34R (aedeagus); Fig. 43R (type) bonadonai Descarpentries & Villiers, 1963 (Agrilus) DESCARPENTRIES & VILLIERS, 1963: 4, 15 [#701] (description). BELLAMY, 2008: 2004 [#8576] (subgenus Sinagrilus; world catalog).

Type material: Agrilus bonadonai Descarpentries & Villiers, 1963 Type locality: Tonkin ... Lao-Kay. Also cited from: Tonkin. - Hanoi; Hoa Binh; Laos - Lakhon. Type specimens: Primary: Holotype (Fig. 43R), ♀, (MNHN): “P Nodis Lao Kai [h] \ ♂ Type [p] [red label] \ Agrilus bonadonai n. sp. Holotype ♂ nob[is]. A. Descarpentries et [h] A. Villiers det. 19 [p] 63 [h]”. Secondary: 3 PT (MNHN). Described from unknown number of specimens (HT, AT, PT). Diagnosis: 4.5-5.5 mm; vertex with subparallel sculpture; body less elongate; elytral apices not expanded. Species group: Subrobustus. Relevant species: A. yunnanicola Obenberger, 1936. Specimens examined: CHINA: Anhui: 1 ♀ (EJCB): “China,

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Anhui, 1998, Dabieshan, 65 km SW Huoshan, 1400 m, Bolm lgt. 21 24 Jun”.

Adult occurence: 6 (n=1). Host plants: Unknown. Distribution: CHINA: Anhui. LAOS. VIETNAM.

Agrilus bonnottei Bourgoin Fig. 26C (habitus); Fig. 39E (aedeagus); Fig. 43S (type) bonnottei Bourgoin, 1925 (Agrilus) BOURGOIN, 1925: 131 [#373] (description). OBENBERGER, 1936: 1076 [#2191] (world catalog). BAUDON, 1962: 68 [#145] (cited as bonnotei; faunal record; Laos). DESCARPENTRIES & VILLIERS, 1963: 57, 62 [#699] (cited as bonnotei; characters; faunal records; Laos; Tonkin; Cambodge (?); Birmanie). BAUDON, 1968: 131, 147 [#162] (note). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2005 [#8576] (world catalog).

Type material: Agrilus bonnottei Bourgoin, 1925 Type locality: Laos ou Cambodge. Type specimens: Holotype by monotypy (Fig. 43S), ♀, (MNHN): “Museum Paris, LAOS OU CAMBODGE, BONNOTTE 1911 [p] \ bonnottei A. Bourgoin Type [h] \ Muséum Paris 1935 Coll. A. Théry [p]”. Described from 1 specimen. Diagnosis: 9.2-11 mm; eyes moderate; apicalmost part of elytra pubescent. Species group: Cyaneovirens. Relevant species: A. cyaneovirens Bourgoin, 1922. Specimens examined: CHINA: Yunnan: 1 ♀ (NMPC): “Yun’nan’, Ust’e Nan’cikhè, 200 m, 11.VI.1956, Khuan Kè zhen’ i dr. [in Russian]”; 1 ♂ (EJCB): “S China, S Yunnan, 24 km N of Jiughong: Nobun, 5.vi.2008, 700 m”. LAOS: Louang Namtha: 1 ♀ (EJCB): “N Laos, 14 22.vi.1996, Louang Namtha pr. 10 30 km NW Louang Namtha, C. Holzschuh leg. 800 m”; 1 ♂ (EJCB): “Laos north, 5 11.V.1997, 20 km NW Louang Namtha, N 21°09.2, E 101°18.7, alt. 900±100 m, E. Jendek & O. Šauša leg.”; 1 ♀ (EJCB): “Laos north, 4.V.1997, Louang Namtha env., N 21°00.3, E 101°24.6, E. Jendek & O. Šauša leg.”; 1 ♂, 1 ♀ (EJCB): “Laos, Louang Namtha pr., 21°09’N, 101°19’E, Namtha Muang Sing, 5 31.v.1997, 900 1200 m, Vit Kubáň leg.”; 1 ♀ (EJCB): “Laos north, 13 24.V.1997, 15 km NW Louang Namtha, N 21°07.5, E 101°21.0, alt. 750±100 m, E. Jendek & O. Šauša leg.”. Phongsali: 1 ♀ (EJCB): “Lao N, Phongsaly prov., 21°41’ 2’N, 102°06’ 08’E, Phongsaly env., 28.v. 20.vi.2003, 1500m, P. Pacholátko leg.”. Vientiane: 1 ♂, 1 ♀ (EJCB): “C Laos, Viang Chan prov. Lao Pako resort, 100 m, 50km NE Vientiane, 2002, M. Štrba leg. 28 30.V.”; 5 (EJCB): “Laos, 35 km NE Vientiane, Lao Pako env., alt. 50 m, 31.V. 4.VI.2004, 18º10’N, 102º52’E, E. Jendek & O. Šauša leg.”; 2 ♀ (EJCB): “Laos centr., Viang Chan pr., Ban PA Kho resort, ca.50 km, NE of Vientiane, ~90 m, 18º10’N, 102º52’E, 9. 14.vi.2007, M. Štrba leg.”. THAILAND: Chanthaburi: 3 (EJCB): “E Thailand, 5. 13.5., Chanthaburi Dist., Khao Soi Dao, 1998, J. Horák leg.”; 2 (EJCB): “E Thailand, 5. 13.5., Chanthaburi Dist., Khao Soi Dao, 1998, M. Knížek leg.”. Mae Hong Son: 1 ♀ (EJCB): “Thai, Mae Hong Son pr., 19°27’N, 98°20’E, 1500 m, Soppong 7 12.V., Vít Kubáň leg., 1996”; 1 ♂ (EJCB): “Thailand, 7 10.V., Mae Hong Son pr., 1997, pass near Soppong, 1500 m, 19°27’N, 98°20’E, lgt. S. Bečvář j. & sen.”. Nan: 1 ♀ (EJCB): “NW THAI, Nan District, Pha Khap, 15 20.V.1992, leg. P. Pacholátko”. VIETNAM: Hoa Binh: 1 (MNHN): “Hoa Binh, Tonkin, H. Perrot”.

55

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Adult occurence: 5-6-7 (n=18). Faunal records published by BAUDON, 1962 [#145] and BAUDON, 1968 [#162] are also included. Altitude range: 50-1500 m (n=12). Host plants: Unknown. Distribution: CAMBODIA. CHINA: Yunnan. LAOS. MYANMAR. THAILAND. VIETNAM.

Agrilus brevinotus Jendek, sp. nov. Fig. 27G (habitus of HT); Fig. 39T (aedeagus)

Description: 3.3-5.0 mm (Holotype 4.4 mm). BODY: Shape: subparallel, Color (dorsally): unicolored, Color (dominant): green. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): deep, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: protruding from head outline, Sculpture on vertex: rugae, Sculpture (intensity): superficial. Eyes: Size: large, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short or moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: transverse, rarely visually square, Sides: arcuate, markedly arcuate, rarely straight, Maximal width: at middle, rarely at anterior margin, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles or at level with anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial only or medial and lateral, Medial impression (shape): anteromedial and posteromedial or posteromedial only, Lateral impressions (deepness): shallow, Lateral impressions (width): narrow. Prehumerus: carinal, Shape: bisinuate, Extent: to anterior pronotal angle, Modifications: rarely obsolete in anterior part, Posterior end: joining posterior pronotal angle or joining posterior pronotal margin, Anterior end: joining pronotal marginal carina, Arc: obvious or moderate. Marginal and submarginal carinae: Interspace: moderate, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: bicolored, complementary with pubescence, rarely unicolored, Alternative color (location): apical portion, Humeral carina: absent. Apices: Arrangement: conjoint, Shape: subtruncate, Truncation: transverse, Modifications: sutural rim spinuliform. Elytral pubescence: absent or proximal and distal, Proximal: like V or upturned T, Distal: preapical, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: narrow or moderate, Anterior margin: arcuately emarginate, Emargination (deepness): shallow. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: acute, Angles (tips): sharp, Disk: flat or impressed. ABDOMEN: Sexual modifications in male: medial sulcus on ventrite 1, Apical

margin of pygidium: arcuate, Apical margin of last ventrite: arcuate or subtruncate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate, Sinuosity (deepness): markedly deep. LEGS: Male sexual modifications: metacoxa medially with cluster of setae and metafemur markedly incrassate. Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: elongate. Diagnosis: frons markedly convex; pronotum distinctly transverse with sides obviously arcuate; aedeagus widest in middle. Species group: Cyaneovirens. Relevant species: A. gussakovskiji Alexeev, 1981. Type locality: China, Shanxi province, Lüliang Shan, road Fangshan-Jiaocheng, Hengjian env., altitude 1000 m. Type specimens: Primary: Holotype (Fig. 27G), ♂, (EJCB): “China, W Shanxi, 9.VI.2000, 37.7N, 111.6E, Lüliang Shan, road Fangshan-Jiaocheng, Hengjian env., ~1000m, Jaroslav Turna leg.”. Secondary: CHINA: Gansu: 1 ♂ PT (EJCB): “China, Kansu, Ponggartang, 30.Jun 1992, M. Bok lgt.”; Shaanxi: 2 ♀ PT (EJCB): “China S, Shaanxi, 14-16.6.1995, Qinling Mts. N. slope, 33.56-59’/108.50’ 800-1200 m, Changan Co., L.+R. Businský lgt.”; 3 ♂ PT (EJCB): “China, Shaanxi prov., 21-23.vi.1998, Quin Ling Shan mts, road Baoji-Taibai vill., pass 35 km S Baoji, O. Šafránek et M. Trýzna lgt.”; Shanxi: 12 PT (EJCB): “China, Shanxi prov, 9.VI.2000, Lüliang Shan, 1000 m, road Fangshan-Jiaocheng, Hengjian env., Zd. Jindra”; 3 PT (EJCB): “China, W Shanxi, 9.VI.2000, 37.7N, 111.6E, Lüliang Shan, road FangshanJiaocheng, Hengjian env., ~1000m, Jaroslav Turna leg.”; Sichuan: 1 ♀ PT (EJCB): “China-Sichuan, JuzhaigouNanping, 7.6.-9.6.2007, leg. E. Kučera”. Adult occurence: 6 (n=6). Altitude range: 800-1200 m (n=3). Host plants: Unknown. Distribution: CHINA: Gansu, Shaanxi, Shanxi, Sichuan. Etymology: The specific name is a combination of the Latin adjective brevis (short) and the Greek noun notos (back). It refers to the transverse shape of pronotum in this species.

Agrilus businskorum Jendek, sp. nov. Fig. 27E (habitus of HT); Fig. 39R (aedeagus)

Description: Holotype 4.0 mm. BODY: Shape: subparallel, Color (dorsally): bicolored, Color (dominant): green. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: protruding from head outline, Sculpture on vertex: rugae, Sculpture (density): dense, Sculpture (intensity): superficial. Eyes: Size: large, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short. PRONOTUM: Shape: visually square, Sides: arcuate or subangulate, Maximal width: at

56 middle, Anterior margin: subequal to posterior. Anterior lobe: obvious, Shape: arcuate, Position to anterior angles: projecting beyond anterior pronotal angles. Posterior angles: Shape: acute or rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial or posteromedial only, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial, Lateral impressions (width): narrow. Prehumerus: carinal, Shape: bisinuate, Extent: to anterior pronotal angle, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: narrow or moderate, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior angles. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: bicolored, complementary with pubescence, Alternative color (location): apical portion, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate or conjoint, Width: wide, Shape: subtruncate, Truncation: transverse. Elytral pubescence: proximal and distal, Proximal: like V or upturned T, Distal: preapical, Color: unicolored or bicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: acute, Angles (tips): sharp, Disk: impressed. ABDOMEN: Sexual modifications in male: medial sulcus on ventrite 1, Apical margin of pygidium: arcuate, Apical margin of last ventrite: subtruncate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate, Sinuosity (deepness): markedly deep. LEGS: Male sexual modifications: metacoxa medially with cluster of setae, lower margin of metafemur with row of hairs and markedly incrassate. Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical. Diagnosis: eyes large; pronotum with feebly arcuate or subangulate sides, widest in middle; elytral apices wide and subtruncate. Species group: Cyaneovirens. Relevant species: A. gussakovskiji Alexeev, 1981. Type locality: China, Shaanxi province, Changan Co., Qinling Mts., north slope, N 33.56-59’, E 108.50’, altitude 800-1200 m. Type specimens: Holotype (Fig. 27E), ♂, (EJCB): “China S, Shaanxi, 14-16.6.1995, Qinling Mts. N. slope, 33.5659’/108.50’ 800-1200 m, Changan Co., L.+R. Businský lgt.”. Adult occurence: 6 (n=1). Altitude range: 800-1200 m (n=1). Host plants: Unknown. Distribution: CHINA: Shaanxi. Etymology: Patronymic; the species was named in honour of Mr. and Mrs. Businský who collected the holotype.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Agrilus caligans Bourgoin Fig. 14K (habitus); Fig. 34O (aedeagus); Figs. 43A, 44A (types) caligans Bourgoin, 1925 (Agrilus) BOURGOIN, 1925: 131 [#373] (description). OBENBERGER, 1936: 1076 [#2191] (world catalog). BAUDON, 1962: 69 [#145] (faunal record; Laos). DESCARPENTRIES & VILLIERS, 1963: 3, 10 [#701] ([Note: An inference of type is not valid lectotype designation because of more syntypes (ICZN, Article 74.3)]; characters; faunal records; Tonkin; Laos). BAUDON, 1968: 108, 115 [#162] (note). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2012 [#8576] (subgenus Sinagrilus; world catalog). assamensis Obenberger, 1936 (Agrilus) syn. nov. OBENBERGER, OBENBERGER, 1936: 117 [#2205] (description). 1936: 1074 [#2191] (world catalog). BELLAMY, 2008: 1978 [#8576] (world catalog).

Type material: Agrilus caligans Bourgoin, 1925 Type locality: Tonkin: Hoa Binh. Type specimens: Primary: Lectotype (Fig. 44A) by present designation, ♂, (MNHN): “Ho Binh de Cooman [h] \ Agrilus caligans Type Bourgoin [h] \ Muséum Paris 1935 coll. A. Théry [p]”. Secondary: 6 PLT (MNHN). Described from unknown number of syntypes. Agrilus assamensis Obenberger, 1936 Type locality: Assam. Type specimens: Primary: Lectotype (Fig. 43A) by present designation, ♀, (NMPC): “Assam [h] \ Typus [p] [red label] \ Agrilus assamensis m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Diagnosis: 4.9-6.6 mm; body robust; pronotum widest around middle; elytra with distal pubescence; elytral apices almost conjointly arcuate; aedeagus with obvious membranous lobes. Species group: Subrobustus. Relevant species: A. subrobustus Saunders, 1873. Specimens examined: CHINA: Yunnan: 1 ♂ (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”; cca 30 (EJCB): “CH Yunnan, 13 14.VI.1995, 100 km W of Kunming, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”; 1 ♂ (EJCB): “Yunnan, 1500 2500 m, 25°22’N, 98°49’E, 17 24 / 5 Gaoligong mts, Vít Kubáň leg., 1995”. INDIA: Meghalaya: 1 ♀ (EJCB): “NE India, Meghalaya, 1999, 9 km NW of Jowai, 1400 m, 25°30’N, 92°10’E, 11 15.v., Dembický & Pacholátko leg.”. Uttarakhand: 2 (EJCB): “Dehra Dun, Cujrara”. West Bengal: 1 ♀ (EJCB): “India, West Bengalen, Darjeelling dist., Kalimpong, 800 m, N. Dangal leg. / Sindaybung 1.5.1989”; 1 (EJCB): “India, West Bengalen, Distr. Darjeelling, Kalimpong, Khani Busty, 10.IV.1991, leg. N. Dangal, 900 m”. LAOS: Borikhamxai: 1 ♂ (EJCB): “Laos, Boli Kham Xai prov., 18º16’N, 103º11’E, 70 km NNE Vientiane, 27 30.iv.1997, 150 m, Vít Kubáň leg.”. Louang Namtha: 1 ♀ (EJCB): “N Laos, 14 22.vi.1996, Louang Namtha pr. 10 30 km NW Louang Namtha, C. Holzschuh leg. 800 m”; 1 ♂ (EJCB): “Laos, Louang Namtha pr., 21°09’N, 101°19’E, Namtha Muang Sing, 5 31.v.1997, 900 1200 m, Vit Kubáň leg.”; 3 ♂, 3 ♀ (EJCB): “Laos north, 24 30.V.1997, 20 km NW Louang Namtha, N 21°09.2, E 101°18.7, alt. 900±100 m, E. Jendek & O. Šauša leg.”. Louangphrabang: 9 (EJCB): “Laos N, 23.iv.1999, Louangphrabang prov. 20°42’ N 102°54’E, 25 km E Muang Ngoy, 1000 m, Vít Kubáň leg.”. Oudomxai: cca 40 (EJCB): “Laos N (Oudomxai), 1 9.v.2002, ~1100m, 20º45’N, 102º09’E, Oudom Xai (17 km NEE), Vít Kubáň leg.”. Phongsali:

57

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA 22 (EJCB): “Lao N, Phongsaly prov., 21°41’ 2’N, 102°06’ 08’E, Phongsaly env., 28.v. 20.vi.2003, 1500m, P. Pacholátko leg.”; 1 ♂, 2 ♀ (EJCB): “Lao N, Phongsaly prov., 21°41’ 2’N, 102°06’ 08’E, 28.v. 20.vi.2003 Phongsaly env., ~1500m, Vít Kubáň leg.”. Vientiane: 1 ♀ (EJCB): “Laos centr., 27.IV. 1.V.1997, 70 km NE Vientiane, Ban Phabat env., 150 m, N 18°16.1, E 103°10.9, E. Jendek & O. Šauša leg.”. NEPAL: 2 ♀ (EJCB): “C Nepal, Gandaki, 1986, Kaski Distr., Chandrakot Hille, 1000 1600 m, leg. C. Holzschuh, 9.VI.”. 1 ♂ (EJCB): “E Nepal, Koshi, Tamur River, Phulvari Waku, 1200 1600 m, 9.VI., leg. Holzschuh, 1985”. 1 ♀ (EJCB): “Nepal Centr., Narayani Zone, Chitawan Distr., Chitawan Nat. P., Sauraha Thati Bagh Mara, 200 500 m, 17 21.v.1996, P. Čechovský leg.”. 1 (EJCB): “E Nepal, Dhankuta, Arun Valley, Lamobagar Gola, 27.5. 3.6.1980, 1000 1400 m, leg. C. Holzschuh”; 11 (EJCB): “W Nepal, NW Pokhara, Khali Gandaki Khola 1100 1300 m, Tatopani, 12 14.V.1994, leg. C. Holzschuh”; 1 (EJCB): “Nepal centr., Saura[e?]ha, 1.5.1994, L. Vojkuvka leg.”. THAILAND: 2 (MNHE): “Thailand, 02.05.2005, Ban O, 17º59’36N/99º31’57E, leg. U. Roosileht”. Chiang Mai: 2 ♂, 2 ♀ (EJCB): “Thailand bor, 19 21.5.1997, Chiang Dao, lgt. M. Snížek”. Kanchanaburi: 4 ♂, 2 ♀ (EJCB): “Thai, 9 13.IV.1991, Thimonghta, 350 m, 15°02’N, 98°35’E, Vít Kubáň leg.”. Mae Hong Son: 1 ♂ (EJCB): “NW Thai 8 18.V.1992, Mae Hong Son, Ban Huai Po, J. Horák leg.”; 1 ♀ (EJCB): “Thailand, 1 8.5.1993, Soppong Pai, 1800 m, Pacholátko & Dembický leg.”; 1 ♀ (EJCB): “Thailand bor.occ., 28.5. 5.6., Pai, Soppong, lgt. M. Snížek, 1997”. Nan: 1 ♂ (EJCB): “Thailand N, Nan prov., Doi Phu Kha N.P. Headq., 19º13’N 101º07’E, 1500m, 22 26.IV.1999, D. Hauck & M. Říha leg.”. VIETNAM: Ninh Binh: 1 ♀ (EJCB): “Vietnam, Cuc phuong, 24 25.V.1986, Han nam ninh prov., Jan Horák lgt.”.

Adult occurence: 4-5-6-7 (n=30). Altitude range: 1502500 m (n=21). Host plants: Unknown. Distribution: CHINA: Yunnan. INDIA. LAOS. NEPAL. THAILAND. VIETNAM.

Agrilus caudicula Jendek, sp. nov. Fig. 28G (habitus of HT); Fig. 40K (aedeagus)

Description: 4.3-5.6 mm (Holotype 4.5 mm). BODY: Shape: subparallel, Build: robust, Color (dorsally): unicolored, Color (dominant): green. HEAD: Medial impression: absent, Epistoma: with raised upper margin. Frons: Shape: moderately convex or markedly convex, Outline: protruding from head outline or not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: punctures or rugae, Sculpture (aspect): irregular. Eyes: Size: large, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short, Sexual modification in male: not apparent. PRONOTUM: Shape: visually elongate or visually square, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior or narrower than posterior. Anterior lobe: obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles or projecting beyond anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): entire or anteromedial and posteromedial, Medial impression (deepness): shallow. Prehumerus: carinal,

Shape: bisinuate, Extent: beyond half of pronotal length or to anterior pronotal angle, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina or joining anterior pronotal angle, Arc: moderate. Marginal and submarginal carinae: Interspace: wide, Convergence: strongly convergent, Junction: present, Narrowest point: at posterior third of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: conjoint, Width: wide, Shape: arcuate, Modifications: sutural rim spinuliform. Elytral pubescence: entire, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: with pit, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate or large, Anterior margin: subtruncate. Prosternal process: moderate, Shape: narrowed, subparallel or dilated, Sides: straight or arcuate, Angles: acute or rectangular, Angles (tips): sharp, Disk: impressed with sides protruding. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate. LEGS: Male sexual modifications: metafemur markedly incrassate. Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): markedly dorsoventrally bent, Ovipositor: uritiform. Diagnosis: apex of elytra with sutural rim projecting to short spinula. Species group: Viridiobscurus. Relevant species: A. foramenifer sp. nov. Type locality: China, Sichuan province, Wudu, 31°41’N, 104°23’E. Type specimens: Primary: Holotype (Fig. 28G), ♂, (EJCB): “China-Sechuan [sic!], Wudu, 31°41’N, 104°23’E, 30.5.2.6.1997, lgt. E. Kučera”. Secondary: CHINA: Guizhou: 1 ♀ PT (EJCB): “China-Guizhou 21-26.V., 60 km N of Kaili, 1995, Shibing-Yuntai Shan, E. Jendek & O. Šauša leg.”; Shaanxi: 1 ♀ PT (EJCB): “China-Shaanxi, Lueang env., 15 km NW, 26.5.-31.5.2001, lgt. E. Kučera”; Sichuan: 1 ♀ PT (EJCB): “China-Sichuan, Pingwu, 32°15’N, 104°16’E, 3-9.6.1997, lgt. E. Kučera”. Adult occurence: 5-6 (n=4). Host plants: Unknown. Distribution: CHINA: Guizhou, Shaanxi, Sichuan. Etymology: The specific name was derived from the Latin diminutive of the noun cauda (tail or tail-like appendage) in apposition. It refers to the sutural rim of the elytral apex projecting in the form of a short spine.

Agrilus chaetocoxalis Jendek, sp. nov. Fig. 22J (habitus of HT); Fig. 37S (aedeagus)

Description: 7.5-8.3 mm (Holotype 8.1 mm). BODY: Shape: subparallel, Build: robust, Color (dorsally): bicolored, Color

58 (dominant): black. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): deep, Epistoma: with raised upper margin. Frons: Shape: flat or moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): irregular, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: large, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square or transverse, Sides: straight or arcuate, Maximal width: at anterior margin or at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate or obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial. Prehumerus: tubercular or carinal, Shape: straight, Extent: close to posterior pronotal angle, Posterior end: joining posterior pronotal angle. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate or submarginal carina divergent at posterior angles. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: conjoint, Width: wide, Shape: subtruncate, Truncation: transverse, Modifications: sutural rim spinuliform. Elytral pubescence: absent. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): moderate. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: rectangular or obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate, Sinuosity (deepness): markedly deep, Sinuosity (width): very narrow. LEGS: Male sexual modifications: metacoxa medially with cluster of setae and metafemur markedly incrassate. Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): feebly asymmetrical, Aedeagus (shape): apex of medial lobe sharply pointed, Ovipositor: elongate. Diagnosis: antennae slightly dilated; prehumerus very short; elytral apices transverselly subtruncate, with spinuliform sutural rim. Species group: Sulcicollis. Relevant species: A. paimpolaiseae Baudon, 1968. Type locality: Central China, Shaanxi province, Qinling Shan, 6 km eastern of Xunyangba, altitude 1000-1300 m. Type specimens: Primary: Holotype (Fig. 22J), ♂, (EJCB): “C-China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000-1300m, 23.V.-13.VI. leg. C. Holzschuh 2000”. Secondary: CHINA: Hubei: 1 ♂ PT (EJCB): “China, central

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Hubei, Dahongshan, 31.5N 113.0E, 30.iv-1.v., Jaroslav Turna leg., 2005”; Shaanxi: 1 ♀ PT (EJCB): “China, 10001300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.13.vi.1998, I. H. Marshal leg.”. Adult occurence: 5-6 (n=3). Altitude range: 1000-1300 m (n=2). Host plants: Unknown. Distribution: CHINA: Hubei, Shaanxi. Etymology: The specific name is a combination of chaeto-, derived from the Greek chaite (long hair), and the Latin noun coxa (hip). It refers to the cluster of setae on the medial side of the metacoxa in the male of this species.

Agrilus charismaticus Jendek Fig. 30C (habitus of HT); Fig. 41F (aedeagus) charismaticus Jendek, 2000 (Agrilus) JENDEK, 2000: 153 154 [#4853] (description). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2023 [#8576] (world catalog).

Type material: Agrilus charismaticus Jendek, 2000 Type locality: China, Fujian prov. SHAOWU env. Type specimens: Primary: Holotype (Fig. 30C), ♂, (EJCB): “China, Fujian prov [ince]. SHAOWU env.”. Described from 1 specimen. Diagnosis: 8.4 mm; dorsally strikingly bicolored; pronotum widest at anterior margin, disk with deep medial impression; prehumerus absent; elytra glabrous with pair of adsutural pubescent spot in apical third; parameres of aedeagus with obvious membranous lobes. Species group: Unassigned. Relevant species: Unknown. Specimens examined: Known only from the holotype.

Host plants: Unknown. Distribution: CHINA: Fujian.

Agrilus chekiangensis Gebhardt Fig. 25F (habitus); Fig. 38Y (aedeagus); Figs. 44I, 52R (types) chekiangensis Gebhardt, 1929 (Agrilus) GEBHARDT, 1929: 33 34 [#1051] (description). OBENBERGER, 1936: 969 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 355 [#2429] (checklist; China). ALEXEEV, 1998: 373 [#4322] (subgenus Quercuagrilus; subgeneric classification). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Zheijang). JENDEK, 2005: 5 6 [#7217] (lectotype designation; synonymy; faunal records; distributional overview). JENDEK, 2006: 391 [#7441] (subgenus Quercuagrilus; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (checklist; Japan). BELLAMY, 2008: 2023 [#8576] (world catalog). semivittatus Tôyama, 1985 (Agrilus) TÔYAMA, 1985: 44 45 [#2924] (description). TÔYAMA, 1985: 22 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 38 [#4091] (checklist; Japan: Kyushu). JENDEK, 2005: 5 6 [#7217] (synonym

59

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA of chekiangensis). JENDEK, 2006: 391 [#7441] (synonym of chekiangensis; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (synonym of chekiangensis). BELLAMY, 2008: 2023 [#8576] (synonym of chekiangensis; world catalog).

Type material: Agrilus chekiangensis Gebhardt, 1929 Type locality: China: Chekiang. Type specimens: Primary: Lectotype (Fig. 44I), ♀, designated by JENDEK (2005: 5-6) [#7217], (NMPC): “China Chekiang [p] \ Typus [p] [red label] \ A. chekiangensis m. Type [h] Det. Dr. A. v. Gebhardt [p] \ Mus. Nat. Pragae Inv. [p] 26477 [h] [orange label]”. Secondary: 1 PLT (RMNH). Described from unknown number of syntypes. Agrilus semivittatus Tôyama, 1985 Type locality: Mt. Konpira, Nagasaki City, Nagasaki Pref. Type specimens: Primary: Holotype (Fig. 52R), ♂, (NSMT): “Mt. Kompira, Nagasaki C. 1. VII. 1954, Coll. S. Ikushima [h] \ Holotype [p] Agrilus semivittatus M. Tôyama, 1985 [h] [red label]”. Secondary: 1 PT (NSMT). Described from 6 specimens (HT, AT, PT). Diagnosis: 3.6-5.7 mm; dorsally black; apical third of elytra more attenuate; prosternal process feebly enlarged; aedeagus with subangulate sides. Species group: Tempestivus. Relevant species: A. fevreae Descarpentries & Villiers, 1963. Specimens examined: CHINA: Fujian: 1 ♂, 1 ♀ (EJCB): “China, Fujian prov., Shaowu env., 23 27.vi.1991, leg. Nikodým” (JENDEK, 2005 [#7217]); 1 ♀ (EJCB): “China, Fujian prov. bor. occ., Shaowu env. 25 km road, Shaowu Taining, 6 10.7., R. Červenka lgt. 1991” (JENDEK, 2005 [#7217]). Hunan: 1 ♂ (EJCB): “China, Hunan SE, 1994, Guidong env. 26 31.V., 26.04N 113.56E, E. Jendek & O. Šauša leg.” (JENDEK, 2005 [#7217]). Jiangxi: 1 ♀ (EJCB): “China, Jiangxi W, Jingggang Shan Ciping, 2 14.VI.1994, E. Jendek & O. Šauša leg.” (JENDEK, 2005 [#7217]). Shaanxi: 1 ♀ (EJCB): “China, Shaanxi pr. Hua Shan, 17 21.6.1991 R. Dunda lgt.” (JENDEK, 2005 [#7217]). Sichuan: 1 ♀ (EJCB): “China: Sichuan, Guanxian 600 m, 12 14.VII.1990, L. & M. Bocák lgt.” (JENDEK, 2005 [#7217]); 1 ♀ (EJCB): “China Sichuan, Pingwu, 32°15’N, 104°16’E, 3 9.6.1997, lgt. E. Kučera” (JENDEK, 2005 [#7217]). Yunnan: 40 (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.” (JENDEK, 2005 [#7217]); 14 (EJCB): “Yunnan 2000m, 25.03N 101.55E, Yipinglang 8 10/6.,Vít Kubáň leg. 1993” (JENDEK, 2005 [#7217]); 1 ♀ (EJCB): “China, Yunnan W, Tengchong env., 10 13.VI.1993, E. Jendek & O. Šauša leg.” (JENDEK, 2005 [#7217]); 2 ♂ (EJCB): “Yunnan 1500 1600m, 23.04N 104.25E, Maguan 25 26.6.1994, Vít Kubáň leg.” (JENDEK, 2005 [#7217]); 2 ♀ (EJCB): “China, Yunnan prov., Lufeng, Yipinglang, 23.VII.1994, leg. Holzschuh” (JENDEK, 2005 [#7217]). JAPAN: Kyushu: 1 ♂ (EJCB): “Mt. Homan San, Dazaifu city, Fukuoka pref., 9.VI.1993, S. Ohmomo leg.” (JENDEK, 2005 [#7217]).

Adult occurence: 5-6-7 (n=14). Altitude range: 600-2000 m (n=3). Host plants: Unknown. Distribution: CHINA: Fujian, Hunan, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang. JAPAN: Kyushu.

Agrilus chlorophyllus Abeille de Perrin Fig. 29E (habitus); Fig. 40T (aedeagus); Figs. 44K, 46B, 50U (types) chlorophyllus Abeille de Perrin, 1904 (Agrilus) ABEILLE DE PERRIN, 1904: 223 [#22] (description). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 658 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 969 [#2191] (world catalog). SCHAEFER, 1949: 412 [#2565] (characters; references; faunal records; biology; notes; ditributional overview; notes). HARIRRI, 1971: 195 [#8105] (checklist; Syria). NIEHUIS & TEZCAN, 1993: 29 30, 46 [#1910] (subgenus Anambus; faunal records from Turkey; distribution and host plants summary; Turkey). FINKEL, CHIKATUNOV & NERO, 2002: 127 [#5708] (distributional summary; adult activity; Israel). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2025 [#8576] (world catalog). esfandiarinus Davatchi, 1958 (Agrilus; subspecies of viridi caerulans cited as viridicerulans) DAVATCHI, 1958: 68 69 [#3658] (description; [Note: Obenberger is cited as the author but he never described this taxon and Davatchi also presents characters]). SALAVATIAN, 1950: 11 [#5627] (subspecies of viridicaerulans; faunal record; Iran). BALACHOWSKY, DAVATCHI & DESCARPENTRIES, 1962: 298 [#96] (subspecies of viridicaerulans; Obenberger is cited as the author; characters; biology; distribution; economic importance; bibliography). GENTRY, 1965: 132 [#8590] (subspecies of viridicaerulans; cited as esfandiarii; pest of crop). ADELI, 1972: 10 [#4871] (subspecies of viridicaerulans; Obenberger is cited as the author; Iran). GURYEVA, 1974: 100 101 [#5428] (subspecies of viridicaerulans; pest of agriculture cultures; references; Iran). RAJABI, 1976: 128 [#8237] (subspecies of viridicaerulans cited as viridicoerulans; checklist; Iran). ABAI & ADELI, 1984: 66 [#5808] (pest; Iran). NIEHUIS & TEZCAN, 1993: 1, 30 [#1910] (synonym of chlorophyllus). MODARRES, 1997: 132 [#8162] (subspecies of viridicaerulans; checklist of agricultural pest; Iran). ABIVARDI, 2001: 675, 678 [#8093] (subspecies of viridicaerulans cited as viridicerulans; bibliography overview of Iranian fauna). BORUMAND, 2002: 50 51 [#8252] (subspecies of roscidus; faunal records; Iran). JENDEK, 2006: 397 [#7441] (synonym of chlorophyllus; Palaearctic catalog). BELLAMY, 2008: 2025 [#8576] (synonym of chlorophyllus; world catalog). pistaciophagus Alexeev & Kulinitsh, 1962 (Agrilus) syn. nov. ALEXEEV & KULINITSH, 1962: 92 94 [#3235] (description). TER MINASYAN, 1955: 452 [#7893] ([Note: Misidentified as nivosus]; host plants; distribution; references). MAKHNOVSKIJ, 1959: 32 [#5829] ([Note: Misidentified as tschitscherini according to Makhnovskij, 1966: 80 [#7356]]; characters; biology; Uzbekistan). KULINITSH, 1963: 154 [#8648] (ecological notes; Tajikistan). ZNAMENSKIJ, 1963: 152 [#8647] (cited as “sp.”; pest; Turkmenistan). MAKHNOVSKIJ, 1966: 80 [#7356] (notes; biology; Kyrgyzstan). KADYROV, 1973: 69 [#8632] (notes; Tajikistan). KADYROV, 1973: 51, 55 [#8631] (biology; Tajikistan). GURYEVA, 1974: 100 [#5428] (pest of agriculture cultures; references; Uzbekistan; southeastern Turkmenia; Tadjikistan). KADYROV, 1976: 52 [#3461] (ecology; Tajikistan). ROMANENKO, 1976: 64 66 [#8635] (characters; larva, biology, harms; eradication; Kyrgyzstan). KRYZHANOVSKII, 1984: 20, 108 [#8665] (pest; Central Asia). VOLKOVITSH & ALEXEEV, 1992: 163 164 [#2973] (faunal records; Turkmenistan). ALEXEEV, 1993: 116 [#4889] (Turkmenistan). NIEHUIS & TEZCAN, 1993: 30 [#1910] (subgenus Anambus; presumed synonym of chlorophyllus). VOLKOVITSH & ALEXEEV, 1994: 437 [#2974] (checklist; Turkmenistan). OVTCHINNIKOV, 1996: 132 [#8621] (cited as pistaceophagus;

60 checklist; Kyrgyzstan). ALEXEEV, 1998: 377 [#4322] (subgenus Rosagrilus; subgeneric classification). JENDEK, 2006: 394 [#7441] (subgenus Rosagrilus; Palaearctic catalog). BELLAMY, 2008: 2237 [#8576] (subgenus Rosagrilus; world catalog).

Type material: Agrilus chlorophyllus Abeille de Perrin, 1904 Type locality: Akbès (Syrie). Type specimens: Primary: Lectotype (Fig. 44K) by present designation, ♂, (MNHN): “Akbès [h] \ Muséum Paris Coll. A. de PERRIN 1919 [p] \ chlorophyllus Abeille [h] Type [p] [red ink]”. Secondary: 4 PLT (MNHN). Described from unknown number of syntypes. Agrilus viridicaerulans esfandiarinus Davatchi, 1958 Type locality: [Iran] régions de Kerman, Kazvin et Fars. Type specimens: Primary: Lectotype (Fig. 46B)by present designation, ♂, (NMPC): “Kazvin, Iran, V.1948, Kiziuchin \ Typus [p] [red label] \ Agrilus viridicoerulans Esfandiarinus m. [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Agrilus pistaciophagus Alexeev & Kulinitsh, 1962 Type locality: Kondara [in Russian]. Type specimens: Primary: Holotype (Fig. 50U), sex not examined, (ZIN, temporarily in AAOR). Secondary: 1 PT (NMPC); 1 PT (ZIN); 1 PT (HNHM). Described from 45 specimens (HT, AT, PT). Diagnosis: 3-5.7 mm; antennomeres 5-11 sharply pointed. Species group: Roscidus. Relevant species: A. roscidus Kiesenwetter, 1857. Specimens examined: IRAN: 1 ♂ (EJCB): “Iran SW, Firuz Abad, 16.5.2005, Dr. J. Romsauer leg.”. TAJIKISTAN: 1 (EJCB): “Tadjikistan, Pervomaysk, vi.1975”; 1 (EJCB): “USSR, Tadzikistan, Javroz, 19.4.1978, Dušanbe 40 km, Fr. Navrátil lgt.”. TURKEY: 1 (EJCB): “Turcia, 14.6.1972, Germik Sinek, Lodos Isl.”. TURKMENISTAN: 5 (ZIN): “Turkmensk. SSR, Ashkhabadsk. obl., Serakhskii r n., yr. Kirlsk, 27 V 1952 [in Russian]”. UZBEKISTAN: 1 (EJCB): “Uzbekistan, Taskent, Aktas, ex larva 1978, ex Pistacia vera, Sv. Bílý leg.”; 1 (EJCB): “26.6.1981, Aktasch, Taschkent, lgt. Ing. Mráček”; 6 (EJCB): “Uzbekistan, W Gissar, Kyzylsu riv., Kaltakol vill., 22.vi.1999”.

Adult occurence: 4-5-6 (n=7). Host plants: Pistacia: SALAVATIAN, 1950 (as esfandiarinus) [#5627]; TER MINASYAN, 1955 (as pistaciophagus) [#7893]; MAKHNOVSKIJ, 1959 (as pistaciophagus) [#5829]; KULINITSH, 1963 (as pistaciophagus) [#8648]; ZNAMENSKIJ, 1963 (as pistaciophagus) [#8647]; GENTRY, 1965 (as esfandiarinus) [#8590]; MAKHNOVSKIJ, 1966 (as pistaciophagus) [#7356]; ADELI, 1972 (as esfandiarinus) [#4871]; KADYROV, 1973 (as pistaciophagus) [#8632]; KADYROV, 1973 (as pistaciophagus) [#8631]; GURYEVA, 1974 (as pistaciophagus) [#5428]; KADYROV, 1976 (as pistaciophagus) [#3461]; ABAI & ADELI, 1984 (as esfandiarinus) [#5808]; VOLKOVITSH & ALEXEEV, 1994 (as pistaciophagus) [#2974]. – P. mutica: DAVATCHI, 1958 (as esfandiarinus) [#3658]; BALACHOWSKY, DAVATCHI & DESCARPENTRIES, 1962 (as esfandiarinus) [#96]; GURYEVA, 1974 (as esfandiarinus) [#5428]. – P. vera: ALEXEEV & KULINITSH, 1962 (as pistaciophagus) [#3235]; ROMANENKO, 1976 (as pistaciophagus) [#8635]; VOLKOVITSH & ALEXEEV, 1992 (as pistaciophagus) [#2973]; ALEXEEV, 1993 (as

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA pistaciophagus) [#4889]; NIEHUIS & TEZCAN, 1993 [#1910]; GURYEVA, 1974 (as esfandiarinus) [#5428]. Distribution: IRAN. ISRAEL. KYRGYZSTAN. SY-RIA. TAJIKISTAN. TURKEY. TURKMENISTAN. UZBEKISTAN. Remarks: Not recorded from East Asia but possible in western China.

Agrilus chujoi Kurosawa Fig. 12E (habitus); Fig. 33O (aedeagus); Fig. 44M (type) chujoi Kurosawa, 1985 (Agrilus) KUROSAWA, 1985: 149 151 [#1640] (description). TÔYAMA, 1985: 25 [#3678] (iconography; Japan). HIRASHIMA, 1989: 322 [#8078] (checklist; Japan). LI JINGKE, 1992: 92 [#6755] (checklist; China: Liaoning). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 31 [#4091] (checklist; Japan: Tsushima; Korea; China: south Manchuria). JENDEK, 2001: 2 3, 10 [#5211] (faunal records; key; North Korea; South Korea; Japan). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2027 [#8576] (world catalog).

Type material: Agrilus chujoi Kurosawa, 1985 Type locality: Nii, N. Tsushima, Japan. Type specimens: Primary: Holotype (Fig. 44M), ♂, (NSMT): “Tsushima [p] N. Nii 14. [h] vii. 1969 Y. Kurosawa [p] \ Holotype [p] Agrilus chujoi Y. Kurosawa, 1985 [h]”. Secondary: PT (NSMT). Described from 17 specimens (HT, AT, PT). Diagnosis: 6.2-7.4 mm; body robust, uniformly goldengreen; elytral apex with outer marginal spine rather obtuse and feebly protruding; basal half of elytral disk densely pubescent. Species group: Plasoni. Relevant species: A. huashanus Jendek, 2001. Specimens examined: JAPAN: Tsushima: 1 ♀ (EJCB): “Mt. Ooboshi, Tsushima Is[lands]. 17.VII.1976, N. Nishikawa leg.” (JENDEK, 2001 [#5211]). KOREA NORTH: 1 (EJCB): “D. P. R. Korea, Okryu valley, Kumgansan Mts. [38°36’N, 128°13’E; M. Kozánek pers. com.], lgt. M. Kozánek” (JENDEK, 2001 [#5211]). KOREA SOUTH: 3 ♂, 2 ♀ (EJCB): “Corée, Kosan, Chass. indigènes” (JENDEK, 2001 [#5211]); 1 ♂ (EJCB): “Keijô [ Soul, 37°34’N, 127°00’E], Corea, 7.VIII.1933” (JENDEK, 2001 [#5211]).

Adult occurence: 7-8 (n=14). Faunal records published by KUROSAWA, 1985 [#1640] are also included. Host plants: Pueraria thunbergiana: KUROSAWA, 1985 [#1640]; AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: CHINA: Guangdong, Liaoning. JAPAN: Tsushima. KOREA NORTH. KOREA SOUTH. Remarks: Character cited for A. chujoi by JENDEK (2001: 10 [#5211]) in the key to species is confused and should be changed from “basal half of pronotal disk” to “basal half of elytral disk”.

61

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA

Agrilus ciping Jendek, sp. nov. Fig. 25L (habitus of HT); Fig. 39D (aedeagus)

Description: 6.7-7.7 mm (Holotype 6.7 mm). BODY: Shape: cuneiform, Build: robust, Color (dorsally): unicolored, Color (dominant): green. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow or deep, Epistoma: in plane with frons. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Sculpture on vertex: rugae, Sculpture (aspect): irregular, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: moderate, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: visually elongate, Sides: straight, rarely arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: acute, rarely rectangular, Apex: sharp. Disk: Convexity: strongly convex, Disk impressions: medial only, rarely medial and lateral, Medial impression (shape): entire or anteromedial and posteromedial, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial, Lateral impressions (deepness): shallow, Lateral impressions (width): narrow. Prehumerus: carinal, Shape: arcuate, Extent: to half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: distant from pronotal marginal carina, Arc: obvious. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Width: narrow, Shape: arcuate. Elytral pubescence: sutural wedge, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): shallow, Emargination (width): wide. Prosternal process: wide, Shape: dilated, Sides: straight, rarely arcuate, Angles: rectangular or obtuse, Angles (tips): sharp, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus or distinctly longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed, Ovipositor: elongate. Diagnosis: antennae in male short; medial pronotal impression deep; pronotal carina sharp and obviously arcuate. Species group: Tempestivus. Relevant species: A. aeneoclarus Bourgoin, 1922. Type locality: China, west Jiangxi province, Jinggang Shan, Ciping.

Type specimens: Primary: Holotype (Fig. 25L), ♂, (EJCB): “China, Jiangxi W, Jinggang Shan, Ciping, 2-14.VI.1994, E. Jendek & O. Šauša leg.”. Secondary: CHINA: Jiangxi: 1 ♀ PT (EJCB): “China, Jiangxi W, Jinggang Shan, Ciping, 2-14. VI.1994, E. Jendek & O. Šauša leg.”. Adult occurence: 6 (n=1). Host plants: Unknown. Distribution: CHINA: Jiangxi. Etymology: The specific name is the noun in apposition. It refers to Ciping, the type locality of the species.

Agrilus compacticornis Jendek, sp. nov. Fig. 14G (habitus of HT); Fig. 34L (aedeagus)

Description: 4.6-6.3 mm (Holotype 5.4 mm). BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): green or brown. HEAD: Impression (extent): vertex, Impression (depth): shallow, Epistoma: in plane with frons. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, rarely punctures, Sculpture (aspect): irregular or subparallel, Sculpture (density): dense, Sculpture (intensity): superficial. Eyes: Size: moderate or large, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short, Shape: stout, Sexual modification in male: antennae longer. PRONOTUM: Shape: transverse, Sides: arcuate or markedly arcuate, Maximal width: at anterior margin, rarely at middle, Anterior margin: wider than posterior. Anterior lobe: absent, vague or moderate, Shape: arcuate, rarely subangulate, Position to anterior angles: not reaching level of anterior pronotal angles or at level with anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: flat or moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow. Prehumerus: obsolete, filamentary, rarely carinal, Shape: arcuate, Extent: to third of pronotal length, Posterior end: joining posterior pronotal angle, joining posterior pronotal margin or distant from pronotal angle or margin, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina divergent at posterior angles. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Shape: arcuate. Elytral pubescence: absent or entire, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: large, Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): narrow. Prosternal process:

62 moderate or wide, Shape: narrowed, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: convex. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): shorter or subequal to 2-3 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe markedly obtuse, Ovipositor: uritiform. Diagnosis: body small; prehumerus less elevated; medial lobe of aedeagus markedly widened apically. Species group: Viduus. Relevant species: A. viduus Kerremans, 1914. Type locality: China, Yunnan province, 100 km west of Kunming, Diaolin Natural Reservation. Type specimens: Primary: Holotype (Fig. 14G), ♂, (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”. Secondary: CHINA: Hebei: 1 PT (EJCB): “China-Hebei. Yi-Xian, 20.7-21.7.2000, leg. E. Kučera”; Henan: 1 ♀ PT (EJCB): “China, N. Henan, 11-12.VI., Jiuligou, 550 m, 35°12’N, 112°26’E, Jaroslav Turna leg. 2007”; Shanxi: 2 PT (EJCB): “China, centr. Shanxi, Chao Shan 37.1N, 112.4E, 25 km SE Pingyao., 4-5.VII., Jaroslav Turna leg. 2001”; 2 PT (EJCB): “China, Shanxi, 4-5.VI.2001, Chao Shan 37.1°N, 112.4°E, 25 km SE Pingyao, O. Šafránek leg.”; Sichuan: 1 ♂ PT (EJCB): “China-Sechuan [sic!], Wudu, 31°41’N, 104°23’E, 30.5.-2.6.1997, lgt. E. Kučera”; 2 ♂ PT (EJCB): “China-Sichuan, Pingwu, 32°15’N, 104°16’E, 3-9.6.1997, lgt. E. Kučera”; 2 PT (EJCB): “China-Sichuan, Pingwu, 2.6.4.6.2001, lgt. E. Kučera”; Yunnan: 1 PT (EJCB): “China, Yunnan, 2000-3000m, 27.20N. 100.11E, Habashan mts., SE slope, 10-13/7., Vít Kubáň leg. [19]92”; 3 PT (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”; 1 PT (ZIN): “China, Yunnan, Loc. 9., Road 214, Dali - Lijiang, 65th-71st km NNW of Xiaguan, 26°03’ N, 100°01’-09’ E, 2350-2528 m, 24.05.2002, Konstantinov & Volkovitsh leg.”; RUSSIA: Primorskiy kray: 1 ♀ PT (EJCB): “USSR, Primorskij kraj Arseneev env. VI.1991 M. Štrba leg.”; 1 ♀ PT (EJCB): “Russia: Primorskij k., Rjazanovka, 4-8.VII.1993, Z. Kletečka leg.”; 1 ♂ PT (ZIN): “Rossia, Primorskii krai, Novokachalinsk, 25 km YuZ Slavyanki, b. Gorshkova, dubnyak, luga, 24.viii.2006, S. Belokobyl’skij [in Russian]”. Adult occurence: 5-6-7-8 (n=13). Altitude range: 5503000 m (n=3). Host plants: Unknown. Distribution: CHINA: Hebei, Henan, Shanxi, Sichuan, Yunnan, RUSSIA: Primorskiy kray. Etymology: The specific name is a combination of the Latin adjective compactus (short, thick) and the noun cornu (horn, antenna in insects). It refers to the short and broad antennae of this species.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Agrilus confinis Faldermann sp. inquirenda confinis Faldermann, 1835 (Agrilus) FALDERMANN, 1835: 24 25 [#927] (description). GEMMINGER & HAROLD, 1869: 1438 [#1057] (catalog). SAUNDERS, 1871: 126 [#2552] (catalog). KERREMANS, 1892: 252 [#1397] (catalog). KERREMANS, 1903: 283 [#1441] (catalog). JAKOBSON, 1913: 796 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 976 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 355 [#2429] (checklist; China). HUA LI ZHONG, 2002: 89 [#6696] (checklist; North China). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). LÖBL & SMETANA, 2007: 29, 30 [#8558] (errata for Palaearctic catalog). BELLAMY, 2008: 2368 2369 [#8576] (species incertae sedis; world catalog).

Type material: Agrilus confinis Faldermann, 1835 Type locality: China boreali[s]. Type specimens: Not located. Described from 1 specimen (Holotype by monotypy). Host plants: Unknown. Distribution: North CHINA. Remarks: This taxon is species inquirenda and its name is nomen dubium.

Agrilus conspersus Jendek, sp. nov. Fig. 14B (habitus of HT); Fig. 34G (aedeagus)

Description: 5.2-6.1mm (Holotype 5.9 mm). BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): black. HEAD: Medial impression: absent, rarely present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): irregular, Sculpture (intensity): rough. Eyes: Size: moderate, Shape: protruding from head outline, Lower margin: above antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: visually square, Sides: arcuate, Maximal width: at middle, rarely at anterior margin, Anterior margin: wider than posterior or subequal to posterior. Anterior lobe: moderate, rarely obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: flat, Disk impressions: medial and lateral, Medial impression (shape): entire, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow. Prehumerus: filamentary or carinal, Shape: bisinuate, Extent: to anterior pronotal angle, Posterior end: joining posterior pronotal angle, rarely distant from pronotal angle or margin, Anterior end: joining anterior pronotal angle, Arc: weak. Marginal and submarginal carinae: Interspace: narrow or moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior third of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Width: narrow, Shape: arcuate, rarely subtruncate, Truncation: oblique with

63

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA projecting pleural angle. Elytral pubescence: entire, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: large, Anterior margin: arcuately emarginate or angulately emarginate, Emargination (deepness): shallow, Emargination (width): narrow. Prosternal process: moderate or wide, Shape: subparallel or dilated, Sides: arcuate, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): distinctly longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical. Diagnosis: body small, dorsally uniformly silky black; elytral pubescence entire, without spots. Species group: Plagiatus. Relevant species: A. plagiatus Ganglbauer, 1889. Type locality: China, Yunnan province, 100 km west of Kunming, Diaolin Natural Reservation. Type specimens: Primary: Holotype (Fig. 14B), ♂, (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”. Secondary: CHINA: Yunnan: 5 ♂ PT (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”, 1 ♂ PT (EJCB): “Yunnan 1800-2000m, 25.04N 101.55E, Yipinglang 17-20.6.1994, Vít Kubáň leg.”.

Diagnosis: 3.5-4.0 mm; eyes large; pronotum visually elongate; prehumerus sharp, bisinuate and long, with anterior end joined with pronotal margin beyond pronotal half; elytra with humeral carina. Species group: Unassigned. Relevant species: Unknown. Specimens examined: CHINA: Yunnan: 1 ♀ (EJCB): “1958.VII., Yunnan, Xishuanbanna”. LAOS: Attapeu: 1 ♀ (EJCB): “Laos south, Attapu prov., Bolaven Plateau, 18 30.IV.1999, 15 km SE of Ban Houaykong, Nong Lom (lake) env., N 15°02’, E 106°35’, alt. 800m, E. Jendek & O. Šauša leg.”. Borikhamxai: 1 ♂, 2 ♀ (EJCB): “Laos centr., Bolikhamsai prov., Ban Nape Kaew Nua Pass, 18.4. 1.5.1998, alt. 600±100 m, N 18°22.3, E 105°09.1 (GPS), E. Jendek & O. Šauša leg.”. Champasak: 1 ♂ (EJCB): “Laos south., Champasak prov., Bolaven Plateau, 10 18.IV.1999, Route (No. 23) Pakse Paksong, Ban Itou env. (km.35), alt. 800m, N 15°10.4’, E 106°05.8’ (GPS), E. Jendek & O. Šauša leg.”. Louang Namtha: 1 ♂ (EJCB): “Laos north, 5 11.V.1997, 20 km NW Louang Namtha, N 21°09.2, E 101°18.7, alt. 900±100 m, E. Jendek & O. Šauša leg.”; 1 ♀ (EJCB): “Laos north, 24 30.V.1997, 20 km NW Louang Namtha, N 21°09.2, E 101°18.7, alt. 900±100 m, E. Jendek & O. Šauša leg.”. THAILAND: Chiang Mai: 1 ♂ (EJCB): “Thailand, 24 29. IV.1993, Doi Suthep, Pacholátko & Dembický leg.”; 1 ♀ (EJCB): “Thailand 1000 1600 m, Chiang Mai prov., 20 km NW from Fang, 2 5.v.1996, lgt. S. & E. Becvar”; 1 ♀ (EJCB): “Thailand, 1000 1600 m, Chiang Mai, 26.iv. 9.v.1996, 18°49’N, 98°54’E, Doi Pui Mt., lgt. S. Becvar”; 2 ♀ (EJCB): “Thailand, Chiang Mai prov., 19°19’N, 98°50’E, 1400 m, Sanpakia vill., 1 15.V.1998, Vít Kubáň leg.”. Kamphaeng Phet: 1 ♂, 1 ♀ (EJCB): “Thailand West, Klong Lan n. p., 50 km SW Kamphaeng Phet, J. Rejsek, 2. 5.7.1997”. Nan: 1 ♀ (EJCB): “Thailand N, Nan prov., Bo Klua, 19°03’N, 101°10’E, 700 m, 28.IV. 1.V.1999, D. Hauck leg.”.

Adult occurence: 5-6 (n=2).

Adult occurence: 2-4-5-7 (n=14). Faunal records published by BAUDON, 1963 [#148] and 1968 [#162] are also included. Altitude range: 500-1600 m (n=9).

Host plants: Unknown.

Host plants: Unknown.

Distribution: CHINA: Yunnan.

Distribution: CHINA: Yunnan. LAOS. THAILAND. VIETNAM.

Etymology: The specific name is the Latin adjective conspersus (sprinkled), referring to the appearance of the elytral pubescence of this species.

Agrilus convictor Descarpentries & Villiers Fig. 30D (habitus); Fig. 41G (aedeagus); Fig. 44R (type) convictor Descarpentries & Villiers, 1963 (Agrilus) DESCARPENTRIES & VILLIERS, 1963: 107, 112 [#700] (description). BAUDON, 1963: 53 [#148] (faunal records; Laos). BAUDON, 1968: 133, 156 [#162] (faunal records; Laos). BELLAMY, 2008: 2038 [#8576] (world catalog).

Type material: Agrilus convictor Descarpentries & Villiers, 1963 Type locality: Tonkin: Hoa-Binh. Type specimens: Primary: Holotype (Fig. 44R), ♂, (MNHN): “Hoa Binh Tonkin de Cooman [p] \ Type [p] ♂ [h] [red label \ Agrilus convictor n. sp. Holotype ♂ nob[is]. A. Descarpentries et [h] A. Villiers det. 19 [p] 62 [h]”. Secondary: 2 PT (MNHN). Described from unknown number of specimens (HT, AT, PT).

Agrilus coomani Bourgoin Fig. 10C (habitus); Fig. 32R (aedeagus); Figs. 44S, 49J (types) coomani Bourgoin, 1925 (Agrilus) BOURGOIN, 1925: 112 [#372] (description). OBENBERGER, 1936: 1078 [#2191] (world catalog). DESCARPENTRIES & VILLIERS, 1963: 57, 62 [#699] (lectotype designation: “holotype au Muséum de Paris”; characters; faunal record; Tonkin). DESCARPENTRIES & VILLIERS, 1967: 146 [#710] (diagnosed with albertianus). PENG ZHONGLIANG, 2002: 269 270, 281 [#6628] (characters; new record for China; Fujian, Guangxi). BELLAMY, 2008: 2038 [#8576] (world catalog). milletae Baudon, 1968 (Agrilus) syn. nov. BAUDON, 1968: 90, 100 102 [#162] (description). BELLAMY, 2008: 2186 [#8576] (world catalog).

Type material: Agrilus coomani Bourgoin, 1925 Type locality: Tonkin: Hoa Binh. Type specimens: Primary: Lectotype (Fig. 44S), ♂, designated by DESCARPENTRIES & VILLIERS (1963: 57, 62)

64 [#699], (MNHN): “Hoo-Binh [sic!] Tonkin [p] \ Agrilus Coomani Type A. Bourgoin [h]”. Secondary: 2 PLT (NMPC). Described from unknown number of syntypes. Agrilus milletae Baudon, 1968 Type locality: Laos: Phonthiou. Type specimens: Primary: Holotype (Fig. 49J), ♂, (MHNB): “Phonthoiu 29.iv.[19]64 [h] Laos (Baudon) [p] \ Agrilus milletae mihi [h] A. Baudon det [p] [blue label] \ Type [p][red label]”. Described from: 1 specimen (HT). Diagnosis: 8.3-10 mm; pronotum visually elongate; eyes moderate; elytra with pair of adsutural pubescent spots just before apices. Species group: Spinipennis. Relevant species: A. fornicatus sp. nov. Specimens examined: THAILAND: Mae Hong Son: 1 ♀ (EJCB): “NW Thailand, 1 15.5., 1991, Mae Hong Son, Ban Huai Po, 800 1600 m, S. Bílý leg.”. VIETNAM: 3 ♂ (MNHN): “Hoa Binh, Tonkin”. 1 (EJCB): “Vietnam N, TamDao 900 m, 16 23.V.1991, Strnad Jan lgt.”.

Adult occurence: 5 (n=2). Altitude range: 800-1600 m (n=2). Host plants: Unknown. Distribution: CHINA: Fujian, Guangxi. LAOS. THAILAND. VIETNAM. Remarks: Records from China cited by PENG ZHONGLIANG, 2002 [#6658] need verification.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Humeral carina: absent. Apices: Arrangement: separate, Shape: cuspidate, Position of dominant cusp (spine): medial, Modifications: sutural rim spinuliform. Elytral pubescence: absent. Tomentum: spots, Spots (locations): postmedial. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): moderate. Prosternal process: moderate or wide, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): distinctly longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Ovipositor: elongate. Diagnosis: prehumerus extending to posterior third of pronotal length; elytral apex with long and sharp medial cusp. Species group: Spinipennis. Relevant species: A. insulicola Kerremans, 1912. Type locality: China, Shaanxi province, Qinling mts., 6 km east Xunyangba, altitude 1000-1300 m. Type specimens: Holotype (Fig. 9K), ♀, (EJCB): “China, 1000-1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.-13.vi.1998, I. H. Marshal leg.”. Adult occurence: 5-6 (n=1). Altitude range: 1000-1300 m (n=1).

Agrilus cordicollis Jendek, sp. nov.

Host plants: Unknown.

Fig. 9K (habitus of HT)

Distribution: CHINA: Shaanxi.

Description: Holotype 11.3 mm. BODY: Shape: subparallel, Build: robust, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: large, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles. Posterior angles: Shape: acute, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow. Prehumerus: carinal, Shape: arcuate, Extent: to third of pronotal length, Posterior end: distant from pronotal angle or margin, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: moderate, Convergence: strongly convergent, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored,

Etymology: The specific name is a combination of the Latin nouns cor (heart) and collum (neck). It refers to the heart-like pronotum of this species.

Agrilus coreanus Obenberger Fig. 8K (habitus of LT); Fig. 32G (aedeagus) coreanus Obenberger, 1935 (Agrilus) OBENBERGER, 1935: 162 163 [#2158] (description). MIWA & CHÛJÔ, 1936: 15 [#1809] (catalog; Japan). OBENBERGER, 1936: 978 [#2191] (world catalog). JENDEK, 1995: 145 [#3768] (lectotype designation; characters). ALEXEEV, 1998: 379 [#4322] (subgenus Sinagrilus; subgeneric classification). JENDEK, 2006: 394 [#7441] (subgenus Sinagrilus; Palaearctic catalog). BELLAMY, 2008: 2039 [#8576] (subgenus Sinagrilus; world catalog).

Type material: Agrilus coreanus Obenberger, 1935 Type locality: Korea: Chemulpo. Type specimens: Primary: Lectotype (Fig. 9K), ♂, designated by JENDEK (1995: 145) [#3768], (NMPC): “Korea Čemulpo [ = Inchon] [h] \ Typus [p] [red label] \ Agrilus coreanus m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24291 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 11 mm; frontovertex with shallower medial impression; anterior pronotal lobe obvious, subangulate;

65

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA prehumerus filamentary, extending to anterior pronotal angle. Species group: Sinensis. Relevant species: A. sinensis Thomson, 1879. Specimens examined: Known only from the lectotype.

Host plants: Unknown. Distribution: KOREA SOUTH.

Agrilus cupes Lewis Fig. 18B (habitus); Fig. 35R (aedeagus); Fig. 44U (type) cupes Lewis, 1893 (Agrilus) LEWIS, 1893: 336 [#1722] (description). KERREMANS, 1903: 284 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 16 [#1809] (catalog; Japan). OBENBERGER, 1936: 979 [#2191] (world catalog). KUROSAWA, 1949: 15 [#5028] (characters). IGA, 1955: 79 [#3828] (iconography; Japan). IGA, 1962: 79 [#3351] (iconography; Japan). KUROSAWA, 1963: 153 [#1618] (characters; Japan). KUROSAWA, 1974: 1 [#4576] (note). KUROSAWA, 1974: 3 [#4575] (characters; notes). TÔYAMA, 1985: 30 [#3678] (iconography; Japan). HIRASHIMA, 1989: 322 [#8078] (checklist; Japan). JENDEK, 1994: 22 [#1355] (lectotype designation; characters; Japan; China: Shaanxi). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 184 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 31 [#4091] (checklist; Japan: Honshu; China: Shaanxi). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). NONNAIZAB et al., 1999: 112 [#8247] (checklist; China: Inner Mongolia). AKIYAMA & OHMOMO, 2000: 88 [#6240] (faunal records; Japan (Hiroshima)). JENDEK, 2006: 393 [#7441] (subgenus Robertius; Palaearctic catalog). BELLAMY, 2008: 2045 [#8576] (subgenus Robertius; world catalog).

Type material: Agrilus cupes Lewis, 1893 Type locality: Main Island. Numata and Chiuzenji. Type specimens: Primary: Lectotype (Fig. 44U), ♂, designated by JENDEK (1994: 22) [#1355], (BMNH): “Type H.T. [p] [round label with red border] \ Japan. G. Lewis 1910-320 [p] \ Numata 29. VIII. [18]81 [h] \ Agrilus cupes Lewis Type [by Lewis hand]”. Described from 2 syntypes. Diagnosis: 6.2-7.5 mm; body ventrally markedly lustrous, apical third of elytra shorter; pronotum with postero-lateral impression; prosternal process attenuate. Species group: Betuleti. Relevant species: A. mio sp. nov. Specimens examined: CHINA: Shaanxi: 1 ♀ (EJCB): “China, Shaanxi, pr. Hua Shan, 17. 21.6.1991, R. Dunda lgt.” (JENDEK, 1994 [#1355]). Shanxi: 22 (EJCB): “China, Shanxi prov, 9.VI.2000, Lüliang Shan, 1000 m, road Fangshan Jiaocheng, Hengjian env., Zd. Jindra”. JAPAN: Honshu: 1 ♀ (EJCB): “Iwate ogasawara, Japan, Y. Miwa” (JENDEK, 1994 [#1355]).

Adult occurence: 6 (n=2). Altitude range: ~1000 m (n=1). Host plants: Staphylea bumalda: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: CHINA: Nei Mongol, Shaanxi, Shanxi. JAPAN: Honshu.

Agrilus cuprescens cuprescens (Ménétriés) Figs. 16K 16L (habitus); Fig. 35F (aedeagus); Figs. 42M, 43Z, 44V, 44Q, 45R, 45W, 46L, 48A, 48I, 49M, 50E, 52F (types) cuprescens Ménétriés, 1832 (Buprestis) MÉNÉTRIÉS, 1832: 154 [#1799] (description). FALDERMANN, 1835: 158 159 [#7916] (redescription). FALDERMANN, 1835: 158 159 [#926] (characters). FALDERMANN, 1838: 75 [#928] (checklist; Transcaucasia). GORY & LAPORTE, 1839: 56 57 [#3738] ([Note: misidentification of sericans according to KIESENWETTER, 1857: 127 [#3015]]). MARSEUL, 1857: 92 [#4371] (checklist; Europe). MARSEUL, 1866: 480 481 [#676] (redescription). GEMMINGER & HAROLD, 1869: 1438 [#1057] (catalog). SAUNDERS, 1871: 121 [#2552] (catalog). MARSEUL, 1887: 252 [#4707] (catalog). KERREMANS, 1892: 253 [#1397] (catalog). ABEILLE DE PERRIN, 1897: 8 [#19] (characters in key). KERREMANS, 1903: 284 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). WEISS, 1915: 134 [#5953] ([Note: Misidentified as viridis var. fagi according to host plant cited]; pest; USA: New Jersey). BEDEL, 1921: 204, 213 214 [#170] ([Note: Misidentified as chrysoderes]; characters in key; references; faunal records; France). FROST, 1924: 27 [#1030] ([Note: Misidentified as viridis subspecies fagi]; collecting record; Massachusetts). CHAMBERLIN, 1926: 85 [#468] ([Note: Misidentified as viridis variety fagi according to host plant cited]; catalog; references; USA: New Yersey; Pennsylvania; Massachusetts). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). LENG & MUTCHLER, 1927: 30 [#1703] ([Note: Misidentified as viridis variety fagi]; supplement to catalog; USA: Massachusetts). VASILEV, 1927: 147 149 [#5834] ([Note: Misidentified as viridis variety fagi]; pest; biology; parasitoids, predators; notes; Ukraine). FISHER, 1928: 13, 209 213 [#980] ([Note: Misidentified as viridis variety fagi according to host plants cited]; characters; bibliography; notes; US). OBENBERGER, 1936: 979 [#2191] (world catalog). GUSSAKOVKII, 1949: 208 [#7805] ([Note: Misidentified as chrysoderes]; notes; synonymy; references). TER MINASYAN, 1955: 450 [#7893] ([Note: Misidentified as cuprescens chrysoderes]; host plants; distribution; references). ALEXEEV, 1957: 139 141, 152 [#3226] ([Note: Misidentified as cuprescens chrysoderes]; biology; pest; eradication; notes; characters in key). ALEXEEV, 1958: 185, 186 188 [#3982] ([Note: Misidentified as chrysoderes]; checklist; biology; larval galleries; Moskovsk). ALEXEEV, 1959: 14 [#3229] (misidentified as cuprescens chrysoderes; characters in key; Latvia; Lithuania; Kalininsk; Moskovsk; Vladimirsk; Gor’kov). BATIASHVILI, 1959: 252 [#8634] ([Note: Misidentified as chrysoderes]; pest). ALEXEEV, 1960: 1508 [#45] ([Note: Misidentified as cuprescens chrysoderes]; larval characters in key). ALEXEEV, 1961: 6 [#3243] ([Note: Misidentified as cuprescens chrysoderes]; larval characters in key). ZAGAIKEVICH, 1962: 106 [#3249] ([Note: Misidentified as chrysoderes]; faunal records; bionomy; Ukraine; Krym). SAMEDOV, 1963: 92, 100 [#6091] ([Note: Misidentified as chrysoderes]; characters; biology; host plants; Azerbaijan). DAVIS & RAGHUVIR, 1964: 154 159 [#669] ([Note: Misidentified as rubicola communis]; biology; Utah). RIKHTER & ALEXEEV, 1965: 300 [#2522] (characters in key; European part of USSR). ANISIMOVA, 1973: 118 124 [#4032] ([Note: Misidentified as chrysoderes]; autoecology; distribution overview in SSSR; Amursk; Kharkovsk; Moskovsk; Tambovsk; Ryazansk; Leningradsk; Saratovsk; Khabarovsk; Primorsk). KOSTIN, 1973: 107 [#3693] (characters in key; Kazakhstan). GURYEVA, 1974: 98 99 [#5428] (pest of agriculture cultures; biology; references; Soviet Union). VOLKOVITSH, 1977: 26 [#2985] (faunal record; notes; USSR: Belgorod, Voronezh; Kursk; Kharkov). ALEXEEV, 1981: 76 [#61] (larval characters in key). KALASHIAN, 1986: 95 [#3865] (note on structure of ovipositor). ALEXEEV, 1989: 488 [#64] (key to species; distributional summary). ALEXEEV, 1990: 8 [#65] (notes). ZEROVA et al., 1991: 179

66 [#6339] (gall forming pest). VOLKOVITSH & ALEXEEV, 1994: 437 [#2974] (checklist; Turkmenistan). LAGUNOV & NOVOZHENOV, 1996: 47 [#7461] (checklist; Russia (Chelyabinsk)). NELSON et al., 1996: 184 [#3975] ([Note: Misidentified as cupreus Redtenbacher according to distribution, host plant, references and synonymy]; notes; USA). OVTCHINNIKOV, 1996: 132 [#8621] (checklist; Kyrgyzstan). ALEXEEV, 1998: 381 [#4322] (subgenus Agrilus; subgeneric classification). TLEPPAEVA, 1999: 184 [#6551] (notes; occurence up to 2500 m; Kazakhstan). BÍLÝ, 2002: 15 [#5877] (bionomy; host plants; distribution). BRECHTEL & KOSTENBADER, 2002: 241 246 [#5766] (biology; ecology; parasitoids; distribution; Germany). JENDEK, 2003: 3 6, 16 [#5878] (lectotype designation; synonymy; faunal records; distribution). MACRAE & NELSON, 2003: 61 [#5977] (faunal records; USA). SIKES, 2004: 38 [#7233] (checklist; USA). TLEPPAEVA & ISHKOV, 2004: 84 [#7211] (subgenus Agrilus; checklist; Kazakhstan). OHMOMO, 2004: 141 [#6850](faunal records; Japan: Hokkaido). BERNHARD et al., 2005: 601, 602 [#7261] (mitochondrial DNA; molecular systematics). DEDYUKHIN, NIKITSKY & SEMENOV, 2005: 303 [#8860] (checklist; Russia: Udmurtia). KOROTYAEV et al., 2005: 251 [#7284] (gall inducing pest). JENDEK, 2006: 388 389 [#7441] (subgenus Agrilus; Palaearctic catalog). WESTCOTT et al., 2006: 5 [#7590] (notes; USA: Oregon). BELLAMY, 2008: 2046 2049 [#8576] (subgenus Agrilus; world catalog). VÉTEK et al., 2008: 54 56 [#8700] (parasitoids). BOLDYREV & KOLESNIKOV, 2009: 2 pp [#8893] (biology; symptoms of attack; Russia). VOLKOVITSH, 2009: 134 [#8850] (subgenus Agrilus; faunal records; Russia: Primorskiy kray). SAGE, 2010: 20 [#8897] (faunal records; notes; Britain: Norfolk, South Essex). aurichalceus Redtenbacher, 1847 (Agrilus) REDTENBACHER, 1847: 286 [#3912] (description). KIESENWETTER, 1849: 40 [#8334] (catalog; Europe). KIESENWETTER, 1857: 126, 157 158 [#3015] (characters in key, redescription; notes; Austria). MARSEUL, 1857: 92 [#4371] (checklist; Europe). MOTSCHULSKY, 1859: 157, 275 [#1832] (catalog; bibliography on larval biology). JACQUELIN DU VAL, 1860: 143 [#8086] (catalog; Europe). MARSEUL, 1866: 436, 483 [#676] (characters in key; redescription). GEMMINGER & HAROLD, 1869: 1437 [#1057] (catalog). SAUNDERS, 1871: 121 [#2552] (catalog). HOCHHUTH, 1873: 159 [#3453] (faunal records; Gouvernements Kiew und Volhynien). RUPERTSBERGER, 1880: 157, 275 [#8550] (commented larval bibliography, checklist). MARSEUL, 1887: 252 [#4707] (catalog). REITTER, 1891: 200 [#5278] (catalog). KERREMANS, 1892: 247 [#1397] (catalog). RUPERTSBERGER, 1894: 161, 287 [#4465] (larval bibliography; catalog). ABEILLE DE PERRIN, 1897: 5, 16 [#19] ([Note: Misidentification of acutangulus according to SCHAEFER, 1949: 393 [#2565]]; characters in key; note). ESCHERICH, 1897: 26 [#8345] (faunal records; Turkey). HEYDEN, 1897: 195 [#5286] (notes). CSIKI, 1901: 89 [#505] (faunal records; Rossia: Kazan). SEMENOV, 1902: 16 18 [#4120] (characters; notes). KERREMANS, 1903: 275 [#1441] (catalog). REITTER, 1911: 194 [#2518] (characters in key). CAILLOL, 1913: 510 [#4053] ([Note: Misidentified as chrysoderes according to SCHAEFER, 1949: 402 [#2565]]). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). BAU, 1914: 257 [#8668] (characters in key). WEISS, 1914: 438 440 [#3920] ([Note: Misidentified as politus according to SCHAEFER, 1949: 402 [#2565]]; bionomy; USA (New Jersey)). OBENBERGER, 1926: 656 [#2057] (Palaearctic catalog). RAGUSA, 1926: 76 [#2496] (notes; Sicilia). OBENBERGER, 1927: 212, 213, 222, 243 [#2064] (note). OGLOBLIN & REICHARDT, 1932: 277 [#8654] (pest). OBENBERGER, 1935: 68 69 [#2178] (catalog; summary of distribution). OBENBERGER, 1936: 954 956 [#2191] (world catalog). THÉRY, 1942: 135, 166 167 [#2873] (subgenus Agrilus). RIKHTER, 1945: 166 [#3507] ([Note: Misidentification of salicis]). SCHAEFER, 1945: 68 [#3583] (note). THÉRY, 1945: 69 [#2884] (remarks on synonymy). RIKHTER, 1948: 394 [#3879] (characters in key; Kievsk; Nizhnee Povolzh’e). SCHAEFER,

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA 1949: 356, 402 405, 491 [#2565] (characters; references; faunal records; biology; notes; ditributional overview; notes). RIKHTER, 1950: 158 [#3881] (characters in key). HORION, 1955: 95 96, 269 270 [#1285] (distribution overview; faunal records; biology). STARK, 1955: 775 [#3533] (faunal record; USSR: Saratov). TER MINASYAN, 1955: 450 [#7893] (host plants; distribution; references). ALEXEEV, 1957: 152 [#3226] ([Note: Misidentification of salicis]; characters in key). ALEXEEV, 1958: 185 [#3982] ([Note: Misidentification of salicis]; checklist; Moskovsk). STEPANOV, 1958: 172 [#3664] ([Note: Misidentification of salicis]). ALEXEEV, 1959: 15 [#3229] ([Note: Misidentification of salicis]). ZAGAIKEVICH, 1960: 257 [#5615] (checklist; Ukraine). ALEXEEV, 1961: 9 [#3243] ([Note: Misidentification of salicis according to host plant cited]; larval characters in key). BALACHOWSKI et al., 1962: 294 296 [#96] (characters; biology; distribution; economic importance; bibliography). ZAGAIKEVICH, 1962: 107 [#3249] ([Note: Misidentification of salicis]). ZAGAIKEVICH, 1962: 60 [#5616] (synonym of acutangulus). SAMEDOV, 1963: 92, 101 [#6091] (characters; biology; host plants; Azerbaijan). POCHON, 1964: 54, 63 64 [#2470] (characters; key to species; faunistic; Switzerland). RIKHTER & ALEXEEV, 1965: 300 [#2522] ([Note: Misidentification of salicis]; characters in key). SIMON, 1966: 301 [#3266] (faunal records; Belgium). COBOS, 1972: 45, 56 [#591] (subgenus Agrilus; faunal records; Mongolia). KOSTIN, 1973: 107 [#3693] ([Note: Misidentification of salicis]). GURYEVA, 1974: 98 [#5428] ([Note: Misidentification of salicis according to host plant cited]; pest of agriculture cultures; references; Soviet Union). KUROSAWA, 1974: 2 [#4575] (notes). ALEXEEV, 1975: 158 [#52] ([Note: Misidentification of salicis according to to JENDEK, 2003 [#5878]]; notes; distribution overview). VOLKOVITSH, 1977: 26 [#2985] ([Note: Misidentification of salicis according to host plant cited]; faunal record; notes; USSR: Belgorod, Voronezh; Kursk; Kharkov). HELLRIGL, 1978: 266 267 [#3442] (critical overview of host plants). LOMPE, 1979: 239 [#3628] (characters in key; Central Europe). BÍLÝ, 1982: 65, 68, 69 70 [#295] (characters in key; distribution overview; biology; Sweden; Finland; Baltic region of the USSR: Estonia; Lithuania; Latvia). COBOS, 1986: 253, 255 256, 317 318 [#609] (subgenus Agrilus; characters in key; distribution; synonymy; bionomy; España). ALEXEEV, 1989: 488 [#64] (synonym of cuprescens). CAMPBELL et al., 1989: 31 32 [#8198] (pest of roses; biology; references; Canada). NIEHUIS & TEZCAN, 1993: 38 39, 48 [#1910] (subgenus Agrilus; faunal records from Turkey; distribution and host plants summary; Turkey (European and Asian part)). BÍLÝ, 1994: 112 [#8200] (larval characters in key). CURLETTI, 1994: 22, 177 178 [#658] (subgenus Agrilus; faunistic data; host plants; Italia; Sicilia; Sardegna). ZIEGLER, 1995: 60 [#6506] (faunal records; Germany). DOWNIE & ARNETT, 1996: 762, 768 769 [#6170] (characters; USA: ON, MA, CT, NY, NJ, PA, MI, IN, PQ, CO). KUROLI, 1999: 257 263 [#5916] (life cycle; parasitoids; Hungary). BÍLÝ, 2002: 15 [#5877] (synonym of cuprescens). BRECHTEL & KOSTENBADER, 2002: 241 [#5766] (synonym of cuprescens). HOU TAOQIAN & ZENG RUI, 2002: 357, 358 [#6626] (checklist; China: Hainan). BELLAMY & WESTCOTT, 2003: 8, 16 [#6153] (synonym of cuprescens; corrections to Nomina Inseta Nearctica). JENDEK, 2003: 6, 16 [#5878] (synonym of cuprescens; notes). LARSON, 2003: 176 178 [#6990] (pest of prairie roses; Canada: Saskatchewan province). OHMOMO, 2004: 141 [#6850] (synonym of cuprescens). JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2046 [#8576] (synonym of cuprescens; world catalog). NIKITSKY et al., 2008: 146 [#8610] (synonym of cuprescens). proximus Rey, 1891 (Agrilus; [preoccupied]) JAKOBSON, 1913: 797 REY, 1891: 19 [#3666] (description). [#4386] (variety of viridis; catalog; Russia and Europe). OBENBERGER, 1916: 272 [#1983] (aberration of viridis; characters

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA in key). OBENBERGER, 1926: 656 [#2057] (aberration of chrysoderes; Palaearctic catalog). OBENBERGER, 1936: 1035 [#2191] (synonym of rubicola; world catalog). SCHAEFER, 1937: 85 [#3570] (synonym of chrysoderes rubicola). BLACKWELDER, 1939: 44 [#340] (synonym of rubicola). THÉRY, 1942: 166 [#2873] (synonym of aurichalceus). SCHAEFER, 1949: 402, 491 [#2565] (synonym of aurichalceus). ALEXEEV, 1959: 14 [#3229] (synonym of chrysoderes). BALACHOWSKI et al., 1962: 294 [#96] (synonym of aurichalceus). COBOS, 1986: 317 [#609] (synonym of aurichalceus). CURLETTI, 1994: 177 [#658] (synonym of aurichalceus). BÍLÝ, 2002: 15 [#5877] (synonym of cuprescens). BRECHTEL & KOSTENBADER, 2002: 241 [#5766] (synonym of cuprescens). JENDEK, 2003: 8, 16 [#5878] (synonym of cuprescens; notes). JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2047 [#8576] (synonym of cuprescens; world catalog). egenus Abeille de Perrin, 1895 (Agrilus; [preoccupied]) ABEILLE DE PERRIN, 1895: CXXII [#15] (description). ABEILLE DE PERRIN, 1895: CCLXXIV [#3905] (synonym of lacrymans). KERREMANS, 1903: 286 [#1441] (synonym of lacrymans). JAKOBSON, 1913: 797 [#4386] (synonym of chrysoderes; catalog). OBENBERGER, 1936: 1038 [#2191] (synonym of rubicola variety lacrymans). HARIRRI, 1971: 195 [#8105] (synonym of chrysoderes variety lacrymans). JENDEK, 2003: 8, 16 [#5878] (synonym of cuprescens; lectotype designation; notes). JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2047 [#8576] (synonym of cuprescens; world catalog). lacrymans Abeille de Perrin, 1895 (Agrilus; replacement name for egenus Abeille de Perrin not Gory) ABEILLE DE PERRIN, 1895: CCLXXIV [#3905] (replacement name proposal). ABEILLE DE PERRIN, 1897: 5, 15 [#19] (variety of chrysoderes; characters in key; note). KERREMANS, 1903: 286 [#1441] (catalog). JAKOBSON, 1913: 797 [#4386] (variety of chrysoderes; catalog; Russia and Europe). OBENBERGER, 1916: 271, 272 [#1983] (variety of viridis; characters in key). OBENBERGER, 1926: 656 [#2057] (variety of chrysoderes; Palaearctic catalog). OBENBERGER, 1927: 200, 211, 220, 241 [#2064] (subspecies of communis). OBENBERGER, 1936: 1038 [#2191] (variety of rubicola; world catalog). THÉRY, 1942: 166, 167 [#2873] (subgenus Agrilus; aberration of aurichalceus). SCHAEFER, 1949: 403 [#2565] (subspecies of aurichalceus; notes). HARIRRI, 1971: 195 [#8105] (variety of chrysoderes; checklist; Syria). COBOS, 1986: 256, 318 [#609] (subgenus Agrilus; subspecies of aurichalceus). NIEHUIS & TEZCAN, 1993: 39 [#1910] (subgenus Agrilus; subspecies of aurichalceus; probably not in Turkey). JENDEK, 2003: 8, 16 [#5878] (synonym of cuprescens). JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2047 [#8576] (synonym of cuprescens; world catalog). epistomalis Abeille de Perrin, 1897 (Agrilus) ABEILLE DE PERRIN, 1897: 4, 13 [#19] (description). HEYDEN, 1897: 194 [#5286] (characters). KERREMANS, 1903: 284 [#1441] (catalog). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 656 [#2057] (Palaearctic catalog). MÉQUIGNON, 1930: 207 209 [#4492] (note). OBENBERGER, 1936: 1037 [#2191] (variety of rubicola; world catalog). SCHMIDT, 1937: 262 [#5323] (aberration of communis; notes; faunal records; Germany). SCHAEFER, 1938: 85 [#3281] (synonym of rubicola). THÉRY, 1942: 166, 167 [#2873] (subgenus Agrilus; aberration of aurichalceus). BALACHOWSKI et al., 1962: 294 [#96] (synonym of aurichalceus). LOMPE, 1979: 239 [#3628] (synonym of aurichalceus). COBOS, 1986: 317 [#609] (synonym of aurichalceus). CURLETTI, 1994: 177 [#658] (synonym of aurichalceus). BÍLÝ, 2002: 15 [#5877] (synonym of cuprescens).

67 BRECHTEL & KOSTENBADER, 2002: 241 [#5766] (synonym of cuprescens). JENDEK, 2003: 9, 16 [#5878] (synonym of cuprescens; notes). JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2047 [#8576] (synonym of cuprescens; world catalog). foveolatus Abeille de Perrin, 1897 (Agrilus; variety of chrysoderes) ABEILLE DE PERRIN, 1897: 4, 15 [#19] (description). JAKOBSON, 1913: 797 [#4386] (variety of chrysoderes; catalog; Russia and Europe). OBENBERGER, 1916: 272 [#1983] (variety of viridis; characters in key). OBENBERGER, 1926: 656 [#2057] (variety of chrysoderes; Palaearctic catalog). OBENBERGER, 1927: 211, 220, 240 241 [#2064] (variety of communis). OBENBERGER, 1935: 66 [#2178] (aberration of communis; catalog). OBENBERGER, 1936: 1038 [#2191] (variety of rubicola; world catalog). THÉRY, 1942: 166 [#2873] (subgenus Agrilus; aberration of aurichalceus). SCHAEFER, 1949: 403 [#2565] (subspecies of aurichalceus; notes). TROPIN, 1965: 457 458 [#8652] ([Note: Misidentification of roscidus according to host plant cited]; biology; Rostov). COBOS, 1986: 256, 318 [#609] (subgenus Agrilus; subspecies of aurichalceus). JENDEK, 2003: 9 10, 16 [#5878] (synonym of cuprescens; notes). JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2047 [#8576] (synonym of cuprescens; world catalog). obtusus Abeille de Perrin, 1897 (Agrilus; variety of chrysoderes; [preoccupied]) ABEILLE DE PERRIN, 1897: 5, 15 [#19] (description). HEYDEN, 1897: 195 [#5286] (variety of chrysoderes; characters). REITTER, 1911: 194 [#2518] (characters in key; Germany). CAILLOL, 1913: 511 [#4053] (variety of chrysoderes; faunal records; France). JAKOBSON, 1913: 797 [#4386] (variety of chrysoderes; catalog; Russia and Europe). OBENBERGER, 1916: 272 [#1983] (aberration of viridis; characters in key). BEDEL, 1921: 213, 214 [#170] (synonym of chrysoderes). OBENBERGER, 1926: 656 [#2057] (aberration of chrysoderes; Palaearctic catalog). OBENBERGER, 1927: 199, 211, 237 [#2064] (synonym of communis aberration rubicola). JAKOBSON & OGLOBLIN, 1931: 162 [#8314] (variety of chrysoderes; characters in key). OGLOBLIN & REICHARDT, 1932: 277 [#8654] (synonym of chrysoderes mokrzeckii). OBENBERGER, 1936: 1035 [#2191] (synonym of rubicola; world catalog). BLACKWELDER, 1939: 44 [#340] (synonym of rubicola). THÉRY, 1942: 166, 167 [#2873] (subgenus Agrilus; aberration of aurichalceus). SCHAEFER, 1949: 402, 491 [#2565] (synonym of aurichalceus aberration obtusifera). ALEXEEV, 1959: 14 [#3229] (synonym of chrysoderes). ZAGAIKEVICH, 1962: 106 [#3249] (synonym of chrysoderes). GURYEVA, 1974: 98 [#5428] (synonym of cuprescens). HELLRIGL, 1978: 266 [#3442] (synonym of aurichalceus). LOMPE, 1979: 239 [#3628] (synonym of aurichalceus). COBOS, 1986: 256 [#609] (subgenus Agrilus; form of aurichalceus). CURLETTI, 1994: 177 [#658] (synonym of aurichalceus). BÍLÝ, 2002: 15 [#5877] (synonym of cuprescens). BRECHTEL & KOSTENBADER, 2002: 241, 313 [#5766] (cited as a synonym of cuprescens on page 241 and with authorship of Obenberger as a synonym of viridis on page 313). JENDEK, 2003: 10, 16 [#5878] (synonym of cuprescens; lectotype designation; notes). JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2047 [#8576] (synonym of cuprescens; world catalog). rubicola Abeille de Perrin, 1897 (Agrilus; variety of chrysoderes) ABEILLE DE PERRIN, 1897: 4, 15 [#19] (description). HEYDEN, 1897: 195 [#5286] (variety of chrysoderes; characters). HEYDEN, 1898: 21 [#5553] (variety of chrysoderes Abeille). CAILLOL, 1913: 510 511 [#4053] (variety of chrysoderes; biology; faunal records; France). JAKOBSON, 1913: 797 [#4386] (variety of chrysoderes;

68 catalog; Russia and Europe). REH, 1913: 488 [#8153] (variety of chrysoderes; biology; references). OBENBERGER, 1916: 272 [#1983] (aberration of viridis; characters in key). BEDEL, 1921: 213, 214 [#170] (variety of chrysoderes; characters in key). KROGERUS, 1922: 112 [#3916] (aberration of viridis chrysoderes). OBENBERGER, 1926: 656 [#2057] (aberration of chrysoderes; cited as rubricola; Palaearctic catalog). OBENBERGER, 1927: 199, 211, 219, 237 239 [#2064] (aberration of communis). FISHER, 1928: 209 [#980] (aberration of communis; subnote). MCDANIEL, 1932: 89 91 [#8682] (subspecies of communis; biology; USA). OGLOBLIN & REICHARDT, 1932: 277 [#8654] (variety of chrysoders; pest). OBENBERGER, 1935: 66 [#2178] (aberration of communis; catalog). MCDANIEL, 1936: 44 46 [#8726] (subspecies of communis; biology; USA). OBENBERGER, 1936: 1032 1039 [#2191] (world catalog). BLACKWELDER, 1939: 44 [#340] (supplement to catalog; North America). GARLICK, 1940: 21 23 [#8683] (subspecies of communis; biology; Canada (Ontario)). MALKIN, 1941: 132 [#1768] (subspecies of communis; faunal record; USA: New York City: Van Cortlandt Park). MUNDINGER, 1941: 532 537 [#5414] (biology; injury; immature stages; phenology; habits; USA: New York). THÉRY, 1942: 166 [#2873] (synonym of aurichalceus). RIKHTER, 1945: 165 [#3507] (aberration of chrysoderes). RIKHTER, 1948: 393 [#3879] (aberration of chrysoderes). SCHAEFER, 1949: 402, 403, 404, 491 [#2565] (aberration of aurichalceus; characters). MÜHLMANN, 1954: 80, 85 [#8152] (variety of chrysoderes on page 80 and variety of communis on page 85; pest in Europe and North America; biology; parasitoid: Tetrastichus rugglesi). HEERING, 1956: 257 [#3716] (variety of communis; notes on American records). ALEXEEV, 1957: 139 [#3226] (aberration of chrysoderes; characters in key). ALEXEEV, 1959: 14 [#3229] (synonym of chrysoderes). BALACHOWSKI et al., 1962: 294 [#96] (synonym of aurichalceus). RIKHTER & ALEXEEV, 1965: 300 [#2522] (synonym of cuprescens). GURYEVA, 1974: 98 [#5428] (synonym of cuprescens). WELLSO et al., 1976: 16 [#3079] (characters; USA: Michigan). HELLRIGL, 1978: 266 [#3442] (synonym of aurichalceus). LOMPE, 1979: 239 [#3628] (synonym of aurichalceus). COBOS, 1986: 256, 317 [#609] (synonym of aurichalceus). ALEXEEV, 1990: 8 [#65] (notes). CURLETTI, 1994: 177 [#658] (synonym of aurichalceus). BÍLÝ, 2002: 15 [#5877] (synonym of cuprescens). BRECHTEL & KOSTENBADER, 2002: 241 [#5766] (synonym of cuprescens). BELLAMY & WESTCOTT, 2003: 9, 16 [#6153] (subspecies of cuprescens; corrections to Nomina Inseta Nearctica). JENDEK, 2003: 10, 16 [#5878] (synonym of cuprescens; lectotype designation; notes). SIKES, 2004: 38 [#7233] (synonym of cuprescens). JENDEK, 2006: 389 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2047 [#8576] (synonym of cuprescens; world catalog). calcicola Obenberger, 1916 (Agrilus; cited as variety of viridis on page 271 and aberration of viridis on page 272) OBENBERGER, 1916: 271, 272 [#1983] (description). OBENBERGER, 1917: 213 [#1997] (subspecies of viridis). KROGERUS, 1922: 110, 111, 112 [#3916] (variety of viridis chrysoderes). RAGUSA, 1922: 8 [#2495] (cited as cascicola; variety of viridis; Sicilia). OBENBERGER, 1926: 656 [#2057] (aberration of chrysoderes; Palaearctic catalog). OBENBERGER, 1927: 200, 202, 210, 211, 218, 234 235 [#2064] (note). MÉQUIGNON, 1930: 209 [#4492] (note). ROUBAL, 1930: 115 [#5172] (notes). OBENBERGER, 1936: 968 969 [#2191] (world catalog). GOBBI, 1983: 40 [#1098] (note). CURLETTI, 1987: 151 152 [#653] (characters; notes; not in Italy). CURLETTI, 1994: 179 [#658] (not in Italy). JENDEK, 2003: 11, 16 [#5878] (synonym of cuprescens; lectotype designation; notes). JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2048 [#8576] (synonym of cuprescens; world catalog).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA communis Obenberger, 1924 (Agrilus; variety of viridis) OBENBERGER, 1924: 41 [#2040] (description). OBENBERGER, 1926: 656 [#2057] (aberration of chrysoderes; Palaearctic catalog). OBENBERGER, 1927: 211, 213, 218, 235 237, 248 [#2064] (note). MÉQUIGNON, 1930: 208 [#4492] (synonym of chrysoderes). OBENBERGER, 1935: 66 68 [#2178] (catalog; summary of distribution; Bulgaria). OBENBERGER, 1936: 1036 [#2191] (variety of rubicola; world catalog). SCHMIDT, 1937: 262 [#5323] (aberration of rubicola; notes). SCHAEFER, 1938: 85 [#3281] (synonym of rubicola). BLACKWELDER, 1939: 44 [#340] (variety of rubicola; supplement to catalog; North America). THÉRY, 1942: 166 [#2873] (synonym of aurichalceus). RIKHTER, 1945: 165 [#3507] (aberration of chrysoderes). RIKHTER, 1948: 393 [#3879] (aberration of chrysoderes). ALEXEEV, 1957: 139 [#3226] (aberration of chrysoderes; characters in key). ALEXEEV, 1959: 14 [#3229] (synonym of chrysoderes). BALACHOWSKI et al., 1962: 294 [#96] (synonym of aurichalceus). DAVIS & RAGHUVIR, 1964: 154 159 [#669] (subspecies of rubicola; [Note: Misidentification of cuprescens]). RIKHTER & ALEXEEV, 1965: 300 [#2522] (synonym of cuprescens). GURYEVA, 1974: 98 [#5428] (synonym of cuprescens). HELLRIGL, 1978: 266 [#3442] (synonym of aurichalceus). LOMPE, 1979: 239 [#3628] (synonym of aurichalceus). COBOS, 1986: 318 [#609] (synonym of aurichalceus). CURLETTI, 1994: 177 [#658] (synonym of aurichalceus). BÍLÝ, 2002: 15 [#5877] (synonym of cuprescens). BRECHTEL & KOSTENBADER, 2002: 241 [#5766] (synonym of cuprescens). JENDEK, 2003: 12, 16 [#5878] (synonym of cuprescens; lectotype designation; notes). JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2048 [#8576] (synonym of cuprescens; world catalog). krasai Obenberger, 1924 (Agrilus; variety of viridis) OBENBERGER, OBENBERGER, 1924: 40 [#2040] (description). 1926: 656 [#2057] (variety of chrysoderes; Palaearctic catalog). OBENBERGER, 1927: 211, 219, 240 [#2064] (aberration of communis). OBENBERGER, 1935: 66 [#2178] (aberration of communis; catalog). OBENBERGER, 1936: 1037 [#2191] (synonym of rubicola variety epistomalis; world catalog). SCHMIDT, 1937: 262 [#5323] (synonym of epistomalis). THÉRY, 1942: 166 [#2873] (subgenus Agrilus; aberration of aurichalceus). SCHAEFER, 1949: 402, 403, 404, 491 [#2565] (aberration of aurichalceus; characters). SIMON, 1966: 301 [#3266] (aberration of aurichalceus; faunal records; Belgium). COBOS, 1986: 256, 318 [#609] (synonym of aurichalceus). CURLETTI, 1994: 177 [#658] (synonym of aurichalceus). BÍLÝ, 2002: 15 [#5877] (synonym of cuprescens). BRECHTEL & KOSTENBADER, 2002: 241 [#5766] (synonym of cuprescens). JENDEK, 2003: 13, 16 [#5878] (synonym of cuprescens; lectotype designation; notes). JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2048 [#8576] (synonym of cuprescens; world catalog). mokrzeckii Obenberger, 1927 (Agrilus; subspecies of communis) OBENBERGER, 1927: 204 209, 210, 211, 220, 241 [#2064] (description). OGLOBLIN & REICHARDT, 1932: 277 [#8654] (subspecies of chrysoderes; pest). OBENBERGER, 1935: 68 [#2178] (catalog; summary of distribution; Bulgaria). OBENBERGER, 1936: 1015 [#2191] (world catalog). THÉRY, 1942: 166 [#2873] (subgenus Agrilus; subspecies of aurichalceus). SCHAEFER, 1949: 405 [#2565] (notes). MÜHLMANN, 1954: 80 [#8152] (cited as mokrzecki; subspecies of communis; pest; Bulgaria). ALEXEEV, 1959: 14 [#3229] (synonym of chrysoderes). BALACHOWSKI et al., 1962: 296 [#96] (notes; Bulgaria; Caucasus; Krym). RIKHTER & ALEXEEV, 1965: 300 [#2522] (synonym of cuprescens). GURYEVA, 1974: 98 [#5428] (synonym of cuprescens). HELLRIGL, 1978: 266 [#3442] (synonym of aurichalceus). JENDEK, 2003: 13 14, 16 [#5878] (synonym of cuprescens; lectotype designation; notes).

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2048 [#8576] (synonym of cuprescens; world catalog). altaicola Obenberger, 1935 (Agrilus) OBENBERGER, 1935: 168 169 [#2158] (description). OBENBERGER, 1936: 941 [#2191] (world catalog). COBOS, 1972: 55 [#591] (subgenus Agrilus; checklist; Mongolia). JENDEK, 2003: 14, 16 [#5878] (synonym of cuprescens; lectotype designation; notes). JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2048 [#8576] (synonym of cuprescens; world catalog). kuznecovi Obenberger, 1935 (Agrilus; subspecies of communis; [preoccupied]) OBENBERGER, OBENBERGER, 1935: 169 [#2158] (description). 1936: 1039 [#2191] (synonym of rubicola kuznecovinus; world catalog). OBENBERGER, 1937: 58 [#2226] (synonym of communis kuznecovinus). COBOS, 1986: 318 [#609] (cited as kurnecovi; synonym of aurichalceus kuznecovinus). JENDEK, 2003: 14, 16 [#5878] (synonym of cuprescens; lectotype designation; notes). JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2048 2049 [#8576] (synonym of cuprescens; world catalog). kuznecovinus Obenberger, 1936 (Agrilus; replacement name for kuznecovi Obenberger, 1935 not Obenberger, 1933) OBENBERGER, 1936: 1038 [#2191] (replacement name proposal). OBENBERGER, 1937: 58 [#2226] (subspecies of communis). THÉRY, 1942: 166 [#2873] (subgenus Agrilus; subspecies of aurichalceus). SCHAEFER, 1949: 403 [#2565] (subspecies of aurichalceus; notes). ALEXEEV, 1975: 157 158 [#52] (subspecies of cuprescens; cited as kuznecovianus; notes; faunal records; distributional overview; Mongolia). COBOS, 1986: 256, 318 [#609] (subgenus Agrilus; subspecies of aurichalceus). ALEXEEV, 1989: 488 [#64] (subspecies of cuprescens; cited as kuznecovianus; key to species; Russian Far East). ALEXEEV & VOLKOVITSH, 1989: 321, 351, 357 [#3241] (subspecies of cuprescens; characters in key; faunal records; Mongolia). JENDEK, 2003: 14, 16 [#5878] (synonym of cuprescens). JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2049 [#8576] (synonym of cuprescens; world catalog). obtusifera Schaefer, 1949 (Agrilus; replacement name for obtusus Abeille de Perrin not Horn) SCHAEFER, 1949: 402, 403, 404, 491 [#2565] (replacement name proposal). COBOS, 1986: 256 [#609] (subgenus Agrilus; form of aurichalceus). JENDEK, 2003: 14, 16 [#5878] (synonym of cuprescens). ARNÁIZ RUIZ & BERCEDO PÁRAMO, 2006: 260 [#7414] (aberration of cuprescens; correct year of publication is 1950). JENDEK, 2006: 388 [#7441] (synonym of cuprescens cuprescens; Palaearctic catalog). BELLAMY, 2008: 2049 [#8576] (synonym of cuprescens; world catalog).

Type material: Buprestis cuprescens Ménétriés, 1832 Type locality: Périmbal, sur les montagnes de Talyche. Type specimens: Primary: Lectotype (Fig. 44V), ♂, designated by JENDEK (2003: 3-6, 16) [#5878], (ZIN): “Taliisch. [p] [red label] \ cuprescens Menet Talysch [h] \ Agrilus cuprescens Mén Type! [h]”. Described from unknown number of syntypes. Agrilus aurichalceus Redtenbacher, 1847 Type locality: not given [Austria is cited in the title of the publication]. Type specimens: Not located. Described from unknown number of syntypes.

69 Agrilus proximus Rey, 1891 Type locality: Lyon. Type specimens: Not located. Described from 1 specimen. Agrilus egenus Abeille de Perrin, 1895 Type locality: Haute-Syrie: Akbès. Type specimens: Primary: Lectotype (Fig. 45R), ♂, designated by JENDEK (2003: 8, 16) [#5878], (MNHN): “Akbès [h] \ Type [p] [red ink] \ A. chrysoderes Ab. var. lacrymans Ab. [h] \ Muséum Paris Coll. Abeille De Perrin 1919 [p]”. Secondary: 1 PLT (ISNB); 2 PLT (MNHN). Described from unknown number of syntypes. Agrilus lacrymans Abeille de Perrin, 1895 Type locality and specimens: See: Agrilus egenus Abeille de Perrin, 1895. Agrilus epistomalis Abeille de Perrin, 1897 Type locality: Moravie. Type specimens: Holotype by monotypy (Fig. 45W), ♂, (MNHN): “Moravia Besciden Reither [p] \ Type [p] [red ink] \ epistomalis Ab. [h] \ Muséum Paris Coll. Abeille De Perrin 1919 [p]”. Described from 1 ♂ specimen. Agrilus chrysoderes variety foveolatus Abeille de Perrin, 1897 Type locality: Saratow. Type specimens: Holotype by monotypy (Fig. 46L), ♂, (MNHN): “Sara-tow [p] \ Typ. Ballion [h] \ Agrilus foveolatus Ball. [h] \ Type [p] [red ink] \ Muséum Paris Coll. Abeille De Perrin 1919 [p] \ A. chrysoderes var. foveolatus Ballion [h]”. Described from 1 specimen. Agrilus chrysoderes variety obtusus Abeille de Perrin, 1897 Type locality: Caucase, Hongrie, Grèce, France: Lyon, N.D. -de -Lure, Digne. Type specimens: Primary: Lectotype (Fig. 50E), sex not examined, designated by JENDEK (2003: 10, 16) [#5878], (MNHN): “N. D. de Lure \ Type [p] [red ink] \ Muséum Paris Coll. Abeille De Perrin 1919 [p] \ A. chrysoderes var. obtusus Ab. [h]”. Secondary: 2 PLT (MNHN). Described from unknown number of syntypes. Agrilus chrysoderes variety rubicola Abeille de Perrin, 1897 Type locality: Turkestan; Syrie: Akbès; France: HautesAlpes, Ain, Landes. Type specimens: Primary: Lectotype (Fig. 52F), ♂, designated by JENDEK (2003: 10, 16) [#5878], (MNHN): “H. Alpes \ Type [p] [red ink] \ Muséum Paris Coll. Abeille De Perrin 1919 [p] \ A. chrysoderes var. rubicola Ab. [h]”. Described from unknown number of syntypes. Agrilus viridis variety calcicola Obenberger, 1916 Type locality: Gr[eece]., Bos[nia]., Herz[egovina]. Type specimens: Primary: Lectotype (Fig. 43Z), ♂, designated by JENDEK (2003: 11, 16) [#5878], (NMPC): “Hercegovina Mostar Dr. Grabowski [p] \ Typus [p] [red label] \ calcicola m. [male sign] [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Agrilus viridis variety communis Obenberger, 1924 Type locality: Bohemia (Závist); Saxonia (Dresden, Prov. Sachsen, Vogtland); Germ[ania] (Mittelfranken) Bavaria; Gallia (Cazaux); Croatia (Fužin); Graecia; Bulgaria (Sofia); Russia m.: Sarepta.

70 Type specimens: Primary: Lectotype (Fig. 44Q), sex not examined, designated by JENDEK (2003: 12, 16) [#5878], (NMPC): “Závist, Bohemia centr. coll. Obenberger [h] \ Typus [p] [red label] \ var. communis m. Type [h] det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24837 [h] [orange label]”. Secondary: ST (NHMB); 5 PLT (NMPC). Described from unknown number of syntypes. Agrilus viridis variety krasai Obenberger, 1924 Type locality: Slovensko: Bratislava. Bohemia: Závist. Type specimens: Primary: Lectotype (Fig. 48A), sex not examined, designated by JENDEK (2003: 13, 16) [#5878], (NMPC): “Bratislava [h] \ Typus [p] [red label] \ viridis var. krasai m. Typus [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24828 [h] [orange label]”. Described from unknown number of syntypes. Agrilus communis mokrzeckii Obenberger, 1927 Type locality: Bulgaria: Kasanlik. Type specimens: Primary: Lectotype (Fig. 49M), ♂, designated by JENDEK (2003: 13-14, 16) [#5878], (NMPC): “Bulgaria Kasanlik [h] \ Typus [p] [red label] \ ssp. Mokrzeckii m. Typ [male sign] [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24842 [h] [orange label]”. Secondary: 1 PLT (NMPC); 1 PLT (NHMB). Described from unknown number of syntypes. Agrilus altaicola Obenberger, 1935 Type locality: Montes Altai. Type specimens: Primary: Lectotype (Fig. 42M), ♂, designated by JENDEK (2003: 14, 16) [#5878], (NMPC): “Altaj mont. [h] \ Typus [p] [red label] \ A. altaicola m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24315 [h] [orange label]”. Described from unknown number of syntypes. Agrilus communis kuznecovi Obenberger, 1935 Type locality: Ussuri mer.: Trojckoje-Chanka; Kamen Rybolov; Irkutskaja Gub.: Statio Padun (in fluvio Tunguska or.; Primorje: Kangou-za, V. Suchodol etc. Type specimens: Primary: Lectotype (Fig. 48I), sex not examined, designated by JENDEK (2003: 14, 16) [#5878], (NMPC): “B. Sukhodol’ Kangou-za Prim. obl. [p] [in Russian] 27. VII. [h] 911 [p] \ Rydzevskii i Vr. Kuznecov’ [p] [in Russian] \ Typus [p] [red label] \ Agrilus communis ssp. Kuzněcovi m. Type [h] Det. Dr Obenberger [p] \ ssp. Kuzněcovinus n. n.! [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae [p] 24830 [h] Inv. [p] [orange label]”. Secondary: 2 PLT (NMPC); 3 PLT (ZIN). Described from unknown number of syntypes. Agrilus kuznecovinus Obenberger, 1936 Type locality and specimens: See: Agrilus communis kuznecovi Obenberger, 1935. Agrilus obtusifera Schaefer, 1949 Type locality and specimens: See: Agrilus chrysoderes variety obtusus Abeille de Perrin, 1897. Diagnosis: 4.8-7.2 mm; body smaller and more robust; eyes moderate; pronotum widest at middle with sides obviously arcuate; apical third of elytra less elongate. Species group: Viridis. Relevant species: A. viridis (Linné, 1758). Specimens examined: ARMENIA: 1 (MDCM): “Arm. SSR,

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Goral [e,s?], 2.VII.1982, M. Danilevski leg.”. AUSTRIA: 3 (ZMAN): “Austria L. Miller” (JENDEK, 2003 [#5878]). AZERBAIJAN: 1 ♂ (EJCB): “Nakhichevan, Alagëz, 3.VI.1955” (JENDEK, 2003 [#5878]); 2 (MDCM): “Altyagach, 18.7.1981, Korostelev” (JENDEK, 2003 [#5878]); 1 ♂ (EJCB): “USSR, Azerb. SSR, 90 km N of Baku, Zarat env. 17.6.1984, E. Jendek leg.” (JENDEK, 2003 [#5878]); 1 (MKCY): “Alty Agach 6 7.5.1987 [in Russian]”; 1 (ZIN): “Azerbaijan: Gr. Caucasus, Pirguli Nat. res., nr. Vill. Sis, forest’s edges 1.vii.2004, K. Nadein leg.”. BELGIUM: Luxembourg: 2 ♂ (EJCB): “Luxembourg, Hallerbach, 29.VI.1980” (JENDEK, 2003 [#5878]). BOSNIA & HERZEGOVINA: 1 (ZMAN): “Radobolje, U. Mostar, Reitter leg.” (JENDEK, 2003 [#5878]); 2 (ZMAN): “Herzegowina” (JENDEK, 2003 [#5878]). BULGARIA: 1 (EJCB): “Bulgaria Arkutino 25.VI.1982, Jendek leg.” (JENDEK, 2003 [#5878]); 1 (EJCB): “Bulgaria Slnčev Briag 1 13.7.1982 Dr. M. Rešl leg.” (JENDEK, 2003 [#5878]); 1 (EJCB): “Bulgaria Kavacite 23 28.VI.1982 Jendek leg.” (JENDEK, 2003 [#5878]); 1 (EJCB): “Bulgaria Asenovgrad, 8.VIII.1985” (JENDEK, 2003 [#5878]); 4 (EJCB): “Bulgaria, Lozenec, 2.7.1987 P. Svoboda” (JENDEK, 2003 [#5878]); 4 (EJCB): “Bulgaria, Sandanski, 17.6.1987 J. Kodada leg.” (JENDEK, 2003 [#5878]); 9 (EJCB): “Bulgaria, Rozhen, 13.6.1987 J. Kodada leg.” (JENDEK, 2003 [#5878]); 2 (EJCB): “Bulgaria, Stara Zagora, 21.6.1987 P. Svoboda” (JENDEK, 2003 [#5878]); 1 (EJCB): “Bulgaria Sofia env. Kutina, 16.VII.1989, O. Šauša leg.” (JENDEK, 2003 [#5878]). CANADA: New Brunswick: 2 ♂ (EJCB): “CAN, New Brunswick, York Co., Charters Settlement, 45.8340ºN, 66.7275ºW, 30 June 2008, R. P. Webster coll. \ Regenerating mixed forest on foliage of Rubus cesius”. CROATIA: 1 ♂ (EJCB): “Dalmatia Umgeb. Labin” (JENDEK, 2003 [#5878]); 1 (EJCB): “Dalm., Stobre 14.6.1970 M. Dvořák lgt.” (JENDEK, 2003 [#5878]); 1 (EJCB): “Juhoslavia, 20.VI.1982 Drovenik Rambousek leg.” (JENDEK, 2003 [#5878]). CZECH REPUBLIC: 3 (EJCB): “ČSSR, Moravia mer. Břeclav 20.7.1979 Jendek leg.” (JENDEK, 2003 [#5878]); 1 ♂ (EJCB): “Bohemia, 17.VI.1993 Praha Baba Z. Kletečka leg.” (JENDEK, 2003 [#5878]). GEORGIA: 1 (EJCB): “Gruzia occ. Gagra 9.6.1966 Voříšek leg.” (JENDEK, 2003 [#5878]). GERMANY: 1 (EJCB): “Thuringia sept. Kyffhausergeb. Kosakenberg 11.8.1963 Dohrn leg.” (JENDEK, 2003 [#5878]). Baden Wurttemberg: 1 (ZMAN): “Karlsruhe” (JENDEK, 2003 [#5878]). Bayern: 1 (ZMAN): “München 18.7.1918” (JENDEK, 2003 [#5878]); 1 (ZMAN): “Beieren Berchtesgaden VI.1969” (JENDEK, 2003 [#5878]). GREECE: 1 ♂ (EJCB): “Graecia, Peloponnes p. Megalopoli env., 10.5.2000 I. Rychlík leg.” (JENDEK, 2003 [#5878]); 1 (EJCB): “GREECE, CRETE Iraklion, KATO SIMI env. 750 1200 m, 35°02’N, 25°29’E, 8 12.V.2003 Jendek, E. leg. 8 12.V.2003” (JENDEK, 2003 [#5878]). IRAN: Gilan: 1 ♂ (DGCC): “N. Iran, Gilan, 5 km E di Tũtkābon, 22.6.2003, m 400, Leg. D. Gianasso”. ITALY: Abruzzi: 1 ♂ (EJCB): “Italia, Cerchio, J. M. Duchon” (JENDEK, 2003 [#5878]). Toscana: 1 (ZMAN): “Toscana, Vallombrosa 6.1924 Righetti” (JENDEK, 2003 [#5878]). JAPAN: Hokkaido: 1 ♂ (SOCI): “2003.7.27, Hokuto, Koshimizu, Shari, Hokkaido, S. Ohmomo leg.”. KAZAKHSTAN: 1 (ZIN): “Kazakh. SSR Aktyubinsk. ob. B. Barsuki 22 V 1950 A. V. Lugovoi”. 1 ♀ (EJCB): “Kazakhstan 22.VII.1965 Saur Mt., Karaungur riv.” (JENDEK, 2003 [#5878]); 2 ♀ (EJCB): “Kazakh. 7.7.1978 Karkaralinsk Vasilenko leg.” (JENDEK, 2003 [#5878]); 1 ♂ (EJCB): “Kazakh. Tarbagatai Mt., r. Kargalinka, 10 km ENE Kirovki, 12 13.VII.1983 Volkovitsh leg.” (JENDEK, 2003 [#5878]); 1 ♂ (EJCB): “Kazakhstan, 21.VII.1985 5 km Pokatilovka, distr. Sopodka, Lopatin leg.” (JENDEK, 2003 [#5878]); 3 (MKCY): “Kazakhstan, 100 km SEE Ust’ Kamenogorsk 5 km NE Panteleymonovka 16 20.06.1993, Napolov leg.”; 1 ♂, 3 ♀ (EJCB): “E KAZAKHSTAN, 800 m, Tarbagatay Mts., Burson, 15 VII 1993 Napolov leg.”; 1 (MKCY): “Kazakh., Tarbagatay Mts., Urdzar Burgon 15.7.[19]93”; 1 ♂ (EJCB): “East Kazakhstan, 6 10.7.1993, 1000 1200 m, Syrianovsk env. R. Andreeva leg.” (JENDEK, 2003 [#5878]); 1

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA (MDCM): “Kazakhstan Astana 10.6.2001, Turgenevka, M. Danilevski leg.” (JENDEK, 2003 [#5878]). KOREA NORTH: 1 (VSCS): “Korea, Jangkangdo, Sinsadong, 1400m, 17.7.1974, Josifov leg.”; 1 (VSCS): “Korea, Jangkangdo, Samdzijôn, 13 19.7.1974, Josifov leg.”; 1 (EJCB): “Korea, Jangkangdo, Samdzijôn, 13 19.VII.1974, Josifov leg.”. LUXEMBOURG: 1 (EJCB): “Luxembourg, Ettelbruck, 26.VI.1977” (JENDEK, 2003 [#5878]). MOLDOVA: 1 ♂ (EJCB): “Mold. SSR, Bendery, 6.VII.1984, Karasev”. MONGOLIA: 1 (ZIN): “Mongoliya, Vostochnyi aimak, r. Numregin Gol, 32 km YuV g. Salkhit, M. Kozlov, 16.vi.976 [in Russian]”; 2 (EJCB): “Mongolia, 5 9 July 2003, Cagan nuga env. 1500 m, Segneger riv. vall. / 35 km NE of Mandal ( Batsumber), cca 70 km N of Ulaanbaatar, Zd. Jindra lgt.”. RUSSIA: Altay Kray: 2 (ZIN): “Gorno Altaisk, 20.vi.[19]89, S. Sipuk [in Russian]”. Amurskaya oblast’: 1 (ZIN): “Amurskaya obl. Skorodinsk. R n, Ulyagir, 24.7.1959 Kabakov [in Russian]”; 2 ♀ (EJCB): “Russia, Amurskaya obl. Shimanovsk reg. Saskal’, 15. VI.1975, Bereya riv., Kuznetsov leg.” (JENDEK, 2003 [#5878]); 1 ♀ (EJCB): “Russia, Amurskaya obl. Shimanovsk reg. Novogeorgievka, 27.VI.1976, Kuznetsov leg.” (JENDEK, 2003 [#5878]). Dagestan Respublika: 7 (MKCY): “Dagestan, okr.s. Glokh, 1800 200m, 24 30.v.1988, Karasev [in Russian]”. Irkutskaya oblast’: 1 (NMPC): “Irkutsk’, V. Yakovlev’, 92 93, [in Russian]”; 20 (ZIN): “Mal’ta Irkutsk. g. D. Smirnov 9.vi.1907”; 1 (ZIN): “Kuz’myka bl. Irkutska Vinogradov 2/VII 1924”; 1 (ZIN): “Alari Balagan u. Irkutsk g. Vinogradov, 10/vi.1924 Vinogradov [in Russian]”. Khabarovskiy kray: 2 ♀ (EJCB): “Sib. or. Chabarovsk 7.6.1977 Voronež Víša leg.” (JENDEK, 2003 [#5878]). Lipetskaya oblast’: 1 (MCCD): “30км В г. Елец, ур. Морозова Гора, 27.5.1996 Цуриков”; 1 (MCCD): “30км В г. Елец, ур. Морозова Гора, 9.8.2003 Цуриков”; 1 (MCCD): “30км В г. Елец, ур. Морозова Гора, 2003 Цуриков”. Magadanskaya oblast’: 4 (ZIN): “r n. Kolyma, Duskasn’ya 20. viii.1979 Migovich [in Russian]”. Novosibirskaya oblast’: 1 (NMPC): “SSSR, VII.1986, Novosibirsk, V. Bíža leg.”. Primorskiy kray: 1 ♂ (EJCB): “SU Ussuri reg. Novovarvarovka, 6 10. VII.1989, R. Červenka leg.” (JENDEK, 2003 [#5878]); 1 ♀ (EJCB): “USSR Sibiria or., Ussuri reg. Jasnoe, 450 m 11 14.7.1989, St. Jákl lgt.” (JENDEK, 2003 [#5878]); 5 (EJCB): “Sib. or. m., Primorje, Sichote Alin Mts., Sokolči, 1 15.7.1990, Kadlec + Voříšek lg.” (JENDEK, 2003 [#5878]); 1 ♀ (EJCB): “Russia Far East, Khanka Lake NW, 4km N Novokachalinsk, Bol’shije Usachi riv., 6. VII.2003, Lg. A. Napolov”. Rostovskaya oblast’: 1 (MKCY): “Novocherkassk 01.07.1990”. Sakha Respublika: 1 (NMPC): “M. Cherepanikha, 25 sev. ot Olekminska, 1.vii.1907g., N. Kharitonov [in Russian]”. Severnaya Osetiya Respublika: 2 (MKCY): “SOACCP Ordzhonikidze, Lysaya gora, na rose, 26.6.1973”. Tyumenskaya oblast’: 1 (ZIN): “Tyumenskaya obl., Uspenka, 10.6.1959”. Tyva Respublika: 1 ♂ (EJCB): “Tuva, Kok Gei, 10 km vyshe Kyzyla, 20.6.1979, na Rosa, Korotyaev leg.” (JENDEK, 2003 [#5878]); 2 ♀ (EJCB): “Tuva, Kyzyl, poima Eniseya, 13.6.1979, na Rosa, Korotyaev leg.” (JENDEK, 2003 [#5878]); 1 (MDCM): “Tuva, 19 22.VI.1979, Ulug Khemskii raion, Ishtin Khem, na plyvushchem murore, S. G. Korolëv leg.” (JENDEK, 2003 [#5878]); 1 (EJCB): “Russia, Tuva, right bank of Biy Khem riv. 800 m, Ujukskiy Mts. 2 5.VII.1999, S. N. Vashchenko leg.” (JENDEK, 2003 [#5878]); 9 (EJCB): “Russia, Tuva, right bank of Biy Khem riv. 800 m, Ujukskiy Mts. 22.V. 6.VI.1999, S. N. Vashchenko leg.” (JENDEK, 2003 [#5878]). Udmurtskaya Respublika: 2 (MKCY): “Udmurt. Votkinsk, Biost. UdGU, r[river]. Siva, 25.6.[19]90 Okulov leg.”. Voronezhskaya oblast’: 1 (MKCY): “Voronezh Tellerman Moravitsev 21.vi.1954”; 1 (MCCD): “Воронежская обл., 30км Ю г. Острогожска, 10.6.1996 Цуриков”; 1 (MCCD): “25км NNO г. Воронежа, Усманский Бор, 11.7.1997 Коротеева”. SERBIA & MONTENEGRO: Serbia: 1 (EJCB): “YU, Serbia Banat, Požarevac, Brančevo 1 5.VI.2002, M. Snížek leg.” (JENDEK, 2003 [#5878]). SLOVAKIA: 3 (EJCB):

71 “Slovakia, Hlohovec 7.1978 E. Jendek leg.” (JENDEK, 2003 [#5878]); 3 (EJCB): “Slovakia, Bratislava 6.6.1978 E. Jendek leg.” (JENDEK, 2003 [#5878]); 3 (EJCB): “Slovakia, Modra 10.6.1982 E. Jendek leg.” (JENDEK, 2003 [#5878]); 2 (EJCB): “Slovakia or, Malý Kamenec 4.7.1992, O. Majzlan lgt.” (JENDEK, 2003 [#5878]); 3 (EJCB): “Slovakia, Košice 1.6.1992 R. Lohaj leg.”; 1 (EJCB): “Slovakia, Durďošík 25.7.1995 Dr. Majzlan lgt.” (JENDEK, 2003 [#5878]); 2 (ISCK): “Slovakia or, Kuzmice env. 8.vi.2001 Smatana I. leg.” (JENDEK, 2003 [#5878]); 1 (EJCB): “Slovakia or, Kuzmice env. 2.VI.2001, I. Smatana lgt.” (JENDEK, 2003 [#5878]); 13 (EJCB): “Slovakia occ., D. Vestenice, DFS 7276c, 1.VII. 12.VIII.2001, Malaise trap, O. Majzlan leg.”; 1 (EJCB): “Slovakia occ., D. Vestenice, DFS 7276c, 27.IV.2001, Malaise trap, O. Majzlan leg.”; 1 (ISCK): “Slovakia or. Sp. Podhradie 28.vi.2002 I. Smatana leg.” (JENDEK, 2003 [#5878]). SLOVENIA: 1 (JVCO): “Slovenia Petrinje, Podgorski Kras mts., 25 26.6.2001 J. Vávra leg.” (JENDEK, 2003 [#5878]). TURKEY: 1 (EJCB): “Turkey mer. Akseki 14.6.1993 Farbiak leg.” (JENDEK, 2003 [#5878]). Ankara: 1 ♂ (EJCB): “Anatolien, Prov. Ankara, Kizilcahamam, 4.6.1975 leg. Holzschuh & Ressl” (JENDEK, 2003 [#5878]). Artvin: 1 (EJCB): “Turc., Artvin distr., Parmak Dagi, 17.VI.2001, 700 m, Parmak Dagi, Artvin, Kafkazor, Voříšek & Dostál” (JENDEK, 2003 [#5878]). Bingol: 1 (EJCB): “Anatolien, 35 km östlich von Bingöl, 10 11.6.1973, 1500 m, leg. C. Holzschuh” (JENDEK, 2003 [#5878]). Icel: 1 (EJCB): “Turkey, vil. Içel, 1200 m, Bolkar Daglari Mts. Çamliyayla 9.6.1991 Bednařík & Kovařík leg.” (JENDEK, 2003 [#5878]). Sivas: 13 (EJCB): “Turc., Sivas distr. Zara env. 9.VI.2001, 1800 m, Kurbagalibeli Geç. Voříšek & Dostál” (JENDEK, 2003 [#5878]). UKRAINE: Crimean: 1 (MKCY): “Ukraine, Crimea, env. Rybach’e, 26.08.1987, Nesterov leg.”; 1 ♂ (EJCB): “S Crymea, Alushta distr., Verknyaja Kutuzovka 21.06.96 Smirnov”; 1 (MSCI): “Krym, r. Alushty okr. p. Verkhnaya Kutuzovka 26.06.96 M. Smirnov”. Dnepropetrovsk: 2 (MSCI): “Dnepropetrovsk, okr. g. Kirovskoe, na ezhevike 15.06.93 Smirnov [in Russian]”. Kherson: 5 ♂, 1 ♀ (EJCB): “Ukraine, Kherson reg. Antonovka vill. 2.VI.1995 Mishustin, R. leg.” (JENDEK, 2003 [#5878]).

Adult occurence: 4-5-6-7-8-9 (n=103). Faunal record published by COBOS, 1972 [#591] (as aurichalceus) is also included. Altitude range: 400-2000 m (n=15). Faunal record published by COBOS, 1972 [#591] (as aurichalceus) is also included. Host plants: Rosa: MÉNÉTRIÉS, 1832 [#1799]; ABEILLE DE PERRIN, 1897 (as rubicola) [#19]; WEISS, 1916 (as viridis) [#6742]; WEISS, 1916 (as viridis) [#3913]; OBENBERGER, 1927 (as mokrzeckii) [#2064]; OGLOBLIN & REICHARDT, 1932 (as mokrzeckii) [#8654]; SAINTE CLAIRE, 1935 (as communis) [#8551]; GUSSAKOVKII, 1949 [#7805]; MÜHLMANN, 1954 (as mokrzeckii) [#8152]; HORION, 1955 (as aurichalceus) [#1285]; TER MINASYAN, 1955 [#7893]; HEERING, 1956 (as rubicola) [#3716]; ALEXEEV, 1957 [#3226]; ALEXEEV, 1958 [#3982]; ALEXEEV, 1959 [#3229]; ALEXEEV, 1961 [#3243]; BALACHOWSKI et al., 1962 (as mokrzeckii) [#96]; RIKHTER & ALEXEEV, 1965 [#2522]; GURYEVA, 1974 [#5428]; ALEXEEV, 1975 (as kuznecovinus) [#52]; WELLSO et al., 1976 (as rubicola) [#3079]; HELLRIGL, 1978 (as aurichalceus) [#3442]; ALEXEEV, 1981 [#61]; BÍLÝ, 1982 (as aurichalceus) [#295]; COBOS, 1986 (as aurichalceus) [#609]; ALEXEEV & VOLKOVITSH, 1989 (as kuznecovinus) [#3241]; RUICANESCU, 1993 (as aurichalceus) [#3257]; CURLETTI, 1994 (as aurichalceus) [#658]; KRÓLIK & MAJEWSKI, 1994 (as aurichalceus) [#3471]; VOLKOVITSH & ALEXEEV, 1994 [#2974]; DOWNIE & ARNETT, 1996 (as aurichalceus) [#6170]; KUROLI, 1999 (as

72 aurichalceus) [#5916]; TLEPPAEVA, 1999 [#6551]; BÍLÝ, 2002 [#5877]; BRECHTEL & KOSTENBADER, 2002 [#5766]; MACRAE & NELSON, 2003 [#5977]; SIKES, 2004 [#7233]; TLEPPAEVA & ISHKOV, 2004 [#7211]; KOROTYAEV et al., 2005 [#7284]. − R. acicularis: LARSON, 2003 (as aurichalceus) [#6990]. − R. alpina: SCHAEFER, 1949 (as aurichalceus) [#2565]; HASTIR & GASPAR, 2002 (as aurichalceus) [#6211]. − R. arkansiana: LARSON, 2003 (as aurichalceus) [#6990]. − R. blanda: WEISS, 1914 (as politus) [#3920]; WEISS, 1914 (as aurichalceus) [#3920]; FISHER, 1928 [#980]. − R. canina: SCHAEFER, 1949 (as aurichalceus) [#2565]; FRANZ, 1974 (as aurichalceus) [#4334]; VOLKOVITSH, 1977 [#2985]; WEIDLICH, 1989 (as calcicola) [#6165]; NIEHUIS & TEZCAN, 1993 (as aurichalceus) [#1910]; HASTIR & GASPAR, 2002 (as aurichalceus) [#6211]; GUTOWSKI & MILKOWSKI, 2008 [#8244]. − R. carolina: WEISS, 1914 (as politus) [#3920]; WEISS, 1914 (as aurichalceus) [#3920]; CHAMBERLIN, 1926 [#468]; FISHER, 1928 [#980]. − R. damascena: ZAGAIKEVICH, 1962 [#3249]; NIEHUIS & TEZCAN, 1993 (as aurichalceus) [#1910]. − R. gallica: ZAGAIKEVICH, 1962 [#3249]. − R. hugonis: FISHER, 1928 [#980]. − R. japonica: SCHAEFER, 1949 (as aurichalceus) [#2565]. − R. multiflora japonica: WEISS, 1914 (as politus) [#3920]; WEISS, 1914 (as aurichalceus) [#3920]; FISHER, 1928 [#980]. − R. nitida: WEISS, 1914 (as politus) [#3920]; WEISS, 1914 (as aurichalceus) [#3920]; FISHER, 1928 [#980]. − R. nutkana var. hispida: NELSON et al., 1996 [#3975]. − R. rubrafolia: WEISS, 1914 (as politus) [#3920]; WEISS, 1914 (as aurichalceus) [#3920]; FISHER, 1928 [#980]. − R. rugosa: WEISS, 1914 (as politus) [#3920]; WEISS, 1914 (as aurichalceus) [#3920]; WEISS, 1915 [#5953]; FROST, 1924 [#1030]; CHAMBERLIN, 1926 [#468]; FISHER, 1928 [#980]; SCHAEFER, 1949 (as aurichalceus) [#2565]; OHMOMO, 2004 [#6850]. − R. setigera: WEISS, 1914 (as politus) [#3920]; WEISS, 1914 (as aurichalceus) [#3920]; FISHER, 1928 [#980]; SCHAEFER, 1949 (as aurichalceus) [#2565]. − R. woodsii: LARSON, 2003 (as aurichalceus) [#6990]. – Rubus: HEYDEN, 1897 (as obtusus) [#5286]; CAILLOL, 1913 (as rubicola) [#4053]; OBENBERGER, 1924 [#2040] (as communis); VASILEV, 1927 [#5834]; OGLOBLIN & REICHARDT, 1932 (as rubicola) [#8654]; SAINTE CLAIRE, 1935 (as rubicola) [#8551]; MUNDINGER, 1941 (as rubicola) [#5414]; GUSSAKOVKII, 1949 [#7805]; RIKHTER, 1950 (as aurichalceus) [#3881]; HORION, 1955 (as aurichalceus) [#1285]; TER MINASYAN, 1955 (as aurichalceus) [#7893]; ALEXEEV, 1957 [#3226]; ALEXEEV, 1959 [#3229]; ALEXEEV, 1961 [#3243]; RIKHTER & ALEXEEV, 1965 [#2522]; SIMON, 1966 (as aurichalceus) [#3266]; FRANZ, 1974 (as aurichalceus) [#4334]; GURYEVA, 1974 [#5428]; ALEXEEV, 1975 (as kuznecovinus) [#52]; WELLSO et al., 1976 (as rubicola) [#3079]; HELLRIGL, 1978 (as aurichalceus) [#3442]; ALEXEEV, 1981 [#61]; BÍLÝ, 1982 (as aurichalceus) [#295]; COBOS, 1986 (as aurichalceus) [#609]; ALEXEEV & VOLKOVITSH, 1989 (as kuznecovinus) [#3241]; RUICANESCU, 1993 (as aurichalceus) [#3257]; CURLETTI, 1994 (as aurichalceus) [#658]; KRÓLIK & MAJEWSKI, 1994 (as aurichalceus) [#3471]; VOLKOVITSH & ALEXEEV, 1994 [#2974]; KUROLI, 1999 (as aurichalceus) [#5916]; TLEPPAEVA, 1999 [#6551]; BÍLÝ, 2002 [#5877]. − R. caesius: BEDEL, 1921 [#170]; SCHAEFER, 1949 (as aurichalceus) [#2565]; NIEHUIS & TEZCAN, 1993 (as aurichalceus) [#1910]; NIEHUIS, 1988

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA (as aurichalceus) [#1912]; BRECHTEL & KOSTENBADER, 2002 [#5766]; HASTIR & GASPAR, 2002 (as aurichalceus) [#6211]. − R. fruticosus: SCHAEFER, 1949 (as aurichalceus) [#2565]; NIEHUIS & TEZCAN, 1993 (as aurichalceus) [#1910]; NIEHUIS, 1988 (as aurichalceus) [#1912]; BRECHTEL & KOSTENBADER, 2002 [#5766]; HASTIR & GASPAR, 2002 (as aurichalceus) [#6211]. − R. idaeus: BEDEL, 1921 [#170]; SCHAEFER, 1949 (as aurichalceus) [#2565]; NIEHUIS & TEZCAN, 1993 (as aurichalceus) [#1910]; NIEHUIS, 1988 (as aurichalceus) [#1912]; BRECHTEL & KOSTENBADER, 2002 [#5766]; HASTIR & GASPAR, 2002 (as aurichalceus) [#6211]. − R. woodsii var. ultramontana: NELSON et al., 1996 [#3975]. Dubious: Hulthemia: ALEXEEV, 1975 (as kuznecovinus) [#52]. Erroneous: Acer: ALEXEEV, 1981 [#61]. − Cornus sanguinea: TROPIN, 1965 (as foveolatus) [#8652]; ALEXEEV, 1981 [#61]; ALEXEEV & VOLKOVITSH, 1989 (as kuznecovinus) [#3241]; MERKL, 1991 (as aurichalceus) [#6564]. − Phlox: MACRAE & NELSON, 2003 [#5977]. − Salix: SAINTE CLAIRE, 1935 (as aurichalceus) [#8551]; ALEXEEV, 1958 (as aurichalceus) [#3982]; ALEXEEV, 1961 (as aurichalceus) [#3243]; RIKHTER & ALEXEEV, 1965 (as aurichalceus) [#2522]; GURYEVA, 1974 (as aurichalceus) [#5428]; VOLKOVITSH, 1977 (as aurichalceus) [#2985]. Distribution: Local: CHINA: Heilongjiang, Jilin, Liaoning. JAPAN: Hokkaido. KOREA NORTH. MONGOLIA. RUSSIA: Amurskaya oblast’, Buryatiya Respublika, Irkutskaya oblast’, Khabarovskiy kray, Primorskiy kray, Sakha Respublika, Tyva Respublika, Zabaykalskiy kray. Global: ASIA: ARMENIA. AZERBAIJAN. CHINA. GEORGIA. JAPAN. KAZAKHSTAN. KOREA NORTH. KYRGYZSTAN. RUSSIA. TURKEY. TURKMENISTAN. EUROPE: ALBANIA. AUSTRIA. BELARUS. BELGIUM. BOSNIA & HERZEGOVINA. BULGARIA. CROATIA. CZECH REPUBLIC. ESTONIA. FINLAND. FRANCE. GERMANY. GREECE. HUNGARY. ITALY. LATVIA. LIECHTENSTEIN. LITHUANIA. LUXEMBOURG. MACEDONIA. MOLDOVA. NETHERLANDS. POLAND. PORTUGAL. ROMANIA. RUSSIA. SERBIA & MONTENEGRO. SLOVAKIA. SLOVENIA. SPAIN. SWEDEN. SWITZERLAND. TURKEY. UKRAINE. UNITED KINGDOM. NORTH AMERICA (introduced): CANADA. UNITED STATES.

Agrilus cyaneomicans Nonfried sp. inquirenda cyaneomicans Nonfried, 1892 (Agrilus) NONFRIED, 1892: 85, 91 [#1924] (description). KERREMANS, 1903: 284 [#1441] (catalog). OBENBERGER, 1936: 980 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 355 [#2429] (checklist; China). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2051 [#8576] (world catalog).

Type material: Agrilus cyaneomicans Nonfried, 1892 Type locality: not given [Südchina, Kwang-Si, Umgebung von Nienghali 24 nördl. Breite und 108 östl. Länge is cited in the introduction].

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Type specimens: Not located. Described from a single specimen. Host plants: Unknown. Distribution: CHINA: Guangxi. Remarks: This taxon is species inquirenda and its name is nomen dubium.

Agrilus cyaneoniger Saunders Figs. 8A, 8B (habitus); Fig. 32A (aedeagus); Figs. 43B, 44W, 44X, 47I, 49D, 49F, 49I (types) cyaneoniger Saunders, 1873 (Agrilus) SAUNDERS, 1873: 515 [#2554] (description). LEWIS, 1879: 15 [#3345] (catalog; Japan). KERREMANS, 1885: 152 [#1381] (catalog). SCHÖNFELDT, 1887: 113 [#8901] (Anambus; catalog; Japan). KERREMANS, 1892: 254 [#1397] (catalog). LEWIS, 1893: 331 332 [#1722] (faunal records; Japan). KERREMANS, 1903: 276 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). YUASA, 1932: 654 [#3369] (characters; Japan). MIWA & CHÛJÔ, 1936: 16 [#1809] (catalog; Japan). OBENBERGER, 1936: 980 981 [#2191] (world catalog). CHÛJÔ & MATUDA, 1940: 65 [#500] (checklist; faunal records; Japan (Kyushu)). IGA, 1947: 15 [#3273] (notes; characters; faunal records; Japan (Shikoku)). KUROSAWA, 1947: 1 2 [#3339] (note). YUASA, 1949: 127 [#8108] (characters; Japan). CHÛJÔ & KUROSAWA, 1950: 8 9 [#501] (catalog; faunal records; Japan (Shikoku)). KUROSAWA, 1950: 1114 [#3274] (note). MÜHLMANN, 1954: 83 [#8152] (notes). IGA, 1955: 78 [#3828] (iconography; Japan). KUROSAWA, 1956: 1114 [#8107] (characters; Japan). OBENBERGER, 1958: 237 [#2389] (faunistic redords; China). IGA, 1962: 78 [#3351] (iconography; Japan). KUROSAWA, 1963: 153 [#1618] (characters; Japan). KUROSAWA, 1974: 1 [#4576] (note). KUROSAWA, 1974: 1, 2 [#4575] (characters; notes). TÔYAMA, 1985: 22 23 [#3678] (iconography; Japan). ANONYMOUS, 1986: 177 [#8848] (insect pest; Korea). HOU, 1987: 589 [#3490] (notes; Yunnan). PENG ZHONGLIANG, 1987: 355 [#2429] (checklist; China). ALEXEEV, 1989: 479 [#64] (subspecies of cyaneoniger; key to species; Primorsk; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). ANONYMOUS, 1991: 106 [#8552] (faunal records; Japan: Aomori). LI JINGKE, 1992: 92 [#6755] (checklist; northeastern China). PENG ZHONGLIANG, 1992: 399 [#2431] (characters; notes; Hunan). JENDEK, 1995: 140 141 [#3768] (lectotype designation; synonymy; characters; Russian Far East; Korea; Japan; China; India). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 184 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 31 [#4091] (checklist; Japan: Hokkaido, Honshu, Shikoku, Kyushu, Yakushima I., Oki). ALEXEEV, 1998: 369 [#4322] (subgenus Anambus; subgeneric classification). NONNAIZAB et al., 1999: 112 [#8247] (checklist; China: Inner Mongolia). JENDEK, 2000: 187 188, 192 [#4852] (faunal records; key). HOU TAOQIAN & ZENG RUI, 2002: 357 [#6626] (checklist; China: Hainan). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Hubei, Jiangxi, Hunan, Sichuan, Yunnan). PENG ZHONGLIANG, 2002: 267 [#6628] (characters; Fujian). FUKUTOMI & HORI, 2004: 14 [#6851] (checklist; faunal records; Japan (Hokkaido)). OHMOMO, 2005: 236 241 [#7253] (phenotypical analysis; faunal records; Russia; Japan; Korea). JENDEK, 2006: 390 [#7441] (subgenus Anambus; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (checklist; Japan). ANONYMOUS, 2007: 32 [#8250] (checklist; Japan: Honshu). BELLAMY, 2008: 2051 2052 [#8576] (subgenus Anambus; world catalog).

73 melanopterus Solsky, 1876 (Agrilus) HEYDEN, SOLSKY, 1876: 277 279 [#2644] (description). 1881: 119 [#5548] (catalog). KERREMANS, 1885: 153 [#1381] (catalog). MARSEUL, 1887: 252 [#4707] (catalog). KERREMANS, 1892: 263 [#1397] (catalog). HEYDEN, 1893: 90 [#3450] (catalog). ABEILLE DE PERRIN, 1897: 4 [#19] (characters in key). KERREMANS, 1903: 287 [#1441] (catalog). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1935: 162 [#2158] (note). OBENBERGER, 1936: 980 [#2191] (variety of cyaneoniger; world catalog). IGA, 1955: 78 [#3828] (variety of cyaneoniger; iconography; Japan). IGA, 1962: 78 [#3351] (variety of cyaneoniger; iconography; Japan). KUROSAWA, 1974: 1 [#4576] (subspecies of cyaneoniger). ALEXEEV, 1989: 479 [#64] (subspecies of cyaneoniger; key to species; Korea; northeastern China). HIRASHIMA, 1989: 323 [#8078] (subspecies of cyaneoniger; checklist; Japan). JENDEK, 1995: 140 [#3768] (synonym of cyaneoniger; lectotype designation). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (subspecies of cyaneoniger; checklist; Japan). AKIYAMA & OHMOMO, 1997: 31 32 [#4091] (subspecies of cyaneoniger; checklist; Japan: Tsushima; Korea; China; Russia: East Siberia; Kashmir). JENDEK, 2000: 187, 188, 192 [#4852] (subspecies of cyaneoniger; faunal records; key). HUA LI ZHONG, 2002: 89 [#6696] (subspecies of cyaneoniger; checklist; China). JENDEK, 2003: 181 [#5823] (subspecies of cyaneoniger; synonymy). OHMOMO, 2005: 236 241 [#7253] (synonym of cyaneoniger; phenotypical analysis). JENDEK, 2006: 390 [#7441] (synonym of cyaneoniger; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (synonym of cyaneoniger). BELLAMY, 2008: 2052 [#8576] (synonym of cyaneoniger; world catalog). VOLKOVITSH, 2009: 134 [#8850] (subgenus Anambus; subspecies of cyaneoniger; faunal records; Russia: Primorskiy kray). cyaneoniger Thomson, 1879 (Agrilus; [preoccupied]) THOMSON, 1879: 71 [#2909] (description). JAKOBSON, 1913: 798 [#4386] (synonym of jamesi; catalog). OBENBERGER, 1936: 980 [#2191] (synonym of cyaneoniger Saunders). JENDEK, 1995: 140 [#3768] (synonym of cyaneoniger; lectotype designation). AKIYAMA & OHMOMO, 1997: 31 [#4091] (synonym of cyaneoniger Saunders). JENDEK, 2006: 390 [#7441] (synonym of cyaneoniger Saunders; Palaearctic catalog). BELLAMY, 2008: 2052 [#8576] (synonym of cyaneoniger; world catalog). impressifrons Kiesenwetter, 1879 (Agrilus) Kiesenwetter in KRAATZ & KIESENWETTER, 1879: 254 255 [#1595] (description). HEYDEN, 1881: 119 [#5548] (catalog). HEYDEN, 1884: 290 [#8482] (faunal record; Amur region). KERREMANS, 1885: 153 [#1381] (catalog). MARSEUL, 1887: 252 [#4707] (catalog). KERREMANS, 1892: 260 [#1397] (catalog). HEYDEN, 1893: 90 [#3450] (catalog). ABEILLE DE PERRIN, 1897: 3 [#19] (characters in key). KERREMANS, 1903: 286 [#1441] (catalog). SEMENOV, 1903: 171 172 [#4114] (synonym of melanopterus). JAKOBSON, 1913: 797 [#4386] (synonym of melanopterus; catalog). FISHER, 1925: 4 [#975] (note). OBENBERGER, 1936: 981 [#2191] (synonym of cyaneoniger variety melanopterus). JENDEK, 1995: 140 [#3768] (synonym of cyaneoniger). AKIYAMA & OHMOMO, 1997: 32 [#4091] (synonym of cyaneoniger melanopterus). JENDEK, 2003: 181 [#5823] (synonym of cyaneoniger melanopterus; lectotype designation). JENDEK, 2006: 390 [#7441] (synonym of cyaneoniger; Palaearctic catalog). BELLAMY, 2008: 2052 [#8576] (synonym of cyaneoniger; world catalog). jamesi Jakobson, 1913 (Agrilus; replacement name for cyaneo niger Thomson not Saunders) JAKOBSON, 1913: 798 [#4386] (replacement name proposal). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 980 [#2191] (synonym of cyaneoniger; world

74 catalog). PENG ZHONGLIANG, 1992: 399 [#2431] (synonym of cyaneoniger). JENDEK, 1995: 140 [#3768] (synonym of cyaneoniger). AKIYAMA & OHMOMO, 1997: 31 [#4091] (synonym of cyaneoniger Saunders). HUA LI ZHONG, 2002: 89 [#6696] (synonym of cyaneoniger). JENDEK, 2006: 390 [#7441] (synonym of cyaneoniger; Palaearctic catalog). BELLAMY, 2008: 2052 [#8576] (synonym of cyaneoniger; world catalog). marquardti Obenberger, 1914 (Agrilus) OBENBERGER, 1914: 41 42 (in Czech), 47 48 (in German) [#1965] (description). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 980 [#2191] (variety of cyaneoniger; world catalog). DESCARPENTRIES & VILLIERS, 1963: 2, 8 [#701] (subspecies of cyaneoniger; characters; faunal records; Tonkin; Chine). PENG ZHONGLIANG, 1987: 355 [#2429] (subspecies of cyaneoniger; checklist; China). JENDEK, 1995: 140 [#3768] (synonym of cyaneoniger; lectotype designation). AKIYAMA & OHMOMO, 1997: 31 [#4091] (synonym of cyaneoniger melanopterus). HUA LI ZHONG, 2002: 89 [#6696] (subspecies of cyaneoniger; checklist; China: Guangxi, Yunnan). JENDEK, 2006: 390 [#7441] (synonym of cyaneoniger; Palaearctic catalog). BELLAMY, 2008: 2052 [#8576] (synonym of cyaneoniger; world catalog). ataman Obenberger, 1924 (Agrilus) OBENBERGER, 1924: 35, 38 [#2040] (description). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1935: 162 [#2158] (subspecies of melanopterus). OBENBERGER, 1936: 981 [#2191] (variety of cyaneoniger; world catalog). JENDEK, 1995: 140, 141 [#3768] (synonym of cyaneoniger; lectotype designation). AKIYAMA & OHMOMO, 1997: 32 [#4091] (synonym of cyaneoniger melanopterus). JENDEK, 2006: 390 [#7441] (synonym of cyaneoniger; Palaearctic catalog). BELLAMY, 2008: 2052 [#8576] (synonym of cyaneoniger; world catalog). mikado Obenberger, 1924 (Agrilus) OBENBERGER, 1924: 35 36, 38 [#2040] (description). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 18 [#1809] (catalog; Japan). OBENBERGER, 1936: 981 [#2191] (synonym of cyaneoniger; world catalog). KUROSAWA, 1963: 153 [#1618] (form of cyaneoniger; characters; Japan). JENDEK, 1995: 140, 141 [#3768] (synonym of cyaneoniger; lectotype designation). AKIYAMA & OHMOMO, 1997: 31 [#4091] (synonym of cyaneoniger Saunders). OHMOMO, 2005: 236 241 [#7253] (synonym of cyaneoniger). JENDEK, 2006: 390 [#7441] (synonym of cyaneoniger; Palaearctic catalog). BELLAMY, 2008: 2052 [#8576] (synonym of cyaneoniger; world catalog).

Type material: Agrilus cyaneoniger Saunders, 1873 Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 44W), ♂, designated by JENDEK (1995: 140-141) [#3768], (BMNH): “Type H.T. [p] [round label with red border] \ Japan. G. Lewis. 1910 - 320. [p] \ cyaneoniger ES. Type [h] [by Saunders hand]”. Described from unknown number of syntypes. Agrilus melanopterus Solsky, 1876 Type locality: sur les bords de l’Oussouri. Type specimens: Primary: Lectotype (Fig. 49F), ♂, designated by JENDEK (1995: 140) [#3768], (ZIN): “Ussuri mer [h] [yellow label] \ A. melanopterus mihi [h] [violet label] \ k. Solskogo [ = collection Solsky] [p] [in Russian]”. Described from unknown number of syntypes. Agrilus cyaneoniger Thomson, 1879 Type locality: Japonia. Type specimens: Primary: Lectotype (Fig. 44X), sex

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA not examined, designated by JENDEK (1995: 140) [#3768], (MNHN): “Th. Type [p] [white label with black border] \ cyaneoniger Thoms. M. ss. Type Ap. I Japon [h] \ EX MUSAEO JAMES THOMSON [p] \ collectio Oberthur [p]”. Described from unknown number of syntypes. Agrilus impressifrons Kiesenwetter, 1879 Type locality: not given [Amur is cited in the title of the publication]. Type specimens: Primary: Lectotype (Fig. 47I), ♀, designated by JENDEK (2003: 181) [#5823], (ZSMC): “Sibirien Amur Christoph [18]77 [p] \ Kiesenwetter [p] \ Sammlung Cl. Müller [p] \ Type von [p] Agrilus impressifrons Kiesw. [h] [red label] \ Holotypus [p] Agrilus impressifrons Kiesenw. [h] Zool. Staatsammlg. München [red label]”. Described from unknown number of syntypes. Agrilus jamesi Jakobson, 1913 Type locality and specimens: See: Agrilus cyaneoniger Thomson, 1879. Agrilus marquardti Obenberger, 1914 Type locality: Kiang-si in China. Type specimens: Primary: Lectotype (Fig. 49D), ♂, designated by JENDEK (1995: 140) [#3768], (NMPC): “Kiangsi [ = Jiangxi province] China [p] \ Typus [p] [red label] \ Agrilus Marquardti m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24785 [h] [orange label]”. Secondary: 1 PLT (DEI); 1 PLT (NMPC). Described from unknown number of syntypes. Agrilus ataman Obenberger, 1924 Type locality: Vladivostok. Type specimens: Primary: Lectotype (Fig. 43B), sex not examined, designated by JENDEK (1995: 140, 141) [#3768], (NMPC): “Vladivostok [h] \ Typus [p] [red label] \ Agrilus Ataman m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24788 [h] [orange label]”. Described from unknown number of syntypes. Agrilus mikado Obenberger, 1924 Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 49I), sex not examined, designated by JENDEK (1995: 140, 141) [#3768], (NMPC): “Japan [p] \ Typus [p] [red label] \ Agrilus Mikado m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24787 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 11-14 mm; body robust, black; eyes moderate; pronotum widest around middle; prehumerus absent. Species group: Cyaneoniger. Relevant species: A. planipennis Fairmaire, 1888. Specimens examined: CHINA: Guizhou: 1 ♀ (EJCB): “China, Guizhou prov., Leigongshan, Xijiang, 29 May 2 June 1997, 1200 1900 m, Bolm lgt.” (JENDEK, 2000 [#4852]). Hebei: 1 ♂ (EJCB): “China, Hebei, Xingtai Taihang Mts., 1300 m, Neiqiu, Xinjiayu, 2002, M. Knížek lgt. 17 20.V. From pheromone trap baited for bark bark beetles Dendoctonus valens LeConte”. Hubei: 1 (EJCB): “P.R. China,, Hubei, Dabieshan, N31˚07’06’’ E115˚48’57”, 11 21. vi.2008, 640 m, V. Grebennikov leg.”. Jiangxi: 1 ♂, 2 ♀ (EJCB): “Kiang Si 1901” (JENDEK, 1995 [#3768]). Jilin: 1 (EJCB): “China, 23.VI.1980 Changbai Shan Forest protective area” (JENDEK, 1995 [#3768]). Shaanxi: 1 ♀ (SMNS): “China: Shaanxi Prov. , Taibai Shan above Houshenzi, 1300 1700 m, 9.VI. 3.VII.1998 leg. P. Jäger

75

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA & J. Martens”; 2 ♂, 7 ♀ (EJCB): “China, 1000 1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v. 13.vi.1998, I. H. Marshal leg.” (JENDEK, 2000 [#4852]); 1 ♀ (EJCB): “China, S Shaanxi, 1700 m, 15 km SW Dongiangkou, 1998, Bolm lgt, 14 17 Jul.” (JENDEK, 2000 [#4852]); 3 (EJCB): “C China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000 1300m, 23.V. 13.VI. leg. C. Holzschuh 2000”. Sichuan: 1 ♂, 2 ♀ (EJCB): “China, Sichuan” (JENDEK, 2000 [#4852]); 15 (EJCB): “China, Sichuan, 8 13.VI.[19]91, Abazhou, 2000 m, Nanping, Jiuzhaigou, 33.15N / 104.15” (JENDEK, 1995 [#3768]); 1 ♀ (EJCB): “CH Sichuan 1992 Gonga Shan Moxi, 50 km S of Luding, Sauer leg.” (JENDEK, 1995 [#3768]); 1 ♀ (EJCB): “Sichuan, Moxi, Gongashan mts., 28.VI. 2.VII.1994, Bolm lgt, 1650 m” (JENDEK, 2000 [#4852]); 1 (EJCB): “China, Sichuan, Gonga Shan, Moxi, 1300 m, 3.VII.1996, 29°13N, 102°10E, Collected by J. Farkač, P. Kabátek and A. Smetana” (JENDEK, 2000 [#4852]). Yunnan: 1 ♀ (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.” (JENDEK, 1995 [#3768]); 4 (EJCB): “CH Yunnan, 13 14.VI.1995, 100 km W of Kunming, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.” (JENDEK, 2000 [#4852]). Zhejiang: 1 ♀ (EJCB): “Qin County, Zhejiang” (JENDEK, 1995 [#3768]). INDIA: Jammu & Kashmir: 2 ♀ (EJCB): “Goorais valley Kasmir 7000 ft, VI.1887” (JENDEK, 1995 [#3768]). JAPAN: Honshu: 2 (EJCB): “Fukushima Pref., Kawauchi VII, 1999” (JENDEK, 2000 [#4852] as cyaneoniger melanopterus); 1 ♀ (JMCZ): “Japan, 10.8.1953, Nikko, J.Aoki leg.”; 1 ♀ (EJCB): “Showa, Okayama pref., Japan, June 4, [19]’67 T. Aono leg.” (JENDEK, 1995 [#3768]); 1 ♂ (JMCZ): “Japan, Ohtaki, Fukushima City, 21.8.1980, S. Ohmomo lgt.”; 2 (EJCB): “Japan, Iwate, Gosho dam, Morioka shi, 16.vi.1993”; 5 (EJCB): “Moniwa, Iizaka Town, Fukushima City, Fukushima Pref., Japan, 25 June 1995, Nobuo Sagiura leg.” (Jendek, 2000 [#4852] as cyaneoniger melanopterus); 2 (EJCB): “Fukushima Pref., Mt. Atsukashi Yama, Kunimi town, 4.VII., 22.VII.1998” (Jendek, 2000 [#4852] as cyaneoniger melanopterus); 1 (EJCB): “Fukushima Pref., Kaida, Kunimi Town, 25.VI, 1999” (Jendek, 2000 [#4852] as cyaneoniger melanopterus). Kyushu: 4 (EJCB): “Kyushu, Japan, 25 26.V.1991, Kurodake, Kujû mts. Oita Pref., Ryô Noda leg.” (Jendek, 2000 [#4852]). Tsushima: 1 ♀ (EJCB): “Tsushima Is., 12.6.1981” (Jendek, 2000 [#4852] as cyaneoniger melanopterus). KOREA NORTH: 1 ♀ (EJCB): “Corée Mirinai” (Jendek, 1995 [#3768]); 1 ♂ (EJCB): “1913 Corée Tyek Syeng” (Jendek, 1995 [#3768]); 1 (ZSCB): “Korea sept. Mjongjiangsan 17.vi.1990 Z. Šustek leg.”; 1 (ZSCB): “Korea sept. Haeju waterfalls 5.vii.1990 Z. Šustek leg.”. RUSSIA: Amurskaya oblast’: 1 (EJCB): “USSR, 15.6.1975 Amurskja Obl., Saskali, Bereja river, leg. V. Kuznetsov” (Jendek, 1995 [#3768]). Khabarovskiy kray: 2 (ZIN): “Khabarovskii krai, Yuzh Bikina 24.7.1958 Kabakov [in Russian]”. Primorskiy kray: 1 (ZIN): “Yu Primor’e Suputinskii zapovednik 10.6.1950 [in Russian]”; 1 (ZIN): “Yu Primor’e Suputinskii zapovednik 10.7.1950 [in Russian]”; 1 (ZIN): “USSR, 29.VI.1983, Primorskij Kraj, Barabat Levada, leg. V. Kuznetsov” (Jendek, 1995 [#3768]); 1 ♂ (JMCZ): “Ross. or., Prim. kr., Termotaežnnaja s., 25.VII.1983, Kuzněcov, V.”; 1 (MNHE): “Russia, 09 11.07.1989, Primorye Distr., Kamenushka, leg. U. Roosileht”; cca 20 (EJCB): “USSR, Ussuri prov. Arsenev env., VII.1990, leg. B. Šiška & R. Čermák” (Jendek, 1995 [#3768]); 1 (EJCB): “Sib. or. m., Primorje, Sichote Alin Mts., Sokolči, 1 15.7.1990, Kadlec + Voříšek lg.” (Jendek, 1995 [#3768]); 1 ♀ (JMCZ): “Ussuri country m., Novočugujevka 15 20.7.1990, leg. D. Boukal”; 1 (EJCB): “SSSR, Primorje, Barabash Levada, VII.1991, leg. Shestopalov” (Jendek, 1995 [#3768]); 1 (EJCB): “Rossia or, 2.7.[19]91 Primorsk, Turij Roh, Kuzněcov lgt.” (Jendek, 2000 [#4852] as cyaneoniger melanopterus); cca 150 (EJCB): “Russia or., Primorskiy kray, 44°10’00’’N, 133°15’00’’E, Arsen’yev env., June 1991, M. Štrba & O. Šauša leg.” (Jendek, 1995 [#3768]); 5 (EJCB): “Sib. or. Primorje, Komarovka flum., Kamenushka env., 300 m, Voříšek, J.

leg. VI.,VII, 1992” (Jendek, 1995 [#3768]); 1 (WBCW): “Russia, Arsenev 25 31.VI.1992 leg. D. Štroň” (Jendek, 1995 [#3768]); 3 (EJCB): “Russia E., Primorskyi reg. Artyom town. env., 3 15. VII.1996, Plutenko leg.” (Jendek, 2000 [#4852] as cyaneoniger melanopterus); 1 (MSCI): “SE Maritime Prov., Khasansky dist., Zanadvorovka env., Kedrovaja riv. 9 17.07.2000 leg. Igor Melnik”; 1 (FACF): “Russia Far East, S Primorie, Hanka lake, 30.vii.2002”; 1 (FACF): “Russia Far East, S Primorie, Ussurijsk, 27.vi.2003”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Preobrazhenie env., 9.vii.2006 leg. М.& L. Smirnov’s”.

Adult occurence: 5-6-7-8 (n=52). Faunal records published by OBENBERGER, 1958 [#2389] and CHÛJÔ & MATUDA, 1940 [#500] are also included. Altitude range: 300-2000 m (n=13). Faunal records published by OBENBERGER, 1958 [#2389] are also included. Host plants: Quercus: MÜHLMANN, 1954 [#8152]; ALEXEEV, 1989 (as melanopterus) [#64]; ALEXEEV, 1989 [#64]; HUA LI ZHONG, 2002 [#6696]. − Q. acutissoma: AKIYAMA & OHMOMO, 1997 [#4091]. − Q. dentata: KUROSAWA, 1947 [#3339]. − Q. mongolica var. grosserrata: AKIYAMA & OHMOMO, 1997 [#4091]. − Q. variabilis: AKIYAMA & OHMOMO, 1997 [#4091]. − Q. serrata: LEWIS, G., 1893 [#1722] (as variety cupreoviridis); AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: CHINA: Guizhou, Hebei, Jiangxi, Mongol, Shaanxi, Shanxi, Sichuan, Yunnan, INDIA. JAPAN: Hokkaido, Honshu, Kyushu, Tsushima. KOREA NORTH. KOREA SOUTH. Amurskaya oblast’, Primorskiy kray. VIETNAM.

Jilin, Nei Zhejiang. Shikoku, RUSSIA:

Agrilus cyaneovirens Bourgoin Fig. 26D (habitus); Fig. 39F (aedeagus); Fig. 45A (type) cyaneovirens Bourgoin, 1922 (Agrilus) BOURGOIN, 1922: 24 [#365] (description). OBENBERGER, 1936: 1079 [#2191] (world catalog). DESCARPENTRIES & VILLIERS, 1963: 108, 117 [#700] ([Note: The original description neither implies not requires that there were syntypes, then Descarpentries & Villiers by the mention about “the holotype” validly fixed the lectotype (ICZN, Article 74.6.)]; lectotype designation; type locality determined by lectotype: “Tonkin: Tuyen Quan (de Galassus)”; characters; faunal records). BELLAMY, 2008: 2052 [#8576] (world catalog).

Type material: Agrilus cyaneovirens Bourgoin, 1922 Type locality: not given [Note: Indo-Chine française is cited in the title of the publication]. Type specimens: Primary: Lectotype (Fig. 45A), ♂, designated by DESCARPENTRIES & VILLIERS (1963: 108, 117) [#700], (MNHN): “Tuyen-Quan G. de Galassus [h] \ Agrilus [h] Type [h] [red ink] cyaneovirens A. Bourgoin [h] \ Muséum Paris 1935 coll. A. Théry [p]”. Described from unknown number of syntypes. Diagnosis: 6.8-8.2 mm; eyes large; apicalmost part of elytra glabrous. Species group: Cyaneovirens. Relevant species: A. bonnottei Bourgoin, 1925. Specimens examined: CHINA: Yunnan: 1 ♀ (NMPC): “Yun’nan’, Damonlun, 700 m, 12.VI.1957, Pu Fu li i dr. [in Russian]”. LAOS: Attapeu: 1 ♂ (EJCB): “Laos south, Attapu prov., Bolaven Plateau, 18 30.IV.1999, 15 km SE of Ban Houaykong, Nong Lom (lake)

76 env., N 15º02’, E 106º35’, alt. 800m, E. Jendek & O. Šauša leg.”. Borikhamxai: 1 ♂ (EJCB): “Laos centr., Bolikhamsai prov., Ban Nape Kaew Nua Pass, 18.4. 1.5.1998, alt. 600±100 m, N 18º22.3, E 105º09.1 (GPS), E. Jendek & O. Šauša leg.”. Champasak: 1 ♂ (EJCB): “Laos south., Champasak prov., Bolaven Plateau, 10 18. IV.1999, Route (No. 23) Pakse Paksong, Ban Itou env. (km.35), alt. 800m, N 15º10.4’, E 106º05.8’ (GPS), E. Jendek & O. Šauša leg.”. Houaphan: 1 ♂, 3 ♀ (EJCB): “Laos NE, Houa Phan prov., 20°12 13.5’N, 103°59’.5 104°01’E, Ban Saluei → Phou Pane Mt., 1340 1870m, 15.iv. 15.v.2008, Lao collectors leg.”. Louang Namtha: 1 ♂ (EJCB): “Laos, Louang Namtha pr., 21°09’N, 101°19’E, Namtha Muang Sing, 5 31.v.1997, 900 1200 m, Vit Kubáň leg.”. VIETNAM: 1 ♀ (MNHN): “Tonkin, Reg. de Hoa Binh, A. de Cooman 1928”. 1 ♀ (MNHN): “Lactho, Tonkin de Cooman”; 2 ♂ (EJCB): “Vietnam, mount. SW, Kui Tchau, 200 m, 15.4.1963, Kabakov”; 7 (EJCB): “S Vietnam, 14.10N 108.30E, 40 km NW Ankhe, Buon Luoi, 620 750 m, 28.3. 12.4.1995, Pacholátko & Dembický leg.”.

Adult occurence: 3-4-5-6 (n=8). Altitude range: 200-1870 m (n=8). Host plants: Unknown. Distribution: CHINA: Yunnan. VIETNAM.

Agrilus cyanescens cyanescens (Ratzeburg) Fig. 19B (habitus); Fig. 36C (aedeagus); Figs. 42B, 42P, 44N, 46G, 47M, 51C, 53R (types) cyanescens Ratzeburg, 1837 (Buprestis) RATZEBURG, 1837: 54 55 [#2498] (description). KIESENWETTER, 1857: 140 [#3015] (synonym of caeruleus). MARSEUL, 1857: 92 [#4371] (Illiger is cited as the author; synonym of cyaneus (Olivier); checklist). MARSEUL, 1866: 452 [#676] (synonym of caeruleus). GEMMINGER & HAROLD, 1869: 1438 [#1057] (synonym of caeruleus; catalog). SAUNDERS, 1870: 10 [#2551] (catalog). SAUNDERS, 1871: 119 [#2552] (catalog). MARSEUL, 1887: 251 [#4707] (synonym of caeruleus). KERREMANS, 1892: 252 [#1397] (synonym of caeruleus; catalog). KERREMANS, 1903: 274 [#1441] (synonym of caeruleus; catalog). JAKOBSON, 1913: 796 [#4386] (synonym of caeruleus; catalog). BEDEL, 1921: 209 [#170] (synonym of caeruleus). ESCHERICH, 1923: 149 [#6717] (Laporte is cited as the author; synonym of caeruleus cited as coeruleus). OBENBERGER, 1926: 654 [#2057] (synonym of caeruleus). FISHER, 1928: 224 [#980] (synonym of caeruleus cited as coeruleus). OBENBERGER, 1936: 975 [#2191] (synonym of caeruleus cited as coeruleus; world catalog). SCHAEFER, 1949: 354, 384 385, 490 [#2565] (characters; references; faunal records; biology; notes; ditributional overview; notes; France). HORION, 1955: 80 81 [#1285] (distribution overview; faunal records; biology; Deutschland; Österreich). TER MINASYAN, 1955: 450 [#7893] (synonym of caeruleus cited as coeruleus). ZAGAIKEVICH, 1962: 102 [#3249] (synonym of caeruleus cited as coeruleus). RIKHTER & ALEXEEV, 1965: 301 [#2522] (synonym of caeruleus; cited as coeruleus). GURYEVA, 1974: 98 [#5428] (synonym of caeruleus cited as coeruleus). HELLRIGL, 1978: 262 263 [#3442] (distribution; commented host plant summary; ecology). KUBÁŇ, 1978: 23 [#3474] (biology; Moravia). LOMPE, 1979: 234 [#3628] (characters in key; Central Europe). BÍLÝ, 1982: 61, 67, 73 74 [#295] (characters in key; distribution overview; biology). COBOS, 1986: 239 240, 316 [#609] (subgenus Anambus; characters in key; distribution; synonymy; bionomy). BRIGHT, 1987: 223, 228, 280 282 [#379] (characters; hosts; distribution; notes; USA (Ontario, Massachusetts, Wisconsin, Michigan); Canada (Ontario)). ALEXEEV, 1989: 483 [#64] (cited as coerulescens Ratzeburg; key to

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA species; Primorsk; south Siberia; Transcaucasia; European USSR; Europe; North America). ALEXEEV, 1990: 8 [#65] (notes). WESTCOTT, 1991: 76 [#3085] (faunal record; Utah). LAUTERBACH, 1992: 241 245 [#4294] (biology; Germany). PENG ZHONGLIANG, 1992: 398 [#2431] (characters; notes; Hunan). NIEHUIS & TEZCAN, 1993: 21 22, 48 [#1910] (subgenus Anambus; faunal records from Turkey; Turkey). BÍLÝ, 1994: 111 [#8200] (larval characters in key). CURLETTI, 1994: 22, 163 164 [#658] (subgenus Agrilus; catalog; faunal records; Italia). NELSON et al., 1996: 184 [#3975] (notes; USA: Colorado, Virginia, West Virginia). DREES, 1999: 125 [#4821] (ecological notes; Germany). NIEHUIS, 1999: 38 [#4829] (key of races; discussion on species group). BÍLÝ, 2002: 16 [#5877] (bionomy; host plants; distribution). BRECHTEL & KOSTENBADER, 2002: 248 251 [#5766] (biology; ecology; distribution; Germany). HUA LI ZHONG, 2002: 89 [#6696] (cited as cyanescence; checklist; China: Jiangxi, Hunan). JENDEK, 2002: 2 5 [#5709] (nomenclatural and taxonomic notes). DEDYUKHIN, NIKITSKY & SEMENOV, 2005: 303 [#8860] (checklist; Russia: Udmurtia). MAIER, 2005: 949 951 [#7225] (notes; faunal records; distributional summary; USA (Connecticut, Pennsylvania, Meine)). JENDEK, 2006: 390 [#7441] (subgenus Dentagrilus; Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (complement to distribution in Palaearctic catalog). OLBERG, 2007: 115 116 [#8058] (faunal record; Norway). BELLAMY, 2008: 2052 2055 [#8576] (subgenus Dentagrilus; world catalog). LELITO et al., 2008: 671 672 [#8728] (effectivity of the visual cue based sticky traps). JENDEK & GREBENNIKOV, 2009: 242 [#8724] (notes). JENDEK & GREBENNIKOV, 2009: 45 52 [#8561] (characters in key; synonymy; faunal records; distributional and biological summary). FÄGERSTRÖM, BYGEBJERG & JONASSON, 2009: 137 140 [#8843] (faunal records; biology; notes; Sweden). VOLKOVITSH, 2009: 134 [#8850] (subgenus Dentagrilus; faunal records; Russia: Primorskiy kray). caeruleus (Rossi, 1792) (Buprestis; [preoccupied]) HELLWIG, 1795: 407 ROSSI, 1792: 62 [#3673] (description). [#3508] (Buprestis; identical redescription). HERBST, 1801: 251 [#1216] (Buprestis; cited as coerulea; [Note: Misidentification of amethystina now in Meliboeus according to SCHÖNHERR, 1817: 258 [#3895] or misidentification of cyanea Olivier according to Dejean, 1821: 30 [#4046]]). KIESENWETTER, 1857: 125, 140 141 [#3015] (cited as coeruleus; characters in key, redescription; notes; Germany). JACQUELIN DU VAL, 1860: 143 [#8086] (cited as coeruleus; catalog; Europe). MARSEUL, 1866: 437, 452 453 [#676] (cited as coeruleus; characters in key; redescription; references; distribution overview). GEMMINGER & HAROLD, 1869: 1438 [#1057] (cited as coeruleus; catalog). SAUNDERS, 1870: 6, 10 [#2551] (synonym of cyanescens). SAUNDERS, 1871: 119 [#2552] (synonym of cyaescens). HEYDEN, 1884: 290 [#8482] (cited as coeruleus; faunal record; Amur region). MARSEUL, 1887: 251 [#4707] (catalog). REITTER, 1891: 200 [#5278] (cited as coeruleus; catalog). KERREMANS, 1892: 252 [#1397] (catalog). HEYDEN, 1893: 90 [#3450] (cited as coeruleus; catalog). RUPERTSBERGER, 1894: 161, 287 [#4465] (cited as coeruleus; larval bibliography; catalog). ABEILLE DE PERRIN, 1897: 3, 9 [#19] (cited as coeruleus; characters in key; note). HEYDEN, 1897: 193 [#5286] (cited as coeruleus; faunal records; Germany). KULIKOWSKI, 1897: 150 [#8666] (cited as coeruleus; checklist; South Russia). KERREMANS, 1903: 274 [#1441] (catalog). REITTER, 1911: 194 [#2518] (cited as coeruleus; characters in key; Germany). CAILLOL, 1913: 511 512 [#4053] (cited as coeruleus; biology; faunal records; France). JAKOBSON, 1913: 796 [#4386] (cited as coeruleus; catalog; Russia and Europe). BEDEL, 1921: 201, 209 [#170] (cited as coeruleus; characters in key; references; faunal records; distribution; France). FROST, 1922: 96 [#4495] (introduction in America; USA (Wisconsin, Sturgeon Bay)). ESCHERICH, 1923: 145; 149 [#6717] (cited as coeruleus; [Note: Misidentification according to host plant cited];

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA characters in key; notes). CHAMBERLIN, 1926: 57 [#468] (cited as coeruleus; catalog; USA: Wisconsin). OBENBERGER, 1926: 654 [#2057] (cited as coeruleus; Palaearctic catalog). RAGUSA, 1926: 76 [#2496] (cited as coeruleus; notes; Sicilia). JAKOBSON, 1927: 179 [#8655] (cited as coeruleus; characters in key). LENG & MUTCHLER, 1927: 30 [#1703] (cited as coeruleus; supplement to catalog; USA: Wisconsin, Massachusetts). FISHER, 1928: 13, 224 226 [#980] (cited as coeruleus; characters; bibliography; notes; US: Massachusetts, Wisconsin). JAKOBSON & OGLOBLIN, 1931: 162 [#8314] (cited as coeruleus; characters in key). OBENBERGER, 1935: 48 49 [#2178] (cited as coeruleus; catalog; summary of distribution; Bulgaria). OBENBERGER, 1936: 972 976 [#2191] (cited as coeruleus; world catalog). RIKHTER, 1945: 166 [#3507] (cited as coeruleus; characters in key; European part of USSR except northernmost and south regions). THÉRY, 1945: 65 [#2884] (synonym of cyanescens). RIKHTER, 1948: 394 [#3879] (cited as coeruleus; characters in key). RIKHTER, 1950: 159 [#3881] (cited as coeruleus; characters in key). MÜHLMANN, 1954: 80 [#8152] (cited as coeruleus; note on biology). STARK, 1955: 775 [#3533] (cited as caeruleus; faunal record; USSR: Saratov). TER MINASYAN, 1955: 450 451 [#7893] (cited as coeruleus; host plants; distribution; references). ALEXEEV, 1957: 154 [#3226] (cited as coeruleus; characters in key). PLAVILSHCHIKOV, 1957: 211 [#8254] (characters; European part of Soviet Union). ALEXEEV, 1958: 185, 188 189 [#3982] (cited as coeruleus; checklist; biology; larva; galleries; Moskovsk). ALEXEEV, 1959: 7 [#3229] (cited as coeruleus; characters in key; European USSR except northernmost parts; Sibera; Primorsk; North China). PORTA, 1959: 197 [#2488] (cited as coeruleus; synonym of cyanescens; notes). ALEXEEV, 1960: 1508 [#45] (cited as coeruleus; larval characters in key). ZAGAIKEVICH, 1960: 257 [#5615] (cited as coeruleus; checklist; Ukraine). ALEXEEV, 1961: 6 [#3243] (cited as coeruleus; larval characters in key). ZAGAIKEVICH, 1962: 102 [#3249] (cited as coeruleus; faunal records; biology; Ukraine). ZAGAIKEVICH, 1962: 60 [#5616] (cited as coeruleus; checklist; Ukraine). RIKHTER & ALEXEEV, 1965: 301 [#2522] (cited as coeruleus; characters in key; European part of USSR). KOSTIN, 1973: 110 [#3693] (cited as coeruleus; characters in key; Kazakhstan). GURYEVA, 1974: 98 [#5428] (cited as coeruleus; pest of agriculture cultures; references; Soviet Union). WELLSO et al., 1976: 16 [#3079] (cited as coeruleus; characters; USA: Michigan). HELLRIGL, 1978: 262 [#3442] (synonym of cyanescens). LOMPE, 1979: 234 [#3628] (cited as caeruleus; synonym of cyanescens). KALASHIAN, 1985: 718 [#1366] (erroneously cited as coerulescens Ratzeburg; faunal records; Armenia). COBOS, 1986: 316 [#609] (synonym of cyanescens). CURLETTI, 1994: 163 [#658] (synonym of cyanescens). ALEXEEV, 1998: 377, 378 [#4322] (subgenus Dentagrilus; cited as coeruleus; genotype of the subgenus Dentagrilus). BÍLÝ, 2002: 16 [#5877] (synonym of cyanescens). BRECHTEL & KOSTENBADER, 2002: 248 [#5766] (synonym of cyanescens). JENDEK, 2002: 3 [#5709] (synonym of cyanescens; taxonomic notes). JENDEK, 2006: 390 [#7441] (synonym of cyanescens cyanescens; Palaearctic catalog). BELLAMY, 2008: 2053 [#8576] (synonym of cyanescens; world catalog). NIKITSKY et al., 2008: 147 [#8610] (synonym of cyanescens). JENDEK & GREBENNIKOV, 2009: 45 [#8561] (synonym of cyanescens; type not traced; notes). amabilis Gory & Laporte, 1839 (Agrilus) GORY & LAPORTE, 1839: 52 53 [#3738] (description). KIESENWETTER, 1849: 40 [#8334] (catalog; Europe). KIESENWETTER, 1857: 141, 142 [#3015] (synonym of caeruleus). MARSEUL, 1857: 92 [#4371] (checklist; Europe). JACQUELIN DU VAL, 1860: 143 [#8086] (synonym of caeruleus cited as coeruleus). MARSEUL, 1866: 452 [#676] (synonym of caeruleus). GEMMINGER & HAROLD, 1869: 1438 [#1057] (synonym of

77 caeruleus cited as coeruleus). SAUNDERS, 1871: 119 [#2552] (synonym of cyanescens). MARSEUL, 1887: 251 [#4707] (synonym of caeruleus). REITTER, 1891: 200 [#5278] (synonym of caeruleus cited as caeruleus). KERREMANS, 1892: 252 [#1397] (synonym of caeruleus; catalog). KERREMANS, 1903: 274 [#1441] (synonym of caeruleus). JAKOBSON, 1913: 796 [#4386] (synonym of caeruleus; catalog). FISHER, 1928: 224 [#980] (synonym of caeruleus cited as coeruleus). OBENBERGER, 1935: 48 [#2178] (aberration of caeruleus cited as coeruleus; catalog). OBENBERGER, 1936: 976 [#2191] (aberration of caeruleus cited as coeruleus; world catalog). THÉRY, 1942: 153 [#2873] (subgenus Agrilus; aberration of cyanescens). SCHAEFER, 1949: 384, 385, 490 [#2565] (aberration of cyanescens; characters). COBOS, 1986: 240, 316 [#609] (synonym of cyanescens). NELSON & BELLAMY, 1993: 304 [#3167] (authorship and publication date). CURLETTI, 1994: 163 [#658] (synonym of cyanescens). JENDEK, 1998: 317 [#4481] (synonym of cyanescens; lectotype designation). BÍLÝ, 2002: 16 [#5877] (synonym of cyanescens). BRECHTEL & KOSTENBADER, 2002: 248 [#5766] (synonym of cyanescens). JENDEK, 2002: 3 [#5709] (synonym of cyanescens; taxonomic notes). JENDEK, 2006: 390 [#7441] (synonym of cyanescens cyanescens; Palaearctic catalog). BELLAMY, 2008: 2054 [#8576] (synonym of cyanescens; world catalog). JENDEK & GREBENNIKOV, 2009: 45 [#8561] (synonym of cyanescens; notes). sulcaticeps Abeille de Perrin, 1869 (Agrilus) ABEILLE DE PERRIN, 1869: 79 80 [#4] (description). ABEILLE DE PERRIN, 1869: 42 [#4493] (note). SAUNDERS, 1871: 121 [#2552] (catalog). MARSEUL, 1872: 338 [#6051] (redescription). KERREMANS, 1885: 154 [#1381] (catalog). REITTER, 1891: 200 [#5278] (synonym of caeruleus cited as caeruleus). KERREMANS, 1892: 252 [#1397] (synonym of caeruleus; catalog). ABEILLE DE PERRIN, 1897: 9 [#19] (synonym of caeruleus). KERREMANS, 1903: 274 [#1441] (synonym of caeruleus). REITTER, 1911: 194 [#2518] (synonym of caeruleus cited as coeruleus). JAKOBSON, 1913: 796 [#4386] (synonym of caeruleus; catalog). BEDEL, 1921: 201 [#170] (variety of caeruleus cited as coeruleus). FISHER, 1928: 224 [#980] (synonym of caeruleus cited as coeruleus). OBENBERGER, 1935: 48 [#2178] (aberration of caeruleus cited as coeruleus; catalog). OBENBERGER, 1936: 976 [#2191] (aberration of caeruleus cited as coeruleus; world catalog). THÉRY, 1942: 153 [#2873] (subgenus Agrilus; aberration of cyanescens). SCHAEFER, 1949: 384, 385, 490 [#2565] (aberration of cyanescens; characters). COBOS, 1986: 240, 316 [#609] (synonym of cyanescens). CURLETTI, 1994: 163 [#658] (synonym of cyanescens). BÍLÝ, 2002: 16 [#5877] (synonym of cyanescens). BRECHTEL & KOSTENBADER, 2002: 248 [#5766] (synonym of cyanescens). JENDEK, 2002: 3 [#5709] (synonym of cyanescens; taxonomic notes). JENDEK, 2006: 390 [#7441] (synonym of cyanescens cyanescens; Palaearctic catalog). BELLAMY, 2008: 2054 [#8576] (synonym of cyanescens; world catalog). JENDEK & GREBENNIKOV, 2009: 45, 46 [#8561] (synonym of cyanescens; type not traced; notes). acuticornis Abeille de Perrin, 1897 (Agrilus) ABEILLE DE PERRIN, 1897: 3, 10 [#19] (description). KERREMANS, 1903: 281 [#1441] (catalog). JAKOBSON, 1913: 796 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 939 [#2191] (world catalog). NIEHUIS & TEZCAN, 1993: 1, 22 [#1910] (synonym of cyanescens; lectotype designation). JENDEK, 2002: 3, 4 [#5709] (synonym of cyanescens; taxonomic notes). JENDEK, 2006: 390 [#7441] (synonym of cyanescens cyanescens; Palaearctic catalog). BELLAMY, 2008: 2054 [#8576] (synonym of cyanescens; world catalog). JENDEK & GREBENNIKOV, 2009: 45, 46 [#8561] (synonym of cyanescens; notes on lectotype designation). fissifrons Abeille de Perrin, 1897 (Agrilus; [preoccupied])

78 ABEILLE DE PERRIN, 1897: 3, 9 10 [#19] (description). HEYDEN, 1897: 193 [#5286] ([Note: Cited as “n. sp.” with Abeille’s authorship and remark: Not in my possession]; characters). KERREMANS, 1903: 285 [#1441] (misspelled as fissiformis; catalog). JAKOBSON, 1913: 796 [#4386] (catalog; Russia and Europe). OBENBERGER, 1924: 50 [#2040] (synonym of kyselyi). OBENBERGER, 1936: 1004 [#2191] (synonym of kyselyi). BÍLÝ, 1982: 74 [#295] (synonym of cyanescens). CURLETTI, 1985: 237 [#649] (synonym of italicus). CURLETTI, 1994: 164 [#658] (synonym of italicus). BORDONI, 1995: 103 [#8274] (synonym of italicus). NIEHUIS, 1999: 39 [#4829] (synonym of italicus; discussion). JENDEK, 2006: 390 [#7441] (synonym of cyanescens cyanescens; Palaearctic catalog). BELLAMY, 2008: 2054 [#8576] (synonym of cyanescens; world catalog). JENDEK & GREBENNIKOV, 2009: 45, 47 [#8561] (synonym of cyanescens; type examined; notes). teriolensis Obenberger, 1916 (Agrilus; subspecies of artemisiae) OBENBERGER, OBENBERGER, 1916: 275 [#1983] (description). 1926: 659 [#2057] (variety of artemisiae; cited as tirolensis; Palaearctic catalog). OBENBERGER, 1936: 950 [#2191] (variety of artemisiae; world catalog). THÉRY, 1942: 147, 148 [#2873] (subgenus Agrilus; aberration of albogularis). SCHAEFER, 1949: 420, 422, 491 [#2565] (aberration of albogularis; characters). HORION, 1955: 104 [#1285] (aberration of albogularis; note). COBOS, 1986: 249, 317 [#609] (synonym of albogularis artemisiae). CURLETTI, 1994: 167 [#658] (synonym of cyanescens). JENDEK, 2002: 3, 4 [#5709] (synonym of cyanescens; taxonomic notes). JENDEK, 2006: 390 [#7441] (synonym of cyanescens cyanescens; Palaearctic catalog). BELLAMY, 2008: 2054 [#8576] (synonym of cyanescens; world catalog). JENDEK & GREBENNIKOV, 2009: 45, 47 [#8561] (synonym of cyanescens; type examined; notes). italicus Obenberger, 1920 (Agrilus) OBENBERGER, OBENBERGER, 1920: 212 [#2012] (description). 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1003 [#2191] (world catalog). GERINI, 1953: 90 91 [#1060] (note). PORTA, 1959: 197 [#2488] (notes). GOBBI, 1983: 40 [#1098] (note). CURLETTI, 1985: 237 [#649] (lectotype designation; synonymy). CURLETTI, 1994: 22, 164 [#658] (subgenus Agrilus; catalog; faunal records; Italia). BORDONI, 1995: 103 [#8274] (notes; Italy). NIEHUIS, 1999: 39, 40 [#4829] (discussion). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2144 [#8576] (world catalog). JENDEK & GREBENNIKOV, 2009: 45, 48 [#8561] (synonym of cyanescens; notes on lectotype designation). kyselyi Obenberger, 1924 (Agrilus; replacement name for fissifrons Abeille not Fairmaire) OBENBERGER, 1924: 50 [#2040] (replacement name proposal). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). ROUBAL, 1930: 115 [#5172] (notes). OBENBERGER, 1936: 1004 [#2191] (world catalog). RIKHTER & ALEXEEV, 1965: 301 [#2522] (cited as kysellii; synonym of caeruleus cited as coeruleus). BÍLÝ, 1982: 74 [#295] (synonym of cyanescens). CURLETTI, 1985: 237 [#649] (synonym of italicus). CURLETTI, 1994: 164 [#658] (synonym of italicus). BORDONI, 1995: 103 [#8274] (synonym of italicus). JENDEK, 2006: 390 [#7441] (synonym of cyanescens cyanescens; Palaearctic catalog). BELLAMY, 2008: 2054 [#8576] (synonym of cyanescens; world catalog). cockerelli Fisher, 1925 (Agrilus) FISHER, 1925: 4 6 [#975] (description). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 972 [#2191] (world catalog). KUROSAWA, 1974: 3 [#4575] (notes). BELLAMY, 1994: 360 [#237] (cited as cockerellei; holotype deposition in USNM). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2031 [#8356] (world catalog). JENDEK & GREBENNIKOV, 2009: 45, 48 [#8561] (synonym of cyanescens;

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA type examined). pooli Théry, 1936 (Agrilus) OBENBERGER, THÉRY, 1936: 120 121 [#2829] (description). 1936: 1022 [#2191] (world catalog). ALEXEEV, 1979: 134 [#57] (subgenus Dentagrilus; characters; notes; faunal records; USSR: south Khabarovsk, Primorsk; Korea). ALEXEEV, 1989: 483 [#64] (key to species; Primorsk; North Korea). ALEXEEV, 1998: 378 [#4322] (subgenus Dentagrilus; subgeneric classification). JENDEK, 2006: 390 [#7441] (subgenus Dentagrilus; Palaearctic catalog). BELLAMY, 2008: 2241 [#8576] (subgenus Dentagrilus; world catalog). JENDEK & GREBENNIKOV, 2009: 45, 50 [#8561] (synonym of cyanescens; type examined).

Type material: Buprestis cyanescens Ratzeburg, 1837 Type locality: not given [Wäldern Preussens are cited in the title of the publication]. Type specimens: Not located. Described from unknown number of syntypes. Buprestis caerulea Rossi, 1792 Type locality: silvae Pisanae. Type specimens: Not located. Described from 4 syntypes. Agrilus amabilis Gory & Laporte, 1839 Type locality: Saxe. Type specimens: Primary: Lectotype (Fig. 42P), ♂, designated by JENDEK (1998: 317) [#4481], (MNHN): “Transcriptio [p] var. Amabilis Gory Typ Saxe [h] \ Muséum Paris 1952 Coll. R. Oberthur [p] [yellow label]”. Secondary: 1 PLT (MNHN). Described from unknown number of syntypes. Agrilus sulcaticeps Abeille de Perrin, 1869 Type locality: Briançon (Hautes-Alpes). Type specimens: ST (MNHN). Described from 2 syntypes. Agrilus acuticornis Abeille de Perrin, 1897 Type locality: Caucase arménien. Type specimens: Primary: Lectotype (Fig. 42B), ♂, designated by NIEHUIS & TEZCAN (1993: 1, 22) [#1910], (MNHN): “Caucasus Armen. Geb..Leder Reitter [p] \ acuticornis Ab. [h] Type [h] [red ink] \ Muséum Paris Coll. Abeille De Perrin 1919 [p] \ Agrilus cyanescens Ratzb. det. Niehuis 1992 [p] \ Muséum Paris Coll. Générale [p] [yellow label]”. Described from unknown number of syntypes. Agrilus fissifrons Abeille de Perrin, 1897 Type locality: Karpathes. Type specimens: Holotype by monotypy (Fig. 46G), ♂, (MNHN): “Karpates ..ostenī-Molda.. [p] \ fissifrons Ab. [h] Type [h] [red ink] \ Muséum Paris 1919 Coll. A. DE PERRIN [p]”. Described from 1 specimen. Agrilus artemisiae teriolensis Obenberger, 1916 Type locality: Tirol. Type specimens: Holotype by monotypy (Fig. 53R), ♀, (NMPC): “Tirolis [p] \ Unicum - Typus! [h] \ Typus [p] [red label] \ Agrilus artemisiae v. teriolensis m. [p] Det. Obenberger [p] \ Mus. Nat. Pragae [p] 24 967 [h] [orange label]”. Described from 1 specimen. Agrilus italicus Obenberger, 1920 Type locality: Italie: Livorno. Type specimens: Primary: Lectotype (Fig. 47M), ♀, designated by CURLETTI (1985: 237) [#649], (NMPC): “Italia

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Livorno [h] \ Typus [p] [red label] \ A. italicus m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 26333 [h] [orange label]”. Described from unknown number of syntypes. Agrilus kyselyi Obenberger, 1924 Type locality and specimens: See: Agrilus fissifrons Abeille de Perrin, 1897. Agrilus cockerelli Fisher, 1925 Type locality: Vladivostok, Siberia. Type specimens: Holotype by monotypy (Fig. 44N), ♀, (USNM): “Vladivostok Siber [sic!] 1923 V. Prinada [p] \ TypeNo. [p] 2756 [h] U. S. N. M. [p] red label] \ Agrilus cockerelli Fisher [h] \ Holotype [p] [red ink] \ Agrilus cockerelli Fisher [h] [white label with red border]”. Described from 1 specimen. Agrilus pooli Théry, 1936 Type locality: Sibérie. Type specimens: Holotype by monotypy (Fig. 51C), ♀, (MNHN): “Siberia [p] \ Agrilus Pooli [h] Type [p] [red ink] \ Muséum Paris 1935 coll. A. Théry [p]”. Described from 1 specimen. Diagnosis: 5.2-7.8 mm; aedeagus as in Fig. 36C; for further characters see JENDEK & GREBENNIKOV, 2009 [#8563]. Species group: Cyanescens. Relevant species: A. thibetanus Obenberger, 1928; A. zhongdian Jendek, 2009. Specimens examined: CANADA: Ontario: 1 (CNC): “Vittoria Ont. 6/12/1956 W. J. Brown” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 (CNC): “Gore Bay, ONT 10.VII.1961 G. Brumpton” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 ♂ (EJCB): “Ontario London, Strathroy, vi.1995, D. Čatloš leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]); 2 ♂, 3 ♀ (EJCB): “Canada, Ontario, Ottawa, on Lonicera, 12.vi.2008, E. Jendek leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]). Quebec: 1 (CNC): “Lac. Taylor, Parc Gatineau, Que. June 22/74, humid soil, col. R. Sexton” (JENDEK & GREBENNIKOV, 2009 [#8561]). CHINA: Shaanxi: 2 ♀ (EJCB): “China Shaanxi, Čun Čan, 26.5. 1.6.2000, leg. E. Kučera” (JENDEK & GREBENNIKOV, 2009 [#8561]). Shanxi: 3 ♀ (EJCB): “China, Shanxi prov, 9.VI.2000, Lüliang Shan, 1000 m, road Fangshan Jiaocheng, Hengjian env., Zd. Jindra” (JENDEK & GREBENNIKOV, 2009 [#8561]). CZECH REPUBLIC: Stredocesky: 1 (EJCB): “Bohemia centr. Krč hospital, 17.vi.1986 lgt. M. Škorpík” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 (EJCB): “Bohemia centr. Prag D. Šárka 18.i.1995, lgt. P. Kabátek ex larvae” (JENDEK & GREBENNIKOV, 2009 [#8561]). FRANCE: Ile-del-France: 3 ♂, 1 ♀ (EJCB): “France Montmorency 29.v.1959” (JENDEK & GREBENNIKOV, 2009 [#8561]). Rhone-Alpes: 8 (EJCB): “Charbonières (Rh) L. Schaefer, Lonicera xylosteum” (JENDEK & GREBENNIKOV, 2009 [#8561]); 5 (EJCB): “Chalamont (Ain) 19.6.1942 L. Schaefer, Lonicera xylosteum” (JENDEK & GREBENNIKOV, 2009 [#8561]). GERMANY: SachsenAnhalt: 1 ♂ (EJCB): “Dessau, Anh. 24.6.1955 E. Heidenreich” (JENDEK & GREBENNIKOV, 2009 [#8561]). ITALY: Toscana: 1 ♂, 1 ♀ (EJCB): “Tosc. Livorno Antignano 26.V.949 F. Gerini” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 ♂ (EJCB): “Tosc. Livorno Antignano 9.V.953 F. Gerini” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 ♂, 1 ♀ (EJCB): “Tosc. Livorno Antignano 13.V.953 F. Gerini” (JENDEK & GREBENNIKOV, 2009 [#8561]). KAZAKHSTAN: 1 ♀ (EJCB): “r. Tainty, 15 km VSV Targyna, Altai, V Kazakh. obl. 1 2.vii.1983 [in Russian]” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 ♂ (EJCB): “Kazakhstan VII.1983 Tarbagatai Mts.” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 ♀ (EJCB): “Kazakhstan NE, 100 km SEE Ust’ Kamenogorsk, 5 km NE Panteleymonovka, Kaiyndy

79 forest, 16 20.6.93, Napolov leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]). KOREA SOUTH: 1 (JLCS): “Mt. Gajisan, Icheon ri, Sangbuk myeon, Ulju gun, Gyeongsangnam do, Korea, 28 30. vi. 1.vii.1990, coll. Myeong ja Kang” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 (JLCS): “Mt. Sangseongsan, Yongtae ri, Hail myeon, Goseong gun, Korea, 5.v.2000, coll. Yeong ju Gu” (JENDEK & GREBENNIKOV, 2009 [#8561]). NETHERLANDS: Gelderland: 1 ♂ (EJCB): “Leuvenum Juli 1926 H. Klaasen” (JENDEK & GREBENNIKOV, 2009 [#8561]). POLAND: 1 (EJCB): “Sebrna Góra 1.vii.1948” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 (EJCB): “LZD Krynica 18.vi.1988 L.ctno Kopciona leg. A. Zabjdzlni” (JENDEK & GREBENNIKOV, 2009 [#8561]). RUSSIA: Ivanovskaya oblast’: 1 ♂ (EJCB): “C Russia, Ivanovo env., 56°59’N, 41°01’E, 2.07.2007, leg. Smirnov”. Kaluzhskaya oblast’: 2 ♂ (EJCB): “C Russia, Kaluzhskaya ar. Malojaroslavetsky dist., Afanasovo vill., 11 19.06.2005, leg. Smirnov”. Primorskiy kray: 1 ♂, 2 ♀ (EJCB): “Russia, Primorsk, Krasnoarm. r n, 14 vi 1951g, V. I. Stepanov, na khimolosti maaka” (JENDEK & GREBENNIKOV, 2009 [#8561]); 3 ♂ (EJCB): “Russia, Primorsk, Krasnoarmeyskiy rayon, 9.vii.1971, Mt. Armu, on Lonicera” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 ♀ (EJCB): “Sib. or. m., Primorje, Sichote Alin Mts., Sokolči, 1 15.7.1990, Kadlec + Voříšek lg.” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 ♂ (EJCB): “Russia, Primorsk. reg., 18 21.7.1992, Rezanovka, E. Kučera leg., ex Lonicera” (JENDEK & GREBENNIKOV, 2009 [#8561]). Sverdlovskaya oblast’: 4 ♂, 1 ♀ (EJCB): “Sverdlovsk obl., st. Khrustal’naya, Sverdl. zh.d. 19 vi 1953 iz khimolosti” (JENDEK & GREBENNIKOV, 2009 [#8561]). Tyva Respublika: 2 ♂ (EJCB): “Tuvinskaja ASSR, Chadyn 29 30.94 Z. Kletečka lgt.” (JENDEK & GREBENNIKOV, 2009 [#8561]). SLOVAKIA: 5 (EJCB): “Slovakia, 29 31.v.1979 Devínska Kobyla, M. Štrba leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 ♂, 1 ♀ (EJCB): “Slovakia, 7.vii.1983 Bystrička na Orave, M. Štrba leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]); 4 (EJCB): “Slovakia occ., D. Vestenice, DFS 7276c, 21.V. 1.VII.2001, Malaise trap, O. Majzlan leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 ♂ (EJCB): “Slovakia Napant, Veľký Bok 23.6.2005, P. Szomody” (JENDEK & GREBENNIKOV, 2009 [#8561]). TURKEY: 1 ♂, 1 ♀ (EJCB): “NE Türkey, Kösedagi Gecidi pass W Bayburt, N40°18 E39°34, 1973 m, 26.VI.2006, lgt. P. Kabátek” (JENDEK & GREBENNIKOV, 2009 [#8561]). Erzincan: 1 ♂ (EJCB): “Anatolien, Prov. Erzincan, 40 km östlich Imranli, Gemecik, 3.6.1975, 1630 m, leg. Holzschuh & Ressl” (JENDEK & GREBENNIKOV, 2009 [#8561]). Ordu: 1 ♂, 1 ♀ (EJCB): “Prov. Ordu 28.5., Melet Fluß n. Mesudiye 800 m, Turkei 1985, leg. Schönmann et Schillhalmmer” (JENDEK & GREBENNIKOV, 2009 [#8561]). UNITED STATES: Massachusetts: 3 (CNC): “Nantick Mass. VI 18 1948 C. A. Frost” (JENDEK & GREBENNIKOV, 2009 [#8561]). Michigan: 1 (CNC): “Midland Co., Michigan, 6 17 45, R. R. Dreisbach” (JENDEK & GREBENNIKOV, 2009 [#8561]); 2 ♀ (EJCB): “Michigan, East Lansing, Inghan Co. VI 4, 1971, J. A. Jackman” (JENDEK & GREBENNIKOV, 2009 [#8561]). New York: 1 (EJCB): “USA 4/673 Commack Fairfield” (JENDEK & GREBENNIKOV, 2009 [#8561]). Ohio: 4 (CNC): “Delaware Co. V 21 o. D.J. & J.J. Knull Collrs.” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 (CNC): “Franklin Co. VI 49 O., E. L. Sleeper Collr.” (JENDEK & GREBENNIKOV, 2009 [#8561]).

Adult occurence: 5-6-7 (n=41). Altitude range: 800-1973 m (n=4). Host plants: Lonicera: HEYDEN, 1863 (as caeruleus) [#8478]; EVERTS, 1903 (as caeruleus) [#6677]; RIKHTER, 1950 (as caeruleus) [#3881]; TER MINASYAN, 1955 (as caeruleus) [#7893]; ALEXEEV, 1958 (as caeruleus) [#3982]; ALEXEEV, 1959 (as caeruleus) [#3229]; ALEXEEV, 1961 (as caeruleus) [#3243]; RIKHTER & ALEXEEV, 1965 (as caeruleus) [#2522]; FRANZ, 1974 [#4334]; GURYEVA, 1974 (as caeruleus) [#5428];

80 HELLRIGL, 1978 [#3442]; KUBÁŇ, 1978 [#3474]; ALEXEEV, 1979 (as pooli) [#57]; NIEHUIS, 1988 [#1912]; ALEXEEV, 1989 [#64]; CURLETTI, 1994 [#658]; BÍLÝ, 2002 [#5877]; HASTIR & GASPAR, 2002 [#6211]; JENDEK & GREBENNIKOV, 2009 [#8561]. − L. alpigena: BRECHTEL & KOSTENBADER, 2002 [#5766]. − L. implexa: BEDEL, 1921 (as caeruleus) [#170]; SCHAEFER, 1949 [#2565]. − L. involucrata: DREES, 1999 [#4821]. − L. morrowii: MAIER, 2005 [#7225]. − L. nigra: ROGER, 1856 (as caeruleus) [#7468]; HEYDEN, 1897 (as caeruleus) [#5286]; SCHAEFER, 1949 [#2565]; BRECHTEL & KOSTENBADER, 2002 [#5766]. − L. periclym; DAEHNE, 1912 (as caeruleus) [#3609]. − L. periclymenum: BRECHTEL & KOSTENBADER, 2002 [#5766]. − L. tatarica: BEDEL, 1921 (as caeruleus) [#170]; SCHAEFER, 1949 [#2565]; WELLSO et al., 1976 (as caeruleus) [#3079]; BRIGHT, 1987 [#379]; DREES, 1999 [#4821]. − L. xylosteum: DAEHNE, 1912 (as caeruleus) [#3609]; SCHAEFER, 1949 [#2565]; BRECHTEL & KOSTENBADER, 2002 [#5766]. Rhamnus: HELLRIGL, 1978 [#3442]; NIEHUIS & TEZCAN, 1993 [#1910]; CURLETTI, 1994 [#658]; BÍLÝ, 2002 [#5877]; JENDEK & GREBENNIKOV, 2009 [#8561]. − R. cathartica: GOBBI, 1973 [#1093]. Symphoricarpos: LAUTERBACH, 1992 [#4294]; NELSON et al., 1996 [#3975]; JENDEK & GREBENNIKOV, 2009 [#8561]. − S. orbiculatus: WESTCOTT, 1991 [#3085]. Erroneous: Acer: CURLETTI, 1994 [#658]. − Alnus: ESCHERICH, 1923 (as caeruleus) [#6717]; HORION, 1955 [#1285]; TER MINASYAN, 1955 (as caeruleus) [#7893]; ALEXEEV, 1989 [#64]; CURLETTI, 1994 [#658]. − Betula: ESCHERICH, 1923 (as caeruleus) [#6717]; HORION, 1955 [#1285]; TER MINASYAN, 1955 (as caeruleus) [#7893]; ALEXEEV, 1989 [#64]; CURLETTI, 1994 [#658]. − Castanea: CURLETTI, 1994 [#658]. − Euonymus: HELLRIGL, 1978 (as dubious) [#3442]. −Fagus: ESCHERICH, 1923 (as caeruleus) [#6717]; HORION, 1955 [#1285]; TER MINASYAN, 1955 (as caeruleus) [#7893]; ALEXEEV, 1989 [#64]; CURLETTI, 1994 [#658]. − Fraxinus: SCHAEFER, 1949 [#2565]; HORION, 1955 [#1285]; ALEXEEV, 1989 [#64]; CURLETTI, 1994 [#658]. − Populus: CURLETTI, 1994 [#658]. − Pyrus pyraster: CROVATO & IZZILLO, 1995 (as italicus) [#3791]. − Quercus: RATZEBURG, 1837 [#2498]; ESCHERICH, 1923 (as caeruleus) [#6717]; SAINTE CLAIRE, 1935 (as caeruleus) [#8551]; SCHAEFER, 1949 [#2565]; HORION, 1955 [#1285]; TER MINASYAN, 1955 (as caeruleus) [#7893]; GOBBI, 1973 [#1093]; HELLRIGL, 1978 (as dubious) [#3442]; ALEXEEV, 1989 [#64]; CURLETTI, 1994 [#658]; HASTIR & GASPAR, 2002 [#6211]. − Rosa: HORION, 1955 [#1285]; TER MINASYAN, 1955 (as caeruleus) [#7893]; ALEXEEV, 1989 [#64]. − Salix: DAEHNE, 1912 (as caeruleus) [#3609]; CURLETTI, 1994 [#658]. − Tilia: CURLETTI, 1994 [#658]. − Ulmus: CURLETTI, 1994 [#658]. Distribution: Local: CHINA: Shaanxi, Shan-xi. KOREA NORTH. KOREA SOUTH. RUS-SIA: Primorskiy kray, Tyva Respublika. Global: ASIA: ARMENIA. CHINA. KAZAKHSTAN. KOREA NORTH. KOREA SOUTH. RUSSIA. TURKEY. EUROPE: ALBANIA. AUSTRIA. BELARUS. BELGIUM. BOSNIA & HERZEGOVINA. BULGARIA. CROATIA. CZECH REPUBLIC. DENMARK. ESTONIA. FRANCE. GERMANY. GREECE. HUNGARY. ITALY. LATVIA. LIECHTENSTEIN. LITHUANIA. LUXEMBOURG. MACEDONIA. MOLDOVA. NETHERLANDS. NORWAY.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA POLAND. PORTUGAL. ROMANIA. RUSSIA. SERBIA & MONTENEGRO. SLOVAKIA. SLOVENIA. SPAIN. SWEDEN. SWITZERLAND. TURKEY. UKRAINE. UNITED KINGDOM. NORTH AMERICA (introduced): CANADA. UNITED STATES. Remarks: Distributional records cited without supporting data from Hunan and Jiangxi (China) by PENG ZHONGLIANG, 1992 [#2431] and HUA LI ZHONG, 2002 [#6696] were omitted.

Agrilus cyanipennis Gory & Laporte Fig. 10G (habitus); Fig. 32V (aedeagus); Figs. 42F, 42Q, 45B, 45D, 50L (types) cyanipennis Gory & Laporte, 1839 (Agrilus) GORY & LAPORTE, 1839: 18 [#3738] (description). GEMMINGER & HAROLD, 1869: 1443 [#1057] (synonym of orientalis). HAROLD, 1869: 124 [#3943] (synonym of orientalis). SAUNDERS, 1871: 122 [#2552] (synonym of insignis). KERREMANS, 1892: 266 [#1397] (synonym of orientalis; catalog). KERREMANS, 1903: 288 [#1441] (synonym of orientalis; catalog). OBENBERGER, 1936: 1096 [#2191] (synonym of orientalis; world catalog). NELSON & BELLAMY, 1993: 303 [#3167] (authorship and publication date). JENDEK, 1998: 320 [#4481] (synonym of orientalis; lectotype designation). JENDEK, 2001: 172 [#5035] (valid name; synonymy; notes). JENDEK, 2005: 8 9 [#7217] (synonymy). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). ornativentris Saunders, 1866 (Agrilus) GEMMINGER & SAUNDERS, 1866: 315 [#2542] (description). HAROLD, 1869: 1443 [#1057] (catalog). SAUNDERS, 1871: 122 [#2552] (catalog). KERREMANS, 1892: 266 [#1397] (catalog). KERREMANS, 1903: 288 [#1441] (catalog). OBENBERGER, 1936: 1096 [#2191] (world catalog). BAUDON, 1963: 52 [#148] (faunal record; Laos). DESCARPENTRIES & VILLIERS, 1963: 52 [#699] (lectotype designation: type: “Laos (British museum)”; characters; faunal record; Laos; Annam). BAUDON, 1968: 72, 74 [#162] (note). JENDEK, 2005: 8 9 [#7217] (synonym of cyanipennis; notes). JENDEK, 2006: 397 [#7441] (synonym of cyanipennis; Palaearctic catalog). BELLAMY, 2008: 2055 [#8576] (synonym of cyanipennis; world catalog). orientalis Harold, 1869 (Agrilus; unnecessary replacement name for cyanipennis Gory & Laporte, 1839) HAROLD, 1869: 124 [#3943] (replacement name proposal). GEMMINGER & HAROLD, 1869: 1443 [#1057] (catalog). KERREMANS, 1892: 266 [#1397] (catalog). KERREMANS, 1903: 288 [#1441] (catalog). OBENBERGER, 1936: 1096 [#2191] (world catalog). JENDEK, 2001: 172 [#5035] (synonym of cyanipennis). JENDEK, 2006: 397 [#7441] (synonym of cyanipennis; Palaearctic catalog). BELLAMY, 2008: 2055 [#8576] (synonym of cyanipennis; world catalog). insignis Saunders, 1871 (Agrilus; unnecessary replacement name for cyanipennis Gory & Laporte, 1839) SAUNDERS, 1871: 122 [#2552] (replacement name proposal). KERREMANS, 1892: 266 [#1397] (synonym of orientalis; catalog). KERREMANS, 1903: 288 [#1441] (synonym of orientalis). OBENBERGER, 1936: 1096 [#2191] (synonym of orientalis). JENDEK, 2001: 172 [#5035] (synonym of cyanipennis). JENDEK, 2006: 397 [#7441] (synonym of cyanipennis; Palaearctic catalog). BELLAMY, 2008: 2055 [#8576] (synonym of cyanipennis; world catalog). amabilis Kerremans, 1893 (Agrilus; [preoccupied]) KERREMANS, 1893: 343 [#1406] (description). KERREMANS, 1903:

81

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA 277 [#1441] (catalog). OBENBERGER, 1936: 91 [#2204] (synonym of amabilicolor). OBENBERGER, 1936: 1072 [#2191] (synonym of amabilicolor). JENDEK, 2005: 8 9 [#7217] (synonym of cyanipennis; notes). JENDEK, 2006: 397 [#7441] (synonym of cyanipennis; Palaearctic catalog). BELLAMY, 2008: 2055 [#8576] (synonym of cyanipennis; world catalog). agilis Kerremans, 1894 (Agrilus) KERREMANS, 1894: 471 472 [#1412] (description). KERREMANS, 1903: 277 [#1441] (catalog). OBENBERGER, 1936: 1071 [#2191] (world catalog). JENDEK, 2005: 8 9 [#7217] (synonym of cyanipennis; lectotype designation; notes). JENDEK, 2006: 397 [#7441] (synonym of cyanipennis; Palaearctic catalog). BELLAMY, 2008: 2055 [#8576] (synonym of cyanipennis; world catalog). dalbergiae Théry, 1930 (Agrilus) OBENBERGER, THÉRY, 1930: 158 160 [#2767] (description). 1936: 1080 [#2191] (world catalog). JENDEK, 2001: 172 [#5035] (synonym of cyanipennis; lectotype designation). JENDEK, 2006: 397 [#7441] (synonym of cyanipennis; Palaearctic catalog). BELLAMY, 2008: 2055 [#8576] (synonym of cyanipennis; world catalog). amabilicolor Obenberger, 1936 (Agrilus; replacement name for amabilis Kerremans not Gory & Laporte) OBENBERGER, 1936: 91 [#2204] (replacement name proposal). OBENBERGER, 1936: 1072 [#2191] (world catalog). JENDEK, 2005: 8 9 [#7217] (synonym of cyanipennis; notes). JENDEK, 2006: 397 [#7441] (synonym of cyanipennis; Palaearctic catalog). BELLAMY, 2008: 2055 [#8576] (synonym of cyanipennis; world catalog).

Type material: Agrilus cyanipennis Gory & Laporte, 1839 Type locality: Indes Orientales. Type specimens: Primary: Lectotype (Fig. 45B), sex not examined, designated by JENDEK, (1998: 320) [#4481], (MNHN): “Cyanipennis Gory type Inde [h] \ Kerremans Vidit 1892 [p] \ Ex-Musaeo Mniszech [p] \ Muséum Paris 1952 Coll. R. Oberthur [p] [yellow label]”. Described from unknown number of syntypes. Agrilus ornativentris Saunders, 1866 Type locality: Laas. Type specimens: Primary: Lectotype (Fig. 50L), ♀, designated by DESCARPENTRIES & VILLIERS (1963: 52) [#699], (BMNH): “Laas [h] Mouhot [p] \ Type [p] \ Saunders 74.18 [p] \ ornativentris in BM [h] C. M. F. von Hayek det. [scratched] 196 [p] 2 [h]”. Described from unknown number of syntypes. Agrilus orientalis Harold, 1869 Type locality and specimens: See: Agrilus cyanipennis Gory & Laporte, 1839. Agrilus insignis Saunders, 1871 Type locality and specimens: See: Agrilus cyanipennis Gory & Laporte, 1839. Agrilus amabilis Kerremans, 1893 Type locality: Kanara. Type specimens: Holotype by monotypy (Fig. 42Q), ♂, (BMNH): “Type [p] [round label with red border] \ amabilis Kerr. [h] \ Kanara Andrewes [h] \ A. amabilis Kerrem. Inde [h] \ Kerremans 1903-59 [p]”. Described from 1 specimen. Agrilus agilis Kerremans, 1894 Type locality: Barway.

Type specimens: Primary: Lectotype (Fig. 42F), sex not examined, designated by JENDEK (2005: 8-9) [#7217], (BMNH): “Type [p] [round label with red border] \ Barway Cardon [h] \ agilis Kerr. Type [h] \ A. agilis Kerrem. Inde [h] \ Kerremans 1903-59 [p]”. Described from unknown number of syntypes. Agrilus dalbergiae Théry, 1930 Type locality: Cantt. Road, Dehra-Dun. Type specimens: Primary: Lectotype (Fig. 45D), ♀, designated by JENDEK (2001: 172) [#5035], (MNHN): “Cantt. Road Dehra Dun, U. P., C. F. C. Beeson 1. VIII. 192 [p] 6 [h] \ R.R.D. 495 B.C.R. 163 Cage 290 [p] \ ex Dalberia Sissoo [p] \ 64 [h] \ Dalbergiae [female sign] [Théry’s MS] Type [p] [red ink] \ coll. Théry [p]”. Described from 4 syntypes. Agrilus amabilicolor Obenberger, 1936 Type locality and specimens: See: Agrilus amabilis Kerremans, 1893. Diagnosis: 10-11.9 mm; pronotal anterior lobe absent; elytra with obvious perisutural impression; metatarsus in male longer than metafemur. Species group: Spinipennis. Relevant species: A. irrorellus Harold, 1869. Specimens examined: CHINA: Yunnan: 1 (NMPC): “Zap. Yun’nan’, okr. Zhuili, 1450 m, 7.VI.1956 [in Russian]”. INDIA: Tamil Nadu: 1 ♂ (EJCB): “Nilgiri Hills, South India, Singara 3400, V.1948”; 9 (EJCB): “India S, Tamil Nadu, Nilgiris, 15 km SE of Kotagiri, Kunchappanai, 900 m, 11.22 N 76.56 E, 7 22.v.2000, leg. P. Pacholátko”. NEPAL: 1 (EJCB): “E Nepal, Dhankuta, Arun Valley, Lamobagar Gola, 27.5. 3.6.1980, 1000 1400 m, leg. C. Holzschuh”; 1 ♀ (EJCB): “W Nepal, Burigandari, Machna Khola, 2915, Kholabenesi, 416.1990, J. Probst leg., 1630 m”. THAILAND: Mae Hong Son: 1 ♂ (EJCB): “Thai, Mae Hong Son pr., 19°27’N, 98°20’E, 1500 m, Soppong 7 12.V., Vít Kubáň leg., 1996”.

Adult occurence: 5-6 (n=6). Faunal record published by BAUDON, 1963 [#148] (as ornativentris) is also included. Altitude range: 900-1630 m (n=5). Host plants: Dalbergia sissoo: THÉRY, 1930 [#2767] (as dalbergiae). Distribution: CHINA: Yunnan. INDIA. LAOS. NEPAL. THAILAND. VIETNAM.

Agrilus daimio Obenberger Fig. 11K (habitus); Fig. 33J (aedeagus); Fig. 45C (type) daimio Obenberger, 1936 (Agrilus) OBENBERGER, 1936: 140 141 [#2207] (description). MIWA & CHÛJÔ, 1936: ADDENDA P. 4 [#1809] (catalog; Japan). OBENBERGER, 1936: 986 [#2191] (world catalog). KUROSAWA, 1963: 153 [#1618] (characters; Japan). KUROSAWA, 1974: 2 [#4577] (note). KUROSAWA, 1974: 1 2 [#4576] (note). TÔYAMA, 1985: 22 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). JENDEK, 1995: 148 [#3768] (lectotype designation; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 32 [#4091] (checklist; Japan: Honshu; Kyushu, Tsushima, Yakushima). FUKUTOMI & HORI, 2004: 10 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). OHMOMO, 2006: 242 245 [#7546] (redescription; faunal records; Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima, Yakushima)). ANONYMOUS, 2007:

82 32 [#8250] (checklist; Japan: Honshu). [#8576] (world catalog).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA BELLAMY, 2008: 2057

Type material: Agrilus daimio Obenberger, 1936 Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 45C), ♀, designated by JENDEK (1995: 148) [#3768], (NMPC): “Japan [h] \ Typus [p] [red label] \ Agrilus daimio m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24890 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 4.2-5.8 mm. Pronotum with distinct medial impression interrupted in middle and deep lateral impressions; elytra with proximal and distal pubescence. Species group: Maculifer. Relevant species: A. maculifer Saunders, 1873. Specimens examined: JAPAN: Honshu: 2 ♂ (EJCB): “1979.7.7., Sadanami, Sekimoto, (Mts. Hanazono), Kita ibaraki, Ibaraki, S. Ohmomo leg.”; 1 ♂ (EJCB): “(Mt. Hanazono), Ibaraki Pref., Japan, 23.vii.1979, A. Niskiyama leg.”; 1 (EJCB): “1982.7.4., Hikawa rindo, Daibosatsu, Yamanashi, S. Ohmomo leg.”; 2 (EJCB): “Japan, Ibaraki prefecture, Mts. Abukuma, Takahashi city, 4.VII.1987” (JENDEK, 1995 [#3768]); 1 ♂ (EJCB): “NGRI, Senbon matsu, Nishi nasuno, Tochigi Pref., 1991.6.16, Nishinasuno machi, Tochigi Pref., S. Ohmomo leg.”.

Adult occurence: 6-7 (n=5).

[#8561]). Sichuan: 1 ♀ PT (EJCB): “China, Sichuan pr., Liziping 28.vi. 3.vii.1991, R. Dunda leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 ♀ PT (EJCB): “Sichuan, VII.1992, Gonggashan Mts., Hailougou vall., Sauer leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 ♀ PT (EJCB): “China, W Sichuan, Ganzi Tibetian Auton. Pref., Daxue Shan, river valley, 15 km S Kangding, 2800 m, 29.56 N, 101.58E, 26.6.1999, leg. A. Pütz” (JENDEK & GREBENNIKOV, 2009 [#8561]). Yunnan: 1 ♂ PT (EJCB): “China N Yunnan Yulongshan Mts. 2500 2800 m Ganhaizi / Lijiang road lgt. D. Král 24 26/7’90” (JENDEK & GREBENNIKOV, 2009 [#8561]); 11 ♂ PT, 7 ♀ PT (EJCB): “China CW, Yunnan prov., Dali, 19 21.v.1993 R. Červenka leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]); 4 ♂ PT, 2 ♀ PT (EJCB): “Yunnan, 2600 3100 m, 25.38N 100.09E, Cangshan Mts., 5 6/6, Vít Kubáň leg. 1993” (JENDEK & GREBENNIKOV, 2009 [#8561]); 2 ♂ PT (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]); 2 ♀ PT (EJCB): “China, Yunnan Dali 1 7.vi.1994 leg. B.Šiška & T. Spevár” (JENDEK & GREBENNIKOV, 2009 [#8561]); 2 ♂ PT (EJCB): “China Yunnan, 28.5. 9.6.1994, Dali, lgt. E. Kučera” (JENDEK & GREBENNIKOV, 2009 [#8561]); 1 ♂ PT (EJCB): “China Yunnan, Ningjing Shan, Wei Shi, 2200 m, 12 13.6.2006, Vladimír Major leg.” (JENDEK & GREBENNIKOV, 2009 [#8561]); 3 ♂, 2 ♀ (EJCB): “China Yunnnan N, Ningjin Shan, 2100 m, Mekong Riv., Tse Kou, N28°01’59”, E98°54’17”, 8 11.6.2007, Major leg.”.

Adult occurence: 5-6-7 (n=11). Altitude range: 2100-3100 m (n=5).

Host plants: Carpinus laxiflora: OHMOMO, 2006 [#7546]. − Euonymus oxyphyllus: OHMOMO, 2006 [#7546]. − Parabenzoin praecox: AKIYAMA & OHMOMO, 1997 [#4091]; OHMOMO, 2006 [#7546]. − Symplocos argutidens: OHMOMO, 2006 [#7546].

Host plants: Unknown.

Distribution: JAPAN: Hokkaido, Honshu, Kyushu, Shikoku, Tsushima. KOREA SOUTH.

Fig. 21H (habitus of HT); Fig. 37E (aedeagus)

Agrilus dali Jendek Fig. 19G (habitus of HT); Fig. 36H (aedeagus)

dali Jendek, 2009 (Agrilus) JENDEK in JENDEK & GREBENNIKOV, 2009: 45, 54 55 [#8561] (description).

Type material: Agrilus dali Jendek, 2009 Type locality: China CW, Yunnan province, Dali (25°41’N, 100°09’E). Type specimens: Primary: Holotype (Fig. 19G), ♂, (EJCB): “China CW, Yunnan prov., Dali, 19–21.v.1993 R. Červenka leg.”. Secondary: 1 ♀ PT (CBCS); 1 ♂ PT (SOCI); 1 PT (MNHN); 33 PT (EJCB). Described from 37 specimens (HT, PT). Diagnosis: 5.1-7.9 mm; pronotum markedly convex with obviously arcuate sides and deep antero-lateral semifoveolate depression; median lobe of aedeagus as in Fig. 36H. Species group: Cyanescens. Relevant species: A. cyanescens (Ratzeburg, 1837). Specimens examined: CHINA: 1 ♂ (MNHN): “Siao Lou Lou Chan, Chasseurs Thibétains 1896” (JENDEK & GREBENNIKOV, 2009

Distribution: CHINA: Sichuan, Yunnan.

Agrilus daur Jendek, sp. nov. Description: 5.5-5.9 mm (Holotype 5.5 mm). BODY: Shape: subparallel, Build: robust, Color (dorsally): unicolored, Color (dominant): green. HEAD: Medial impression: present, Impression (extent): vertex, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): irregular. Eyes: Size: small, Shape: not protruding from head outline or protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short. PRONOTUM: Shape: visually square or transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: absent or vague, Shape: arcuate or subangulate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: acute, rectangular or obtuse, Apex: sharp. Disk: Convexity: flat, Disk impressions: medial and lateral or lateral only, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow. Prehumerus: absent or obsolete. Marginal and submarginal carinae: Interspace: wide, Convergence: subparallel, Junction: absent. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate,

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Width: wide, Shape: arcuate. Elytral pubescence: entire, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): moderate. Prosternal process: moderate, Shape: subparallel, Sides: straight or arcuate, Angles: rectangular or obtuse, Angles (tips): sharp or blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): shorter or subequal to 2-3 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): markedly dorsoventrally bent. Diagnosis: antennae in male very short, reaching only to anterior pronotal angles; tarsi shorter; aedeagus more robust. Species group: Albogularis. Relevant species: A. ecarinatus Marseul, 1866. Type locality: China, Nei Mongol province, Bayan Gol. Type specimens: Primary: Holotype (Fig. 21H), ♂, (EJCB): “China-Nei Mongol, Bayan Gol, 14.5.-20.5.1996, E. Kučera leg.”. Secondary: CHINA: Nei Mongol: 2 ♂ PT (EJCB): “China-Nei Mongol, Bayan Gol, 14.5.-20.5.1996, E. Kučera leg.”. Adult occurence: 5 (n=1). Host plants: Unknown. Distribution: CHINA: Nei Mongol. Etymology: Noun in apposition; the specific name refers to the ethnic group Daur inhabiting the Inner Mongolia, Heilongjiang and Xinjiang provinces of China.

Agrilus decoloratus Kerremans Figs. 9A, 9B (habitus); Fig. 32I (aedeagus); Figs. 42E, 42H, 42J, 45E, 48R, 50F, 51P, (types) decoloratus Kerremans, 1892 (Agrilus) KERREMANS, 1892: 213 214 [#1402] (description). KERREMANS, 1903: 284 [#1441] (catalog). OBENBERGER, 1936: 1080 [#2191] (world catalog). DESCARPENTRIES & VILLIERS, 1963: 55, 58 59 [#699] (characters; faunal records; Tonkin; Laos; Bouthan). BAUDON, 1968: 92, 102 [#162] (note). JENDEK, 1995: 143 [#3768] (synonymy; characters; North India; Nepal; West Bengal; Bhutan; Laos; North Vietnam; China: Yunnan, Sichuan). AKIYAMA & OHMOMO, 1997: 32 [#4091] (checklist; Taiwan; north China; Vietnam; Laos; Bhutan; Nepal; West and North India). MÜHLE, 2003: 46 [#6155] (checklist). JENDEK, 2005: 9 [#7217] (subspecies of decoloratus; synonymy). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2060 [#8576] (world catalog). alazon Lewis, 1893 (Agrilus) syn. nov. LEWIS, 1893: 333 [#1722] (description). KERREMANS, 1903: 276 [#1441] (catalog). JAKOBSON, 1913: 796 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 15 [#1809] (catalog; Japan). OBENBERGER, 1936: 940 [#2191] (world catalog). IGA, 1947: 14

83 15 [#3273] (notes; characters; faunal records; Japan (Shikoku)). KUROSAWA & IGA, 1947: 27 29 [#1604] (characters; notes). CHÛJÔ & KUROSAWA, 1950: 8 [#501] (cited as alazan; catalog; faunal records; Japan (Shikoku)). KUROSAWA, 1950: 1116 [#3274] (note). IGA, 1955: 78 [#3828] (iconography; Japan). KUROSAWA, 1956: 1116 [#8107] (characters; Japan). IGA, 1962: 78 [#3351] (iconography; Japan). KUROSAWA, 1963: 152 [#1618] (characters; Japan). KUROSAWA, 1974: 2 [#4575] (characters; notes). TÔYAMA, 1985: 23 24 [#3678] (iconography; Japan). HIRASHIMA, 1989: 322 [#8078] (checklist; Japan). JENDEK, 1995: 143 [#3768] (subspecies of decoloratus; lectotype designation; synonymy; characters; Japan; Taiwan; China: Zheijang). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 184 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 32 [#4091] (subspecies of decoloratus; checklist; Japan: Honshu, Shikoku, Kyushu, Tsushima). AKIYAMA & OHMOMO, 2000: 88 [#6240] (subspecies of decoloratus; faunal records; Japan (Hiroshima)). JENDEK, 2000: 504 [#4851] (subspecies of decoloratus; synonymy). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China). MÜHLE, 2003: 46 [#6155] (subspecies of decoloratus; checklist; Taiwan). JENDEK, 2006: 397 [#7441] (subspecies of decoloratus; Palaearctic catalog). BELLAMY, 2008: 2060 [#8576] (subspeies of decoloratus; world catalog). ajax Kerremans, 1912 (Agrilus) KERREMANS, 1912: 207 [#1377] (description). MIWA, 1931: 124 [#6101] (catalog; Formosa). MIWA & CHÛJÔ, 1936: 15 [#1809] (catalog; Japan). OBENBERGER, 1936: 1072 [#2191] (world catalog). IGA, 1947: 15 [#3273] (notes; Japan (Shikoku)). KUROSAWA & IGA, 1947: 27 29 [#1604] (subspecies of alazon; notes; characters). KUROSAWA, 1974: 2 [#4575] (subspecies of alazon; characters; notes). PENG ZHONGLIANG, 1987: 354 [#2429] (cited as aiax; checklist; China). JENDEK, 1995: 143 [#3768] (synonym of decoloratus alazon). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Taiwan). PENG ZHONGLIANG, 2002: 268 [#6628] (subspecies of alazon; characters; Fujian). JENDEK, 2006: 398 [#7441] (synonym of decoloratus alazon; Palaearctic catalog). BELLAMY, 2008: 2060 [#8576] (synonym of decoloratus alazon; world catalog). pyrrhus Kerremans, 1913 (Agrilus) KERREMANS, 1913: 112 113 [#1471] (description). MIWA, 1931: 124 [#6101] (catalog; Formosa). MIWA & CHÛJÔ, 1936: 19 [#1809] (catalog; Japan). OBENBERGER, 1936: 1099 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 357 [#2429] (checklist; China). JENDEK, 2000: 504 [#4851] (synonym of decoloratus alazon; lectotype designation). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Taiwan). JENDEK, 2006: 398 [#7441] (synonym of decoloratus alazon; Palaearctic catalog). BELLAMY, 2008: 2060 [#8576] (synonym of decoloratus alazon; world catalog). octostictus Bourgoin, 1922 (Agrilus) BOURGOIN, 1922: 24 [#365] (description). OBENBERGER, 1936: 1095 [#2191] (world catalog). DESCARPENTRIES & VILLIERS, 1963: 58 [#699] (synonym of decoloratus; [Note: An inference of type is not valid lectotype designation because there were two syntypes in MNHN (ICZN, Article 74.3)]). JENDEK, 1995: 143 [#3768] (synonym of decoloratus; lectotype designation). JENDEK, 2006: 398 [#7441] (synonym of decoloratus aldecoloratus; Palaearctic catalog). BELLAMY, 2008: 2060 [#8576] (synonym of decoloratus; world catalog). lunlingensis Obenberger, 1958 (Agrilus; subspecies of alazon) OBENBERGER, 1958: 237 [#2389] (description). KUROSAWA, 1974: 2 [#4575] (subspecies of alazon; notes). PENG ZHONGLIANG, 1987: 354 [#2429] (subspecies of alazon; checklist; China). YANG XINGKE et al., 1991: 31 [#6756] (subspecies of alazon; type specimen in Institute of Zoology, Academica Sinica, Beijing).

84 JENDEK, 1995: 143 [#3768] (synonym of decoloratus; lectotype designation). HUA LI ZHONG, 2002: 89 [#6696] (subspecies of alazon; cited as alzon; checklist; China: Yunnan). JENDEK, 2006: 398 [#7441] (synonym of decoloratus aldecoloratus; Palaearctic catalog). BELLAMY, 2008: 2060 [#8576] (synonym of decoloratus; world catalog). afghanistanicus Alexeev, 1992 (Agrilus) ALEXEEV in ALEXEEV, VOLKOVITSH & KABAKOV, 1992: 378 379, 389 [#72] (description). ALEXEEV, 1998: 369 [#4322] (subgenus Anambus; subgeneric classification). JENDEK, 2005: 9 [#7217] (synonym of decoloratus subspecies decoloratus). JENDEK, 2006: 398 [#7441] (synonym of decoloratus aldecoloratus; Palaearctic catalog). BELLAMY, 2008: 2060 [#8576] (synonym of decoloratus; world catalog).

Type material: Agrilus decoloratus Kerremans, 1892 Type locality: not given [Inde is cited in the title of the publication]. Type specimens: Holotype by monotypy (Fig. 45E), ♀, (MNHN): “India [h] [round label] \ Ex Musaeo Ed. Brown [p] \ decoloratus Kerr. Type [h] [by Kerremans hand] \ Kerremans Vidit 1892 [p]”. Described from 1 specimen. Agrilus alazon Lewis, 1893 Type locality: Kiushiu ... from Yuyama in Higo. Type specimens: Primary: Lectotype (Fig. 42J), ♂, designated by JENDEK (1995: 143) [#3768], (BMNH): “Higo [h] \ Japan G. Lewis. 1910 - 320 [p] \ Agrilus alazon Lewis Type [h] [by Lewis hand] \ Type H. T. [round label with red border]”. Secondary: 3 PLT (BMNH). Described from unknown number of syntypes. Agrilus ajax Kerremans, 1912 Type locality: [Formosa] Kosempo, Alikang. Type specimens: Primary: Lectotype (Fig. 42H), ♂, designated by JENDEK (1995: 143) [#3768], (MNHN): “Kosempo [ = Chiahsien (Kaohsiung)] Formosa [= Taiwan] H. Sauter [p] IX. [19]09. [h] \ Ajax Kerr. Type [h] [by Kerremans hand] \ Museum Paris Coll. Ch. Kerremans 1923 [p]”. Secondary: 1 PLT (ISNB); 2 PLT (MNHN); 1 PLT (DEI). Described from unknown number of syntypes. Agrilus pyrrhus Kerremans, 1913 Type locality: [Formosa] Kosempo. Type specimens: Primary: Lectotype (Fig. 51P), ♀, designated by JENDEK (2000: 504) [#4851], (DEI): “Kosempo Formosa H. Sauter, 1909 [p] \ IX. [p] \ Syntypus [p] [red label] \ Kerremanns [sic!] det. [p] \ Agrilus Pyrrhus Kerr. Type [h] [Kerremans’ MS] \ Coll. DEI Eberswalde [p]”. Described from unknown number of syntypes. Agrilus octostictus Bourgoin, 1922 Type locality: not given [Indo-Chine française is cited in the title of the publication]. Type specimens: Primary: Lectotype (Fig. 50F), sex not examined, designated by JENDEK (1995: 143) [#3768], (MNHN): “Ban Nam Mo [ in Laos] 4-IV-1918 [h] \ Agrilus octostictus A. Bourgoin [h] [black ink] Type [h] [red ink] \ Muséum Paris 1935 Coll. A. Théry [p]”. Secondary: 1 PLT (MNHN). Described from unknown number of syntypes. Agrilus alazon lunlingensis Obenberger, 1958 Type locality: Chine: Yunnan, Lunling; Yunnan: 30 km SW of Monshih, Santaishan, 1200 m; Szechuan: Omei Mountain,

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Omeishan. [Note: Type locality is determined by lectotype (ICZN, Article 76.2.)] Type specimens: Primary: Lectotype (Fig. 48R), sex not examined, designated by JENDEK (1995: 143) [#3768], (NMPC): “1955 V. 20 [three lines, first and third in Chinese] [p] \ Lunling 1600 m Yunnan 20.V.1955 V. Popov [p] [in Russian] \ Yunnan, Lunling [h] \ Cotype [p] [red label] \ A. Alazon ssp. lunlingensis m. Cotype [h] Det. Dr. Obenberger [p]”. Secondary: PLT (IZAS); 2 PLT (NMPC). Described from unknown number of syntypes. Agrilus afghanistanicus Alexeev, 1992 Type locality: Kunar: N Waygal, 2200 m [in Russian]. Type specimens: Primary: Holotype (Fig. 42E), ♂, (ZIN): “Afganistan Nurestan C Waygal, 2200, 10 VII 72 Kabakov [h] \ Pr Af-Agr-115-74 ♂ [h] [in Russian] \ Holotypus [p] [red label] \ Agrilus afganistanicus [sic!] [h] Alexeev det. [p] sp. n. [h]”. Described from 1 specimen. Diagnosis: 9-14 mm; elytra with 4 pairs of obvious tomentose spots; elytral apex narrowly subangulately arcuate. Species group: Biguttatus. Relevant species: A. biguttatus (Fabricius, 1777); A. ventricosus Fairmaire, 1888. Specimens examined: CHINA: Taiwan: 1 ♂ (EJCB): “Taiwan, Kaohsiung Hsien, Mt. Daynan shan, 22.VIII.1988, leg. Ch. Wenlong” (JENDEK, 1995 [#3768]); 3 (EJCB): “Taiwan, Kaohsiung Hsien, Mt. Daynan shan, 2.VIII.1988, leg. Ch. Wenlong” (JENDEK, 1995 [#3768]); 1 ♂ (NMW): “Taiwan: Taichung Ku Kwang, 10.4.1992, leg. C. L. Li”; 3 ♀ (EJCB): “Taiwan, Nantou, 1997, Lien Hua Chih, 4 km S Wuchen, 23 27 VI., B. Šiška & T. Spevár lgt.”. Xizang: 1 (EJCB): “China E Tibet, Tomi (Tangmai) 2075 m, 30 km W Donjung, leg. A. Wrzecionko, 11.6.2005”. Yunnan: 1 ♂ (EJCB): “Yun’nan’. Syaomon’yan. 810 m, 4.iv.1957, Pu Fu di [in Russian]”; 13 (EJCB): “Yunnan, 1500 2500 m, 25°22’N, 98°49’E, 17 24 / 5 Gaoligong mts, Vít Kubáň leg., 1995”. Zhejiang: 1 ♀ (EJCB): “Chekiang, Chusan [ Zhonshan Qundao] 12.VI.1936” (JENDEK, 1995 [#3768]). INDIA: Arunachal Pradesh: 4 ♀ (EJCB): “India, W Arunachal Pr., Dirang vicinity, 1500 1800m, 27°21’ 23’N 92°13’ 16’E, L. Dembický leg.; 1. 10.vi.2004”. Assam: 1 (MNHN): “Assam, Patkai \ Doherty” (JENDEK, 1995 [#3768]). Jammu & Kashmir: 1 ♂, 2 ♀ (EJCB): “India, Jammu u. Kashmir, Jammu, district Kishtwar, Ekala Sonder, 1700 2000 m, 8.7.1980, leg. C. Holzschuh” (JENDEK, 1995 [#3768]). Meghalaya: 1 (EKCS): “NE INDIA, ASSAM, 1999, 5 km N of UMRONGSO; 700m, 25°27’N, 92°43’E, 17. 25.v. Zd. Koštál leg.”. Uttarakhand: 3 (EJCB): “N India, Uttaranchal state, ca 30 km N of Bageshwar, Khati village env., 2100 2300 m, 27 30.vi.2003, Z. Kejval & M. Tryzna lgt.”. West Bengal: 1 (MNHN): “Maria Basti, L. Durel” (JENDEK, 1995 [#3768]); 1 (MNHN): “Padong, L. Durel 1914” (JENDEK, 1995 [#3768]). JAPAN: Honshu: 1 ♂ (EJCB): “Japan, Hyogo prefecture, Akasai, 27.VII.1973, leg. M. Yuma” (JENDEK, 1995 [#3768]); 1 ♀ (EJCB): “Gongen forest, Nagara Cho, Chiba pref., 16 V 1996, M. M. Maruyama leg.”; 2 (EJCB): “Japan, Yamagata, Myozawa, Oguni town, 19.vii.2005, N. Sugiura leg.”. Kyushu: 1 (EJCB): “Shiroyama, Fukuoka Pref., 2.VI.1984, leg. S. Ogata” (JENDEK, 1995 [#3768]); 1 ♂ (JMCZ): “Japan, Kyushu, Oita pref., Yufuin cho, Tsukahara, 14.vii.1985”. Tsushima: 1 ♂ (EJCB): “Japan, Tsushima Island, Nagasaki pref., 7.VI.1981, A. Torikata leg.” (JENDEK, 1995 [#3768]). KOREA NORTH: 1 ♂ (EJCB): “Nordkorea, Jong Won, Mt. Huang Yonglan, 12 26.6.2004, ex Coll. E. v. d. Berghe”. LAOS: Borikhamxai: 7 (EJCB): “Laos centr., Bolikhamsai prov., Ban Nape Kaew Nua Pass, 18.4. 1.5.1998, alt. 600±100 m, N 18º22.3, E 105º09.1 (GPS), E. Jendek & O. Šauša leg.”. Louangphrabang: 2 ♂, 1 ♀ (EJCB): “Laos N, 24.iv. 16.v.1999, Louang Phrabang prov. 20°33 4’N 102°14’E, Ban Song

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Cha (5km W) ± 1200 m, Vít Kubáň leg.”. Oudomxai: 8 (EJCB): “Laos N (Oudomxai), 1 9.v.2002, ~1100m, 20º45’N, 102º09’E, Oudom Xai (17 km NEE), Vít Kubáň leg.”. Phongsali: 3 (EJCB): “Lao N, Phongsaly prov., 21°41’ 2’N, 102°06’ 08’E, Phongsaly env., 28.v. 20.vi.2003, 1500m, P. Pacholátko leg.”. NEPAL: 1 (EJCB): “E. Nepal, Arun V.[aley], M. Brancucci, Lamobagar Gola, 1400 m, 9 14.VI.1983” (JENDEK, 1995 [#3768]). THAILAND: Chiang Mai: 1 (EJCB): “N Thailand, 100km N Chiang Mai, Chiang Dao hill resort, 20 30.vi.2008, S. Murzin”.

Adult occurence: 4-5-6-7-8 (n=24). Altitude range: 5002500 m (n=14). Faunal record published by BAUDON, 1968 [#162] is also included. Host plants: Celtis sinensis var. japonica: KUROSAWA & IGA, 1947 [#1604]; AKIYAMA & OHMOMO, 1997 [#4091]. – Quercus: ALEXEEV in ALEXEEV, VOLKOVITSH & KABAKOV, 1992 [#72] (as afghanistanicus) Distribution: AFGHANISTAN. BHUTAN. CHINA: Sichuan, Taiwan, Xizang, Yunnan, Zhejiang. INDIA. JAPAN: Honshu, Kyushu, Shikoku, Tsushima. KOREA NORTH. LAOS. NEPAL. THAILAND. VIETNAM. Remarks: Specimens from Japan and Taiwan cited as subspecies alazon Lewis differ from the nominotypical subspecies only by predominate blue color of the dorsal side. This character is considered to be of infrasubspecific value.

Agrilus delphinensis Abeille de Perrin Fig. 17H (habitus); Fig. 35M (aedeagus); Figs. 45F, 45V (types) delphinensis Abeille de Perrin, 1897 (Agrilus) ABEILLE DE PERRIN, 1897: 4, 12 13 [#19] (description). KERREMANS, 1903: 284 [#1441] (catalog). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). BEDEL, 1921: 202, 210 [#170] (synonym of pseudocyaneus; characters in key; faunal records; distribution; biology; France). OBENBERGER, 1926: 656 [#2057] (synonym of pseudocyaneus). OBENBERGER, 1936: 1024 1025 [#2191] (synonym of pseudocyaneus; world catalog). RIKHTER, 1945: 167 [#3507] ([Note: Misidentified as pseudocyaneus]; characters in key; Lithuania; Latvia; Estoniya; North European part of USSR). RIKHTER, 1948: 394 [#3879] ([Note: Misidentified as pseudocyaneus]; characters in key). STEPANOV, 1948: 243 [#3535] ([Note: Misidentified as pseudocyaneus]; checklist; faunal record; Chelyabinskaya oblast’). SCHAEFER, 1949: 355, 389 391, 490 [#2565] (subspecies of pseudocyaneus; characters; references; faunal records; biology; notes; ditributional overview; notes; France). HORION, 1955: 91 92 [#1285] ([Note: Misidentified as pseudocyaneus according to host plants cited]; distribution overview; faunal records; biology; Deutschland; Österreich). ALEXEEV, 1959: 8 [#3229] (synonym of pseudocyaneus; characters in key; Latvia; Lithuania; Estoniya; north of European USSR to Primorsk). ZAGAIKEVICH, 1962: 103 [#3249] (synonym of pseudocyaneus; faunal records; biology; Ukraine). RIKHTER & ALEXEEV, 1965: 301 [#2522] ([Note: Misidentified as pseudocyaneus]; characters in key; European part of USSR; Lugansk). FRANZ, 1974: 125 [#4334] ([Note: Misidentified as pseudocyaneus according to host plant cited]; faunal records; Austria). VOLKOVITSH, 1977: 26 [#2985] ([Note: Misidentified as pseudocyaneus]; faunal record; notes; USSR: Belgorod; Lugansk). KETTERING, 1978: 81 83 [#3623] (subspecies of pseudocyaneus; Germany). LOMPE, 1979: 237 [#3628] (subspecies of pseudocyaneus; characters in key; Central Europe). BÍLÝ, 1982: 64, 79 80 [#295] ([Note: Misidentified as pseudocyaneus according to host plant]; characters in key;

85 distribution overview; bionomy). GEISER, 1984: 90 [#6841] ([Note: Misidentified as pseudocyaneus]; red list; Germany). ALEXEEV, 1989: 487 [#64] ([Note: Misidentified as pseudocyaneus]; key to species; Khabarovsk; Amursk; Primorsk; Chita; Buryatia; Tuva; North Korea; northeastern China). ALEXEEV & VOLKOVITSH, 1989: 318, 346, 356 [#3241] ([Note: Misidentified as pseudocyaneus]; characters in key; faunal records; Mongolia). GUTOWSKI, 1993: 302 [#3638] (characters). CURLETTI, 1994: 165 [#658] (subgenus Agrilus; [Note: Misidentified as pseudocyaneus]; catalog; faunal records; Italia). JENDEK, 1995: 173 174, 177 [#3767] (lectotype designation; synonymy; key; Eurosiberian; west to France and Germany; north to Finland; south to Italy, Serbia and Bosnia; central and southern Russia; east to Russian Far East; Mongolia). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). BÍLÝ, 2002: 16 17 [#5877] (bionomy; host plants; distribution). BRECHTEL & KOSTENBADER, 2002: 251 253 [#5766] (biology; ecology; distribution; Germany). JENDEK, 2006: 393 [#7441] (subgenus Robertius; Palaearctic catalog). BELLAMY, 2008: 2063 [#8576] (subgenus Robertius; world catalog). VOLKOVITSH, 2009: 135 [#8850] (subgenus Robertius; faunal records; Russia: Primorskiy kray). eogenes Obenberger, 1930 (Agrilus; subspecies of pseudo cyaneus) OBENBERGER, 1930: 112 [#2088] (description). JENDEK, 1995: 173, 174 [#3767] (synonym of delphinensis; lectotype designation). BRECHTEL & KOSTENBADER, 2002: 251 [#5766] (synonym of cyanescens). JENDEK, 2006: 393 [#7441] (synonym of delphinenis; Palaearctic catalog). BELLAMY, 2008: 2063 [#8576] (synonym of delphinensis; world catalog).

Type material: Agrilus delphinensis Abeille de Perrin, 1897 Type locality: France: Voiron. Type specimens: Primary: Lectotype (Fig. 45F), ♂, designated by JENDEK (1995: 173-174, 177) [#3767], (MNHN): “Voiron osiers E. ABEILLE [h] \ delphinensis typ. 97 Ab. [h] \ Museum Paris Coll. L. BEDEL 1922 [p]”. Secondary: 9 PLT (MNHN). Described from unknown number of syntypes. Agrilus pseudocyaneus eogenes Obenberger, 1930 Type locality: Siberia or.: Primorje. Type specimens: Primary: Lectotype (Fig. 45V), sex not examined, designated by JENDEK (1995: 173, 174) [#3767], (NMPC): “Primorje [h] \ Typus [p] [red label] \ Agrilus pseudocyaneus ssp. eogenes m. Type [h] det. Dr. Obenberger [p]”. Described from unknown number of syntypes. Diagnosis: 3.8-5.3 mm; subparallel; dorsally blue; sides of pronotum less arcuate. Species group: Betuleti. Relevant species: A. betuleti (Ratzeburg, 1837). Specimens examined: CHINA: Heilongjiang: 2 (ZIN): “Kharbin Kitai 29.vi.1935 [in Russian]”; 1 (HMMG): “6.vii.1954, Heilongjiang, Harbin”. KAZAKHSTAN: 1 (ZIN): “khr. Tarbagatai, 30km ZSZ p. Tarbagatai, Kazakhstan, Belokabyl’skii 9 20.vii.1983 [in Russian]”. MONGOLIA: 1 (EJCB): “Mongolia, Sukhbaatar, Darichanga, 17.vi.1975, 1200 m”; 1 (EJCB): “Mongolia, Central Ajmak, river Tola, 230 km NW Ulan Bator, 13 16.VI.1975” (JENDEK, 1995 [#3767]); 1 ♂ (EJCB): “Mongolia, Central Ajmak, Ulan Bator, 26.VI.1988, leg. O. Majzlan” (JENDEK, 1995 [#3767]). RUSSIA: Amurskaya oblast’: 2 (ZIN): “Amurskaya obl., Arkharinskii r n, s. Antonovka, S. Vinter 5 9.vi.1976 [in Russian]”. Buryatiya Respublika: 1 (ZIN): “Buryatia r. Temnik 13.vii.1961 [in Russian]”; 1 (ZIN): “Yu Buryatiya, V. Torey 15.7.1985 Sokolov [in Russian]”. Irkutskaya oblast’: 2 (ZIN): “Irkutsk, Yakovlev

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[in Russian]”. Krasnoyarskiy kray: 1 (ZIN): “Krasnoyar. kr., Usinskii maralosovkoz 22.6.1979, Korotyaev [in Russian]”. Leningradskaya oblast’: 1 (ZIN): “Yu Peterburga S. Novinki, 20.VII.1992 Kadyrov [in Russian]”. Primorskiy kray: 2 (ZIN): “Y Primor’e, sr. ter. r. Suchan, Kabakov, 10 20.7.1952 [in Russian]”; 2 ♀ (ZIN): “Prim. kr., Khankajsk. r n, 20 km NW poc. Kamen’ Rybolov, 28 29.VI.1974, leg. Loginova” (JENDEK, 1995 [#3767]); 1 ♂ (ZIN): “Primorskii krai, Lyalitchi, r. Ilistaja, les, 4.VI.1990, leg. Belokobylskij [in Russian]” (JENDEK, 1995 [#3767]); 2 ♀ (EJCB): “Sib. or. m., Primorje, Sichote Alin Mts., Sokolči, 1 15.7.1990, Kadlec + Voříšek lg.” (JENDEK, 1995 [#3767]); 1 (ZIN): “Primorskii krai, Spassk, 13.VI.1990, leg. Belokobylskij [in Russian]” (JENDEK, 1995 [#3767]); 1 ♀ (ZIN): “Primor’e, Merkushevka, 13 km V Dmitrijevki, 5.06.1990, Belokobylskij [in Russian]” (JENDEK, 1995 [#3767]); 1 (MSCI): “NW Primorskii kr., zh./d. stanciya Ruzhino na kustarnikovykh Ulmus, 16.06.06, leg. М.& L. Smirnov’s [in Russian]”; 1 (EJCB): “Russia, Primorsky Kray, Kaymanovka vill., N43°37’, E132°14’, 7 11.vii.2007, 100 m, V. Grebennikov”. Yevreyskaya avtonomnaya oblast’: 4 (EJCB): “Russia, ES Siberia, Khabarovsk., Oblutchie distr.[Remark: district Oblutchie belongs to Yevreyskaya avtonomnaya oblast’], S. Chingan Mts., 500 800 m, 15.VI. 17.VII.2001, R. Misustin & S. Duz leg.”.

Type material: Agrilus densetomentosus Obenberger, 1940 Type locality: China: Fokien: Tainingli. Type specimens: Primary: Lectotype (Fig. 45H), ♀, designated by JENDEK (2005: 9) [#7217], (NMPC): “Fokien: Tainingli [h] \ Typus [p] [red label] \ A. densetomentosus m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24943 [h] [orange label]”. Described from unknown number of syntypes.

Adult occurence: 4-6-7 (n=25). Altitude range: 100-1200 m (n=3).

Host plants: Unknown.

Host plants: Salix: ABEILLE DE PERRIN, 1897 [#19]; BEDEL, 1921 [#170]; ALEXEEV, 1959 [#3229]; ZAGAIKEVICH, 1962 [#3249]; RIKHTER & ALEXEEV, 1965 [#2522]; BÍLÝ, 1982 [#295]; ALEXEEV, 1989 [#64]; ALEXEEV & VOLKOVITSH, 1989 [#3241]; BÍLÝ, 2002 [#5877]; GUTOWSKI & MILKOWSKI, 2008 [#8244]. − S. aurita: BRECHTEL & KOSTENBADER, 2002 [#5766]. − S. caprea: SCHAEFER, 1949 [#2565]; HORION, 1955 [#1285]; FRANZ, 1974 [#4334]; JENDEK, 1995 [#3767]; HASTIR & GASPAR, 2002 [#6211]. − S. viminalis: SCHAEFER, 1949 [#2565]; HORION, 1955 [#1285]; KETTERING, 1978 [#3623]; CURLETTI, 1994 [#658]; JENDEK, 1995 [#3767]; BRECHTEL & KOSTENBADER, 2002 [#5766]; HASTIR & GASPAR, 2002 [#6211]. Distribution: Local: CHINA: Heilongjiang, Jilin, Liaoning. KOREANORTH. MONGOLIA. RUS-SIA: Amurskaya oblast’, Buryatiya Respublika, Irkutskaya oblast’, Khabarovskiy kray, Primorskiy kray, Tyva Respublika, Zabaykalskiy kray. Global: ASIA: CHINA. KAZAKHSTAN. KOREA NORTH. RUSSIA. EUROPE: AUSTRIA. BELARUS. BELGIUM. BOSNIA & HERZEGOVINA. CZECH REPUBLIC. ESTONIA. FINLAND. FRANCE. GERMANY. HUNGARY. ITALY. LATVIA. LITHUANIA. POLAND. ROMANIA. RUSSIA. SERBIA & MONTENEGRO. SLOVAKIA. SWITZERLAND. UKRAINE.

Agrilus densetomentosus Obenberger Fig. 28C (habitus); Fig. 40G (aedeagus); Fig. 45H (type) densetomentosus Obenberger, 1940 (Agrilus) PENG OBENBERGER, 1940: 184 185 [#2273] (description). ZHONGLIANG, 1987: 355 [#2429] (checklist; China). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China). PENG ZHONGLIANG, 2002: 270 [#6628] (notes). JENDEK, 2005: 9 [#7217] (lectotype designation). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2063 [#8576] (world catalog).

Diagnosis: 4.5-5.1 mm; body more robust; humeri with obvious impression. Species group: Densetomentosus. Relevant species: A. uenoi Kurosawa, 1963. Specimens examined: CHINA: Fujian: 1 ♀ (EJCB): “China, Fujian prov. Shaowu env., 5 10.VII.1991”. Hunan: 1 ♂ (EJCB): “China, Hunan SE, 1994, Guidong env. 26 31.V., 26.04N 113.56E, E. Jendek & O. Šauša leg.”. Jiangxi: 1 ♀ (EJCB): “China, Jiangxi W, Jingggang Shan Ciping, 2 14.VI.1994, E. Jendek & O. Šauša leg.”.

Adult occurence: 5-6-7 (n=3).

Distribution: CHINA: Fujian, Hunan, Jiangxi.

Agrilus dianthus Kerremans Fig. 12C (habitus); Fig. 33M (aedeagus); Figs. 44O, 45J, 45O, 50B, (types) dianthus Kerremans, 1892 (Agrilus) KERREMANS, KERREMANS, 1892: 823 [#1398] (description). 1903: 284 [#1441] (catalog). OBENBERGER, 1936: 1081 [#2191] (world catalog). BAUDON, 1961: 75 [#143] ([Note: Misidentified as acutus according to JENDEK, 2004: 8 [#6675]]; faunal records; characters; Laos). DESCARPENTRIES & VILLIERS, 1963: 52, 53 54 [#699] (subspecies of acutus; lectotype designation: “Birmanie (type de dianthus au Musée de Gênes)”; characters; faunal records; Birmanie; Tonkin; Laos; Cambodge; Cochinchine; Annam; Thailand). DESCARPENTRIES & VILLIERS, 1967: 144 [#710] (subspecies of acutus; ornatus species group; faunal records; Cochinchine; Annam). BAUDON, 1968: 71, 73 [#162] (subspecies of acutus). JENDEK, 2004: 7 9, 16 [#6675] (characters; synonymy; faunal records; China (Yunnan province); Myanmar; Thailand; Laos; Vietnam; Cambodia; Indonesia: Kalimantan). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2068 2069 [#8576] (world catalog). nodosus Kerremans, 1894 (Agrilus) KERREMANS, 1894: 470 471 [#1412] (description). KERREMANS, 1903: 277 [#1441] (catalog). OBENBERGER, 1936: 1094 [#2191] (world catalog). DESCARPENTRIES & VILLIERS, 1963: 53, 54 [#699] (synonym of acutus dianthus; [Note: An inference of type is not valid lectotype designation because there were two syntypes in BMNH (ICZN, Article 74.3)]). JENDEK, 2004: 7 [#6675] (synonym of dianthus; lectotype designation). JENDEK, 2006: 398 [#7441] (synonym of dianthus; Palaearctic catalog). BELLAMY, 2008: 2068 [#8576] (synonym of dianthus; world catalog). coelicus Bourgoin, 1922 (Agrilus) BOURGOIN, 1922: 24 [#365] (description). OBENBERGER, 1936: 1078 [#2191] (world catalog). DESCARPENTRIES & VILLIERS, 1963: 54 [#699] (synonym of acutus dianthus; lectotype designation: “Louang Prabang (type de coelicus, Muséum de Paris)”). JENDEK, 2004: 7 8 [#6675] (synonym of dianthus; notes).

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA JENDEK, 2006: 398 [#7441] (synonym of dianthus; Palaearctic catalog). BELLAMY, 2008: 2069 [#8576] (synonym of dianthus; world catalog). dorri Bourgoin, 1925 (Agrilus) BOURGOIN, 1925: 131 [#373] (description). OBENBERGER, 1936: 1081 [#2191] (world catalog). DESCARPENTRIES & VILLIERS, 1963: 54 [#699] (synonym of acutus dianthus; lectotype designation: “Long Xuyen (type de dorri au Muséum de Paris)”). JENDEK, 2004: 7, 8 [#6675] (synonym of dianthus; notes). JENDEK, 2006: 398 [#7441] (synonym of dianthus; Palaearctic catalog). BELLAMY, 2008: 2069 [#8576] (synonym of dianthus; world catalog).

Type material: Agrilus dianthus Kerremans, 1892 Type locality: Bhamò. Type specimens: Primary: Lectotype (Fig. 45J), ♀, designated by DESCARPENTRIES & VILLIERS (1963: 52, 53-54) [#699], (MCSN): “Bhamò Birmania Fea [leg.] [p] X [h] 188 [p] 6 [h] \ Typus [p] [red ink, red border] \ dianthus Kerr. [h] [black border] \ A. dianthus Kerr. typus ! [h] [yellow label] \ di Described from unknown number of syntypes. Agrilus nodosus Kerremans, 1894 Type locality: Bima. Type specimens: Primary: Lectotype (Fig. 50B), ♂, designated by JENDEK (2004: 7) [#6675], (BMNH): “SYNType [p] [round label with blue border] \ Bima Boucard [leg.] [Kerremans’ manuscript] \ nodosus Kerr. Type [Kerremans’ manuscript] \ Kerremans 1903-59. [p]”. Secondary: 1 PLT (BMNH). Described from unknown number of syntypes. Agrilus coelicus Bourgoin, 1922 Type locality: not given. Type specimens: Primary: Lectotype (Fig. 44O), ♀, designated by DESCARPENTRIES & VILLIERS (1963: 54) [#699], (MNHN): “Vieng Vai 12-VI-1918 Luang Prab. [h] \ Agrilus coelicus Type Bourg. [h] \ Type [p] [red ink] \ Muséum Paris 1935 coll. A. Théry [p]”. Secondary: 1 PLT (ISNB); 1 PLT (BMNH). Described from unknown number of syntypes. Agrilus dorri Bourgoin, 1925 Type locality: Cochinchine: Long Xuyen. Type specimens: Primary: Lectotype (Fig. 45O), ♀, designated by DESCARPENTRIES & VILLIERS (1963: 54) [#699], (MNHN): “Long Xuyen Cochinchine Dorr [p] \ Agrilus Dorri A. Bourgouin [black ink] Type [red ink] [h]”. Described from unknown number of syntypes. Diagnosis: 6.6-10.2 mm; pronotal wide/length ratio = 1.41.55; elytral apex with small outer spine. Species group: Acutus. Relevant species: A. acutus (Thunberg, 1787). Specimens examined: CHINA: Yunnan: 1 (EJCB): “Yun’nan’, 10 km N Czinduna [ ?Jindun, 26°30’N, 100°12’E], 1250 m, 28.VI.1956, A. Shnitnikov [in Russian]” (JENDEK, 2004 [#6675]); 2 (NMPC): “Yun’nan’, 30 km SW Czin’lina, 420 m, 21.IV.1956 Khuan Kèzhen’ i dr. [in Russian]” (JENDEK, 2004 [#6675]); 1 (HMMG): “28.vi.1957, Yunnan, Jinghong”; 1 (HMMG): “9.vii.1958, Yunnan, Xishuangbanna”; 1 (HMMG): “6.vii.1958, Yunnan, Jinghong”; 1 (HMMG): “6.ix.1958, Yunnan, Jinghong”; 1 (HMMG): “18. vi.1959, Yunnan, Jinghong”; 1 ♀ (EJCB): “S. China, Jinghong, 10 14.7.1990, pr. Yunnan, lgt. S. Bečvář” (JENDEK, 2004 [#6675]); 1 (MNCA): “China: S Yunnan, 25km N Jinghong, vic. Man Dian, 10.xii.2007, leg. Weigel KL”. INDONESIA: Kalimantan: 1 ♂

87 (EJCB): “Indonesia, Kalimantan W, 4 10.VIII.1993, Nanga Ela env. Nanga Nyuruh, 700 m, J. Schneider leg.”.” (JENDEK, 2004 [#6675]). LAOS: Khammouan: 2 (EJCB): “LAOS C., Khammouan pr., 20 29.V.2004, alt. 250m, Ban Khoun Ngeun env., 18º07’N, 104º29’E, E. Jendek & O. Šauša leg.”. Louang Namtha: 5 ♀ (EJCB): “Laos north, 24 30.V.1997, 20 km NW Louang Namtha, N 21°09.2, E 101°18.7, alt. 900±100 m, E. Jendek & O. Šauša leg.” (JENDEK, 2004 [#6675]); 1 ♂, 1 ♀ (EJCB): “Laos north, 13 24.V.1997, 15 km NW Louang Namtha, N 21°07.5, E 101°21.0, alt. 750±100 m, E. Jendek & O. Šauša leg.” (JENDEK, 2004 [#6675]); 1 ♀ (EJCB): “Laos north, 5 11.V.1997, 20 km NW Louang Namtha, N 21°09.2, E 101°18.7, alt. 900±100 m, E. Jendek & O. Šauša leg.” (JENDEK, 2004 [#6675]). Louangphrabang: 1 ♂, 1 ♀ (EJCB): “Laos N (Louangphrabang) 11 21.V.2002, 19°35’N, 101°58’E, Thong Khan, ~ 750 m, Vít Kubáň leg.” (JENDEK, 2004 [#6675]). Phongsali: 1 ♀ (EJCB): “Lao N, Phongsaly prov., 21°41’ 2’N, 102°06’ 08’E, Phongsaly env., 28.v. 20.vi.2003, 1500m, P. Pacholátko leg.”; 3 (EJCB): “Lao, Phongsaly prov. 21°41’N, 102°06’E Phongsaly env. 6 17.v.2004, 1500 m, P. Pacholátko leg.”. Vientiane: 1 ♂, 1 ♀ (EJCB): “Laos centr., 27.IV. 1.V.1997, 70 km NE Vientiane, Ban Phabat env., 150 m, N 18°16.1, E 103°10.9, E. Jendek & O. Šauša leg.” (JENDEK, 2004 [#6675]). MALAYSIA: Johor: 1 ♂ (EJCB): “W Malaysia, Johor, 30 km NE of Labis, Mt. Besar, [2°31’N, 103°09’E], 800 m, 25 27.IV.1997, P. Čechovský leg.” (JENDEK, 2004 [#6675]). MYANMAR: 2 (NMPC): “Tenasserim” (JENDEK, 2004 [#6675]); 1 ♂ (EJCB): “Burma, Dawna Berge [16°50’N, 98°15’E], 7.V.1994, erw. Steinicke” (JENDEK, 2004 [#6675]). THAILAND: Chiang Mai: 3 (EJCB): “Thailand, 10 16.V.1991, Chiang Dao, 600 m, 19°25’N, 98°52’E, Vít Kubáň leg.” (JENDEK, 2004 [#6675]); 1 ♀ (EJCB): “Thai, 26 28.V.1991, Palong, 750 m, 19°55’N, 99°06’E, Vít Kubáň leg.” (JENDEK, 2004 [#6675]); 28 (EJCB): “Thailand N, 21.V. 4.VI.1995, Chiang Dao env., M. Snížek leg.” (JENDEK, 2004 [#6675]); 1 (EJCB): “Thailand, Chiang Dao, 1.VII.1996, M. Kondler leg.” (JENDEK, 2004 [#6675]). Kamphaeng Phet: 2 ♂ (EJCB): “W Thailand, 2 5.VII.1997, Khlong Lan Nat. Park [16°11’N, 99°12’E] 50 km SW Khamphaeng Phet, Kaláb, J. & Rejsek, J. leg.” (JENDEK, 2004 [#6675]). Kanchanaburi: 1 ♀ (EJCB): “Thai, 9 13.IV.1991, Thimonghta, 350 m, 15°02’N, 98°35’E, Vít Kubáň leg.” (JENDEK, 2004 [#6675]). Mae Hong Son: 1 (EJCB): “NW Thailand, Mae Hong Son, 17 23.V.1991, Ban Huai Po, 19°19’N, 97°59’E, 1600 2000 m, L. Dembický leg.” (JENDEK, 2004 [#6675]); 1 (EJCB): “Thailand, Mae Hong Son, [19°18’N, 97°58’E], 1000 m, 16 23.VI.1993, J. Schneider leg.” (JENDEK, 2004 [#6675]); 4 (EJCB): “Thailand bor., prov. Mae Hong Son, Pai, 16 24.VII.1995, R. Šigut leg.” (JENDEK, 2004 [#6675]); 1 (EJCB): “Thailand bor., prov. Mae Hong Son, Pai, 24 30.IV.1997, R. Šigut leg.” (JENDEK, 2004 [#6675]). Nan: 1 ♀ (EJCB): “Thailand, 11 15.V.1993, Nan Pha Khab [site unlocated] P. Pacholátko & L. Dembický leg.” (JENDEK, 2004 [#6675]); 1 ♀ (EJCB): “Thailand N, Nan prov., Bo Klua, 19°08’N, 101°10’E, 700 m, 28.IV. 1.V.1999, M. Říha leg.” (JENDEK, 2004 [#6675]); 1 ♂ (EJCB): “Thailand N, Nan prov., Bo Klua, 19°03’N, 101°10’E, 700 m, 28.IV. 1.V.1999, D. Hauck leg.” (JENDEK, 2004 [#6675]); 1 ♂, 2 ♀ (EJCB): “Thailand, Nan prov., Ban Bo Klua env. [19°03’N, 101°10’E ] 13 26.V.2002, Průdek, P. & Obořil, M. leg.” (JENDEK, 2004 [#6675]); 1 ♂, 1 ♀ (EJCB): “Thailand, Nan prov., Ban Huay Kon [19°34’N, 101°05’E,] env., 27.V. 10.VI.2002, Průdek, P. & Obořil, M. leg.” (JENDEK, 2004 [#6675]). Yala: 4 ♂, 1 ♀ (EJCB): “S Thailand, Betong [5°47’N, 101°04’E], Gunung Cang Dung vill., Yala dist. 25.III. 22.IV.1993, J. Horák leg.”.” (JENDEK, 2004 [#6675]). VIETNAM: 5 ♂, 3 ♀ (EJCB): “S Vietnam, 1 15.V.1994, Nam Cat Tien Nat. Park, [11°27’N, 107°19’E] P. Pacholátko & L. Dembický leg.” (JENDEK, 2004 [#6675]). Dong Nai: 1 ♂, 1 ♀ (EJCB): “Vietnam, Dong Nai prov., Ma Da forest, 1.VI.1991, Vít Kubáň leg.” (JENDEK, 2004 [#6675]); 2 ♀ (EJCB): “Vietnam, Dong Nai prov., Ma Da forest, 1.VI.1991, Vít Kubáň leg.” (JENDEK, 2004

88 [#6675]). Hoa Binh: 1 (EJCB): “Vietnam, Hoa Binh [20°50’N, 105°20’E], 26 28.V.1989 J. Secký leg.” (JENDEK, 2004 [#6675]). Ninh Binh: 1 (EJCB): “Vietnam, Ha nam ninh prov., Cuc Phuong, 24 25.V.1986, J. Víša leg.” (JENDEK, 2004 [#6675]).

Adult occurence: 3-4-5-6-7-9-12 (n=42). Altitude range: 150-2000 m (n=20). Faunal record published by BAUDON, 1968 [#162] is also included. Host plants: Unknown. The following host plants were published from the very close, possibly consubspecific taxon A. acutus (Thundberg, 1787): Abelmoschus: KALSHOVEN, 1951 [#3718].− Corchorus: LEEFMANS, 1922 [#8158]; Kabir, 1995 [#7048]. C. olitorius: DUTT, 1969 [#3659]. − Hibiscus cannabinus: MÜHLMANN, 1954 [#8152]; DUTT & BHATTACHERJEE, 1960 [#7046]; DUTT, 1969 [#3659]; DUTT, 1971 [#7047]; BHUIYAN & KABIR, 1985 [#7045]; PANDIT, 1998 [#6287]; AHMAD et al., 2001 [#7043]. − H. esculentus: DE, 1956 [#8155]. − H. sabdariffa: DUTT, 1969 [#3659]. − Urena: KALSHOVEN, 1951 [#3718]. − U. lobata: BHATTACHARYA et al., 1977 [#4209]. Distribution: CAMBODIA. CHINA: Yunnan. INDONESIA. LAOS. MALAYSIA. MYANMAR. THAILAND. VIETNAM.

Agrilus diao Jendek, sp. nov. Fig. 25K (habitus of HT)

Description: 5.7-5.9 mm (Holotype 5.9 mm). BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): green. HEAD: Medial impression: present, Impression (extent): vertex, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (density): dense, Sculpture (intensity): superficial. Eyes: Size: large, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: visually elongate or visually square, Sides: arcuate, Maximal width: at anterior margin, Anterior margin: subequal to posterior. Anterior lobe: obvious, rarely moderate, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles or projecting beyond anterior pronotal angles. Posterior angles: Shape: acute or rectangular, Apex: sharp. Disk: Convexity: flat or moderately convex, Disk modifications: with obvious tomentose spots, Disk impressions: medial and lateral or posterolateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow. Prehumerus: carinal, Shape: bisinuate, Extent: beyond half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina, Arc: obvious. Marginal and submarginal carinae: Interspace: narrow or moderate, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior angles or at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: obsolete. ELYTRA: Color: unicolored with darker hue in apical half, Humeral carina: absent. Apices: Arrangement: conjoint, Shape: subtruncate,

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Truncation: oblique with projecting pleural angle. Elytral pubescence: proximal and distal, Proximal: like V or upturned T, Distal: apical, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: large, Anterior margin: angulately emarginate, Emargination (deepness): deep, Emargination (width): moderate. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: rectangular, Angles (tips): sharp, Disk: impressed, rarely impressed with sides protruding. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Ovipositor: elongate. Diagnosis: elytral apices obliquely subtruncate; prosternal lobe markdly deeply subangulately emarginate. Species group: Tempestivus. Relevant species: A. malloti Théry, 1930. Type locality: Central China, Shaanxi province, Qinling Shan, 6 km eastern of Xunyangba, altitude 1000-1300 m. Type specimens: Primary: Holotype (Fig. 25K), ♀, (EJCB): “C-China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000-1300m, 23.V.-13.VI. leg. C. Holzschuh 2000”. Secondary: CHINA: Shaanxi: 1 ♀ PT (EJCB): “China, 1000-1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.-13.vi.1998, I. H. Marshal leg.”. Adult occurence: 5-6 (n=2). Altitude range: 1000-1300 m (n=2). Host plants: Unknown. Distribution: CHINA: Shaanxi. Etymology: The specific name is the Chinese verb diao (to engrave). It refers to the deeply subangulately emarginate prosternal lobe of this species.

Agrilus diaolin Jendek Fig. 12H (habitus of HT); Fig. 33Q (aedeagus) diaolin Jendek, 2001 (Agrilus) JENDEK, 2001: 5 6, 10 [#5211] (description). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2069 [#8576] (world catalog).

Type material: Agrilus diaolin Jendek, 2001 Type locality: China, Yunnan, 100 km W of Kunming, Diaolin Nat. Reservation. Type specimens: Primary: Holotype (Fig. 12H), ♂, (EJCB): “China, Yunnan, 22. V. - 2. VI., 100km W of KUNMING, 1993, DIAOLIN Nat. Reservation, E. Jendek & O. Šauša leg.”. Secondary: 11 PT (EJCB). Described from 12 specimens (HT, PT). Diagnosis: 6.9-8.6 mm; body dorsally black, silky lustrous; elytra with O-like ornament of whitish pubescence in apical

89

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA third. Species group: Plasoni. Relevant species: A. plasoni Obenberger, 1917. Specimens examined: CHINA: Yunnan: 4 ♂ PT, 4 ♀ PT (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.” (JENDEK, 2001 [#5211]); 3 ♀ PT (EJCB): “Yunnan, 2000 m, 25.03N, 101.55E, Yipinglang, 8 10/6, 1993” (JENDEK, 2001 [#5211]). Adult occurence: 5 6 (n 2). Altitude range: ~2000 m (n 1). Host plants: Unknown. Distribution: CHINA: Yunnan.

Agrilus discalis Saunders Fig. 13K (habitus); Fig. 34E (aedeagus); Fig. 45L (type) discalis Saunders, 1873 (Agrilus) SAUNDERS, 1873: 514 515 [#2554] (description). LEWIS, 1879: 15 [#3345] (catalog; Japan). KERREMANS, 1885: 152 [#1381] (catalog). SCHÖNFELDT, 1887: 113 [#8901] (Anambus; catalog; Japan). KERREMANS, 1892: 255 [#1397] (catalog). LEWIS, 1893: 335 [#1722] (faunal records; Japan). KERREMANS, 1900: 81 [#1436] (faunal records; India: Cachemire). KERREMANS, 1903: 276 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). YUASA, 1932: 653 [#3369] (characters; Japan). MIWA & CHÛJÔ, 1936: 16 [#1809] (catalog; Japan). OBENBERGER, 1936: 989 [#2191] (world catalog). CHÛJÔ & MATUDA, 1940: 65 [#500] (checklist; faunal records; Japan (Kyushu)). MIWA, 1940: 72 [#4586] (checklist; Japan). YUASA, 1949: 128 [#8108] (characters; Japan). CHÛJÔ & KUROSAWA, 1950: 9 [#501] (catalog; faunal records; Japan (Shikoku)). KUROSAWA, 1950: 1115 [#3274] (note). IGA, 1955: 79 [#3828] (iconography; Japan). KUROSAWA, 1956: 1115 [#8107] (characters; Japan). OBENBERGER, 1958: 237 [#2389] (faunistic redords; China). IGA, 1962: 79 [#3351] (iconography; Japan). KUROSAWA, 1963: 153 [#1618] (characters; Japan). KUROSAWA, 1974: 2 [#4576] (note). TÔYAMA, 1985: 29 [#3678] (iconography; Japan). HOU, 1987: 589 [#3490] (notes; Yunnan). PENG ZHONGLIANG, 1987: 355 [#2429] (checklist; China). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). PENG ZHONGLIANG, 1992: 400 [#2431] (characters; notes; Hunan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 185 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 32 [#4091] (checklist; Japan: Honshu, Shikoku, Kyushu, Izu Island, Tsushima; Taiwan; China; India (Kashmir)). AKIYAMA & OHMOMO, 2000: 88 [#6240] (faunal records; Japan (Hiroshima)). NAKAMURA, KAMEYAMA & KATAYAMA, 2001:66 (faunal records; Japan) [#8761] HOU TAOQIAN & ZENG RUI, 2002: 357 [#6626] (checklist; China: Hainan). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Hubei, Taiwan, Hunan, Yunnan). PENG ZHONGLIANG, 2002: 268 [#6628] (characters; Fujian). MÜHLE, 2003: 46 [#6155] (checklist; Taiwan). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2071 2072 [#8576] (world catalog).

Type material: Agrilus discalis Saunders, 1873 Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 45L) by present designation, sex not examined, (BMNH): “Type H. T. [p] [round label with red border] \ Japan, G. Lewis, 1910-320. [p] \ Grassy Banks April 1871 [h] \ Agr. discalis Es. Type [h] [Saunders’ MS]”. Described from unknown number of syntypes.

Diagnosis: 6.2-7.9 mm; Elytra carmineous, with with ornamental pubescence and obvious glabrous spot in distal half. Species group: Plagiatus. Relevant species: A. plagiatus Ganglbauer, 1889. Specimens examined: CHINA: Guizhou: 7 ♂, 1 ♀ (EJCB): “China Guizhou 21 26.V., 60 km N of Kaili, 1995, Shibing Yuntai Shan, E. Jendek & O. Šauša leg.”. Sichuan: 1 ♂ (EJCB): “China N Sechuan, Shangliang, 24 29.5.2002, lgt. E. Kučera leg.”. Taiwan: 1 (EJCB): “Lu Shan, Formosa, 6.v.1973, K. Matsuda”. JAPAN: Honshu: 1 (JMCZ): “Zenkotlonsen (Nagano) Japan, 10 V 1940”; 1 (JMCZ): “Japan, 2.5.[19]67, Sôia, Okayama Pr. T. Aono lgt.”; 1 (EJCB): “Japan, 3.5.1967, Sôja, Okayama pn., T. Aono lgt.”; 1 (EJCB): “Japan, Atsugi, 1976, Kanagawa pref., K. Akiyama leg., 10.2”; 1 (JMCZ): “Japan, 10.5.1979, Shishizuka, Tsuchiura city, K. Akiyama lgt.”; 1 ♂ (EJCB): “Yamanaka cho, Yokosuka shi, Kanagawa pref., 24 26.vii.1996, K. Ishida et al. leg.”; 2 ♂ (EJCB): “Japan, April 2001, Tsukuba, L. Roller leg.”; 1 (EJCB): “Japan, Honshu, Ibaraki pref., Tsukuba city, 25.xi.2004, L. Roller leg.”. Kyushu: 2 ♂, 4 ♀ (EJCB): “Medoro, Kami Agata cho, Tsushima Isls., Nagasaki Pref., Kyushu, Japan, 5.v.1993, Ryô Noda leg.”.

Adult occurence: 4-5-6-7-11 (n=15). Faunal records published by CHÛJÔ & MATUDA, 1940 [#500] and OBENBERGER, 1958 [#2389] are also included. Altitude range: ~1200 m (n=1) OBENBERGER, 1958 [#2389]. Host plants: Zelkova serrata: AKIYAMA & OHMOMO, 1997 [#4091]. − Celtis sinensis var. japonica: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: CHINA: Guizhou, Hainan, Hubei, Hunan, Sichuan, Taiwan, Yunnan. INDIA: Jammu and Kashmir. JAPAN: Honshu, Kyushu, Shikoku, Tsushima.

Agrilus diversornatus Jendek, sp. nov. Fig. 30E (habitus of HT)

Description: 6.8-7.1 mm (Holotype 7.1 mm). BODY: Shape: subparallel, Build: robust, Color (dorsally): bicolored, Color (dominant): black. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): deep, Epistoma: with raised upper margin. Frons: Shape: flat, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline. Eyes: Size: small, Shape: protruding from head outline, Lower margin: in line with antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at posterior margin, Anterior margin: narrower than posterior. Anterior lobe: moderate or obvious, Shape: subangulate, Position to anterior angles: at level with anterior pronotal angles or projecting beyond anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: blunt. Disk: Convexity: flat, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): deep, Anteromedial and posteromedial impressions: subequal in size. Prehumerus: obsolete or carinal, Shape: straight or bisinuate, Extent: to third of pronotal length, Modifications: with rudiment at anterior pronotal angle, Posterior end: distant from pronotal angle or margin, Anterior end: distant from pronotal marginal carina. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent,

90 Junction: absent or present, Narrowest point: at posterior angles or at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present or obsolete. ELYTRA: Color: bicolored, complementary with pubescence, Alternative color (location): apical portion, Humeral carina: absent. Apices: Arrangement: separate, Width: wide, Shape: arcuate. Elytral pubescence: ornamental, Color: bicolored. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): shallow, Emargination (width): moderate. Prosternal process: wide, Shape: subparallel, Sides: straight or arcuate, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. Diagnosis: frons flat; eyes very small; pronotum strongly transverse, moderately convex; anterior pronotal lobe angulate; elytra with bichromatic pubescence; elytral apices golden. Species group: Unassigned. Relevant species: Unknown, but somewhat similar to species around A. grisator Kerremans, 1893. Type locality: Eastern Russia, south Primorskiy kray, Lazovskii zapovednik, kordon Korpad’, 43°15’17’’N, 134°07’59’’E. Type specimens: Primary: Holotype (Fig. 30E), ♀, (ZIN): “Yu Primor’e, Lazovskii zapovednik, kordon Korpad’, 43°15’17’’N, 134°07’59’’E, 13-19.vii.2005, leg. K. Makarov [in Russian]”. Secondary: RUSSIA: Primorskiy kray: 1 ♀ PT (EJCB): “Prim. kr., s. Nadepedinskaya okr., 232km shirokolistvennyi les, 31.8.[19]76, V. Kuznecov [in Russian]”; 1 ♀ PT (ZIN): “Zap. Kedrovaya pad’, Khasansk. r-n, Primor’e, Sinëv, 9.08.[1]988 [in Russian]”; 1 PT (IMCM): “Primorskii krai, Lazovskii zapovednik, ~10 km SW of Sokolchi vill., kordon Korpad’, 11-13.VIII.2009, leg. V. Shokhrin (in Russian). I. Melnik collection”.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): semispherical, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: large, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: visually elongate or visually square, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate or obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles or projecting beyond anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: strongly convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow. Prehumerus: filamentary or carinal, Shape: bisinuate, Extent: beyond half of pronotal length, Posterior end: distant from pronotal angle or margin, Anterior end: close to pronotal marginal carina or joining pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length or at posterior third of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Shape: subangulate. Elytral pubescence: entire except glabrous part at apical half, Color: unicolored, Character: with spots of denser pubescence. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: large, Anterior margin: arcuate. Prosternal process: wide, Shape: narrowed, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: medial sulcus on ventrite 1, Apical margin of pygidium: extended into short protrusion, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: elongate.

Host plants: Sorbaria.

Diagnosis: elytral disk pubescent except for medial portion at apical half, elytral spots pubescent; prosternal lobe very large and markedly declivous. Species group: Spinipennis. Relevant species: A. zhui sp. nov.

Distribution: RUSSIA: Primorskiy kray.

Type locality: China, Shanxi province, Ming county.

Etymology: The specific name was derived from the Latin adjectives diversus (different) and ornatus (adorned); it refers to the bichromatic elytral pubescence of the species.

Type specimens: Primary: Holotype (Fig. 9J), ♂, (EJCB): “Ming county, Shangxi, 1992”. Secondary: CHINA: Shanxi: 1 ♀ PT (EJCB): “Ming county, Shangxi [sic!], 1992”; INDIA: West Bengal: 1 ♀ PT (MNHN): “British Bootang, Maria Basti, L. Durel”.

Adult occurence: 7-8 (n=4).

Agrilus dureli Jendek, sp. nov. Fig. 9J (habitus of HT); Fig. 32N (aedeagus)

Adult occurrence and Altitude range: Unknown. Host plants: Unknown.

Description: 8.9-9.9 mm (Holotype 8.9 mm). BODY: Shape: cuneiform, Build: robust, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Medial impression: present, Impression (extent): vertex and frons, Epistoma: with raised upper margin. Frons: Shape: flat, Outline: not protruding

Distribution: CHINA: Shanxi. INDIA: West Bengal. Etymology: Patronymic; the species is named in honor of L. Durel, the collector of the type specimens.

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA

Agrilus ecarinatus Marseul Fig. 21A (habitus); Fig. 36W (aedeagus); Figs. 45Q, 46S (types) ecarinatus Marseul, 1866 (Agrilus) MARSEUL, 1866: 439, 484 485 [#676] (description). GEMMINGER SAUNDERS, 1871: & HAROLD, 1869: 1439 [#1057] (catalog). 121 [#2552] (catalog). HEYDEN, 1881: 119 [#5548] (catalog). MARSEUL, 1887: 252 [#4707] (catalog). KERREMANS, 1892: 255 [#1397] (catalog). ABEILLE DE PERRIN, 1897: 7, 8 [#19] (characters in key). KERREMANS, 1903: 276 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 658 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 989 [#2191] (world catalog). COBOS, 1968: 390 [#580] (subgenus Agrilus; faunal records; Mongolia). COBOS, 1972: 46, 56 [#591] (subgenus Agrilus; faunal records; Mongolia). ALEXEEV, 1975: 157 [#52] (notes; faunal records; distributional overview; Mongolia). ALEXEEV, 1989: 481 [#64] (key to species; Khabarovsk; Amursk; Primorsk; Chita; Buryatia; Tuva; steppe Altay; North Korea; northeastern China; northeastern Mongolia). ALEXEEV & VOLKOVITSH, 1989: 317, 344, 356 [#3241] (characters in key; faunal records; Mongolia). ALEXEEV, 1998: 375 [#4322] (subgenus Xeragrilus; subgeneric classification). JENDEK, 1998: 321 [#4481] (lectotype designation). NONNAIZAB et al., 1999: 112 [#8247] (checklist; China: Inner Mongolia). JENDEK, 2005: 11 [#7217] (synonymy). JENDEK, 2006: 396 [#7441] (subgenus Xeragrilus; Palaearctic catalog). BELLAMY, 2008: 2078 2079 [#8576] (subgenus Xeragrilus; world catalog). VOLKOVITSH, 2009: 137 [#8850] (subgenus Xeragrilus; faunal records; Russia: Primorskiy kray). gracilicornis Ganglbauer, 1889 (Agrilus) GANGLBAUER, 1889: 22, 35 36 [#3427] (description). HEYDEN, 1893: 90 [#3450] (catalog). KERREMANS, 1903: 285 [#1441] (synonym of grassator; catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 658 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 991 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 355 [#2429] (checklist; China). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Gansu, Sichuan). JENDEK, 2005: 11 [#7217] (synonym of ecarinatus). JENDEK, 2006: 396 [#7441] (synonym of ecarinatus; Palaearctic catalog). BELLAMY, 2008: 2079 [#8576] (synonym of ecarinatus; world catalog). grassator Kerremans, 1903 (Agrilus; unnecessary replacement name for gracilicornis Ganglbauer) KERREMANS, 1903: 285 [#1441] (replacement name proposal). JAKOBSON, 1913: 798 [#4386] (synonym of gracilicornis; catalog). OBENBERGER, 1936: 991 [#2191] (synonym of gracilicornis). JENDEK, 2006: 396 [#7441] (synonym of ecarinatus; Palaearctic catalog). BELLAMY, 2008: 2079 [#8576] (synonym of ecarinatus; world catalog).

Type material: Agrilus ecarinatus Marseul, 1866 Type locality: Sibérie orientale. Type specimens: Primary: Lectotype (Fig. 45Q), sex not examined, designated by JENDEK (1998: 321) [#4481], (MNHN): “Transcriptio [p] Ecarinatus Mars. Type Sibérie or. [h] \ Muséum Paris 1952 Coll. R. Oberthur [p] [yellow label]”. Secondary: 1 PLT (MNHN). Described from unknown number of syntypes. Agrilus gracilicornis Ganglbauer, 1889 Type locality: Sze-tschuan [Dorf Hunnei-hu (Grenze der Provinzen Kan-ssu und Sze-tschuan)] [Note: Locality was complemented from the list of localities on page 21-22].

91 Type specimens: Holotype by monotypy (Fig. 46S), ♂, (ZIN): “Sze-Tchňan [sic!] 1885 G. Patanin [sic!] [p] 2224. VII. 85 [h] [underside] \ Agril. gracilicornis Gglb. Typ, [h] \ Coll. Semenov-Tian-Shansky [p] \ Holotypus [p] [red label]”. Described from 1 specimen. Agrilus grassator Kerremans, 1903 Type locality and specimens: See: Agrilus gracilicornis Ganglbauer, 1889. Diagnosis: 4.8-6.9 mm; body without tomentum; prehumerus missing; antennae and tarsi long; aedeagus very thin. Species group: Albogularis. Relevant species: A. macrotatopus Obenberger, 1930. Specimens examined: CHINA: Beijing: 2 (AGCM): “China, Beijing, Fragrant Hill, Bot. Garden, 2.VII.1992, L. Borowiec leg.”. Gansu: 1 ♀ (MNHN): “Kansou, Louochan, 9.VI.[19]18”; 19 (EJCB): “China Kansu Ponggartang 30.Jun 1992 M. Bok lgt.”; 16 (EJCB): “China Gansu prov. 120 km SW Lanzhou Ponggartang 30.VI. 2. VII.1992 Jaroslav Turna leg.”; 1 ♀ (EJCB): “China (Gansu), ca 25 km e. Xiahe, 2700 2900 m (Wagatan Timber), 30.VII. 2.VIII.1993, Heinz leg.”; 3 (EJCB): “China Gansu, Wudu, 1 7.6.1996, E. Kučera leg.”; 1 ♀ (EJCB): “China/Gansu, Tewo #227, 2800 3100 m, 21 24.VI.[19]88, leg. Eckweiler”. Hebei: 1 (EJCB): “China, NE Hebei, 30.V., Chengde env., 2000, 41.0N, 117.9E, Jaroslav Turna leg.”; 1 ♀ (EJCB): “China, Hebei prov., Chengde env., 30 May 2000, 300 km NNE of Beijing, Zd. Jindra lgt.”. Qinghai: 1 ♂ (MNHN): “Thibet, Kuku Nor, 3200 m, F. Hauser 1898”. Shaanxi: 2 ♂ (EJCB): “China, Shaanxi pr., Hua Shan, 17 21.VI.1991, R. Sauer lgt.”; 2 ♂ (EJCB): “China, Shaanxi, 1992, Huashan, 120 km E of, 3. 4.Jun 1992, Xian, M. Bok lgt.”; 1 ♀ (EJCB): “China, Shaanxi, 1992, 120 km E Xi an, Hua Shan, 3 4.vi., Jaroslav Turna”; 3 ♀ (JMCZ): “China c., 17 18.V., Ganguyi 110,3’ 36,8’, 35 km NE of Yanan, leg. J. Halada 1996”; 1 ♀ (EJCB): “China Shaanxi, Baiche, 32º42’N, 110º05’E, 13 17.6.[19]97, lgt. E. Kučera”; 1 ♂, 1 ♀ (EJCB): “China, Shaanxi, 26.VI.1998, Qing Ling Shan mts., 1500 m, Hou Zen Zi vill. env., 30 km SE of Taibai Shan Mts., O. Šafránek et M. Trýzna lgt.”; 2 (EJCB): “China Shaanxi, Wutai Shan, 4 7.7.2000, leg. E. Kučera”; 1 (EJCB): “China Shaanxi, Hunyuan, 16 18.7.2000, leg. E. Kučera”. Shanxi: 17 (EJCB): “China, Shanxi, 1200 m, Taibaishan, 20 km S Wangzhuangbu, 1998, Bolm lgt, 23 26 Jul”. Sichuan: 6 (EJCB): “China / NW Sichuan, 103.50, 33.10, Min Shan, 2500 4500 m, 14. 16.VII.1990, leg. J. Kolibáč”; 3 (EJCB): “China, NW Sichuan prov., Maowen, 1000 m, 31º30’N 103º50’E, 10 18.vii.1990, J. Kolibáč leg.”; 1 ♂ (EJCB): “CHINA N, Sichuan, Jiuzhaigou, 2000m, 16.Jun 1992, M. Bok lgt.”; 2 ♂ (EJCB): “China, N Sichuan 30 km W Nanping 11.,16.VI.1992 Jiuzhaigou, 2100 m, Jaroslav Turna leg.”; 1 (AGCM): “China, N. Sichuan, env. Nanping, h 1800 m, 15.7.2002 leg. A. Gorodinski”. Xizang: 1 ♂ (EJCB): “E Tibet, Qamdo E env., 31º09’N, 97º11’E, cca 3500 m, 15.VII.1997, cultural steppe, M. Trýzna et O.Šafránek lgt.”. KAZAKHSTAN: 1 ♀ (JMCZ): “Ilijsk, 26.V.1958 g, Zhelokhovcev [in Russian]”. KOREA NORTH: 1 ♀ (ZIN): “S Koreya, Saryon, prov. Khoank, 21.VII.1950 [in Russian]”; 1 (VSCS): “Korea, Onpha ri, Kreis Kjôngsông, 12.6.1975, Josifov leg.”. KOREA SOUTH: 2 (JLCS): “Mt. Tuwibong, Gangwon Prov., Korea, 21.vi.2000, S. L. An leg.”. KYRGYZSTAN: 1 ♂ (ZIN): “ushch. r. Agamedin, boki Yu Frunze, Kirgiziya, Volkovich, 7 8.VII.1984 [in Russian]”. MONGOLIA: 2 (NMPC): “Mongolia, Chilgant, 29.7.1975”; 5 (EJCB): “Mongolia, Calgant, 10.vii.1975”; 1 (NMPC): “Mongolia, Ubur. Chong aj., Nugrin Els, 19.8.1979 [in Russian]”; 1 (EJCB): “Mongolia Central aimak, Tuul valley, Nalaikha, 1600 1800 m, 6 8.7.1984, Leg. Černý”; 3 (EJCB): “Mongolia or., Chentejský ajmak, dol. Mörön gol, B. Malec lgt. / 5 31.7.1985”; 1 ♂ (EJCB): “Mongolia bor., Džargalant, 10.7.[19]87, O. Majzlan lgt.”; 1 ♂, 1 ♀ (EJCB):

92 “Mongolia bor., Džargalant, 2.7.[19]87, O. Majzlan lgt.”; 1 ♂ (EJCB): “Mongolia bor., Džargalant, 8.VII.1987, O. Majzlan lgt.”; 4 (EJCB): “Mongolia, 5 9 July 2003, Cagaan nuga env., 1500 m, Segneger riv. vall., 35 km NE of Mandal ( Batsumber), cca 70 km N of Ulaanbaatar, Zd. Jindra lgt.”. Bayanhongor: 1 (NMPC): “Mongolia, Bayankhongor, 6.7.1975, 20 km SW lake Bon Cagan”. Hovd: 1 ♂ (NMPC): “Mongolia, Khovd ajmak, Bajtag Bogd Nuru, 25.vii.1975”. Suhbaatar: 1 ♂ (ZIN): “Mongoliya, Sukhè Bator. aim., peski Ongon èls, 15 km YUV Khongora, 5 6.VII.[1]971 [in Russian]”. RUSSIA: Altay Kray: 1 ♂, 1 ♀ (ZIN): “Altaisk. kr., okr. pos. Ko[?] Agan, Nuiskaya step’, 700, Kamenist. pustiny, Pisarenko, 18.VI.1989”. Amurskaya oblast’: 1 ♀ (ZIN): “Samodon, 100 km W Svobodnovo, Amur. obl., Zinovjev, 4.vii.1959”. Irkutskaya oblast’: 1 ♂ (JMCZ): “USSR, Bajkal mer., Nikola, 1 7.viii. K, Deneš lgt., 1984”; 1 ♀ (JMCZ): “USSR, Baical, 1977, Listvjanka, 26.6.”; 1 ♂ (ZIN): “Irkutskaya obl., Irkutsk. r n., 28.6.[19]87g, V. Kuznetcov [in Russian]”; 1 (FACF): “Russia Siberia S Baikal lake, Listvianka, 5.vIi.1990”; 1 (FACF): “Russia Siberia W Baikal lake, Sarma, 20.vi.1997”. Khabarovskiy kray: 1 (AGCM): “Russia, Khabarovsk reg., near Tarmanchukan st. 30.6.1997, leg. Pavlov. M.”. Primorskiy kray: 1 ♂, 1 ♀ (ZIN): “Nikolsk Ussurijsk, 16.IX.[19]31 Ryssakov”; 2 (ZIN): “Y Primor’e, sr. ter. r. Suchan, Kabakov, 10 20.7.1952 [in Russian]”; 1 (ZIN): “USSR, 29.VI.1983, Primorskij Kraj, Barabat Levada, leg. V. Kuznetsov”; 1 ♀ (EJCB): “SU Ussuri reg. Jasnoe 400 m 12 19.7.1989 M. Nikodým lgt.”; 1 ♂ (EJCB): “USSR Sibiria or., Sikhotealin range, Terney env., 23 30.7., St. Jakl lgt., 1989”; 1 ♀ (EJCB): “Russia, Primorsk, 21.VII., Benevskoe, 1989, R. Červenka leg.”; 1 ♂ (EJCB): “USSR Sibiria or., Primorski region, Dalnerechensk, 31.7., St. Jakl lgt., 1989”; 1 ♀ (EJCB): “SU Ussuri reg., Benevskoe, 20 21.7., lgt. S. Bečvář, [19]89”; 1 ♂ (EJCB): “SU Ussuri reg., Benevskoe, 20.7.1989, M. Nikodým leg.”; 3 ♂ (JMCZ): “SSSR or., 29 31.7., Primorskij Kraj, Pograničnyj, Sergejevka, 1990, lgt. M. Snížek”; 1 ♂, 1 ♀ (EJCB): “Sib. or. m., Primorje, Ussuri res., 20.7.1990, Kadlec + Voříšek lg.”; 2 ♂, 1 ♀ (EJCB): “USSR, Primorsk. kraj, Novocugujevka env., 16 18.VII.1990, leg. Snížek”; 1 ♂ (EJCB): “SSSR or., 20 28.7, Primorskij kraj, Tigrovij, 1990, Lgt. Snížek”; 1 ♂ (JMCZ): “SSSR or., 20 28.7, Primorskij kraj, Tigrovij, 1990, Lgt. Snížek”; 2 ♂ (EJCB): “Russia, Primorsky kray, Troitskoye, 2.VII.1991, V. Kuznetsov leg.”; 1 ♀ (JMCZ): “Ross. or. , Prim. kr., Slavjanka / Rjazanovka/ VII.1992, Snížek leg.”; 1 ♂, 1 ♀ (JMCZ): “Russia Far East. 19. 20.VII.1993, Primorsk region, Lazo, lgt. M. Trýzna”; 1 ♀ (EJCB): “Russia, Primorski kray, Khankajski distr. 1995, lake Khanka, 17 19.vii., Kamen’ Rybolov”. Tyva Respublika: 2 ♀ (EJCB): “Russia, C Tuva, Kyzyl env., 800 m, 14 15.VII.200[?], Vaschchenko A. N. leg.”; 2 ♂, 1 ♀ (EJCB): “Tuvinskaja ASSR, Saryg Sep, 23.6.[19]94, Z. Kletečka lgt.”; 5 (EJCB): “Tuva, 18 20. vi.1994, W of Tanu Ola, 30 km N Samagaltaj, Z. Kletečka leg.”; 1 ♂, 3 ♀ (EJCB): “Russia, Tuva, right bank of Biy Khem riv. 800 m, Ujukskiy Mts. 2 5.VII.1999, S. N. Vashchenko leg.”.

Adult occurence: 5-6-7-8-9 (n=71). Altitude range: 4004500 m (n=17). Host plants: Artemisia: ALEXEEV, 1975 [#52]. − A. rutifolia: ALEXEEV & VOLKOVITSH, 1989 [#3241]. − A. santolinifolia: ALEXEEV & VOLKOVITSH, 1989 [#3241]. − Nitraria: ALEXEEV & VOLKOVITSH, 1989 [#3241]. Distribution: CHINA: Beijing, Gansu, Hebei, Heilongjiang, Qinghai, Jilin, Liaoning, Nei Mongol, Shaanxi, Shanxi, Sichuan, Xizang. KAZAKHSTAN. KOREA NORTH. KOREA SOUTH. MONGOLIA. RUSSIA: Altay kray, Amurskaya oblast’, Buryatiya Respublika, Irkutskaya oblast’, Khabarovskiy kray, Primorskiy kray, Tyva Respublika, Zabaykalskiy kray.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Agrilus emmerichi Obenberger Fig. 11B (habitus of LT); Fig. 33B (aedeagus) emmerichi Obenberger, 1935 (Agrilus) OBENBERGER, 1935: 164 [#2158] (description). OBENBERGER, 1936: 989 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 355 [#2429] (checklist; China). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Sichuan). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2082 [#8576] (world catalog).

Type material: Agrilus emmerichi Obenberger, 1935 Type locality: Tatsienlu, Kiulung, China. Type specimens: Primary: Lectotype (Fig. 11B) by present designation, ♂, (NMPC): “China [p] \ Tatsienlu [p] \ Kiulung [p] \ Reitter [p] \ Typus [p] [red label] \ Agrilus Emmerichi m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 26478 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 8.3 mm (lectotype); prosternal process protruding (lateral view), distinctly impressed on disk, with elevated and denticulate sides. Species group: Quadrisignatus. Relevant species: A. quadrisignatus Marseul, 1866. Specimens examined: Known only from the lectotype.

Host plants: Unknown. Distribution: CHINA: Sichuan.

Agrilus erythrostictus Bourgoin Fig. 10E (habitus); Fig. 32T (aedeagus); Figs. 43I, 45X, (types) erythrostictus Bourgoin, 1922 (Agrilus) BOURGOIN, 1922: 169 [#366] (description). OBENBERGER, 1936: 1082 [#2191] (world catalog). DESCARPENTRIES & VILLIERS, 1963: 55, 58 [#699] ([Note: The original description neither implies not requires that there were syntypes, then Descarpentries & Villiers by the mention about “the holotype” validly fixed the lectotype (ICZN, Article 74.6.)]; lectotype designation; type locality determined by lectotype: “Tonkin: Nghia Lo (holotype au Muséum Paris)”; characters; Tonkin; Birmanie). JENDEK, 2005: 11 [#7217] (synonymy; notes). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2084 [#8576] (world catalog). barmensis Obenberger, 1936 (Agrilus; subspecies of poeta) OBENBERGER, 1936: 44 [#2200] (description). OBENBERGER, 1936: 1098 [#2191] (subspecies of poeta; world catalog). JENDEK, 2005: 11 [#7217] (synonym of erythrostictus; lectotype designation). JENDEK, 2006: 398 [#7441] (synonym of erythrosticus; Palaearctic catalog). BELLAMY, 2008: 2084 [#8576] (synonym of erythrostictus; world catalog).

Type material: Agrilus erythrostictus Bourgoin, 1922 Type locality: not given [Note: Indo-Chine française is cited in the title of the publication]. Type specimens: Primary: Lectotype (Fig. 45X), ♀, designated by DESCARPENTRIES & VILLIERS (1963: 55, 58) [#699], (MNHN): “Nghia Lo [h] \ Agrilus erythrostictus Bourgoin [h] Type [h] [red ink] \ Muséum Paris 1936 coll. A. Théry [p]”. Secondary: 1 PLT (BMNH). Described from unknown number of syntypes.

93

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Agrilus poeta barmensis Obenberger, 1936 Type locality: Burmah. Type specimens: Primary: Lectotype (Fig. 43I), ♀, designated by JENDEK (2005: 11) [#7217], (NMPC): “Barmah [h] \ Typus [p] [red label] \ A. poëta ssp. barmensis m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Diagnosis: 8.5-10.5 mm; head markedly large; frons conspicuously protruding from head outline; lateral sides of pronotum without obvious tomentose spots. Species group: Spinipennis. Relevant species: A. hmong sp. nov. Specimens examined: CHINA: Yunnan: 1 ♂ (NMPC): “Yun’nan’. Ganlanba, 540 m, 20.iii.1957, Pu Fu di [in Russian]”; 1 ♂ (EJCB): “China, Yunnan, Menglian County, Mt. Ka bie ke, 1 19.vi.2009, local collector”; 2 (CBCS): “Yunnan, Jing Gu County, Mt. Yingpantou, 7 14.vi.2009 local collector”. LAOS: Houaphan: 7 (EJCB): “NE Laos, Hua Phan prov., Ban Saluei, 1300 2000m, 20°13’N, 103°59’E, 6 18.iv., Phu Phan Mt. env. 2004, F. & L. Kantner leg.”; 3 ♂ (EJCB): “NE Laos, Hua Phan prov., 2004, 20°13’N, 103°59’E, 6 18.iv., Ban Saluei, Phu Phan Mt. env., 2004, J. Bezděk leg., 1300 2000m. Louangphrabang: 1 ♀ (EJCB): “Laos N, 24.iv. 16.v.1999, Louang Phrabang prov. 20°33 4’N 102°14’E, Ban Song Cha (5km W) ± 1200 m, Vít Kubáň leg.”. THAILAND: Mae Hong Son: 1 ♂ (EJCB): “NW Thailand, Mae Hong Son pr., 1500m, Soppong, 9 12.V., Sv. Bílý leg., 1996”. VIETNAM: Hoa Binh: 1 ♀ (MNHN): “Tonkin, region de Hoa Binh”.

Adult occurence: 4-5-6 (n=7). Altitude range: 570-2000 m (n=5). Host plants: Unknown. Distribution: CHINA: Yunnan. LAOS: Houaphan, Louangphrabang. MYANMAR, THAILAND: Mae Hong Son. VIETNAM: Hoa Binh.

1 ♀: “Hatsuno, Amami Is[land] [p] 26. V. [p] 19 [p] 60 [h] T. Shibata [p]”. Described from 10 specimens (HT, PT). Examined PT (Fig. 46A). See also Remarks. Agrilus toyoshimai Tôyama, 1988 Type locality: Hida-kanayama, Gifu Pref., Japan. Type specimens: Primary: Holotype (Fig. 53Y), ♀, (NSMT): “Hida - Kanayama Gifu pref. 27. vi. 1976 Toyoshima & Katō [h] \ Holotype [p] Agrilus toyoshimai Tôyama, 1987 [h] [red label]”. Described from 1 specimen (HT). Diagnosis: 5.8-7.3 mm; prehumerus absent; anterior pronotal lobe not extending beyond anterior pronotal angles; aedeagus (Fig. 39A) enlarged apically. Species group: Tempestivus. Relevant species: A. shibatai Kurosawa, 1964. Specimens examined: JAPAN: Honshu: 1 ♀ (SOCI): “1967.5.31., Nippara, Tokyo, C Japan, H.Ooki leg.”. Ryukyu isl. (Okinawa incl.): 4 (NSMT): “Inutabudake Mt., Tokunoshima, 10.V.~ 19.VII.1972, M. Takakuwa [leg.]”; 1 (EJCB): “Mt. Inokawa dake, Tokunoshima Is., 15.vi.1986 em, K. Hasokawa”; 1 ♂, 1 ♀ (EJCB): “Ohkuni Rindō, Okinawa hontō Is., iv.1989, M. Takeda \ Host: Machilus thunbergii”.

Adult occurence: 4-5-6-7 (n=11). Faunal records published by KUROSAWA, 1964 [#1619] are also included. Distribution: JAPAN: Honshu, Ryukyu Islands. Host plants: Castanopsis luchuensis: AKIYAMA & OHMOMO, 1997 [#4091]. − Machilus thunbergii (see Specimens examined). Remarks: Only two paratypes of A. esakii (see Type specimens) with a comment that the holotype has not been located in NSMT were sent for examination by S. Nomura in March, 2001.

Agrilus eunuchus Jendek, sp. nov. Agrilus esakii Kurosawa Fig. 25H (habitus); Fig. 39A (aedeagus); Figs. 46A, 53Y (types) esakii Kurosawa, 1964 (Agrilus) KUROSAWA, 1964: 323 325 [#1619] (description). KUROSAWA, 1974: 2 [#4576] (note). TÔYAMA, 1985: 29 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 32 [#4091] (checklist; Japan: Ryukyus (Amami oshima I., Tokunoshima I., Okinawa I.)). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2084 [#8576] (world catalog). toyoshimai Tôyama, 1988 (Agrilus) syn. nov. TÔYAMA, 1988: 752 753 [#2932] (description). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 232 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 41 [#4091] (checklist; Japan: Honshu). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2332 [#8576] (world catalog).

Type material: Agrilus esakii Kurosawa, 1964 Type locality: Gusuku-Nishinakama, Amami-Oshima. Type specimens: Primary: Holotype, ♂, not in NSMT (S. Nomura pers. com.). Secondary: 2 PT (NSMT): 1 ♀: “Ikari, Amami Is[land] [p] 17. V. [p] 19 [p] 60 [h] T. Shibata [p]”;

Fig. 29F (habitus of HT); Fig. 40U (aedeagus)

Description: 4.0-4.9 mm (Holotype 4.2 mm). BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): brown, rarely green. HEAD: Medial impression: absent or present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: in plane with frons. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (density): dense. Eyes: Size: moderate or large, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at anterior margin, Anterior margin: subequal to posterior. Anterior lobe: vague or moderate, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles or at level with anterior pronotal angles. Posterior angles: Shape: acute or rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial or subequal in size.

94 Prehumerus: filamentary or carinal, Shape: bisinuate, Extent: to half of pronotal length or beyond half of pronotal length, Modifications: with rudiment at anterior pronotal angle or obsolete in anterior part, Posterior end: joining posterior pronotal angle, Anterior end: distant from pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior third of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Shape: arcuate or subangulate. Elytral pubescence: entire except epipleura, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): shallow, Emargination (width): wide. Prosternal process: narrow or moderate, Shape: narrowed, Sides: straight or arcuate, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed, Ovipositor: elongate. Diagnosis: prehumerus filamentary or feebly carinal, extending almost to half of pronotal length; aedeagus markedly small. Species group: Roscidus. Relevant species: Agrilus ulmicola sp. nov. Type locality: China, Yunnan province, 100 km west of Kunming, Diaolin Natural Reservation. Type specimens: Primary: Holotype (Fig. 29F), ♂, (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”. Secondary: CHINA: Yunnan: 6 PT (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”. Adult occurence: 5-6 (n=1). Host plants: Unknown. Distribution: CHINA: Yunnan. Etymology: The specific name is the Latin noun eunuchus (castrated man) in apposition. It refers to the tiny aedeagus of the species.

Agrilus euonymi Tôyama Fig. 26E (habitus); Fig. 39G (aedeagus); Fig. 46C (type) euonymi Tôyama, 1985 (Agrilus) TÔYAMA, 1985: 28 30 [#2924] (description). TÔYAMA, 1985: 21 [#3678] (iconography; Japan). AKIYAMA, 1986: 25 [#3199] (faunal records; Japan: Kanagawa prefecture). HIRASHIMA, 1989: 323

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 32 33 [#4091] (checklist; Japan: Honshu, Kyushu, Oki Island). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2086 [#8576] (world catalog). yagii Tôyama, 1985 (Agrilus) TÔYAMA, 1985: 30 [#2924] (description). AKIYAMA & OHMOMO, 1997: 32 [#4091] (synonym of euonymi). JENDEK, 2006: 398 [#7441] (synonym of euonymi; Palaearctic catalog). BELLAMY, 2008: 2086 [#8576] (synonym of euonymi; world catalog).

Type material: Agrilus euonymi Tôyama, 1985 Type locality: Kinuta, Setagaya, Tokyo. Type specimens: Primary: Holotype (Fig. 46C), ♂, (NSMT): “Kinuta, Setagaya Tokyo [p], 18. VI. [p] 19 [p] 78 [h] leg. Y. Kurosawa [p] \ Holotype [p] Agrilus euonymi M. Tôyama, 1985 [h] [red label]”. Secondary: 6 PT (NSMT). Described from 93 specimens (HT, AT, PT).Agrilus yagii Tôyama, 1985 Type locality: Mt. Takahachi, Tottori Pref. Type specimens: Not located. Described from 2 specimens (HT, AT). Diagnosis: 4.8-6.8 mm; body more robust; elytral apical third broader; prosternal process more enlarged; basal ventrite in male with medial longitudinal impression. Species group: Cyaneovirens. Relevant species: A. tsushimanus Kurosawa, 1963. Specimens examined: CHINA: Henan: 1 ♂ (EJCB): “China, N. Henan, 11 12.VI., Jiuligou, 550 m, 35°12’N, 112°26’E, Jaroslav Turna leg. 2007”; Shaanxi: 5 (EJCB): “China, Shanxi [should be Shaanxi], Xian City, Peng leg.”; 1 ♂, 1 ♀ (EJCB): “China, Shaanxi pr., Hua Shan, 17 21.VI.1991, R. Dunda lgt.”; 2 (EJCB): “China, Shaanxi prov., 21 23.June 1998 \ Qing Ling Shan mts, 1000 m road Baoji Taibai vill., pass 40 km S Baoji, Zd. Jindra lgt.”; 12 (EJCB): “China, Shaanxi prov., 21 23.vi.1998, Quin Ling Shan mts, road Baoji Taibai vill., pass 35 km S Baoji, O. Šafránek et M. Trýzna lgt.”. Shanghai: 1 ♂ (EJCB): “Shanghai”. JAPAN: Honshu: 1 (EJCB): “K. Tsuji, 1968.6.22”; 2 (EJCB): “Kinuta, Setagaya, Tokyo, 2.vi.1974, leg. Y. Kurosawa”. KOREA NORTH: 1 (EJCB): “Korea, Mts. Mjohjang san, leg. M. Mroczkowski et Riedel, distr. Hjang san, Hjangam ri, 22.vi.1965”; 1 (EJCB): “N. Korea, 29.7. Sinpkho env., S. Murzin, 1990”; 1 ♀ (JMCZ): “N. Korea, 29.7. Sinpkho env., S. Murzin, 1990”. KOREA SOUTH: 1 (JLCS): “Mt. Baekun, Gyeonggi Prov., Korea, 20.vii.1986, S. L. An leg.”; 2 (JLCS): “Korea, Chungnam Prov. Gongiu City, Sangsin ri, Mt. Gyeryongsan, N 36°22’03.2’’ E 127°12’50’’, 31.v. 18.vi.2004, SM Choi, JS Park ex FIT”; 1 (JLCS): “Hyeonnae ri, Mupung myeon, Muju gun, Jeollabuk do, Korea, 13.v.2005, coll. Jung gu Lee”. RUSSIA: Primorskiy kray: 1 ♂ (EJCB): “Russia, Primorsk reg., 30.7. 8.8.1992, Kajmanovka, E. Kučera leg. ex Euonymus”.

Adult occurence: 5-6-7-8 (n=13). Faunal record published by TÔYAMA, 1985 [#2924] (as yagii) is also included. Altitude range: 550-1000 m (n=2). Host plants: Euonymus japonicus AKIYAMA & OHMOMO, 1997 [#4091]. − Prunus yedoensis: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: CHINA: Henan, Shaanxi, Shanghai. JAPAN: Honshu, Kyushu. KOREA NORTH. KOREA SOUTH. RUSSIA: Primorskiy kray.

95

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA

Agrilus fareastensis Jendek Fig. 17F (habitus of HT); Fig. 35K (aedeagus) fareastensis Jendek, 1995 (Agrilus) JENDEK, 2006: 398 JENDEK, 1995: 177 [#3767] (description). [#7441] (Palaearctic catalog). BELLAMY, 2008: 2091 [#8576] (world catalog). VOLKOVITSH, 2009: 136 [#8850] (subgenus Robertius; faunal records; Russia: Primorskiy kray).

Type material: Agrilus fareastensis Jendek, 1995 Type locality: Russia, Khabarovskiy region, 30 km S Khabarovsk, Khekhcir Mt. Type specimens: Primary: Holotype (Fig. 17F), ♂, (EJCB): “Sib. or. 7.1977 Chechcir chrebet Ing. Gotwald lgt.”. Secondary: 3 PT (EJCB). Described from 4 specimens (HT, PT). Diagnosis: 5.4-7.1 mm; pronotum widest at middle; less transverse; elytral apices separately subtruncate. Species group: Betuleti. Relevant species: A. betuleti (Ratzeburg, 1837). Specimens examined: CHINA: Hebei: 1 (EJCB): “China, Hebei/ Nei Mongol, pass Chengde Chifeng, 41.6N, 118.2E, 14 16.VI.2001, O. Šafránek leg.”; 2 ♀ (EJCB): “China, Hebei/Nei Mongol, pass Chengde Chifeng, 41.6N, 118.2E, 14 16.VI., Jaroslav Turna leg. 2001”. RUSSIA: Primorskiy kray: 1 ♀ PT (HMMG): “Primorje” (JENDEK, 1995 [#3767]); 1 ♀ (EJCB): “DVK, Sedanka, 25 vii (?vi) 57, Filimonov [in Russian]”; 1 ♀ PT (EJCB): “Primorie zap. Kedrovaja Pad 30.V.1983 Zlobin [leg.]” (JENDEK, 1995 [#3767]); 1 ♀ PT (EJCB): “Sib. or. m. Primorje Sichote Alin Mts. Sokolči 1 15.7.1990 Kadlec + Voříšek lg.” (JENDEK, 1995 [#3767]); 1 ♀ (EJCB): “Rossia, Primorsk, Kraskino env. mer., 13 16.VII.1992, P. Smrž leg.”; 1 ♂ (EJCB): “Russia, Primorski kray, Khankajski distr. 1995, lake Khanka, 17 19.vii., Kamen’ Rybolov”; 1 ♀ (EJCB): “Russia E, Primorje reg, Anisimovka, 7 13.vii.1996, Mt. Litovka, 200 m, Plutenko leg.”; 1 (MNCA): “Russia, Ussuri, Primor’e region, Pedan Mts., 11 17.7.2006, leg. K. Hadulla”; 1 ♂ (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Preobrazhenie env. 9 11.07.06 leg. М. & L. Smirnov’s”.

Adult occurence: 5-6-7 (n=9). Altitude range: ~200 m (n=1). Host plants: Unknown. Distribution: CHINA: Hebei, Nei Mongol. RUSSIA: Khabarovskiy kray, Primorskiy kray.

Agrilus fevreae Descarpentries & Villiers Fig. 25E (habitus); Fig. 38X (aedeagus); Fig. 46E (type) fevreae Descarpentries & Villiers, 1963 (Agrilus) DESCARPENTRIES & VILLIERS, 1963: 107, 114 [#700] (description). BELLAMY, 2008: 2093 [#8576] (world catalog).

Type material: Agrilus fevreae Descarpentries & Villiers, 1963 Type locality: Tonkin: Hoa Binh. Type specimens: Primary: Holotype (Fig. 46E), ♂, (MNHN): “Tonkin Région de Hoa-Binh \ Type [p] ♂ [h] [red label] \ Agrilus fevreae n. sp. Holotype ♂ nob. A. Descarpentries et [h] A. Villiers det. 19 [p] 62 [h]”. Described from 2 specimens (HT, PT).

Diagnosis: 3.7-4.2 mm; body dark-brown; elytra with proximal and distal perisutural pubescence; groove on apex of last ventrite arcuate; prosternal process strongly enlarged, with short erect pubescence in male; aedeagus with oval sides. Species group: Tempestivus. Relevant species: A. chekiangensis Gebhardt, 1929; A. malloti Théry, 1930. Specimens examined: CHINA: Guizhou: 4 ♂, 2 ♀ (EJCB): “CH Guizhou NE, 27.V. 3.VI., 20 km NW of Jiangkou, 1995, Fanjing Shan Kuaichang, E. Jendek & O. Šauša leg.”. LAOS: Houaphan: 1 ♀ (EJCB): “NE Laos, Hua Phan prov., 2004, 20°13’N, 103°59’E, 6 18.iv., Ban Saluei, Phu Phan Mt. env., 2004, J. Bezděk leg., 1300 2000m”; 3 ♀ (EJCB): “NE Laos, Hua Phan prov., Ban Saluei, 1300 2000m, 20°13’N, 103°59’E, 6 18.iv., Phu Phan Mt. env. 2004, F. & L. Kantner leg.”; 31 (EJCB): “Laos NE, Houa Phan prov., 20°12 13.5’N, 103°59’.5 104°01’E, Ban Saluei → Phou Pane Mt., 1340 1870m, 15.iv. 15.v.2008, Lao collectors leg.”. VIETNAM: Gia Lai: 1 ♂ (EJCB): “S Vietnam, 14.10N 108.30E, 40 km NW Ankhe, Buon Luoi, 620 750 m, 28.3. 12.4.1995, Pacholátko & Dembický leg.”.

Adult occurence: 3-4-5-6 (n=5). Altitude range: 620-2000 m (n=4). Host plants: Unknown. Distribution: CHINA: Guizhou. LAOS. VIETNAM.

Agrilus fissus Obenberger Fig. 24A (habitus); Fig. 38H (aedeagus); Fig. 46H (type) fissus Obenberger, 1917 (Agrilus) OBENBERGER, 1917: 214 215 [#1997] (description). OBENBERGER, 1924: 47 [#2040] (note). OBENBERGER, 1926: 660 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 990 [#2191] (world catalog). KUROSAWA, 1974: 4 [#4577] (synonym of rotundicollis). COBOS, 1986: 231 [#609] (notes). PENG ZHONGLIANG, 1987: 355 [#2429] (checklist; China). JENDEK, 1995: 146 147 [#3768] (lectotype designation; synonymy; characters; Russian Far East; northeastern and central China). AKIYAMA & OHMOMO, 1997: 36 [#4091] (synonym of moerens). ALEXEEV, 1998: 373 [#4322] (subgenus Quercuagrilus; subgeneric classification). HUA LI ZHONG, 2002: 89 [#6696] (checklist; E China). JENDEK, 2006: 391 [#7441] (subgenus Quercuagrilus; Palaearctic catalog). BELLAMY, 2008: 2095 [#8576] (subgenus Quercuagrilus; world catalog). VOLKOVITSH, 2009: 135 [#8850] (subgenus Quercuagrilus; faunal records; Russia: Primorskiy kray). LEE, KIM & AHN, 2010: 264 266 [#8939] (characters; faunal records; South Korea). lubricus Théry, 1942 (Agrilus) THÉRY, 1942: 260 262 [#2875] (description). OBENBERGER, 1943: 10 [#2333] (synonym of fissus). JENDEK, 1995: 146 [#3768] (synonym of fissus). JENDEK, 2006: 391 [#7441] (synonym of fissus; Palaearctic catalog). BELLAMY, 2008: 2095 [#8576] (synonym of fissus; world catalog).

Type material: Agrilus fissus Obenberger, 1917 Type locality: China. Type specimens: Primary: Lectotype (Fig. 46H), ♂, designated by JENDEK (1995: 146-147) [#3768], (NMPC): “Kiantschou [= former colony of Germany near Qingdao, Shandong province] China [h] \ Typus [p] [red label] \ Agrilus fissus m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24877 [h] [orange label]. Described from unknown number of syntypes.

96 Agrilus lubricus Théry, 1942 Type locality: Erzendjanzcy, Mandchourie. Type specimens: Not located. Described from 1 specimen (Holotype by monotypy). Diagnosis: 3.7-4.2 mm; pronotum with weak lateral impressions; elytral apices broad; aedeagus strongly asymmetric Species group: Sulcicollis. Relevant species: A. truncatus Jendek, 2007. Specimens examined: CHINA: Beijing: 5 (EJCB): “China, Beijing distr., Yan Shan, Dongling Mts., 39°58’N, E 115°26’E, 1400 m, Xiaolongmen, 15 16.VI.2001”. Shaanxi: 1 ♂ (EJCB): “China S, Shaanxi, Qinling Mts. N. slope, Changan Co., 33.56 59’ / 108.50’ 800 1200 m, 14 16.6.1995, L.+R. Businský lgt.”; 1 ♀ (EJCB): “China Yunnan [sic!][should be Shaanxi], Lüyeang, 25.7. 30.7.1996, E. Kučera leg.”; 1 (EKCS): “China Shaanxi, Lüeang 33°07’N 106°05’E, 18.6. 24.6.[19]97 lgt. E. Kučera”; 5 ♀ (EJCB): “China, 1000 1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v. 13.vi.1998, I. H. Marshal leg.”; 3 ♂, 1 ♀ (EJCB): “China, 1150 1300 m, Shaanxi, Qinling mts., Foping (6 km N), 20 21.VI.1998, I. H. Marshal leg.”; 1 ♂ (EJCB): “China, Shaanxi prov., 18.July 1998, Qing Ling Shan mts, road Baoji Taibai vill., cca 35 km S Baoji, Zd. Jindra lgt.”; 1 ♀ (EKCS): “China Shaanxi, Lueyang, 18.5. 21.5.2000, lgt. E. Kučera”; 1 ♂ (EJCB): “China Shaanxi, Lüeang, 22.5. 25.5.2000, leg. E. Kučera”; 1 ♂, 2 ♀ (EJCB): “C China, Shaanxi, Qinling, Shan, 6 km N of Foping, 1150 1300m, 20 21.VI.2000, leg. C. Holzschuh”; 1 ♀ (EJCB): “China Shaanxi, Čun Čan, 26.5. 1.6.2000, leg. E. Kučera”; 2 ♂ (EJCB): “China Shaanxi, Lueang env., 15 km NW, 26.5. 31.5.2001, lgt. E. Kučera”; 1 (EKCS): “China Shaanxi, Lueyang, 23.6. 26.6.2004, lgt. E. Kučera”; 7 (EKCS): “China Shaanxi, Lueyang, 29.5. 2.6.2005, lgt. E. Kučera”; 1 (EKCS): “China Shaanxi, Lueyang, 23 30.5.2009, lgt. E. Kučera”. Shanxi: 1 ♂, 1 ♀ (JMCZ): “China c. 29 30.v., Jingangling, 111.7’ 50km W Linfen, 36.2’, leg. J. Halada 1996”; 13 (EJCB): “China, W Shanxi, 9.VI.2000, 37.7N, 111.6E, Lüliang Shan, road Fangshan Jiaocheng, Hengjian env., ~1000m, Jaroslav Turna leg.”; 10 (EJCB): “China, Shanxi prov, 9.VI.2000, Lüliang Shan, 1000 m, road Fangshan Jiaocheng, Hengjian env., Zd. Jindra”. Sichuan: 1 ♀ (EJCB): “China / NW Sichuan, 103.50, 33.10, Min Shan, 2500 4500 m, 14. 16.VII.1990, leg. J. Kolibáč” (JENDEK, 1995 [#3768]); 2 ♂ (EKCS): “China N, Sechuan, Nanjiang, 21 23.5.2002, lgt. E. Kučera”. KOREA SOUTH: 1 (JLCS): “Geumhyeon ri, Soheul eup, Pocheon gun, Gyeongi do, Korea, 11.vi.2006, coll. Jung gu Lee”; 1 (JLCS): “Gwansan dong, Deokyang gu, Goyang si, Gyeonggi do, Korea, 3.viii.2006, coll In seong Yoo”. RUSSIA: Khabarovskiy kray: 1 (EJCB): “Sib. or., Chechcir chrebet, 7.1977, leg. Gottwald” (JENDEK, 1995 [#3768]). Primorskiy kray: 1 (EJCB): “Russia, Primorskiy region, Vladivostok, 20.VI.1927, leg. V. Belov” (JENDEK, 1995 [#3768]); 1 ♂, 1 ♀ (EJCB): “USSR, Primorsk. kraj, Arsenev env., VI.1991, 44.11N, 133.19E, leg. O. Šauša” (JENDEK, 1995 [#3768]); 1 ♂ (EJCB): “USSR, Primorsk. kraj, Arsenev env., VI.1991, 44.11N, 133.19E, leg. M. Štrba” (JENDEK, 1995 [#3768]); 1 ♀ (EJCB): “Russia, Primorski reg., Lazovskiy reserve, 11 km S Staraya Kamenka , 10.07.2002, Filimonov leg.”.

Adult occurence: 5-6-7-8 (n=27). Altitude range: 8004500 m (n=8). Host plants: Unknown. Distribution: CHINA: Beijing, Heilongjiang, Shaanxi, Shandong, Shanxi, Sichuan. KOREA NORTH. KOREA SOUTH. RUSSIA: Khabarovskiy kray, Primorskiy kray.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Agrilus fleischeri Obenberger Fig. 9G (habitus); Fig. 32M (aedeagus); Figs. 46I, 48F, 54B (types) fleischeri Obenberger, 1925 (Agrilus) OBENBERGER, 1925: 33 34 [#2048] (description). OBENBERGER, 1926: 653 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 990 [#2191] (world catalog). STEPANOV, 1954: 117 118, 119 [#2662] (subgenus Epinagrilus; redescription; characters in key; Zabaikal’e). KUROSAWA, 1963: 152 [#1618] (characters; Japan). KUROSAWA, 1974: 2 [#4576] (note). TÔYAMA, 1985: 24 [#3678] (iconography; Japan). VOLKOVITSH & ALEXEEV, 1988: 44 [#2972] (subspecies of ater; cited as fleisheri). ALEXEEV, 1989: 482 [#64] (subspecies of ater; cited as fleisheri; key to species; Khabarovsk; Amursk; Primorsk; Chita; Buryatia). LI JINGKE, 1992: 92 [#6755] (checklist; northeastern China). AKIYAMA & OHMOMO, 1997: 33 [#4091] (checklist; Korea; North China; Russia: Sakhalin, East Siberia). HUA LI ZHONG, 2002: 89 [#6696] (cited as fleisheri; checklist; northeastern China). JENDEK, 2005: 11 [#7217] (lectotype designation; synonymy). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2095 2096 [#8576] (world catalog). VOLKOVITSH, 2009: 136 [#8850] (subgenus Uragrilus; faunal records; Russia: Primorskiy kray). kurosawai Obenberger, 1940 (Agrilus; subspecies of fleischeri) OBENBERGER, 1940: 26 [#2287] (description). MIWA & CHÛJÔ, 1940: 74 [#1810] (subspecies of fleischeri; faunal records; Sakhalin: Konuma). KUROSAWA, 1974: 2 [#4576] (subspecies of fleischeri). JENDEK, 2005: 11 [#7217] (synonym of fleischeri; lectotype designation). JENDEK, 2006: 398 [#7441] (synonym of fleischeri fleischeri; Palaearctic catalog). BELLAMY, 2008: 2096 [#8576] (synonym of fleischeri; world catalog). kochi Théry, 1942 (Agrilus) THÉRY, 1942: 253 254 [#2875] (description). OBENBERGER, 1943: 9 [#2333] (synonym of fleischeri). JENDEK, 2006: 398 [#7441] (synonym of fleischeri fleischeri; Palaearctic catalog). BELLAMY, 2008: 2096 [#8576] (synonym of fleischeri; world catalog). tscherepanovi Stepanov, 1954 (Agrilus; subgenus Epinagrilus) STEPANOV, 1954: 115 116, 119 [#2662] (description). ALEXEEV, 1998: 372 [#4322] (subgenus Uragrilus; subgeneric classification). JENDEK, 2005: 11 [#7217] (synonym of fleischeri). JENDEK, 2006: 398 [#7441] (synonym of fleischeri fleischeri; Palaearctic catalog). BELLAMY, 2008: 2096 [#8576] (synonym of fleischeri; world catalog). nipponicola Kurosawa, 1963 (Agrilus; subspecies of fleischeri) syn. nov. KUROSAWA, 1963: 111 [#1617] (description). KUROSAWA, 1954: 92 [#1612] (as fleischeri; faunal records; Japan). KUROSAWA, 1963: 152 [#1618] (subspecies of fleischeri; characters; Japan). KUROSAWA, 1974: 2 [#4576] (subspecies of fleischeri). TÔYAMA, 1985: 24 [#3678] (subspecies of fleischeri; iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (subspecies of fleischeri; checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (subspecies of fleischeri; checklist; Japan). AKIYAMA & OHMOMO, 1997: 33 [#4091] (subspecies of fleischeri; checklist; Japan: Hokkaido, Honshu). FUKUTOMI & HORI, 2004: 11 [#6851] (subspecies of fleischeri; checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 398 [#7441] (subspecies of fleischeri; Palaearctic catalog). BELLAMY, 2008: 2096 [#8576] (subspecies of fleischeri; world catalog).

Type material: Agrilus fleischeri Obenberger, 1925 Type locality: Berezovka (Transbaikalia).

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TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Type specimens: Primary: Lectotype (Fig. 46I), ♂, designated by JENDEK (2005: 11) [#7217], (NMPC): “Verozovka Transbaik. [h] \ Typus [p] [red label] \ A. fleischeri m. Type [h] Det. Dr. Obenberger [p]”. Described from unknown number of syntypes. Agrilus fleischeri kurosawai Obenberger, 1940 Type locality: Sachalin. Type specimens: Primary: Lectotype (Fig. 48F), ♂, designated by JENDEK (2005: 11) [#7217], (NMPC): “Saghalien K. Tamanuku [p] [reverse side] Konuma 19.vii.1930 [h] \ Typus [p] [red label] \ A. Fleischeri ssp. Kurosawai m. Type [h] Det. Dr. Obenberger [p]”. Described from unknown number of syntypes. Agrilus kochi Théry, 1942 Type locality: Mandchourie, Weischadre. Type specimens: Not located. Described from 5 syntypes. Agrilus tscherepanovi Stepanov, 1954 Type locality: v Tuvinskoi oblasti v urochische Chaa-Khol’ [in Russian]. Type specimens: Primary: Not located. Secondary: 1 PT (Fig. 54B),(SZMN). Described from 2 specimens (HT, PT). Agrilus fleischeri nipponicola Kurosawa, 1963 Type locality: Aomori Pref.: Lake-side of Towada-ko. Type specimens: Not located. Described from 2 specimens (HT, PT). Diagnosis: 7.3-11.7 mm; body darker, less lustrous; prehumerus shorter, with anterior apex distant from lateral pronotal margin; elytral spots prolonged and less tomentose; adsutural spots at apical third of elytra in form of “V”; elytral apex more pubescent; elytral apex subangulate or acuminate with top shifted laterally. Species group: Spinipennis. Relevant species: A. ater (Linné, 1767). Specimens examined: CHINA: Beijing: 1 (EJCB): “China, Beijiing Shi, 28.v., 60 km NW Beijing, 2000, Badaling, 40.3N 116.0E, Jaroslav Turna leg.”; Heilongjiang: 1 ♀ (MNHN): “Pingchiang, N. Manchuria, vii.1939, 2000 ft”; 1 (ZIN): “st. Gaolinczy, vostochnaya liniya KV zhd., Kitai, 30.vi.1940 [in Rusian]”; 1 ♀ (ZIN): “China, Harbin, 1.vii.1994, V. Kuznetsov”. Shaanxi: 1 ♀ (EJCB): “China, Shaanxi prov., 21 23.vi.1998, Quin Ling Shan mts, road Baoji Taibai vill., pass 35 km S Baoji, O. Šafránek et M. Trýzna lgt.”. Sichuan: 2 ♂ (EJCB): “China Sechuan, Juizhaigou, 11.6. 16.6.2001 leg. E. Kučera”. JAPAN: Hokkaido: 1 ♂ (EJCB): “1998.7.15, Kasuga, Asahikowa, Hokkaido, N Japan, K. Kume leg.”; 1 ♂ (EJCB): “1998.7.15, Kasuga, Asahikowa, Hokkaido, N Japan, K. Kume leg.”. KAZAKHSTAN: 1 ♂ (EJCB): “Kazakhstan S Altai Mts, Ognevka, 19.vi.2001”. MONGOLIA: 2 ♀ (EJCB): “Mongolia, 14 20.7.2003, 20km E Terelzh, 1200 1400m, 60km NE of Ulaanbaatar, Tuul riv.valley Terelzh N.P., Z. Jindra lgt.”; 1 ♂, 3 ♀ (EJCB): “Mongolia, 100 km E Ulanbatar, 20 km NE, Tereltz, Tuul riv., 15 21.vii.2003, J. Halada leg.”. RUSSIA: Primorskiy kray: 1 ♂ (ZIN): “Primor’e Pivan’ 20.6.1976 Kabakov [in Russian]”; 1 ♀ (JMCZ): “Ross. or., Prim. kr., Partizansk, 20.vi.1981, V. Navišov leg.”; 11 (EJCB): “USSR, Primorskij kraj Arseneev env. 300 400 m 11. 19.VII.1990 B. Šiška & R. Čermák leg.”; 1 ♀ (EJCB): “Sib. or. Primorje, Komarovka flum., Kamenushka env. 300 m, Voříšek, J. leg. VII.1992”; 4 (EJCB): “Russia, Primorsk. prov. Arsenev env., VII.1992 B. Šiska leg.”; 1 (EJCB): “SSSR, Primorskij k., Kajmanovka 10 22.vii.1993, Z. Kletečka leg.”; 3 (EJCB): “Russia, Primorski kray, Chassanski district, Res. Kedrovja Padj, 22 27. vi.1995”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., 12 кm S Staraja Kamenka, 7.07.04 leg. М.& L. Smirnov’s”. Tyva

Respublika: 2 ♀ (EJCB): “Tuva, 25.6.1979, Ishtii Khem, S. Korolev leg.”.

Adult occurence: 5-6-7-8 (n=22). Faunal records published by Kurosawa, 1963 [#1617] (as fleischeri nipponicola) and MIWA & CHÛJÔ, 1940 [#1810] (as fleischeri kurosawai) are also included. Altitude range: 300-1400 m (n=3). Host plants: Populus: STEPANOV, 1954 [#2662]; ALEXEEV, 1989 [#64]. – P. laurifolia: STEPANOV, 1954 [#2662] (as tscherepanovi). – P. sieboldi: AKIYAMA & OHMOMO, 1997 [#4091]. − Salix ALEXEEV, 1989 [#64]. Erroneous: Quercus ALEXEEV, 1989 [#64]. Distribution: CHINA: Beijing, Heilongjiang, Shaanxi, Sichuan. JAPAN: Hokkaido, Honshu. KAZAKHSTAN. KOREA NORTH. KOREA SOUTH. MONGOLIA. RUSSIA: Amurskaya oblast’, Buryatiya Respublika, Khabarovskiy kray, Primorskiy kray, Sakhalinskaya oblast’, Tyva Respublika, Zabaykalskiy kray. Remarks: KUROSAWA, 1963 [#1617] proposed the subspecies nipponicola for the specimens from Japan (Hokkaido, Honshu) having “sides of pronotum slightly but distinctly sinuate just before the base causing the posterior angles rectangular but not obtuse; prehumeral carinae obsolete, not so strongly elevated”. These characters are well within the species variability as seen among the mainland specimens.

Agrilus foliatus Jendek Fig. 23H (habitus of HT); Fig. 38C (aedeagus) foliatus Jendek, 2007 (Agrilus) JENDEK, 2007: 40 41 [#8006] (description). [#8576] (world catalog).

BELLAMY, 2008: 2097

Type material: Agrilus foliatus Jendek, 2007 Type locality: China, Guizhou NE, 20 km NW of Jiangkou, Fanjing Shan-Kuaichang. Type specimens: Primary: Holotype (Fig. 23H), ♂, (EJCB): “CH-Guizhou NE 27.V.-3.VI. 20 km NW of Jiangkou 1995 Fanjing Shan-Kuaichang”. Secondary: 9 PT (EJCB). Described from 10 specimens (HT, PT). Diagnosis: 3.9-5.5 mm; elytra with pubescence in proximal half; basal ventrite in male with tubercles. Species group: Sulcicollis. Relevant species: A. takahashii Tôyama, 1988. Specimens examined: CHINA: Guizhou: 2 ♂ PT, 4 ♀ PT (EJCB): “CH Guizhou NE, 27.V. 3.VI., 20 km NW of Jiangkou, 1995, Fanjing Shan Kuaichang” (JENDEK, 2007 [#8006]). Shaanxi: 2 ♀ PT (EJCB): “China Shaanxi, Lüeang 33°07’N, 106°05’E, 18.6. 24.6.[19]97” (JENDEK, 2007 [#8006]). Sichuan: 1 ♂ PT (EJCB): “China, N Sechuan, Shangliang, 24 29.5.2002” (JENDEK, 2007 [#8006]).

Adult occurence: 5-6 (n=3). Host plants: Unknown. Distribution: CHINA: Guizhou, Shaanxi, Sichuan.

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Agrilus foramenifer Jendek, sp. nov. Fig. 28H (habitus of HT); Fig. 40L (aedeagus)

Description: 3.6-4.7 mm (Holotype 4.4 mm). BODY: Shape: subparallel, Build: robust, Color (dorsally): unicolored, Color (dominant): green, brown, rarely blue. HEAD: Shape: markedly convex, Medial impression: absent, rarely present, Impression (depth): shallow. Frons: Shape: moderately convex or markedly convex, Outline: protruding from head outline or not protruding from head outline. Vertex: Outline: protruding from head outline or not protruding from head outline, Sculpture on vertex: punctures or rugae. Eyes: Size: large, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short or moderate, Sexual modification in male: not apparent or antennae longer. PRONOTUM: Shape: visually elongate or visually square, Sides: arcuate, Maximal width: at middle, rarely at anterior margin, Anterior margin: subequal to posterior or narrower than posterior. Anterior lobe: moderate or obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles, rarely not reaching level of anterior pronotal angles or projecting beyond anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp or blunt. Disk: Convexity: moderately convex, Disk impressions: medial and lateral or lateral only, Medial impression (shape): entire or anteromedial and posteromedial, Lateral impressions (deepness): shallow. Prehumerus: filamentary or carinal, Shape: bisinuate, Extent: beyond half of pronotal length or to anterior pronotal angle, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina or joining anterior pronotal angle, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate or wide, Convergence: strongly convergent, Junction: present, Narrowest point: at posterior third of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: conjoint, Width: wide, Shape: arcuate. Elytral pubescence: entire, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: with pit, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: large, Anterior margin: arcuate or subtruncate. Prosternal process: narrow, Shape: subparallel, Sides: straight, Angles: rectangular, Angles (tips): sharp, Disk: impressed with sides protruding. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate. LEGS: Male sexual modifications: metafemur markedly incrassate. Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): markedly dorsoventrally bent, Ovipositor: uritiform. Diagnosis: head markedly convex; eyes smaller; pronotal sides more arcuate; prosternal process subparallel and deeply impressed; metasternal process with deep pit. Species group: Viridiobscurus. Relevant species: A. pusillesculptus Obenberger, 1940.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Type locality: China, Yunnan province, 100 km west of Kunming, Diaolin Natural Reservation. Type specimens: Primary: Holotype (Fig. 28H), ♂, (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”. Secondary: CHINA: Shaanxi: 1 ♀ PT (EKCS): “ChinaShaanxi, Lueyang, 15.7.-22.7.2005, lgt. E. Kučera”; Sichuan: 1 ♂ PT (EJCB): “China, W Sichuan, 1993, Moxi, Luding Co., 25-31/6, Gonga Shan env., M. Häckel leg.”; Yunnan: 2 ♂ PT, 1 ♀ PT (EJCB): “China, Yunnan, 6-10. VII, Ninglang env. 27.19N, 100.55E, E. Jendek leg., 1992”; 1 ♂ PT (EJCB): “China, Yunnan, 27-28.VI., Daju (50 km N of Lijiang), 27.21N, 100.19E, E. Jendek, 1992”; 2 ♂ PT (EJCB): “China, Yunnan, 2000-3000m, 27.20N. 100.11E, Habashan mts., SE slope, 10-13/7., Vít Kubáň leg. [19]92”; 1 ♀ PT (EJCB): “China, Yunnan, 1800 m, Lijiang, 23.6.-21.7., 26.53N, 100.18E, E. Jendek leg. 1992”; 1 ♀ PT (EJCB): “Yunnan, 2000-2800m, 25.11N, 100.24E, Weibaoshan mts., W slope, 25-28/6.1992, Vít Kubáň”; 1 ♀ PT (EJCB): “CH, Yunnan, vii.1993, Weishan (County), Weibaoshan, 26002900 m”; 5 PT (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”; 1 ♂ PT (EJCB): “Yunnan 2000m, 25.03N 101.55E, Yipinglang 8-10/6.,Vít Kubáň leg. 1993”; 1 ♂ PT (EJCB): “China-Yunnan, 24-29.6.[19]94, Daju, lgt. E. Kučera”; 1 ♀ PT (EJCB): “China, Yunnan, Daju, 17.VI.1995, Šiška & Spevár & Pekarovič”; 1 ♂ PT (EJCB): “China, Yunnan, Lijiang, 19.VI.1995, Šiška & Spevár & Pekarovič”; 2 PT (ZIN): “China, Yunnan, Loc. 2, Hway 320, Kunming-Xiaguan, 8 km WNW Chuxiong, 25°07’30.9’’N, 101°24’04.2’’E, 1874 m, 19.05.2002, Konstantinov & Volkovitsh leg.”. Adult occurence: 5-6-7 (n=14). Altitude range: 1800-3000 m (n=6). Host plants: Unknown. Distribution: CHINA: Shaanxi, Sichuan, Yunnan. Etymology: The specific name was derived from the Latin noun foramen (hole, opening) and the verb fero (bear or carry); it refers to the deep pit on the metasternal process of the species.

Agrilus fornicatus Jendek, sp. nov. Fig. 10D (habitus of HT); Fig. 32S (aedeagus)

Description: 10.2-11.2 mm (Holotype 10.2 mm). BODY: Shape: subparallel, Color (dorsally): unicolored or bicolored, Color (dominant): blue, green or brown. HEAD: Medial impression: present, Impression (extent): frons, Epistoma: with raised upper margin. Frons: Shape: flat, Outline: protruding from head outline. Vertex: Outline: protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): subparallel, Sculpture (density): sparse. Eyes: Size: large, Shape: not protruding from head outline or protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate or long, Shape: slender, Sexual modification in male: antennae longer. PRONOTUM:

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TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Shape: visually elongate, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles or at level with anterior pronotal angles. Posterior angles: Shape: acute or rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): deep, Lateral impressions (width): narrow. Prehumerus: carinal, Shape: arcuate or bisinuate, Extent: beyond half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina, Arc: obvious. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: absent or present, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate, Shape: subangulate, cuspidate or spinose, Position of dominant cusp (spine): medial. Elytral pubescence: distal only, Distal: apical, Color: unicolored. Tomentum: spots, Spots (locations): humeral, premedial or postmedial. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): moderate. Prosternal process: moderate, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: convex. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): distinctly longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: elongate. Diagnosis: prehumerus in posterior part very sharply elevated; elytral apices subangulate or subcuspidate; prosternum in both sexes obviously convex; basal ventrite in male markedly attenuate. Species group: Spinipennis. Relevant species: A. albertianus Descarpentries & Villiers, 1967. Type locality: China, west Jiangxi province, Jinggang ShanCiping. Type specimens: Primary: Holotype (Fig. 10D), ♂, (EJCB): “China, Jiangxi W, Jinggang Shan-Ciping, 2-14.VI.1994, E. Jendek & O. Šauša leg.”. Secondary: CHINA: Fujian: 1 ♀ PT (EJCB): “China, Fujian prov. Shaowu env. -Taining, 5-10. VII.1991, M. Nikodým”; 1 ♀ PT (EJCB): “China, Fujian prov. Shaowu env., 5-10.VII.1991”; 1 ♀ PT (EJCB): “China, Fujian prov., Shaowu env., 23-27.vi.1991, leg. Nikodým”; 1 ♂ PT, 1 ♀ PT (EJCB): “China, Fujian prov., Shaowu env., 13-16.6.1991”. Adult occurence: 6-7 (n=5). Host plants: Unknown.

Distribution: CHINA: Fujian, Jiangxi. Etymology: The specific name is the Latin adjective fornicatus (vaulted, arched). It refers to the obviously convex prosternum of the species.

Agrilus fortunatus Lewis Fig. 11A (habitus); Fig. 33A (aedeagus); Fig. 46K (type) fortunatus Lewis, 1893 (Agrilus) LEWIS, 1893: 333 334 [#1722] (description). KERREMANS, 1903: 285 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 16 [#1809] (catalog; Japan). OBENBERGER, 1936: 990 [#2191] (world catalog). MIWA, 1940: 72 [#4586] (checklist; Japan). IGA, 1947: 15 16 [#3273] (notes; characters; faunal records; Japan (Shikoku)). KUROSAWA, 1949: 13 14 [#5028] (characters). CHÛJÔ & KUROSAWA, 1950: 8 [#501] (catalog; faunal records; Japan (Shikoku)). IGA, 1955: 78 [#3828] (iconography; Japan). IGA, 1962: 78 [#3351] (iconography; Japan). KUROSAWA, 1963: 152 [#1618] (characters; Japan). KUROSAWA, 1974: 2 [#4576] (cited as fortunei). KUROSAWA, 1974: 1, 2 [#4575] (characters; notes). KUROSAWA, 1975: 2 [#4578] (note). TÔYAMA, 1985: 26 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 185 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 33 [#4091] (checklist; Japan: Hokkaido, Okushuri I., Honshu, Shikoku, Kyushu). FUKUTOMI & HORI, 2004: 10 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). ANONYMOUS, 2007: 32 [#8250] (checklist; Japan: Honshu). BELLAMY, 2008: 2098 [#8576] (world catalog).

Type material: Agrilus fortunatus Lewis, 1893 Type locality: Main island. Kiga, Nikko, and Chiuzenji. Type specimens: Primary: Lectotype (Fig. 46K) by present designation, ♂, (BMNH): “SYN-Type [p] [round label with blue border] \ Japan, G. Lewis, 1910-320 [p] \ 6. ‘80 [h] \ Agrilus fortunatus Lewis Type [h]”. Secondary: 1 PLT (BMNH). Described from unknown number of syntypes. Diagnosis: 7.8-9 mm; prehumerus short, carinal; prosternal process in male with transverse carina. Species group: Quadrisignatus. Relevant species: A. sospes Lewis, 1893. Specimens examined: JAPAN: Honshu: 1 (EJCB): “Konsei Touge (Nikko), 4.viii.1963, K. Kume leg.”; 1 (EJCB): “Mts. Daibusatsu, Yamanashi Pref., 20.VII.1979, K. Masumoto”; 1 (EJCB): “Kogenazawa, Yamanashi, Japan, 13.v.1982, leg. K. Ikeda”; 1 (EJCB): “Mts. Daibusatsu, Yamanashi Pref., 21.VI.1985 S. Ohmomo”; 1 ♀ (EJCB): “Japan: Honshu, Nara ken, Shakagatakeyama, E slope, 28 Jun 2002, Bolm lgt., 1600m, 34.06.30N, 135.53.40E”; 1 ♂ (EJCB): “Japan, Honshu, Osaka fu, Kongosan, 300 m, 2 km E of Amami, 12. May 2002, Bolm Leg.”. Adult occurence: 5 6 7 8 (n 6). Altitude range: 300 1600 m (n 2). Host plants: Fagus crenata: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: JAPAN: Hokkaido, Honshu, Kyushu, Shikoku.

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Agrilus friebi Obenberger Fig. 24I (habitus); Fig. 38P (aedeagus); Figs. 46N, 47U, 49T, 54Q, (types) friebi Obenberger, 1922 (Agrilus) OBENBERGER, 1922: 25 26 [#2021] (description). OBENBERGER, 1924: 47 [#2040] (note). OBENBERGER, 1926: 656 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 990 991 [#2191] (world catalog). THÉRY, 1942: 253 [#2875] (note). KUROSAWA, 1963: 154 [#1618] (characters; Japan). KUROSAWA, 1974: 2 [#4576] (note). KUROSAWA, 1974: 2 [#4575] (characters; notes). TÔYAMA, 1985: 26 [#3678] (iconography; Japan). ANONYMOUS, 1986: 177 [#8848] (insect pest; Korea). ALEXEEV, 1989: 480 [#64] (key to species; Primorsk; Korean peninsula; northeastern China). JENDEK, 1994: 19 20 [#1355] (lectotype designation; synonymy; characters; Russian Far East; Korea; Northeast China; Japan). AKIYAMA & OHMOMO, 1997: 33 [#4091] (checklist; Russia: East Siberia; North China). ALEXEEV, 1998: 373 [#4322] (subgenus Quercuagrilus; subgeneric classification). HUA LI ZHONG, 2002: 89 [#6696] (cited as friebei; checklist; China: Jiangsu). MÜHLE, 2003: 46 [#6155] (checklist; Taiwan). FUKUTOMI & HORI, 2004: 7 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 391 [#7441] (subgenus Quercuagrilus; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (checklist; Japan). BELLAMY, 2008: 2100 [#8576] (subgenus Quercuagrilus; world catalog). VOLKOVITSH, 2009: 135 [#8850] (subgenus Quercuagrilus; faunal records; Russia: Primorskiy kray). kiangsuanus Obenberger, 1935 (Agrilus; subspecies of friebi) OBENBERGER, 1935: 171 [#2158] (description). OBENBERGER, 1936: 991 [#2191] (subspecies of friebi; world catalog). OBENBERGER, 1943: 10 [#2333] (variety of friebi). KUROSAWA, 1974: 2 [#4576] (subspecies of friebi). PENG ZHONGLIANG, 1987: 355 [#2429] (subspecies of friebi; checklist; China). JENDEK, 1994: 19, 20 [#1355] (synonym of friebi; lectotype designation). HUA LI ZHONG, 2002: 89 [#6696] (cited twice, as a species as kangsuanus [sic!] and as a subspecies of friebi as friebe kiangsueanus [sic!]; checklist; China: Jiangsu, Gansu. JENDEK, 2006: 392 [#7441] (synonym of friebi friebi; Palaearctic catalog). BELLAMY, 2008: 2100 [#8576] (synonym of friebi; world catalog). vodaki Obenberger, 1936 (Agrilus) OBENBERGER, 1936: 141 [#2207] (description). MIWA & CHÛJÔ, 1936: ADDENDA P. 4 [#1809] (catalog; Japan). OBENBERGER, 1936: 1069 [#2191] (world catalog). IGA, 1955: 79 80 [#3828] (iconography; Japan). IGA, 1962: 79 80 [#3351] (iconography; Japan). KUROSAWA, 1963: 154 [#1618] (subspecies of friebi; characters; Japan). KUROSAWA, 1974: 2 [#4576] (related to A. friebi). KUROSAWA, 1975: 3 4 [#4591] (note). ALEXEEV, 1989: 480 [#64] (synonym of friebi). JENDEK, 1994: 19, 20 [#1355] (synonym of friebi; lectotype designation; remark on type locality). AKIYAMA & OHMOMO, 1997: 33 [#4091] (synonym of friebi friebi). JENDEK, 2006: 392 [#7441] (synonym of friebi friebi; Palaearctic catalog). BELLAMY, 2008: 2100 [#8576] (synonym of friebi; world catalog). ainu Miwa & Chûjô, 1940 (Agrilus) MIWA & CHÛJÔ, 1940: 55, 64 65 [#1810] (description). KUROSAWA, 1974: 2 [#4576] (subspecies of friebi). TÔYAMA, 1985: 26 [#3678] (subspecies of friebi; iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (subspecies of friebi; checklist; Japan). JENDEK, 1994: 19, 20 [#1355] (synonym of friebi; notes). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (subspecies of friebi; checklist; Japan). AKIYAMA & OHMOMO, 1997: 5, 33 [#4091] (subspecies of friebi; checklist; Japan: Hokkaido, Honshu, Shikoku, Kyushu, Tsushima). JENDEK, 2006: 391 [#7441] (subgenus Quercuagrilus; subspecies of friebi; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (synonym of friebi). BELLAMY, 2008: 2100 [#8576] (subspecies of friebi; world catalog).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA enisus Théry, 1942 (Agrilus) THÉRY, 1942: 264 266 [#2875] (description). OBENBERGER, 1943: 10 [#2333] (synonym of friebi kiangsuanus). JENDEK, 1994: 19 [#1355] (synonym of friebi). JENDEK, 2006: 392 [#7441] (synonym of friebi friebi; Palaearctic catalog). BELLAMY, 2008: 2100 [#8576] (synonym of friebi; world catalog). nalajchanus Cobos, 1968 (Agrilus; subgenus Agrilus) syn. nov. COBOS, 1968: 391 393 [#580] (description). COBOS, 1972: 47, 56 [#591] (subgenus Agrilus; faunal record; Mongolia). ALEXEEV, 1975: 155 [#52] (notes; faunal records; distributional overview; Mongolia). ALEXEEV, 1989: 480 [#64] (cited as nalaichanus; key to species; ?Chita; ?Buryatia, Tuva; Mongolia). ALEXEEV & VOLKOVITSH, 1989: 314, 343, 356 [#3241] (characters in key; faunal records; Mongolia). ALEXEEV, 1998: 374 [#4322] (subgenus Quercuagrilus; subgeneric classification). JENDEK, 2006: 392 [#7441] (subgenus Quercuagrilus; Palaearctic catalog). BELLAMY, 2008: 2198 [#8576] (subgenus Quercuagrilus; world catalog).

Type material: Agrilus friebi Obenberger, 1922 Type locality: Siberia orientalis: Vladivostok. Type specimens: Primary: Lectotype (Fig. 46N), ♀, designated by JENDEK (1994: 19-20) [#1355], (NMPC): “COLLECTIO DR.OBENBERGER MUS. PRAGENSE [p] \ Vladivostok [h] \ Typus [p] [red label] \ Agrilus Friebi m. Type [h] Det. Dr Obenberger [p]\ Mus. Nat. Pragae Inv.[p] 24 881[h] [orange label]”. Secondary: 1 PLT (NMW); 1 PLT (NMPC); 1 PLT (NHMB). Described from 6 syntypes. Agrilus friebi kiangsuanus Obenberger, 1935 Type locality: China: Kiang-su. Type specimens: Primary: Lectotype (Fig. 47U), ♀, designated by JENDEK (1994: 19, 20) [#1355], (NMPC): “China: Kiangsu [h] \ Typus [p] [red label] \ Mus. Nat. Pragae Inv. [p] 24 305 [h] [orange label] \ Agrilus Friebi ssp. Kiangsuanus m. Type [h] Det. Dr Obenberger [p]”. Secondary: ST (NHMB). Described from unknown number of syntypes. Agrilus vodaki Obenberger, 1936 Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 54Q), ♀, designated by JENDEK (1994: 19, 20) [#1355], (NMPC): “Ussuri [h] \ Typus [p] [red label] \ Agrilus Vodaki m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv.[p] 24 891 [h] [orange label]”. Described from unknown number of syntypes. Agrilus ainu Miwa & Chûjô, 1940 Type locality: Mt. Moiwa in the vicinity of Sapporo, Hokkaidô, Japan. Type specimens: Not located. Described from 1 specimen. Agrilus enisus Théry, 1942 Type locality: Maoerschan, Mandchourie. Type specimens: Not located. Described from 1 specimen. Agrilus nalajchanus Cobos, 1968 Type locality: Central aimak, cca 30 km W von somon Nalajch, 1530 m. alt. Type specimens: Primary: Holotype (Fig. 49T), ♀, (HNHM): “Mongolia, Central aimak, cca 30 km O [sic!] von Somon Nalajch, 1530 m. Exp. Dr. Z. Kaszab, 1996 [p] \ Nr. 523 14. VI. 1966 [p] \ Holotypus A. Cobos [p] [red label]

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TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA \ Agrilus nalajchanus nov. sp. Holotypus [h] A. Cobos det. 196 [p] 8 [p]”. Described from 1 specimen. Diagnosis: 3.5-5.2 mm; pronotum widest at base; elytra with proximal and distal pubescence; aedeagus symmetrical. Species group: Sulcicollis. Relevant species: A. obscurosericeus Obenberger, 1940. Specimens examined: CHINA: Anhui: 1 ♂ (EJCB): “China, Anhui, 1998, Dabieshan, 65 km SW Huoshan, 1400 m, Bolm lgt. 21 24 Jun”. Beijing: 1 ♂, 2 ♀ (EJCB): “N China, 7.vi. 1991, Beijing, Xiang Shan, leg. M. Nikodým”; 1 ♀ (EJCB): “China, Beijing distr., Yan Shan, Dongling Mts., 39º58’N, E 115º26’E, 1400 m, Xiaolongmen, 15 16.VI.2001”. Gansu: 1 ♀ (EJCB): “China, Gansu reg. Xia He 17 18.7., Hubička leg. 1990” (JENDEK, 1994 [#1355]). Jilin: 2 (ZIN): “Hsiaoling, Kirin [ Jilin] Province, China 15.vi.1939 [in Russian]”. Shaanxi: 5 ♂, 1 ♀ (EJCB): “China, Shaanxi, pr. Hua Shan, 17. 21.6.1991, R. Dunda lgt.”; 1 ♂, 1 ♀ (EJCB): “China, Shaanxi, 1992, 120 km E Xi an, Hua Shan, 3 4.vi., Jaroslav Turna”; 1 ♂ (EJCB): “China, Shaanxi, 1992, Huashan, 120 km E of, 3. 4.Jun 1992, Xian, M. Bok lgt.”; 1 (EJCB): “China c., 17 18.V., Ganguyi 110,3’ 36,8’, 35 km NE of Yanan, leg. J. Halada 1996”; 1 ♂ (EJCB): “China Shaanxi, Lüeang, 18.5. 21.5.2000, leg. E. Kučera”. Shanxi: 1 (EJCB): “China, Shanxi, 9.June.2000, Lüliang Shan, 1000m, road Fangshan Jiaocheng, Hengjian env., Zd. Jindra”. Sichuan: 1 ♂ (EKCS): “China N, Sechuan, Nanjiang, 21 23.5.2002, lgt. E. Kučera”. JAPAN: Honshu: 1 ♀ (JMCZ): “Numata shi, Gumma pref., Honshu, Japan, 31.5.1955, coll. T. Takei”; 1 (EJCB): “Japon, Gunma, 4 km SW Tsumagoi, 1050 m, Löbl, 18.7.1980”; 1 (EJCB): “Kawakami Mura, Minamisaku Gun, Nagano Pref., vi.29.1980, T. Hattori leg.”; 1 (EJCB): “Tateiwamura vill., Minami Aizu, Fukushima pref., 27.vi.1992, S. Ohmomo”; 2 ♂ (MNCA): “Tateiwamura, Minami Aizu Fukushima Pref. 27.VI.1992 S. Ohmomo leg.” (JENDEK, 1994 [#1355]). KOREA NORTH: 1 ♀ (EJCB): “N Korea, 19.7.1990, Toxong env., Omdouri”. KOREA SOUTH: 1 (JLCS): “Chilyeonpokpo, Mt. Deokyusan, Korea, 15.v.1999, S. L. An leg.”; 1 (JLCS): “Sagimakgol, Jungwon gu, Seongnam si, Gyeongg do, Korea, 1.vi.2001, coll. Hee ah Lee”. MONGOLIA: 1 ♀ (EJCB): “Mongolia, Arkhangal, 5 km S of lake Bulgana, 2.7.1975”; 1 ♂ (ZIN): “Mongoliya, Khèntèiskii aimak, 8 km Yu Norovlina, Kerzhner, 2.vii.1976 [in Russian]”; 1 ♂, 1 ♀ (ZIN): “Mongoliya, Khèntèiskii aimak, 8 km Yu Norovlina, M. Kozlov, 2.vii.1976 [in Russian]”. RUSSIA: Primorskiy kray: 1 ♂ (EJCB): “Vladivostok 27.VI.1927 B. Bjelov” (JENDEK, 1994 [#1355]); 2 ♂ (ZIN): “Primorskii kraj, 2.VI., Lazovskii raion, 1951, d.Tatchik goze, Stepanov leg.” (JENDEK, 1994 [#1355]); 1 (ZIN): “Y Primor’e, sr. ter. r. Suchan, Kabakov, 10 20.7.1952 [in Russian]”; 1 ♂ (ZIN): “Primorskii kr., g. Vladivostok, Uchal’naya r., Peschanna, 10 xii [19]60 \ vyv. iz kul’turn. vinograda [in Russian]”; 1 (EJCB): “Primorskii kr. g. Vladivostok, s. Novickoe 16 1 61 L. A. Anufriev \ vyv. iz kul’turn. vinograda [in Russian]” (JENDEK, 1994 [#1355]); 1 (ZIN): “SSSR, Primorje, Barabasch Levada 7.VI.1980 C. V. Konovalov lgt.” (JENDEK, 1994 [#1355]); 1 ♂ (EJCB): “UdSSR, Primorskij kraj Vladivostok 26.VII.1990 leg. A. Pütz” (JENDEK, 1994 [#1355]); 2 (ZIN): “Primorskii kraj Spassk. les 13.06.1990 Belokobylskij [in Russian]” (JENDEK, 1994 [#1355]); 5 ♂ (EJCB): “USSR, Primorskij kraj Arseneev env. 300 400 m 11 19.VII.1990 B. Šiška & R. Čermák leg.” (JENDEK, 1994 [#1355]); 1 ♂ (EJCB): “Sib.or. m., Primorje Sichote Alin Mts. Sokolči 1. 15.7.1990 Kadlec + Voříšek lg.” (JENDEK, 1994 [#1355]); 1 (EJCB): “Prim. Khasansk Rjazanovka 5.6.1991 Filimonov” (JENDEK, 1994 [#1355]); 2 ♀ (EJCB): “USSR, Primorsk. kraj, Arsenev env., VI.1991, 44.11N, 133.19E, O. Šausa leg.” (JENDEK, 1994 [#1355]); 1 ♀ (ZIN): “Russia, Primorskiy Terr. Spassk Dal’niy env., forest edge, glades, 27,30.VI.2006, S. Belokobylskiy leg.”; 1 ♀ (ZIN): “Primorskii kr., Lazovskii z k, b. Proselochnaya, 2 5.VII.2007 Y. Sundukov, V. Khokhrin [in Russian]”.

Adult occurence: 5-6-7 (n=36). Faunal record published by COBOS, 1972 [# 591] is also included. Altitude range: 3001530 m (n=7). Host plants: Quercus: MÜHLE, 2003 [#6155]. − Q. acutissima; AKIYAMA & OHMOMO, 1997 (as ainu) [#4091]. − Q. serrata; AKIYAMA & OHMOMO, 1997 (as ainu) [#4091]. − Q. crispula; AKIYAMA & OHMOMO, 1997 (as ainu) [#4091]. − Rhamnus japonica: AKIYAMA & OHMOMO, 1997 (as ainu) [#4091]; MÜHLE, 2003 [#6155]. − Vitis: ALEXEEV, 1989 [#64]. Distribution: CHINA: Anhui, Beijing, Gansu, Heilongjiang, Jiangsu, Jilin, Liaoning, Shaanxi, Sichuan. JAPAN: Hokkaido, Honshu, Kyushu, Shikoku, Tsushima. KOREA NORTH. KOREA SOUTH. MONGOLIA. RUSSIA: Primorskiy kray, Tyva Respublika.

Agrilus fusciapex Jendek, sp. nov. Fig.16B (habitus of HT); Fig. 35B (aedeagus)

Description: Holotype 7.7 mm. BODY: Shape: cuneiform, Color (dorsally): bicolored, Color (dominant): green or brown. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: in plane with frons. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): subparallel, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: visually square, Sides: subangulate, Maximal width: at middle, Anterior margin: wider than posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk modifications: with two black longitudinal stripes, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Anteromedial and posteromedial impressions: subequal in size, Lateral impressions (deepness): deep. Prehumerus: absent. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior angles or at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: bicolored, Alternative color (location): perisutural stripe, pleural portions or apical portion, Humeral carina: absent. Apices: Arrangement: separate, Shape: arcuate. Elytral pubescence: absent. Tomentum: spots, Spots (locations): premedial or postmedial. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): shallow or moderate, Emargination (width): narrow or moderate. Prosternal process: narrow or moderate, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: convex, Modification: disk in male with transverse rugae. ABDOMEN: Sexual modifications in male: not apparent,

102 Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed. Diagnosis: apical half of elytra reddish-brown; elytra with two pairs of pubescent spots; prosternal process in male with protuberance (lateral vierw); aedeagus symmetrical. Species group: Smaragdinus. Relevant species: A. odetteae Baudon, 1968. Type locality: China, Guizhou province, Leishan Co., southestern Kaili, 15 km east of Leishan, Leigong Shan, eastern slope, 26°23.39’N, 108°13.33’E, about 2.5 km east of pass between Leishan and Fangxiang village, altitude about 1600 m. Type specimens: Holotype (Fig. 16B), ♂, (EJCB): “China: Guizhou, Leishan Co., SE Kaili, 15 km E Leishan, Leigong Shan, E-slope, 26°23.39’N, 108°13.33’E / ca 2.5 km E of pass between Leishan and Fangxiang vill., ca 1600 m, 23/24.6.2001, leg. Schillhammer (17)”. Adult occurence: 6 (n=1). Altitude range: 1600 m (n=1). Host plants: Unknown. Distribution: CHINA: Guizhou. Etymology: The specific name was derived from the Latin adjective fuscus (of brown color) and the noun apex (tip); it refers to the brown color of the pubescence on the elytral apex of the species.

Agrilus fusiformis Jendek, sp. nov. Fig. 28E (habitus of HT); Fig. 40I (aedeagus)

Description: 3.5-4.7 mm (Holotype 4.1 mm). BODY: Shape: fusiform, Color (dorsally): bicolored, Color (dominant): green. HEAD: Shape: markedly convex, Medial impression: absent, Epistoma: with raised upper margin. Frons: Shape: markedly convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): irregular, Sculpture (density): dense. Eyes: Size: large, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short or moderate, Sexual modification in male: antennae longer. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at posterior margin, rarely at middle, Anterior margin: narrower than posterior. Anterior lobe: obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles or projecting beyond anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: strongly convex, Disk impressions: medial and lateral, Medial impression (shape): entire or anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow, Lateral impressions (width): narrow. Prehumerus: carinal, Shape: bisinuate, Extent: beyond half

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina, Arc: obvious. Marginal and submarginal carinae: Interspace: wide, Convergence: strongly convergent, Junction: present, Narrowest point: at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: bicolored, complementary with pubescence, Alternative color (location): transverse band before apex, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: conjoint, Width: wide, Shape: arcuate. Elytral pubescence: proximal and distal, Proximal: like V or upturned T, Distal: apical, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: large, Anterior margin: subtruncate. Prosternal process: moderate, Shape: dilated, Sides: straight, Angles: obtuse, Angles (tips): sharp, Disk: impressed. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate or angulately sinuate. LEGS: Male sexual modifications: metafemur markedly incrassate. Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: uritiform. Diagnosis: pronotum with anterior lobe obvious and markedly projecting beyond anterior angles. Species group: Densetomentosus. Relevant species: A. bernardii Descarpentries & Villiers, 1963. Type locality: China, Yunnan province, Yipinglang, altitude 2000 m, 25.03N 101.55E. Type specimens: Primary: Holotype (Fig. 28E), ♂, (EJCB): “Yunnan 2000m, 25.03N 101.55E, Yipinglang 8-10/6.,Vít Kubáň leg. 1993”. Secondary: CHINA: Yunnan: 3 ♀ PT (EJCB): “Yunnan 2000m, 25.03N 101.55E, Yipinglang 8-10/6.,Vít Kubáň leg. 1993”; 3 PT (EJCB): “Yunnan 18002000m, 25.04N 101.55E, Yipinglang 17-20.6.1994,Vít Kubáň leg.”. Adult occurence: 6 (n=2). Altitude range: 1800-2000 m (n=2). Host plants: Unknown. Distribution: CHINA: Yunnan. Etymology: The specific name was derived from the Latin noun fusus (spindle) and the suffix -formis (having the form of); it refers to the body shape of the species.

Agrilus ganglbaueri Semenov Fig. 30F (habitus); Fig. 41H (aedeagus); Figs. 42K, 46O (types) ganglbaueri Semenov, 1891 (Agrilus) SEMENOV, 1891: 263, 344 345 [#3527] (description). KERREMANS, 1892: 258 [#1397] (catalog). HEYDEN, 1893: 90 [#3450] (catalog). KERREMANS, 1903: 285 [#1441] (catalog). SEMENOV, 1903: 172 [#4114] (notes on type locality). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 657 [#2057]

103

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA (Palaearctic catalog). OBENBERGER, 1936: 991 [#2191] (world catalog). SINADSKIJ, 1957: 40 [#7355] (biology; Amu Darya estuary). VORONCOV & SINADSKIJ, 1957: 85 [#8645] (notes; Amu Daria and Syr Daria river basins). KULINITSH, 1963: 154 [#8648] (notes; Tajikistan). SINADSKIJ, 1963: [#5038] (notes; biology). PRIPISNOVA, 1965: 61, 101 [#8633] (pest; biology; Tajikistan). KOSTIN, 1973: 107 [#3693] (characters in key; Kazakhstan). AIBASOV, 1974: 144 [#7212] (biology; distribution; Kazakhstan). YAGDYEV, 1974: 257 [#8644] (pest; Turkmenistan). ALEXEEV, 1975: 155 [#52] (notes; faunal records; distributional summary; Mongolia; Uzbekistan; Afganistan; China). YAGDYEV, 1975: 60, 61 [#7775] (biology; Turkmenia). KADYROV, 1976: 52 [#3461] (cited as ganglebaueri; biology; Tajikistan). YAGDYEV, 1984: 70 [#6196] (notes; Turkmenistan; Tajikistan). ALEXEEV & VOLKOVITSH, 1989: 320, 349, 357 [#3241] (characters in key; faunal records; Mongolia). SOYUNOV, 1991: 112 [#8657] (ecology; Turkmenistan). VOLKOVITSH & ALEXEEV, 1992: 164 [#2973] (faunal records; Turkmenistan). JENDEK, 1993: 38 [#1354] (lectotype designation). PENG ZHONGLIANG, 1994: 19 [#2432] (faunal records; China). VOLKOVITSH & ALEXEEV, 1994: 437 [#2974] (checklist; Turkmenistan). ALEXEEV, 1998: 374, 375 [#4322] (subgenus Micragrilus; genotype of the subgenus Micragrilus). NONNAIZAB et al., 1999: 112 [#8247] (checklist; China: Inner Mongolia). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Xinjiang). TLEPPAEVA & ISHKOV, 2004: 84 [#7211] (subgenus Micragrilus; checklist; Kazakhstan). HUANG RENXIN, 2005: 77 [#8248] (checklist; China: Xinjiang). JENDEK, 2006: 391 [#7441] (subgenus Micragrilus; Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2105 [#8576] (subgenus Micragrilus; world catalog). albifrons Reitter, 1897 (Agrilus) REITTER, 1897: 249 250 [#3506] (description). HEYDEN, 1898: 21 [#5553] (catalog). KERREMANS, 1903: 281 [#1441] (catalog). SEMENOV, 1903: 15 [#4116] (synonym of ganglbaueri). JAKOBSON, 1913: 797 [#4386] (synonym of ganglbaueri; catalog). OBENBERGER, 1936: 991 [#2191] (synonym of ganglbaueri). JENDEK, 2006: 391 [#7441] (synonym of ganglbaueri; Palaearctic catalog). BELLAMY, 2008: 2105 [#8576] (synonym of ganglbaueri; world catalog).

Type material: Agrilus ganglbaueri Semenov Tian Shanskij, 1891 Type locality: Prov. Transcaspica: Tedshen. Type specimens: Primary: Lectotype (Fig. 46O), ♂, designated by JENDEK (1993: 38) [#1354], (ZIN): “Tedzhen, 11.VI.[18]88 A. Semenov [h] [in Russian] \ Agrilus Ganglbaueri m. typ AS.XI.90 [h] \ coll. Semenov-Tian-Shansky [p]”. Secondary: 2 PLT (ZIN). Described from 5 syntypes. Agrilus albifrons Reitter, 1897 Type locality: Sefir-Kuh, nördlich von Herat. Type specimens: 1 ST (Fig. 42K), (MNHN); 1 ST (HNHM); 1 ST (BMNH). Described from unknown number of syntypes. Diagnosis: 2.9-4 mm; eyes smaller, elytral pubescence entire and long, with indications of denser pattern. Species group: Unassigned. Relevant species: A. lituratus (Klug, 1829). Specimens examined: MONGOLIA: 4 (EJCB): “Mongolia mer., Šara chulany Bul., 7.7.1988, O. Majzlan lgt.”. Adult occurence: 7 (n=1). Host plants: Populus: VORONCOV & SINADSKIJ, 1957 [#8645]; SINADSKIJ, 1963 [#5038]; PRIPISNOVA, 1965 [#8633]; KOSTIN,

1973 [#3693]; AIBASOV, 1974 [#7212]; KADYROV, 1976 [#3461]; VOLKOVITSH & ALEXEEV, 1992 [#2973]; VOLKOVITSH & ALEXEEV, 1994 [#2974]. − P. ariana: SINADSKIJ, 1957 [#7355]; ALEXEEV, 1975 [#52]. − P. diversifolia: YAGDYEV, 1974 [#8644]; YAGDYEV, 1975 [#7775]; ALEXEEV & VOLKOVITSH, 1989 [#3241]; TLEPPAEVA & ISHKOV, 2004 [#7211]. − P. pruinosus: SINADSKIJ, 1957 [#7355]; ALEXEEV, 1975 [#52]. Distribution: CHINA: Nei Mongol, Xinjiang. MONGOLIA. AFGHANISTAN. KAZAKHSTAN. TAJIKISTAN. TURKMENISTAN. UZBEKISTAN.

Agrilus gaoligong Jendek Fig. 13A (habitus of HT); Fig. 33V (aedeagus) gaoligong Jendek, 2000 (Agrilus) JENDEK, 2000: 152 153 [#4853] (description). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2105 [#8576] (world catalog).

Type material: Agrilus gaoligong Jendek, 2000 Type locality: Yunnan 1500-2500 m, 25.22N 98.49E, 1724/5 Gaoligong mts. Type specimens: Primary: Holotype (Fig. 13A), ♂, (EJCB): “Yunnan 1500-2500 m, 25.22N 98.49E, 17-24/5 Gaoligong mts. 1995”. Secondary: 4 PT (EJCB). Described from 5 specimens (HT, PT). Diagnosis: 9.4-10.8 mm; body unicolor, golden-bronze; elytra with wide perisutural stripes reaching to sutural margin; prosternum in male with bunch of erect setae in middle and row of erect setae on prosternal process. Species group: Imitans. Relevant species: A. blatteicollis Bourgoin, 1922; A. nepalensis Tôyama, 1988; A. sichuanus sp. nov. Specimens examined: CHINA: Yunnan: 1 ♀ PT (NMPC): “Yunnan, Tszindun, 1200 m, 15.V.1957, A. Monchadskii [in Russian] [p] \ [1.line] [Chinese text] [2.line] 1957 V.15 [3.line] A. [Chinese text] [p]” (JENDEK, 2000 [#4853]); 2 ♂ PT, 1 ♀ PT (EJCB): “Yunnan, 1500 2500 m, 25°22’N, 98°49’E, 17 24 / 5 Gaoligong mts, Vít Kubáň leg., 1995” (JENDEK, 2000 [#4853]).

Adult occurence: 5 (n=2). Altitude range: 1200-2500 m (n=2). Host plants: Unknown. Distribution: CHINA: Yunnan.

Agrilus grisator Kerremans Fig. 20D (habitus); Fig. 36Q (aedeagus); Fig. 46V (type) grisator Kerremans, 1893 (Agrilus) KERREMANS, 1893: 343 344 [#1406] (description). KERREMANS, 1903: 278 [#1441] (catalog). MAXWELL LEFROY & HOWLETT, 1909: 331 [#8091] (notes; India). OBENBERGER, 1936: 1084 [#2191] (world catalog). QUAYLE, 1938: 314 [#7340] (recorded from Citrus; India). MÜHLMANN, 1954: 83 [#8152] (notes). BAUDON, 1961: 75 [#143] (faunal records; Laos). BALACHOWSKI et al., 1962: 287 [#96] (pest on Citrus). DESCARPENTRIES & VILLIERS, 1963: 105, 112 [#700] (lectotype designation: “Inde: Kanara (lectotype, British Museum)”; characters; faunistic data; Inde (Kanara); Laos;

104 Cambodge). BAUDON, 1968: 131, 147 [#162]. DESCARPENTRIES & CHŰJÔ, 1968: 15 [#3559] (faunal record; Laos). KUROSAWA, 1974: 3 [#4575] (characters; notes). OHMOMO, 2002: 23 [#5751] (Thailand). BELLAMY, 2008: 2117 [#8576] (world catalog).

Type material: Agrilus grisator Kerremans, 1893 Type locality: Kanara. Type specimens: Primary: LT (Fig. 46V), ♂, designated by Descarpentries & Villiers (1963: 105, 112) [#700], (BMNH): “Kanara Andrewes [Kerremans’ MS] \ grisator Kerr. Type [Kerremans’ MS] \ Kerremans 1903-59 [p] Lecto [h] Type [p]”. Secondary: 1 PLT (MNHN); 9 PLT (BMNH). Described from unknown number of syntypes. Diagnosis: 7-10.5 mm; body generally larger and thicker; pubescent elytral fasciae less wide and often disrupted, with larger interspaces. Species group: Angulatus. Relevant species: A. olivaceidorsis Obenberger, 1917. Specimens examined: CHINA: Hong Kong: 1 ♂, 2 ♀ (EJCB): “Hong Kong, New Territories, Clear Water Bay, VII 4 1963, C. C. Lindt”.

Adult occurence: 5-6-7 (n=7). Faunal records published by BAUDON, 1961 [#143], 1968 [#162]; DESCARPENTRIES & VILLIERS, 1963: [#700] and DESCARPENTRIES & CHŰJÔ, 1968 [#3559] are also included. Host plants: Citrus: MAXWELL LEFROY & HOWLETT, 1909 [#8091]; MÜHLMANN, 1954 [#8152]; BALACHOWSKI et al., 1962 [#96]. Distribution: CHINA: Hong Kong. CAMBODIA. INDIA. LAOS. THAILAND.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Diagnosis: 3.5-4.7 mm; body dark-green; prosternal lobe narrow; prosternal process feebly enlarged; metasternal projection slightly impressed. Species group: Cyaneovirens. Relevant species: A. sagittifer sp. nov.; A. tsushimanus Kurosawa, 1963. Specimens examined: CHINA: Shaanxi: 3 (EJCB): “China, 1150 1300 m, Shaanxi, Qinling mts., Foping (6 km N), 20 21.VI.1998, I. H. Marshal leg.”; 1 ♂ (EJCB): “China, Shaanxi, 1992, 120 km SW Xi An, Foping, 7.VI., Jaroslav Turna leg.”; 2 ♂ (EJCB): “C China, Shanxi [sic! should be Shaanxi], Qinling Shan, 6km E of Xunyangba, 1000 1300 m, 23.v. 13.vi., leg. C. Holzschuh, 2000”; 1 ♂ (EJCB): “China Shaanxi, Čun Čan, 26.5. 1.6.2000, leg. E. Kučera”. Xinjiang: 1 ♂ (EJCB): “77.7.15, Xinjiang, Ürümqi”. RUSSIA: Primorskiy kray: 1 ♂ (EJCB): “Sib. or. m., Primorje, Ussuri res., 20.7.1990, Kadlec + Voříšek lg.”; 1 ♂ (EJCB): “Sib. or. m., Primorje, Sichote Alin Mts., Sokolči, 1 15.7.1990, Kadlec + Voříšek lg.”; 1 ♀ (EJCB): “USSR, Primorsk. kraj, Novocugujevka env., 18.VII.1990, Kantner leg.”; 1 ♂ (EJCB): “Russia, Primorsk. reg., 18 21.7.1992, Rezanovka, E. Kučera leg. ex Lonicera”; 1 ♀ (JMCZ): “Russia Far East, 19 20.VII.1993, Primorsk region, Lazo, lgt. M. Trýzna”; 8 (EJCB): “Rossia, Primorskij k., Rjazanovka 4 8. vii.1993, Z. Kletečka leg.”; 1 (EJCB): “Russia, Primorsky kray, Vladivostok, N43º12’, E131º59’, 6.vii.2007, 50m, Grebennikov”.

Adult occurence: 5-6-7 (n=15). Faunal records published by ALEXEEV, 1981 [#62] are also included. Altitude range: 50-1300 m (n=3). Host plants: Juglans mandshurica: ALEXEEV, 1981 [#62]; ALEXEEV, 1989 [#64]. − Ulmus: ALEXEEV, 1981 [#62]. Distribution: CHINA: Shaanxi, Xinjiang. RUSSIA: Primorskiy kray.

Remarks: This species will be examined in more detail in the revision of the whole species group.

Agrilus han Jendek, sp. nov.

Agrilus gussakovskiji Alexeev

Description: Holotype 4.1 mm. BODY: Shape: subparallel, Color (dorsally): unicolored, Color (dominant): green. HEAD: Shape: markedly convex, Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: markedly convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture (density): dense. Eyes: Size: large, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate or obvious, Shape: subangulate, Position to anterior angles: projecting beyond anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow. Prehumerus: carinal, Shape: arcuate, Extent: beyond half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: close to pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior third of pronotal

Fig. 27A (habitus); Fig. 39N (aedeagus), Fig. 46W (type) gussakovskiji Alexeev, 1981 (Agrilus; subgenus Dentagrilus) ALEXEEV, 1981: 3 7 [#62] (description). ALEXEEV, 1989: 484 [#64] (cited as gussakovskii; key to species; Yu Primorsk). ALEXEEV, 1998: 378 [#4322] (subgenus Dentagrilus; cited as gussakovskii; subgeneric classification). JENDEK, 2006: 390 [#7441] (subgenus Dentagrilus; Palaearctic catalog). JENDEK, 2007: 90 [#8008] (remarks on correct original spelling). BELLAMY, 2008: 2119 [#8576] (world catalog). VOLKOVITSH, 2009: 135 [#8850] (subgenus Dentagrilus; faunal records; Russia: Primorskiy kray).

Type material: Agrilus gussakovskiji Alexeev, 1981 Type locality: Sokolovskii r-n, bukh. Proselochnaya [in Russian]. Type specimens: Primary: Holotype (Fig. 46W), ♂, (ZIN): “Holotypus [p] [red label] \ Primorskii kr. Sokolovskii r. n. bukh Tochingaza 14-IX-48 V. V. Gussakovskii [in Russian] [h] na manchzhurs. orekhe [h] [underside] \ Agrilus gussakovskii m. sp. n. [h] V. Stepanov det. [p] \ Agrilus gussakovskii sp. n. [h] Alexeev det. [p]”. Secondary: 2 PT (ZIN). Described from 7 specimens (HT, PT).

Fig. 25G (habitus of HT); Fig. 38Z (aedeagus)

105

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate, Width: wide, Shape: arcuate. Elytral pubescence: entire. Tomentum: absent. STERNUM: Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: large, Anterior margin: arcuately emarginate, Emargination (deepness): moderate. Prosternal process: wide, Shape: dilated, Sides: arcuate, Angles: rectangular, Angles (tips): sharp, Disk: impressed with sides protruding. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Male sexual modifications: metafemur markedly incrassate. Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical. Diagnosis: body suparallel; elytra entirely pubescent; prosternum medially with erect pubescence; sternal groove on apex of last ventrite arcuate. Species group: Tempestivus. Relevant species: A. fevreae Descarpentries & Villiers, 1963. Type locality: China, north Henan province, Jiuligou, altitude 550 m, 35°12’N, 112°26’E. Type specimens: Holotype (Fig. 25G), ♂, (EJCB): “China, N. Henan, 11-12.VI., Jiuligou, 550 m, 35°12’N, 112°26’E, Jaroslav Turna leg. 2007”. Adult occurence: 6 (n=1). Altitude range: 550 m (n=1). Host plants: Unknown. Distribution: CHINA: Henan. Etymology: Noun in apposition; the specific name refers to the ethnic group Han, native to China and, by most recent estimation, the largest single ethnic group in the world.

Agrilus hani Jendek, sp. nov. Fig. 27H (habitus of HT); Fig. 40A (aedeagus)

Description: 4.4-5.4 mm (Holotype 5.1 mm). BODY: Shape: cuneiform, Color (dorsally): bicolored, Color (dominant): green. HEAD: Shape: markedly convex, Medial impression: present, Impression (extent): vertex, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): semispherical, Sculpture (density): dense. Eyes: Size: large, Shape: not protruding from head outline or protruding from head outline, Lower margin: in line with antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent or antennae longer. PRONOTUM: Shape: visually square, Sides: straight or arcuate, Maximal width: at anterior margin, rarely at middle, Anterior margin: subequal to posterior. Anterior lobe: obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior

angles: Shape: acute or rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial or posteromedial only, Medial impression (deepness): shallow, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial. Prehumerus: carinal, Shape: bisinuate, Extent: beyond half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior angles or at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate or submarginal carina divergent at posterior angles. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: bicolored, complementary with pubescence, Alternative color (location): apical portion, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate or conjoint, Width: wide, Shape: arcuate. Elytral pubescence: proximal and distal, Proximal: like V or upturned T, Distal: preapical, Color: bicolored. Tomentum: stripes or spots, Spots (locations): humeral or premedial. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate or large, Anterior margin: subtruncate or arcuately emarginate, Emargination (deepness): shallow. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: acute or rectangular, Angles (tips): sharp, Disk: impressed. ABDOMEN: Sexual modifications in male: medial sulcus on ventrite 1 and last ventrite impressed on apex, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate, Sinuosity (deepness): markedly deep. LEGS: Male sexual modifications: metacoxa medially with cluster of setae and metafemur markedly incrassate. Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: elongate. Diagnosis: elytra and elytral pubescence markedly bicolored; elytra with tomentose spots. Species group: Cyaneovirens. Relevant species: A. gussakovskiji Alexeev, 1981. Type locality: China, Yunnan province, Yipinglang, altitude 2000 m, 25.03N 101.55E. Type specimens: Primary: Holotype (Fig. 27H), ♂, (EJCB): “Yunnan 2000m, 25.03N 101.55E, Yipinglang 8-10/6.,Vít Kubáň leg. 1993”. Secondary: CHINA: Yunnan: 2 ♀ PT (EJCB): “Yunnan 2000m, 25.03N 101.55E, Yipinglang 8-10/6.,Vít Kubáň leg. 1993”; 3 ♀ PT (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”; 1 ♂ PT (EJCB): “Yunnan, 1500-2500 m, 25°22’N, 98°49’E, 17-24 / 5 Gaoligong mts, Vít Kubáň leg., 1995”. Adult occurence: 5-6 (n=3). Altitude range: 1500-2500 m (n=2). Host plants: Unknown.

106 Distribution: CHINA: Yunnan. Etymology: Noun in apposition; the specific name refers to the ethnic group Hani living mostly in the southwestern Chinese province of Yunnan and scattered across the Ailao Mountains between the Mekong River and the Red River (Yuanjiang).

Agrilus hattorii Nakane Fig. 30G (habitus); Fig. 41I (aedeagus); Fig. 47A (type) hattorii Nakane, 1983 (Agrilus) NAKANE, 1983: 111 112 [#4589] (description). TÔYAMA, 1985: 30 [#3678] (iconography; Japan). AKIYAMA, 1986: 25 [#3199] (faunal records; Japan: Kanagawa prefecture). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 185 [#3962] (faunal record; Japan: Hiroshima Prefecture). AKIYAMA & OHMOMO, 1997: 33 [#4091] (checklist; Japan: Honshu, Shikoku, Kyushu, Tsushima). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2122 [#8576] (world catalog).

Type material: Agrilus hattorii Nakane, 1983 Type locality: Shiomi-cho, Nagoya, Honshu, Japan. Type specimens: Primary: Holotype (Fig. 47A), ♂, (EIHU): “[Japanese grapheme] [h] (Nagoya) 12-5-1938 H. Hattori [h] \ Agrilus hattorii Nakane [h] det. T. Nakane [p] [pink label]”. Described from 15 specimens (HT, PT). Diagnosis: 3.8-6.0 mm; body almost subparallel, elytral apices very broad; pronotum widest at anterior margin; prehumerus absent. Species group: Unassigned. Relevant species: A. lisu sp. nov. Specimens examined: JAPAN: Honshu: 1 (EJCB): “Mt. Senage Seto, Aichi Pref., May 18. 1969, Coll. H. Yamada”; 1 (EJCB): “Sakurakubo, Kuroiso City, Tochigi pref., 2.vi.1991, S. Ohmomo leg.”; 1 ♂, 1 ♀ (EJCB): “Nishinasuno, Tochigi Pref., 28.v.1991, S. Ohmomo”. Kyushu: 1 ♀ (EJCB): “Mt. Kurodake, Mts. Kuysan, Oita Pref., 1.vi.1991, S. Ohmomo”.

Adult occurence: 5-6 (n=4). Host plants: Quercus acutissima: AKIYAMA & OHMOMO, 1997 [#4091]. − Q. serrata: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: JAPAN: Hokkaido, Honshu, Kyushu, Shikoku, Tsushima.

Agrilus hmong Jendek, sp. nov. Fig. 10F (habitus of HT); Fig. 32U (aedeagus)

Description: 9.2-12.9 mm (Holotype 12.8 mm). BODY: Shape: subparallel, Color (dorsally): unicolored, Color (dominant): black. HEAD: Medial impression: present, Impression (extent): vertex and frons, Impression (depth): deep. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): semispherical, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: large, Shape: not protruding from head outline or protruding from head outline, Lower

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA margin: below antennal socket. Antennae: Length: short or moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: visually elongate or visually square, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate or obvious, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles or at level with anterior pronotal angles. Posterior angles: Shape: acute, Apex: sharp. Disk: Convexity: moderately convex, Disk modifications: with obvious tomentose spots, Disk impressions: medial and lateral, Medial impression (shape): entire or anteromedial and posteromedial, Medial impression (deepness): deep, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial, Lateral impressions (deepness): deep. Prehumerus: carinal, Shape: bisinuate, Extent: to half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina, Arc: obvious or moderate. Marginal and submarginal carinae: Interspace: narrow or moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed or impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate, rarely conjoint, Width: wide, Shape: subangulate, subtruncate or cuspidate, Truncation: oblique with projecting sutural angle. Elytral pubescence: absent. Tomentum: spots, Spots (locations): postmedial. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate, Anterior margin: subtruncate. Prosternal process: wide, Shape: narrowed or subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: impressed. ABDOMEN: Sexual modifications in male: impression at intercoxal projection, medial sulcus on ventrite 1 and ventrite 1 attenuate, Apical margin of pygidium: extended into long spine, Apical margin of last ventrite: subtruncate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate, Sinuosity (deepness): very shallow. LEGS: Male sexual modifications: lower margin of metafemur with row of hairs. Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): markedly dorsoventrally bent, Ovipositor: elongate. Diagnosis: pronotal sides with obvious golden-yellow tomentose spots; prehumerus sharply carinal; elytral apices subarcuate or cuspidate or obliquely truncate with projecting sutural angle. Species group: Spinipennis. Relevant species: A. erythrostictus Bourgoin, 1922. Type locality: Northeastern Laos, Hua Phan province, 20°12’N, 104°01’E, Phu Phan Mt., altitude 1500-1900 m. Type specimens: Primary: Holotype (Fig. 10F), ♂, (EJCB): “Lao-NE, Hua Phan pr., ~20°12’N, 104°01’E, Phu Phan Mt., 1500-1900m, 17.v.-3.vi.2007, Vít Kubáň leg.”. Secondary: CHINA: Yunnan: 1 ♀ PT (EJCB): “S China, S Yunnan, 24 km N Jinghong, Naban, 700m, 6.vi.2008”; 1 ♀ PT (EJCB):

107

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA “Yunnan, Jing-Gu County, Mt. Yingpantou, 7-14.vi.2009 local collector”; INDIA: West Bengal: 1 ♂ PT (MNHN): “British Bootang, Maria Basti”; LAOS: Houaphan: 1 ♀ PT (EJCB): “Laos-NE, Houa Phan prov., 20°12-13.5’N, 103°59’.5-104°01’E, Ban Saluei → Phou Pane Mt., 13401870m, 15.iv.-15.v.2008, Lao collectors leg.”. Adult occurence: 4-5-6 (n=4). Altitude range: 700-1900 m (n=3). Host plants: Unknown. Distribution: CHINA: Yunnan. INDIA: West Bengal. LAOS: Houaphan. Etymology: Noun in apposition; the specific name refers to the Asian ethnic group Hmong from the mountainous regions of Vietnam, Laos, Thailand and Myanmar.

Agrilus holzschuhi Jendek Fig. 28B (habitus of HT) holzschuhi Jendek, 1994 (Agrilus) JENDEK, 1994: 31 32 [#1356] (description). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2126 [#8576] (world catalog).

Type material: Agrilus holzschuhi Jendek, 1994 Type locality: China, W Yunnan - Yunlong. Type specimens: Primary: Holotype (Fig. 28B), ♀, (EJCB): “China 30.VI.1993 W Yunnan - Yunlong”. Described from 1 specimen (HT). Diagnosis: 7.4 mm; body with violet tinge; vertex with deep medial sulcus. Species group: Barrati. Relevant species: A. meliboeiformis Descarpentries & Villiers, 1967. Specimens examined: Known only from the holotype.

Host plants: Unknown. Distribution: CHINA: Yunnan.

Agrilus huashanus Jendek Fig. 12F (habitus of HT); Fig. 33P (aedeagus) huashanus Jendek, 2001 (Agrilus) JENDEK, 2001: 3, 10 [#5211] (description). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2128 [#8576] (world catalog).

Type material: Agrilus huashanus Jendek, 2001 Type locality: China, Shaanxi pr., Hua Shan. Type specimens: Primary: Holotype (Fig. 12F), ♂, (EJCB): “China, Shaanxi pr., Hua Shan 17.-21.VI.1991, R. Sauer lgt.”. Secondary: 4 PT (EJCB). Described from 5 specimens (HT, PT). Diagnosis: 6-6.8 mm; body slender, bicolored; pronotum golden-orange, golden-yellow, elytra golden-green or golden-blue; elytral apices with outer marginal spine sharply protruding; basal half of elytral disk only with adsutural strip

and indication of two humeral strips of narrow pubescence. Species group: Plasoni. Relevant species: A. chujoi Kurosawa, 1985. Specimens examined: CHINA: Shaanxi: 2 ♂ PT (EJCB): “China, Shaanxi prov., Hua Shan, 28.7.1990 R. Dunda lgt.” (JENDEK, 2001 [#5211]); 1 ♀ PT (EJCB): “China, Shaanxi pr., Hua Shan, 17 21. VI.1991, R. Dunda lgt.” (JENDEK, 2001 [#5211]); 1 ♂ (EJCB): “China, Shaanxi, Hua Shan, 18 27.7.1990, R. Sauer lgt.”. Sichuan: 1 (EJCB): “China, NW Sichuan prov, Guanxian, 700 m, 31°00’N, 103°36’E, 8 20.VII.1990, J. Kolibáč leg.”; 1 ♂ PT (EJCB): “China, Sichuan, 16.VII.1995, Huatan env., (Shpshin), cca 50 km SW of Yaan, 29”47’N, 102”41’E [sic!], M. Tryzna et O. Safranek lgt.” (JENDEK, 2001 [#5211]).

Adult occurence: 6-7 (n=5). Altitude range: ~700 m (n=1). Host plants: Unknown. Distribution: CHINA: Shaanxi, Sichuan. Remarks: Character cited for A. huashanus by JENDEK (2001: 10 [#5211]) in the key to species is confused and should be changed from “basal half of pronotal disk” to “basal half of elytral disk”.

Agrilus hui Jendek, sp. nov. Fig. 27D (habitus of HT); Fig. 39Q (aedeagus)

Description: 3.3-4.3 mm (Holotype 3.8 mm). BODY: Shape: subparallel, Color (dorsally): bicolored, Color (dominant): green or brown. HEAD: Medial impression: present, Impression (extent): vertex, Epistoma: with raised upper margin. Frons: Shape: flat or moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae. Eyes: Size: large, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short or moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square, Sides: straight or arcuate, Maximal width: at anterior margin or at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate or obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial only, medial and lateral or lateral only, Medial impression (shape): posteromedial only, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow, Lateral impressions (width): narrow. Prehumerus: carinal, Shape: bisinuate, Extent: to anterior pronotal angle, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: bicolored, complementary with pubescence, Alternative color (location): apical portion, Humeral carina: absent. Apices: Arrangement: separate or conjoint, Shape: arcuate or subtruncate. Elytral pubescence: proximal and distal,

108 Proximal: like V or upturned T, Distal: preapical, Color: bicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate or large, Anterior margin: arcuately emarginate, Emargination (deepness): moderate or deep. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: acute, Angles (tips): sharp, Disk: flat or convex. ABDOMEN: Sexual modifications in male: medial sulcus on ventrite 1, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate, Sinuosity (deepness): markedly deep. LEGS: Male sexual modifications: metacoxa medially with cluster of setae and metafemur markedly incrassate. Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: elongate. Diagnosis: sides of pronotum very slightly arcuate; pronotum widest in middle; elytra and elytral pubescence bicolored; aedeagus subparallel. Species group: Cyaneovirens. Relevant species: A. gussakovskiji Alexeev, 1981. Type locality: China, north Henan province, Jiuligou, altitude 550 m, 35°12’N, 112°26’E. Type specimens: Primary: Holotype (Fig. 27D), ♂, (EJCB): “China, N. Henan, 11-12.VI., Jiuligou, 550 m, 35°12’N, 112°26’E, Jaroslav Turna leg. 2007”. Secondary: CHINA: Henan: 9 PT (EJCB): “China, N. Henan, 11-12. VI., Jiuligou, 550 m, 35°12’N, 112°26’E, Jaroslav Turna leg. 2007”; Shaanxi: 1 ♂ PT (EJCB): “China S, Shaanxi, 14-16.6.1995, Qinling Mts. N. slope, 33.56-59’/108.50’ 8001200 m, Changan Co., L.+R. Businský lgt.”; 1 ♂ pT (EJCB): “China, 1000-1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.-13.vi.1998, I. H. Marshal leg.”. Adult occurence: 5-6 (n=3). Altitude range: 550-1300 m (n=3). Host plants: Unknown. Distribution: CHINA: Henan, Shaanxi. Etymology: Noun in apposition; the specific name refers to the Chinese ethnic group Hui typically distinguished by their practice of Islam.

Agrilus hunanus Jendek Fig. 20J (habitus of HT); Fig. 36U (aedeagus)

hunanus Jendek, 2009 (Agrilus) JENDEK in JENDEK & GREBENNIKOV, 2009: 5, 16 17 [#8563] (description).

Type material: Agrilus hunanus Jendek, 2009 Type locality: China, Hunan province, Huitong (= Lincheng, 109°42’E, 26°52’N), 400–600 m. Type specimens: Primary: Holotype (Fig. 20J), ♂, (EJCB): “China: Hunnan province, Huitong, 400–600 m”. Described from 1 specimen (HT).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Diagnosis: 4.1 mm. aedeagus as in Fig. 36U; for further characters see JENDEK & GREBENNIKOV, 2009 [#8563]. Species group: Muscarius. Relevant species: A. apicaureus Jendek, 2009. Specimens examined: Known only from the holotype. Host plants: Unknown. Distribution: CHINA: Hunan.

Agrilus imasakai Tôyama Fig. 11L (habitus) imasakai Tôyama, 1985 (Agrilus) TÔYAMA, 1985: 23, 26 [#2924] (description). TÔYAMA, 1985: 21 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 34 [#4091] (checklist; Japan: Kyushu). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2134 [#8576] (world catalog).

Type material: Agrilus imasakai Tôyama, 1985 Type locality: Mt. Mayuyama, Shimabara City, Nagasaki Pref. Type specimens: Holotype not in NSMT (S. Nomura pers. com. 2001, 2003). Described from 1 ♀ (HT). Diagnosis: 5.1-5.6 mm; elytra with proximal and distal pubescence interrrupted by oval glabrous spot. Species group: Maculifer. Relevant species: A. sudai Kurosawa, 1985. Specimens examined: JAPAN: Kyushu: 1 ♀ (SOCI): “2003.8.2, Tsukahara, Yufuin, Oita, Kyushu, S Jpn, Y. Tsutsumiuchi leg.”.

Adult occurence: 8 (n=1). Host plants: Unknown. Distribution: JAPAN: Kyushu. Remarks: The concept of this taxon was judged from examination of a single female specimen borrowed from the collection of S. Ohmomo.

Agrilus imitans Lewis Fig. 12I (habitus); Fig. 33R (aedeagus); Fig. 47G (type) imitans Lewis, 1893 (Agrilus) LEWIS, 1893: 332 333 [#1722] (description). KERREMANS, 1903: 286 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). OBENBERGER, 1935: 163 [#2158] (note). MIWA & CHÛJÔ, 1936: 17 [#1809] (catalog; Japan). OBENBERGER, 1936: 1000 [#2191] (world catalog). CHÛJÔ & MATUDA, 1940: 65 [#500] (checklist; faunal records; Japan (Kyushu)). IGA, 1947: 11, 16 [#3273] (notes; characters; faunal record; Japan (Shikoku)). KUROSAWA, 1949: 13 [#5028] (characters). CHÛJÔ & KUROSAWA, 1950: 8 [#501] (catalog; faunal records; Japan (Shikoku)). IGA, 1955: 78 [#3828] (iconography; Japan). IGA, 1962: 78 [#3351] (iconography; Japan). KUROSAWA, 1963: 152 [#1618] (characters; Japan). KUROSAWA, 1974: 2 [#4576] (note). KUROSAWA, 1975: 2 [#4578] (note). TÔYAMA, 1985: 23 [#3678] (iconography; Japan).

109

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 34 [#4091] (checklist; Japan: Hokkaido, Honshu, Shikoku, Kyushu, Tsushima). FUKUTOMI & HORI, 2004: 11 12 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). ANONYMOUS, 2007: 32 [#8250] (checklist; Japan: Honshu). BELLAMY, 2008: 2134 [#8576] (world catalog).

Type material: Agrilus imitans Lewis, 1893 Type locality: Main Island ... at Kashiwagi. Type specimens: Primary: Lectotype (Fig. 47G) by present designation, ♂, (BMNH): “Type H. T. [p] [round label with red border] \ Japan, G. Lewis, 1910-320 [p] \ Higo [h] \ Agrilus imitans Lewis Type [h]”. Described from 5 syntypes. Diagnosis: 7.9-10 mm; body wider; male with cluster of short erect setae in middle of prosternum. Species group: Imitans. Relevant species: A. blatteicollis Bourgoin, 1922; A. nepalensis Tôyama, 1988; A. sichuanus sp. nov. Specimens examined: JAPAN: Shikoku: 1 (EJCB): “Mt. Ishizuchisan (Johjusha), Ehime pref., 1.vii.1984, Hikaru Kan leg.”. Tsushima: 1 (EJCB): “Mt. Ohboshiyama, Tsushima Is., 2.vii.1989, K. Adachi”; 1 (EJCB): “Mt. Ohboshiyama, Tsushima Is., 12.vii.1989, K. Adachi”; 1 (EJCB): “Mt. Ohboshiyama, Tsushima Is., 28.v.1989, K. Adachi”; 1 ♂ (EJCB): “Tsutsuzaki, Tsushima Isls., Nagasaki Pref., Kyushu, Japan, 3.V.1993, Ryô Noda leg.”.

Adult occurence: 5-7 (n=6). Faunal record published by CHÛJÔ, M. & MATUDA, 1940 [#500] is also included. Host plants: Celtis sinensis var. japonica: AKIYAMA & OHMOMO, 1997 [#4091]. − Ulmus davidiana var. japonica: AKIYAMA & OHMOMO, 1997 [#4091]. − Zelkova serrata: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: JAPAN: Hokkaido, Honshu, Kyushu, Shikoku, Tsushima.

Shape: bisinuate, Extent: beyond half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: absent or present, Narrowest point: at posterior angles or at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate or conjoint, Width: wide, Shape: arcuate or subtruncate, Truncation: transverse. Elytral pubescence: entire, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: large, Anterior margin: subtruncate. Prosternal process: wide, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): sharp, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Male sexual modifications: metafemur markedly incrassate. Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): medial lobe distinctly wider than parameres. Diagnosis: body smaller and wider; prehumerus bisinuate carinal, reaching beyond half of pronotum, anterior end joining pronotal marginal carina; aedeagus markedly robust. Species group: Unassigned. Relevant species: A. pilosovittatus Saunders, 1873. Type locality: China, Yunnan province, 100 km west of Kunming, Diaolin Natural Reservation.

Agrilus improcerus Jendek, sp. nov.

Type specimens: Holotype (Fig. 30H), ♂, (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”.

Fig. 30H (habitus of HT); Fig. 41J (aedeagus)

Adult occurence: 5-6 (n=1).

Description: Holotype 3.3 mm. BODY: Shape: subparallel, Build: robust, Color (dorsally): unicolored, Color (dominant): black. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: flat or moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture (density): sparse, Sculpture (intensity): superficial. Eyes: Size: small, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short or moderate, Width: slender. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: absent. Posterior angles: Shape: obtuse, Apex: blunt. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow. Prehumerus: carinal,

Host plants: Unknown. Distribution: CHINA: Yunnan. Etymology: The specific name is the Latin adjective improcerus (small, low of stature), referring to the tiny size of the species.

Agrilus inamoenus Kerremans Fig. 19L (habitus); Fig. 36M (aedeagus); Fig. 47J (type) inamoenus Kerremans, 1892 (Agrilus) KERREMANS, 1892: 824 825 [#1398] (description). KERREMANS, 1903: 286 [#1441] (catalog). OBENBERGER, 1936: 1086 [#2191] (world catalog). BAUDON, 1961: 74 [#143] (faunal records; Laos). DESCARPENTRIES & VILLIERS, 1963: 104, 109 [#700] (lectotype designation: “Birmanie (Type, Musée de Gênes)”; characters; faunistic data; Birmanie; Laos; Tonkin; Annam). BAUDON, 1968:

110 131, 143 [#162] (note). PENG ZHONGLIANG & HUANG BANGKAN, 1995: 98 101 [#6309] (characters; biology; China: Fujian). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Fujian [Note: Misidentification]). PENG ZHONGLIANG, 2002: 270 [#6628] (characters; Fujian). JENDEK, 2003: 181 182 [#5823] (remark on lectotype designation). BELLAMY, 2008: 2136 [#8576] (world catalog).

Type material: Agrilus inamoenus Kerremans, 1892 Type locality: Carin Cheba, 900-1100 m. Type specimens: Primary: Lectotype (Fig. 47J), ♀, designated by DESCARPENTRIES & VILLIERS (1963: 104, 109) [#700], (MCSN): “Carin Cheba 900-1100 m L. Fea V XII[18]88 [p] \ Typus [p] [red ink, red border] \ inamoenus Kerr. [h] [black border] \ A. inamoenus Kerr. typus! [h] [yellow label] \ inamoenus Kerr. typus! [h] [yellow label] \ inamoenus Kerr. Type [Kerremans’ MS] \ Museo Civico di Genova [p].” Described from unknown number of syntypes. Diagnosis: 7.8-9.8 mm; elytral pubescent fasciae fragmented into mosaic. Species group: Occipitalis. Relevant species: A. auriventris Saunders, 1873; A. tonkineus Kerremans, 1895. Specimens examined: CHINA: Yunnan: 1 ♀ (NMPC): “Yun’nan’, 30 km SW Czin’lina, 420 m, 26.IV.1956 Khuan Kèzhen’ i dr. [in Russian]”; 1 ♀ (NMPC): “Yun’nan, Cznidun, 1170 m, 2.vii.1956, Kryzhanovskii [in Russian]”. LAOS: Khammouan: 1 ♀ (EJCB): “Laos centr., Khammouan prov., Nakai env., 17º43’N, 105º09’E, 22.V. 8.VI.2001, alt. 500 600 m, E. Jendek & O. Šauša leg.”; 1 ♂ (EJCB): “Laos centr., Khammouan prov., Nakai env., 17º43’N, 105º09’E, 7 25.V.2002, alt. 560 m, M. Štrba leg.”. Louang Namtha: 1 ♂, 3 ♀ (EJCB): “Laos north, 13 24.V.1997, 15 km NW Louang Namtha, N 21°07.5, E 101°21.0, alt. 750±100 m, E. Jendek & O. Šauša leg.”. THAILAND: Chiang Mai: 1 ♂ (EJCB): “Thailand 1000 1600 m, Chiang Mai prov., 20 km NW from Fang 2 5.v.1996, lgt. S. & E. Becvar”. VIETNAM: 3 ♂ (NMPC): “Tonkin, reg. de Hoa Binh, A. de Cooman, 1928”. 2 (EJCB): “S Vietnam Gia Lai Kon Tum, 60 km N Ankhé, 900 m, Con Hanung 19 27.vi., Kannack, 1983”.

Adult occurence: 4-5-6-7 (n=9). Faunal records published by BAUDON, 1961 [#143] are also included. Altitude range: 4201600 m (n=8). Faunal record published by DESCARPENTRIES & VILLIERS, 1963 [#700] is also included. Host plants: Citrus: HUA LI ZHONG, 2002 [#6696]. Distribution: CHINA: Yunnan. LAOS, THAILAND, MYANMAR, VIETNAM. Remarks: Distributional records from Fujian (China) cited by Chinese authors needs verification due to the possible misidentification with similar species.

Agrilus ineptus Kerremans Fig. 14L (habitus); Fig. 34P (aedeagus); Fig. 47K (type) ineptus Kerremans, 1892 (Agrilus) KERREMANS, 1892: 218 [#1402] (description). KERREMANS, 1903: 286 [#1441] (catalog). OBENBERGER, 1936: 1087 [#2191] (world catalog). BELLAMY, 2008: 2138 [#8576] (world catalog).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Type material: Agrilus ineptus Kerremans, 1892 Type locality: Haute Birmanie. Type specimens: Holotype by monotypy (Fig. 47K), ♂, (MNHN): “Hte Birmanie Mines des Rubis 1200m 2300m Doherty 1890 [p] \ ineptus Kerr. Type [h] \ Kerremans vidit 1892 [p] \ collectio Oberthür [p]”. Described from 1 specimen. Diagnosis: 4.2-5.5 mm; body more elongate; pronotum greenish, elytra dark brown; prehumerus absent; aedeagus convex, without angulate sides. Species group: Subrobustus. Relevant species: A. quentini Descarpentries & Villiers, 1963, A. subrobustus Saunders, 1873. Specimens examined: CHINA: Yunnan: 2 ♂, 2 ♀ (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”; 1 ♂ (EJCB): “Yunnan 2000m, 25.03N 101.55E, Yipinglang 8 10/6.,Vít Kubáň leg. 1993”; 12 (EJCB): “Yunnan 1800 2000m, 25.04N 101.55E, Yipinglang 17 20.6.1994,Vít Kubáň leg.”.

Adult occurence: 5-6 (n=3). Altitude range: 1800-2000 m (n=2). Host plants: Unknown. Distribution: CHINA: Yunnan. MYANMAR.

Agrilus insulicola Kerremans Fig. 9L (habitus); Fig. 32O (aedeagus); Fig. 53K (type) insulicola Kerremans, 1912 (Agrilus) KERREMANS, 1912: 205 [#1377] (description). MIWA, 1931: 123 [#6101] (catalog; Formosa). MIWA & CHÛJÔ, 1936: 17 [#1809] (catalog; Japan). OBENBERGER, 1936: 1088 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 356 [#2429] (checklist; China). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Taiwan). MÜHLE, 2003: 47 [#6155] (checklist; Taiwan). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2141 [#8576] (cited as insulicolus; world catalog). suensoni Obenberger, 1940 (Agrilus) syn. nov. PENG OBENBERGER, 1940: 176 177 [#2273] (description). ZHONGLIANG, 1987: 358 [#2429] (checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2315 [#8576] (world catalog).

Type material: Agrilus insulicola Kerremans, 1912 Type locality: Taihorin. Type specimens: Lectotype by present designation, sex not examined, (DEI): “Formosa, Taihorin [see Remarks]”. Secondary: 1 PLT (MNHN) from “Formosa, Kosempo”. Described from unknown number of syntypes. Agrilus suensoni Obenberger, 1940 Type locality: China: Chinkiang. Type specimens: Primary: Lectotype (Fig. 53K) by present designation, ♂, (NMPC): “Nordwestl. China Chinkiang Col. Reitter [p] \ Typus [p] [red label] \ Agrilus Suensoni m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24299 [h] [orange label]”. Secondary: 1 PLT (NMPC). Described from unknown number of syntypes.

111

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Diagnosis: 8.5-10.2 mm; prehumerus extending beyond half of pronotal length; elytral apex with shorter and less sharp median cusp. Species group: Spinipennis. Relevant species: A. cordicollis sp. nov. Specimens examined: CHINA: Taiwan: 1 (EJCB): “Gōkankei, Formosa, 20.ix.1980, K. La leg.”; 13 (EJCB): “Taiwan, Nantou, 1997, Lien Hua Chih, 4 km S Wuchen, 23 27 VI., B. Šiška & T. Spevár lgt.”.

Adult occurence: 6-9 (n=2). Host plants: Unknown. Distribution: CHINA: Jiangsu, Taiwan. Remarks: The lectotype of A. insulicola has been examined in 1996 and complete label data is not currently available. The locality of the paralectotype differs from the originally published type locality, but the specimen is labeled as “Type” by Kerremans’ handwriting, thus it is included in the type series.

Agrilus jaminae Baudon Fig. 10I (habitus); Fig. 32X (aedeagus); Fig. 47N (type) jaminae Baudon, 1968 (Agrilus) BAUDON, 1968: 76, 80 82 [#162] (description). 2145 [#8576] (world catalog).

BELLAMY, 2008:

Type material: Agrilus jaminae Baudon, 1968 Type locality: Laos: Muong Panh. Type specimens: Primary: Holotype (Fig. 47N), ♂, (MHNB): “Muong Panh 20.III.[19]64 [h] Laos (Baudon) [p] \ Type [p] [red label] \ Agrilus jaminae mihi Type [h] A. Baudon det. [p] [blue label]”. Secondary: 7 PT (MHNB); 1 PT (HHCC). Described from unknown number of specimens (HT, AT, PT). Diagnosis: 4.2-5.6 mm; antennae normal, elytral tomentose spots small and rounded, elytral apices sharply acuminate. Species group: Niveoguttatus. Relevant species: A. madeci Baudon, 1968. Specimens examined: BHUTAN: 2 (NHMB): “km 87 von Phuntsholing, 22.5. \ Nat. Hist. Museum Basel Bhutan, Expedition, 1972”. CHINA: Shaanxi: 1 ♂, 2 ♀ (EJCB): “China, S Shaanxi, Qinling Mts. S slope, Xunyangba S+W env., 33°28 37’ / 108°23 33’, 1400 2100m, 5 9.6.[19]95, L.+R. Businský lgt.”; 1 ♂, 3 ♀ (EJCB): “China, 1000 1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v. 13.vi.1998, I. H. Marshal leg.”. Yunnan: 1 ♂ (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”. INDIA: Arunachal Pradesh: 58 (EJCB): “India, W Arunachal Pr., Dirang vicinity, 1500 1800m, 27°21’ 23’N 92°13’ 16’E; L. Dembický leg. 1 10.vi.2004”; 1 ♂ (EJCB): “India, W Arunachal Pr., betw. Dirang & Bomdila Pass, 27°19’N 92°22’E; 1900±300m L. Dembický leg. 12. 16.vi.2004”; 42 (EJCB): “India, W Arunachal Pr., Dirang vicinity, 1800±100m, 27°21’, 92°13’, 2006, P. Pacholátko leg., 8 22.v.”. THAILAND: Chiang Mai: 74 (EJCB): “Thailand N, 21.V. 4.VI.1995, Chiang Dao env., M. Snížek leg.”. Mae Hong Son: 1 ♂ (EJCB): “Thailand, 1 8.5.1993, Soppong Pai, 1800 m, Pacholátko & Dembický leg.”.

Adult occurence: 4-5-6 (n=9). Altitude range: 1000-2100 m (n=6).

Host plants: Unknown. Distribution: BHUTAN. CHINA: Shaanxi, Yunnan. INDIA: Arunachal Pradesh. LAOS: Xiangkhoang. THAILAND: Chiang Mai.

Agrilus japanocarinatus Ohmomo Fig. 29B (habitus of PT); Fig. 40R (aedeagus) japanocarinatus Ohmomo, 2002 (Agrilus) OHMOMO, 2002: 158 160 [#5750] (description). FUKUTOMI & HORI, 2004: 6 7 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2145 [#8576] (world catalog).

Type material: Agrilus japanocarinatus Ohmomo, 2002 Type locality: Masuzawa, Tateiwa-mura, Minami-Aizu, Fukushima Pref. Type specimens: Primary: Holotype, ♂, (NSMT). Secondary: 2 PT (EJCB). Described from 46 specimens (HT, PT). The PT from EJCB collection is illustrated. Diagnosis: 4.2-5.6 mm; elytra with entire pubescence except for postero-pleural portions; basal ventrite in male deeply excavate. Species group: Japanocarinatus. Relevant species: A. bocaki sp. nov., A. leii sp. nov. Specimens examined: Known to us from the two paratypes. Host plants: Fraxinus mandshurica: OHMOMO, 2002 [#5750]. Distribution: JAPAN: Hokkaido, Honshu, Kyushu.

Agrilus jindrai Jendek, sp. nov. Fig. 18C (habitus of HT); Fig. 35S (aedeagus)

Description: 5.7-7.6 mm (Holotype 5.7 mm). BODY: Shape: subparallel, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Medial impression: present, Impression (extent): vertex, rarely vertex and frons, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Shape: rarely markedly convex, Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): irregular. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate or long, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square or transverse, Sides: straight, arcuate or subangulate, Maximal width: at anterior margin, Anterior margin: wider than posterior or subequal to posterior. Anterior lobe: absent or vague, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles. Posterior angles: Shape: obtuse, rarely rectangular, Apex: sharp, rarely blunt. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): deep,

112 Lateral impressions (width): wide. Prehumerus: carinal, Shape: arcuate, rarely bisinuate, Extent: to third of pronotal length, Posterior end: joining posterior pronotal angle, rarely joining posterior pronotal margin, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: narrow or moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior third of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: sometimes spatulate. Apices: Arrangement: conjoint, Width: wide, Shape: subtruncate, Truncation: transverse, Modifications: rarely sutural rim spinuliform. Elytral pubescence: absent. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): moderate. Prosternal process: moderate, Shape: dilated, rarely subparallel, Sides: straight or arcuate, Angles: obtuse, Angles (tips): sharp, Disk: impressed with sides protruding, rarely impressed. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: subangulate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: elongate. Diagnosis: body dorsally bronze; eyes not projecting beyond head outline; pronotum widest at anterior margin; prosternal process with sharp lateral angles. Species group: Betuleti. Relevant species: A. sibiricus Obenberger, 1912. Type locality: China, Shanxi province, Lüliang Shan, road Fangshan-Jiaocheng, Hengjian env., altitude 1000 m. Type specimens: Primary: Holotype (Fig. 18C), ♂, (EJCB): “China, Shanxi, 9.June.2000, Lüliang Shan, 1000m, road Fangshan-Jiaocheng, Hengjian env., Zd. Jindra”. Secondary: CHINA: Shanxi: 1 ♀ PT (EJCB): “China, W Shanxi, 9.VI.2000, 37.7N, 111.6E, Lüliang Shan, road Fangshan-Jiaocheng, Hengjian env., ~1000m, Jaroslav Turna leg.”; 4 ♀ PT (EJCB): “China, Shanxi, 9.June.2000, Lüliang Shan, 1000m, road Fangshan-Jiaocheng, Hengjian env., Zd. Jindra”. Adult occurence: 6 (n=2). Altitude range: 1000 m (n=2). Host plants: Unknown. Distribution: CHINA: Shanxi. Etymology: Patronymic; the species was named in honor of entomologist Zdeněk Jindra, Czech Republic, who collected the holotype.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Agrilus kaluganus Obenberger Fig. 24H (habitus); Fig. 38O (aedeagus); Figs. 47S, 51L (types) kaluganus Obenberger, 1940 (Agrilus; subspecies of olivicolor) OBENBERGER, 1940: 60 [#2292] (description). ALEXEEV, 1964: 22 [#3847] (larval characters; USSR: Voronezh). ALEXEEV, 1964: 15 17 [#3846] (redescription; notes; bionomy; faunal records; Voronezhskaya oblast’; Tul’skaya oblast’). RIKHTER & ALEXEEV, 1965: 297 [#2522] (characters in key; European part of USSR). ALEXEEV, 1968: 881 882 [#50] (redescription; USSR: Voronezh; Tula; Svyatogorsk). ALEXEEV, 1981: 73 [#61] (larval characters in key). COBOS, 1986: 236, 316 [#609] (subgenus Anambus; subspecies of olivicolor). ALEXEEV, 1998: 374 [#4322] (subgenus Quercuagrilus; subgeneric classification). JENDEK, 2005: 12 [#7217] (lectotype designation). JENDEK, 2006: 392 [#7441] (subgenus Quercuagrilus; Palaearctic catalog). BELLAMY, 2008: 2148 2149 [#8576] (subgenus Quercuagrilus; world catalog). pseudoussuricola Alexeev, 1979 (Agrilus; subgenus Quercu agrilus) syn. nov. ALEXEEV, 1979: 130 131 [#57] (description). ALEXEEV, 1989: 479 [#64] (key to species; Amursk; Primorsk; Chita). ALEXEEV & VOLKOVITSH, 1989: 314, 343, 356 [#3241] (characters in key; faunal records; Mongolia). ALEXEEV, 1998: 374 [#4322] (subgenus Quercuagrilus; subgeneric classification). JENDEK, 2006: 392 [#7441] (subgenus Quercuagrilus; Palaearctic catalog). BELLAMY, 2008: 2251 [#8576] (subgenus Quercuagrilus; world catalog).

Type material: Agrilus olivicolor kaluganus Obenberger, 1940 Type locality: Rossia: Kaluga. Type specimens: Primary: Lectotype (Fig. 47S), ♂, designated by JENDEK (2005: 12) [#7217], (NMPC): “Okr. Kalugi [h] [in Russian] \ Typus [p] [red label] \ A. olivicolor ssp. Kaluganus m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24930 [h] [orange label]”. Described from unknown number of syntypes. Agrilus pseudoussuricola Alexeev, 1979 Type locality: Vladimiro-Aleksandrovo, Partizanskii r-n [in Russian]. Type specimens: Primary: Holotype (Fig. 51L), ♂, (ZIN). Secondary: 2 PT (ZIN). Described from 5 specimens (HT, PT). Diagnosis: 3.8-5.4 mm; antennomeres in male not enlarged; ventral side in male glabrous. Species group: Sulcicollis. Relevant species: A. latipalpis Jendek, 2007. Specimens examined: CHINA: Beijing: 2 ♀ (EJCB): “China, Beijing distr., Yan Shan, Dongling Mts., 39º58’N, E 115º26’E, 1400 m, Xiaolongmen, 15 16.VI.2001”. Hebei: 1 ♀ (EJCB): “China, Hebei/Nei Mongol, pass Chengde Chifeng, 41.6N, 118.2E, 14 16. VI., Jaroslav Turna leg. 2001”. RUSSIA: Buryatiya Respublika: 1 ♀ (EJCB): “USSR Baikal, Chamar Daban, Babuschkino, 3 9. VIII.1990, R. Sauer lgt.”. Moskovskaya oblast’: 2 ♀ (ZIN): “Glavn. Bot. Sad, 2 Moskva 6.6.1963 K. Arnol’di [in Russian]”; 2 ♂, 2 ♀ (ZIN): “Uzkoe, Moskovsk., dubrava, K. Arnol’di, 1968 [in Russian]”. Primorskiy kray: 1 ♀ (EJCB): “SU Ussuri reg. Jasnoe 400 m 12 19.7.1989 M. Nikodým lgt.”; 1 ♀ (EJCB): “USSR, Ussuri prov. Arsenev env., VII.1990, leg. B. Šiška & R. Čermák”; 2 ♀ (ZIN): “Primorskii krai, Spassk, les, 15.06.1990, Belokobyl’skij [in Russian]”; 1 (AGCM): “Russia, Primorskij kraj, Ussurijsk, 13.8.1992, Leg. J. Sawoniewicz”; 2 ♂ (EJCB): “Rossia, Primorskii krai, Novokachalinsk, 3 ber. oz. Khanka, dubnyak, luga, 4 7. viii.2006, S. Belokobyl’skij [in Russian]”; 1 ♀ (EJCB): “Russia,

113

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Primorsky Kray, Kaymanovka vill., N43°37’, E132°14’, 7 11. vii.2007, 100 m, V. Grebennikov”. Sakhalinskaya oblast’: 1 ♀ (EJCB): “Russland, S. Sakhalin Novoaleksandrovska, 25.06.1991, Putchkov leg.,”. Tyva Respublika: 1 (ZIN): “Tuva, 14.vii.1949”.

Adult occurence: 6-7-8 (n=15). Faunal records published by ALEXEEV, 1979 [#57] (as pseudoussuricola) are also included. Altitude range: 100-1400 m (n=3). Host plants: Corylus: ALEXEEV, 1964 [#3847]; RIKHTER & ALEXEEV, 1965 [#2522]; ALEXEEV, 1981 [#61]. – ALEXEEV, 1989 (as pseudoussuricola) [#64]. − C. avellana: ALEXEEV, 1964 [#3846]; ALEXEEV, 1968 [#50]. Distribution: CHINA: Beijing, Hebei, Nei Mongol. JAPAN: Tsushima. MONGOLIA. RUSSIA: Amurskaya oblast’, Buryatiya Respublika, Kaluzhskaya oblast’, Moskovskaya oblast’, Omskaya oblast’, Primorskiy kray, Sakhalinskaya oblast’, Tul’skaya oblast’, Tyva Respublika; Voronezhskaya oblast’, Zabaykalskiy kray. UKRAINE. Remarks: The holotype of A. pseudoussuricola in Alexeev’s collection was examined during the first author’s visit to Alexeev in 1994.

Agrilus kawarai Kurosawa Fig. 9H (habitus of HT) kawarai Kurosawa, 1963 (Agrilus) KUROSAWA, 1963: 107 108 [#1617] (description). KUROSAWA, 1963: 152 [#1618] (characters; Japan). KUROSAWA, 1974: 1 [#4577] (note). TÔYAMA, 1985: 24 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 34 [#4091] (checklist; Japan: Honshu, Shikoku, Kyushu). NONNAIZAB et al., 1999: 112 [#8247] (checklist; China: Inner Mongolia). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2150 [#8576] (world catalog).

Type material: Agrilus kawarai Kurosawa, 1963 Type locality: Shizuoka Pref.: Foot of Mt. Hijiri-dake. Type specimens: Primary: Holotype (Fig. 9H), ♂, (NSMT): “Foot of Mt. Hijiridake S-Alps Shizuoka Pref. 3. VIII. 1956 M. Kawara [h] \ Holotype [p] Agrilus kawarai Y. Kurosawa, 1959 [last two numbers line through] 63 [h] [red label with black border]”. Secondary: 3 PT (NSMT). Described from 7 specimens (HT, AT, PT). Diagnosis: 8.5-8.7 mm; elytra with 3 pairs of whitish spots; elytral apices separately arcuate; male without sexual modifications on ventral side; sternal lobe narrow, arcuately emarginate; prosternal process subparallel; last ventrite arcuate. Species group: Spinipennis. Relevant species: A. spinipennis Lewis, 1893. Specimens examined: JAPAN: Honshu: 1 ♀ (SOCI): “1987.7.1, Maki, Kamuushi, Hiroshima, W Jpn, M. Akiyama leg.”.

Adult occurence: 7 (n=4). Faunal records published by KUROSAWA, 1963 [#1617] are also included. Host plants: Unknown. Distribution: JAPAN: Honshu, Kyushu, Shikoku.

Remarks: The distributional record cited without supporting data from Inner Mongolia (China) by NONNAIZAB et al., 1999 [#8247] was omitted.

Agrilus komareki komareki Obenberger Fig. 30I (habitus); Fig. 41K (aedeagus); Figs. 47Y, 52C (types) komareki Obenberger, 1926 (Agrilus) OBENBERGER, 1926: 102 103 [#2054] (description). OBENBERGER, 1926: 656 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 17 [#1809] (catalog; Japan). OBENBERGER, 1936: 1004 [#2191] (world catalog). KUROSAWA, 1963: 154 [#1618] (characters; Japan). KUROSAWA, 1974: 1 [#4575] (characters; notes). KUROSAWA, 1974: 1 [#4577] (note). TÔYAMA, 1985: 22 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). LI JINGKE, 1992: 92 [#6755] (checklist; China: Jilin). JENDEK, 1994: 19 [#1355] (lectotype designation; synonymy; characters; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 185 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 34 [#4091] (checklist; Japan: Hokkaido, Honshu, Shikoku, Kyushu). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). FUKUTOMI & HORI, 2004: 14 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 393 [#7441] (subgenus Robertius; Palaearctic catalog). BELLAMY, 2008: 2152 [#8576] (subgenus Robertius; world catalog). ronino Obenberger, 1935 (Agrilus) OBENBERGER, 1935: 167 [#2158] (description). OBENBERGER, 1936: 1029 [#2191] (world catalog). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). JENDEK, 1994: 19 [#1355] (synonym of komareki; lectotype designation). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 34, 36 [#4091] (synonym of komareki on page 34 as well as synonym of nicolanus on page 36). NAKAMURA, KAMEYAMA & KATAYAMA, 2001:66 (faunal records; Japan) [#8761] JENDEK, 2006: 393 [#7441] (synonym of komareki; Palaearctic catalog). BELLAMY, 2008: 2152 [#8576] (synonym of komareki; world catalog).

Type material: Agrilus komareki Obenberger, 1926 Type locality: Japan: Nakano, Tokyo. Type specimens: Primary: Lectotype (Fig. 47Y), ♀, designated by JENDEK (1994: 19) [#1355], (NMPC): “Nakano, Tokyo Japan, 12.VI.23 [h] \ Typus [p] [red label] \ Agrilus Komareki m type m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Agrilus ronino Obenberger, 1935 Type locality: Japan: Hitoyoshi. Type specimens: Primary: Lectotype (Fig. 52C), ♀, designated by JENDEK (1994: 19) [#1355], (NMPC): “Hitoyoshi Japan [h] \ Typus [p] [red label] \ Agrilus Ronino m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Diagnosis: 4.2-5.1 mm; body almost subparallel, elytral apices broad; aedeagus robust. Species group: Unassigned. Relevant species: Unknown. Specimens examined: JAPAN: Honshu: 1 ♂ (EJCB): “Japan, Honshu, Gumma Pref., Numata shi, 1.vi.1955, Coll. T. Takei”; 1 (MNCA): “Ikutaryokuchi, Kawasaki City, Kanagawa Pref. 31.V.[19]79 leg. T. Horiguchi” (JENDEK, 1994 [#1355]); 1 ♂ (EJCB): “Henokara, Yamanashi Pref., 21.V.[19]81 leg. N. Kobayachi”

114 (JENDEK, 1994 [#1355]); 2 (EJCB): “Sakuramura, Ibaraki Pref., 19.V.1984 S. Ohmomo” (JENDEK, 1994 [#1355]); 3 (EJCB): “Ishidojyuku, Kitamoto City, Saimata Pref., Japan 11 13.V. 1986 K. Akiyama leg.” (JENDEK, 1994 [#1355]); 1 ♀ (EJCB): “Japan, c., Tochigi pref., Oashi gawa, Kanuma shi, 29.vii.1992”.

Adult occurence: 5-6-7 (n=7). Host plants: Morus bombycis: AKIYAMA & OHMOMO, 1997 [#4091]. − Symplocos prunifolia: AKIYAMA & OHMOMO, 1997 [#4091]. − Ulmus davidiana var. japonica: AKIYAMA & OHMOMO, 1997 [#4091]. Erroneous: Salix: HUA LI ZHONG, 2002 (as sapporoensis) [#6696]. Distribution: JAPAN: Honshu, Kyushu, Shikoku. Remarks: Distributional records from Anhui and Jilin (China) cited without supporting data by LI JINGKE, 1992 [#6755]; HUA LI ZHONG, 2002 (as sapporoensis) [#6696] and MENG XU WU, 2003 (as sapporoensis) [#8101] were omitted.

Agrilus komareki sapporoensis Obenberger nom rev. Fig. 30J (habitus); Fig. 41L (aedeagus); Figs. 45U, 52O (types) sapporoensis Obenberger, 1935 (Agrilus) nom. rev. OBENBERGER, 1935: 168 [#2158] (description). MIWA & CHÛJÔ, 1936: 20 [#1809] (catalog; Japan). OBENBERGER, 1936: 1039 [#2191] (world catalog). KUROSAWA, 1949: 15 [#5028] (cited as sapoensis; characters). CHÛJÔ & KUROSAWA, 1950: 10 11 [#501] (catalog; faunal records; Japan (Shikoku)). KUROSAWA, 1950: 1114 [#3274] (note). IGA, 1955: 80 [#3828] (iconography; Japan). KUROSAWA, 1956: 1114 [#8107] (characters; Japan). IGA, 1962: 80 [#3351] (iconography; Japan). KUROSAWA, 1963: 154 [#1618] (subspecies of komareki; characters; Japan). KUROSAWA, 1974: 1, 4 [#4577] (notes). JENDEK, 1994: 19 [#1355] (synonym of komareki; lectotype designation). AKIYAMA & OHMOMO, 1997: 34, 36 [#4091] (synonym of komareki on page 34 as well as synonym of moerens on page 36). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Henan). MENG XU WU, 2003: 45 [#8101] (checklist; China: Anhui). JENDEK, 2006: 393 [#7441] (synonym of komareki; Palaearctic catalog). BELLAMY, 2008: 2152 [#8576] (synonym of komareki; world catalog). emisi Ohmomo, 2005 (Agrilus; subspecies of komareki) syn. nov. OHMOMO, 2005: 238 239 [#7253] (description). BELLAMY, 2008: 2152 [#8576] (subspecies of komareki; world catalog).

Type material: Agrilus sapporoensis Obenberger, 1935 Type locality: Japan: Sapporo. Type specimens: Primary: Lectotype (Fig. 52O), ♀, designated by JENDEK (1994: 19) [#1355], (NMPC): “Japan: Sapporo [h] \ Typus [p] [red label] \ Agrilus sapporoënsis m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Agrilus komareki emisi Ohmomo, 2005 Type locality: Higashikawa-cho, Kamikawa-gun, Hokkaido. Type specimens: Primary: Holotype (Fig. 45U), ♂ (NSMT). Secondary: 1 PT (EJCB). Described from 17 specimens (HT, PT).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Diagnosis: 4.8-6.6 mm, “prehumeral carinae indistinct, sometimes absent in middle” (OHMOMO, 2005 [#7253]). Species group: Unassigned. Relevant species: A. komareki Obenberger, 1926. Specimens examined: Known to us only from the type specimens.

Host plants: See A. komareki komareki. Distribution: JAPAN: Hokkaido. Remarks: OHMOMO, 2005 [#7253] proposed the new subspecies A. komareki emisi from Hokkaido, but he overlooked the priority of the name sapporoensis Obenberger, 1935.

Agrilus kostali Jendek Fig. 13H (habitus of HT); Fig. 34C (aedeagus) kostali Jendek, 2007 (Agrilus) JENDEK, 2007: 42 [#8006] (description). [#8576] (world catalog).

BELLAMY, 2008: 2153

Type material: Agrilus kostali Jendek, 2007 Type locality: China, W Sichuan, Jiulong Co., Taka He valley, 2500-4000 m, 28º45’-29º09’ & 101º42’-50’. Type specimens: Primary: Holotype (Fig. 13H), ♂, (EJCB): “China, W Sichuan, Jiulong Co., Taka He valley, 25004000 m, 28º45’-29º09’ & 101º42’-50’, 2-10.VII. 2001”. Secondary: 7 PT (EJCB). Described from 8 specimens (HT, PT). Diagnosis: 6.9-8.6 mm; prehumerus filamentary; elytra longer, with more elongate apical third; aedeagus longer and slender. Species group: Plagiatus. Relevant species: A. plagiatus Ganglbauer, 1889. Specimens examined: CHINA: Sichuan: 4 ♂ PT, 3 ♀ PT (EJCB): “China, W Sichuan, Jiulong Co., Taka He valley, 2500 4000 m, 28°45’ 29°09’ & 101°42’ 50’, 2 10.VII. 2001” (JENDEK, 2007 [#8006]).

Adult occurence: 7 (n=1). Altitude range: 2500-4000 m (n=1). Host plants: Unknown. Distribution: CHINA: Sichuan.

Agrilus koyoi Ohmomo Fig. 30K (habitus of PT); Fig. 41M (aedeagus) koyoi Ohmomo, 2002 (Agrilus) OHMOMO, 2002: 162 163 [#5750] (description). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2153 [#8576] (world catalog).

Type material: Agrilus koyoi Ohmomo, 2002 Type locality: Masuzawa, Tateiwa-mura, Minamiazu, Fukushima Pref. Type specimens: Primary: Holotype, ♂, NSMT (OHMOMO, 2002 [#5750]). Secondary: 3 PT (EJCB). Described from

115

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA 21 specimens (HT, PT). Paratype (Fig. 30K) from EJCB collection is illustrated. Diagnosis: 5.1-6.2 mm; dorsally unicolored; anterior pronotal lobe not reaching beyond anterior pronotal angles. Species group: Unassigned. Relevant species: A. sugiurai Jendek, 2007. Specimens examined: Known to us from three paratypes.

Host plants: Unknown. Distribution: JAPAN: Honshu.

Agrilus kuangcenensis Obenberger sp. inquirenda kuangcenensis Obenberger, 1940 (Agrilus) OBENBERGER, 1940: 114 [#2297] (description). HUA LI ZHONG, 2002: 90 [#6696] (cited as kuangcensis; checklist; northeastern China). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2154 [#8576] (world catalog).

Type material: Agrilus kuangcenensis Obenberger, 1940 Type locality: Mandjuria: Kuang-čen. Type specimens: Not located. Type is not in NMPC. Described from unknown number of syntypes. Host plants: Unknown. Distribution: CHINA: Jilin. Remarks: This taxon is species inquirenda and its name is nomen dubium.

posterior pronotal angle, Anterior end: distant from pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior third of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Shape: arcuate. Elytral pubescence: entire except glabrous part at apical half, Color: unicolored, Character: homogenous. STERNUM: Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: subtruncate or arcuately emarginate, Emargination (deepness): shallow, Emargination (width): narrow. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: acute or rectangular, Angles (tips): sharp, Disk: impressed or impressed with sides protruding. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate. LEGS: Male sexual modifications: metafemur markedly incrassate. Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed, Ovipositor: elongate. Diagnosis: prehumerus extending to half of pronotal length with anterior end widely distant from lateral pronotal margin. Species group: Cyaneovirens. Relevant species: A. yamabusi Miwa & Chûjô, 1940. Type locality: China, Yunnan province, Lijiang.

Agrilus kucerai Jendek, sp. nov. Fig. 26L (habitus of HT); Fig. 39M (aedeagus)

Description: 4.3-5.6 mm (Holotype 5.5 mm). BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): green. HEAD: Shape: markedly convex, Medial impression: present, Impression (extent): vertex, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (density): dense. Eyes: Size: moderate, Shape: not protruding from head outline, rarely protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square or transverse, Sides: straight or arcuate, Maximal width: at middle or at posterior margin, Anterior margin: narrower than posterior. Anterior lobe: moderate or obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles or projecting beyond anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow. Prehumerus: carinal, Shape: arcuate, Extent: beyond half of pronotal length, Posterior end: joining

Type specimens: Primary: Holotype (Fig. 26L), ♂, (EJCB): “China-Shaanxi pr. Hua Shan 17-21.VI.1991 R. Dunda & R. Sauer lgt.”. Secondary: CHINA: Hubei: 1 PT (EJCB): “P.R. China, Hubei, Dabieshan, N31˚07’06’’ E115˚48’57”, 11-21.vi.2008, 640 m, V. Grebennikov leg.”; Shaanxi: 2 PT (EJCB): “China, Shaanxi pr., Hua Shan, 17-21.VI.1991, R. Dunda lgt.”; 9 PT (EJCB): “China-Shaanxi pr. Hua Shan 1721.VI.1991 R. Dunda & R. Sauer lgt.”; 9 PT (EJCB): “ChinaShaanxi, Hua-Shan, 30.6.1.7.2000”; 1 PT (EJCB): “China S, Shaanxi, 14-16.6.1995, Qinling Mts. N. slope, 33.5659’/108.50’ 800-1200 m, Changan Co., L.+R. Businský lgt.”; 13 PT (EJCB): “China, Shaanxi, 110 km E of Xi’An, HuaShan, ~500 m, 23-24.vi.1998, I. H. Marshal leg.”. Adult occurence: 6 (n=6). Altitude range: 500-1200 m (n=3). Host plants: Unknown. Distribution: CHINA: Hubei, Shaanxi. Etymology: Patronymic; the species was named in honour of Emil Kučera, Czech Republic, who collected some of the type specimens.

116

Agrilus kurumi Kurosawa Fig. 20L (habitus); Fig. 36V (aedeagus); Fig. 48G (type) kurumi Kurosawa, 1957 (Agrilus) KUROSAWA, 1957: 187 189 [#1615] (description). KUROSAWA, 1963: 154 [#1618] (characters; Japan). KUROSAWA, 1974: 1, 2 [#4577] (note). ALEXEEV, 1979: 136 137 [#57] (subgenus Austragrilus; notes; faunal records; Russia: Primorsk). TÔYAMA, 1985: 20 [#3678] (iconography; Japan). ALEXEEV, 1989: 482 [#64] (key to species; Primorsk; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). LI JINGKE, 1992: 92 [#6755] (cited as kulumi; checklist; northeastern China). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 35 [#4091] (checklist; Japan: Hokkaido, Honshu; Russia: southeastern Siberia). ALEXEEV, 1998: 374 [#4322] (subgenus Austragrilus; genotype of the subgenus Austragrilus). FUKUTOMI & HORI, 2004: 5 6 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 390 [#7441] (subgenus Austragrilus; Palaearctic catalog). BELLAMY, 2008: 2154 [#8576] (subgenus Austragrilus; world catalog). JENDEK & GREBENNIKOV, 2009: 5, 28 29 [#8563] (characters in key; HT examined; faunal records; distributional summary; hosts; North China (Shaanxi), Japan (Hokkaido, Honshu), eastern Russia (Primorye)). VOLKOVITSH, 2009: 134 [#8850] (subgenus Austragrilus; faunal record; Russia: Primorskiy kray).

Type material: Agrilus kurumi Kurosawa, 1957 Type locality: Komanoyu, near Kiso-Fukushima, Nagano Pref., Japan. Type specimens: Primary: Holotype (Fig. 48G), sex not examined, (NSMT): “[Japanese grapheme] Kiso 5. VII. 1947 T. Nakane [h] \ Holotype [p] Agrilus kurumi Y. Kurosawa, 1957 [h] [red label]”. Described from 14 specimens (HT, AT, PT). Diagnosis: 4.1-5.0 mm; pronotum flat, with sides less arcuate; elytral pubescence obsolete. See revision of JENDEK & GREBENNIKOV, 2009 [#8563]. Species group: Muscarius. Relevant species: A. mirei Descarpentries & Villiers, 1963. Specimens examined: CHINA: Shaanxi: 1 (EJCB): “China, Shaanxi prov., 21 23.June 1998 \ Qing Ling Shan mts, 1000 m road Baoji Taibai vill., pass 40 km S Baoji, Zd. Jindra lgt.” (JENDEK & GREBENNIKOV, 2009 [#8563]). JAPAN: Honshu: 1 ♂, 1 ♀ (EJCB): “Mt. Hanazono, Ibaraki Pref., Japan, 23.VII.1977, A. Nishiyama leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). RUSSIA: Primorskiy kray: 1 (EJCB): “SSR, Primorsk, Pogranichnyi, Pisarenko” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 (EJCB): “Prim. kr. Chuguevskii r n., s. Zavetnoe, kedrovo shirokol. les, 26.8.76g, V. Kuznetsov [in Russian]” (JENDEK & GREBENNIKOV, 2009 [#8563]); 2 (ZIN): “Pogranichnyi D.V., Primor’e, na list. Juglans, 6.viii.1980 S. Konovalov [in Russian]”; 1 ♀ (EJCB): “Sib. or. m., Primorje, Ussuri res., 20.7.1990, Kadlec + Voříšek lg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 2 (JMCZ): “SU or, 18.7.1990, Primorskij reg., Novočugujevka, Ing. Kantner lgt.”; 1 ♀ (EJCB): “Russia, Primorskij k., Kajmanovka, 10 22.vii.1993, Z. Kletečka leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 ♂ (EJCB): “SSSR, Primorskij k., Kajmanovka 10 22.vii.1993, Z. Kletečka leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Lazo env., 10.07.04 leg. М.& L. Smirnov’s” (JENDEK & GREBENNIKOV, 2009 [#8563]).

Adult occurence: 6-7-8 (n=13). Faunal records published by KUROSAWA, 1957 [#1615] are also included. Altitude range: ~1000 m (n=1).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Host plants: Juglans sieboldiana: KUROSAWA, 1957 [#1615]; AKIYAMA & OHMOMO, 1997 [#4091]. − J. mandshurica: ALEXEEV, 1979 [#57]; ALEXEEV, 1989 [#64]. Distribution: CHINA: Shaanxi. JAPAN: Hokkaido, Honshu. RUSSIA: Primorskiy kray.

Agrilus lacrima Jendek, sp. nov. Fig. 16A (habitus of HT)

Description: Holotype 7.1 mm. BODY: Shape: cuneiform, Color (dorsally): bicolored, Color (dominant): green. HEAD: Medial impression: absent, Epistoma: in plane with frons. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): semispherical, Sculpture (intensity): superficial. Eyes: Size: moderate, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: visually square, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles. Posterior angles: Shape: rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk modifications: with two black longitudinal stripes, Disk impressions: medial and lateral or lateral only, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial, Lateral impressions (deepness): deep, Lateral impressions (width): narrow. Prehumerus: absent. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: bicolored, Alternative color (location): perisutural drop or pleural portions, Humeral carina: absent, Apical part: markedly attenuate. Apices: Arrangement: separate, Width: narrow, Shape: arcuate. Tomentum: spots, Spots (locations): postmedial. STERNUM: Metasternal projection: flat. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): shallow, Emargination (width): moderate. Prosternal process: moderate, Shape: narrowed, Sides: straight, Angles: absent, Disk: flat. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Ovipositor: elongate. Diagnosis: frons moderately convex, narrower; two black longitudinal pronotal stripes divided by wide green interspace; ventral side except for prosternum golden carmine. Species group: Smaragdinus. Relevant species: A. odetteae Baudon, 1968. Type locality: China, Yunnan province, Gaoligong mts., 24°57’N, 98°45’E, altitude 2200-2500 m.

117

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Type specimens: Holotype (Fig. 16A), ♀, (EJCB): “Yunnan, 2200-2500 m, 24°57’N, 98°45’E, 8-16/5 Gaoligong mts, Vít Kubáň leg., 1995”.

Agrilus lahu Jendek, sp. nov.

Adult occurence: 5 (n=1). Altitude range: 2200-2500 m (n=1).

Description: 4.7-4.9 mm (Holotype 4.9 mm). BODY: Shape: subparallel, Build: slender, Color (dorsally): bicolored, Color (dominant): green. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): deep, Epistoma: with raised upper margin. Frons: Shape: flat, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: protruding from head outline, Sculpture on vertex: rugae, Sculpture (intensity): superficial. Eyes: Size: large, Shape: not protruding from head outline, Lower margin: in line with antennal socket or below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: visually square or transverse, Sides: arcuate or subangulate, Maximal width: at anterior margin or at middle, Anterior margin: subequal to posterior. Anterior lobe: obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial or posteromedial only. Prehumerus: obsolete or carinal, Shape: arcuate, Extent: beyond half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: distant from pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: narrow or moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior angles or at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: bicolored, complementary with pubescence, Alternative color (location): apical portion, Humeral carina: absent, Apical part: spatulate. Apices: Arrangement: separate, Width: wide, Shape: arcuate or subtruncate, Truncation: transverse. Elytral pubescence: proximal and distal, Proximal: like V or upturned T, Distal: preapical, Color: bicolored. Tomentum: stripes or spots, Spots (locations): humeral. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): shallow. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: acute or rectangular, Angles (tips): blunt, Disk: impressed. ABDOMEN: Sexual modifications in male: medial sulcus on ventrite 1, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate, Sinuosity (deepness): markedly deep. LEGS: Male sexual modifications: metacoxa medially with cluster of setae and metafemur markedly incrassate. Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical.

Host plants: Unknown. Distribution: CHINA: Yunnan. Etymology: The specific name is the Latin noun lacrima (tear) in apposition. It refers to the tear-like elytral spot of the species.

Agrilus laetecyanescens Obenberger Fig. 18I (habitus); Fig. 35Y (aedeagus); Fig. 48J (type) laetecyanescens Obenberger, 1940 (Agrilus) OBENBERGER, 1940: 178 [#2273] (description). PENG ZHONGLIANG, 1987: 356 [#2429] (checklist; China). JENDEK, 2000: 504 [#4851] (lectotype designation). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2156 [#8576] (world catalog).

Type material: Agrilus laetecyanescens Obenberger, 1940 Type locality: China: Yunnan. Type specimens: Primary: Lectotype (Fig. 48J), ♀, designated by JENDEK (2000: 504) [#4851], (NMPC): “China: Yunnan [province] [h] \ Typus [p] [red label] \ Agrilus laetecyanescens m. Type [h] [Obenberger’s MS] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24796 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 6.2-7.5 mm; elytra glabrous; elytral apices separately arcuate. Species group: Laetecyanescens. Relevant species: A. yunnanus Obenberger, 1927. Specimens examined: CHINA: Sichuan: 1 (HMMG): “Sichuan Xiangchen 3000 m, 22.vi.1982”; 1 (HMMG): “Sichuan Xiangchen 2700 m, 27.vi.1982”. Yunnan: 1 ♀ (EJCB): “China, Yunnan, Mt. Jizushan, 2000m, 1 3.vii.1993, Bolm leg.”; 1 (EJCB): “China Yunnan, 10 15.6.[19]94, Litiang , lgt. E. Kučera”; 10 (EJCB): “China, Yunnan, Lijiang, 19.VI.1995, Šiška & Spevár & Pekarovič”; 6 (EJCB): “China Yunnan, Lijiang, 11.6. 15.6.1995, lgt. E. Kučera”; 9 (EJCB): “China, Yunnan, Lijiang, 8 24.VI.1994, Šiška & Spevár”. LAOS: Houaphan: 1 ♂ (EJCB): “NE Laos, Hua Phan prov., 2004, 20°13’N, 103°59’E, 6 18.iv., Ban Saluei, Phu Phan Mt. env., 2004, J. Bezděk leg., 1300 2000m”. Louangphrabang: 1 ♂ (EJCB): “Laos N, 24.iv. 16.v.1999, Louang Phrabang prov. 20°33 4’N 102°14’E, Ban Song Cha (5km W) ± 1200 m, Vít Kubáň leg.”. NEPAL: 1 ♂ (EJCB): “W Nepal, 1992, Tiknedhanga, 1500 m, 8.vi., leg. Jiří Moravec”.

Adult occurence: 4-5-6-7 (n=10). Altitude range: 12003000 m (n=6). Host plants: Unknown. Distribution: CHINA: Sichuan, Yunnan. LAOS. NEPAL.

Fig. 27I (habitus of HT); Fig. 40B (aedeagus)

Diagnosis: elytra with tomentose ornament; elytral apices distinctly dilated. Species group: Cyaneovirens. Relevant species: A. gussakovskiji Alexeev, 1981. Type locality: China, Yunnan province, 100 km west of Baoshan, Gaoligongshan nat. res.

118 Type specimens: Primary: Holotype (Fig. 27I), ♂, (EJCB): “China, Yunnan, 14-21.VI.1993, 100 km W of Baoshan, Gaoligongshan nat. res., E. Jendek & O. Šauša leg.”. Secondary: CHINA: Yunnan: 1 ♀ PT (EJCB): “China, Yunnan, 14-21.VI.1993, 100 km W of Baoshan, Gaoligongshan nat. res., E. Jendek & O. Šauša leg.”. Adult occurence: 6 (n=1). Host plants: Unknown. Distribution: CHINA: Yunnan. Etymology: Noun in apposition; the specific name refers to the Lahu, an ethnic group of Southeast Asia and Yunnan province of China.

Agrilus laticornis (Illiger) Fig. 24F (habitus); Fig. 38M (aedeagus); Figs. 42AA, 52V (types) laticornis (Illiger, 1803) (Buprestis) ILLIGER, 1803: 243 244 [#1310] (description). SCHÖNHERR, 1817: 245 [#3895] (Buprestis). DEJEAN, 1821: 30 [#4046] (Buprestis; catalog). DEJEAN, 1833: 83 [#689] (catalog). DEJEAN, 1836: 94 [#692] (catalog). MANNERHEIM, 1837: 116 [#3018] (characters). GORY & LAPORTE, 1839: 54 55 [#3738]. RATZEBURG, 1844: unpaged [#8764] (Buprestis; checklist of pests). SCHAUM, 1852: 45 [#8335] (catalog). KIESENWETTER, 1857: 125, 142 143 [#3015] (cited as laticornis on page 125 and laticollis on page 142 143; characters in key, redescription; notes; Germany). MARSEUL, 1857: 92 [#4371] (checklist; Europe). REDTENBACHER, 1857: 477 [#4134] (characters in key; Austria). MARSEUL, 1866: 437, 458 459 [#676] (characters in key; redescription; references; distribution overview). GEMMINGER & HAROLD, 1869: 1441 [#1057] (catalog). SAUNDERS, 1870: 19 [#2551] (catalog). SAUNDERS, 1871: 120 [#2552] (catalog). REDTENBACHER, 1874: 515 [#3784] (characters in key; Austria). MARSEUL, 1887: 251 [#4707] (catalog). KERREMANS, 1892: 246, 262 [#1397] (cited as angustulus; catalog). RUPERTSBERGER, 1894: 161, 287 [#4465] (larval bibliography; catalog). ABEILLE DE PERRIN, 1897: 3 [#19] (characters in key). KERREMANS, 1903: 275 [#1441] (catalog). CAILLOL, 1913: 514 [#4053] (biology; faunal records; France). JAKOBSON, 1913: 796 [#4386] (catalog; Russia and Europe). BEDEL, 1921: 203, 215 [#170] (characters in key; references; France). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1005 1008 [#2191] (world catalog). RIKHTER, 1945: 167 [#3507] (characters in key; Latvia; Lithuania; Ukraine; Krym; Caucasia). RIKHTER, 1948: 394 [#3879] (characters in key; Kharkovsk; Krym; Caucasia). SCHAEFER, 1949: 353, 364 366, 373, 490 [#2565] (characters; references; faunal records; biology; notes; ditributional overview; notes; France). RIKHTER, 1950: 159 [#3881] (characters in key). HORION, 1955: 81 [#1285] (distribution overview; faunal records; biology; Deutschland; Österreich). TER MINASYAN, 1955: 452 [#7893] (host plants; distribution; references). ALEXEEV, 1959: 18 [#3229] (characters in key; Pribaltika; Ukraine; Krym; Caucasia). RIKHTER & ALEXEEV, 1965: 296 [#2522] (characters in key; Pribaltika; west Ukraine; Krym; Caucasia). LOMPE, 1979: 232 [#3628] (characters in key; Central Europe). ČAPEK, HLADIL & ŠEDIVÝ, 1982: 350 [#6200] (hymenopterous parasitoids). COBOS, 1986: 228, 315 [#609] (subgenus Anambus). NIEHUIS, 1988: 65 66 [#1912] (faunal records; biology; Germany). NIEHUIS & TEZCAN, 1993: 11, 48 [#1910] (subgenus Anambus; faunal records from Turkey; distribution and host plants summary; Turkey). CURLETTI, 1994: 22, 155 156 [#658] (subgenus Agrilus; catalog; faunal records; Italia; Sicilia; Corsica). ALEXEEV, 1998:

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA 374 [#4322] (subgenus Quercuagrilus; subgeneric classification). BÍLÝ, 2002: 20 21 [#5877] (bionomy; host plants; distribution). BRECHTEL & KOSTENBADER, 2002: 273 275 [#5766] (biology; ecology; distribution; Germany). FARRUGIA, 2002: 22, 69 [#6511] (characters in key; France). HASTIR & GASPAR, 2002: 13, 33 [#6211] (characters; synonymy; host plants; Belgium). BERNHARD, FRITZSCH, GLÖCKNER & WURST, 2005: 601, 602 [#7261] (mitochondrial DNA; molecular systematics). JENDEK, 2006: 392 [#7441] (subgenus Quercuagrilus; Palaearctic catalog). BELLAMY, 2008: 2159 [#8576] (subgenus Quercuagrilus; world catalog). NIKITSKY et al., 2008: 148 [#8610] (faunal record; Krasnodar). scaberrimus (Ratzeburg, 1837) (Buprestis) RATZEBURG, 1837: 55 [#2498] (description). GAUBIL, 1849: 104 [#4037] (catalog). KIESENWETTER, 1849: 40 [#8334] (catalog; Europe). SCHAUM, 1852: 45 [#8335] (catalog). DOHRN, 1855: 47 [#8329] (catalog; Europe). GISTEL, 1857: 338 [#8063] (checklist). KIESENWETTER, 1857: 125, 143 144 [#3015] (characters in key, redescription; notes). MARSEUL, 1857: 92 [#4371] (checklist; Europe). REDTENBACHER, 1857: 479 [#4134] (characters; Austria). CALWER, 1858: 379 [#8204] (subgenus Agrilus; checklist). DOHRN, 1858: 51 [#8331] (catalog; Europe). JACQUELIN DU VAL, 1860: 143 [#8086] (catalog; Europe). MARSEUL, 1866: 437, 490 [#676] (cited as scaberrimus on page 437 and scaberimus on page 490; characters in key; redescription; distribution overview). BACH, 1867: 398, 401 [#8077] (characters in key; Germany). GEMMINGER & HAROLD, 1869: 1445 [#1057] (catalog). SAUNDERS, 1871: 121 [#2552] (catalog). REDTENBACHER, 1874: 517 [#3784] (characters; Austria). BAUDUER, 1876: xxi [#5293] (Agrilus). BRISOUT, 1876: xxi [#8420] (form of angustulus). STEIN & WEISE, 1877: 91 [#2660] (catalog; Europe). KRAATZ, 1882: 255 [#8477] (notes). BAUDUER, 1883: 21, 22 [#5143] (characters in key; notes; France). MARSEUL, 1887: 252 [#4707] (catalog). REITTER, 1891: 200 [#5278] (catalog). KERREMANS, 1892: 246, 271 [#1397] (synonym of angustulus; catalog). ABEILLE DE PERRIN, 1897: 11 [#19] (notes). HEYDEN, 1897: 193 [#5286] (form of angustulus; characters). KERREMANS, 1903: 275 [#1441] (synonym of angustulus; catalog). PIC, 1909: 165, 189 [#5295] (catalog; France). REITTER, 1911: 195 [#2518] (synonym of angustulus). JAKOBSON, 1913: 796 [#4386] (variety of angustulus; catalog; Russia and Europe). BEDEL, 1921: 214 [#170] (synonym of angustulus). OBENBERGER, 1926: 654 [#2057] (aberration of angustulus; Palaearctic catalog). FUENTE, 1930: 152 [#1036] (morph of angustulus; catalog; Península Ibérica). THÉRY, 1930: 429 [#2754] (synonym of angustulus). PORTEVIN, 1931: 330 [#3832] (synonym of angustulus). OBENBERGER, 1935: 52 [#2178] (catalog; summary of distribution; Bulgaria). OBENBERGER, 1936: 1039 1040 [#2191] (world catalog). THÉRY, 1942: 160, 169 [#2873]. SCHAEFER, 1945: 147 148 [#3584] (synonym of laticornis). THÉRY, 1945: 66 68 [#2884] (taxonomic remarks). PORTA, 1949: 249 [#2487] (synonym of laticornis). HORION, 1955: 82, 269 [#1285] (distribution overview; faunal records; biology; Deutschland; Österreich). LOMPE, 1979: 232 [#3628] (synonym of laticornis). COBOS, 1986: 315 [#609] (synonym of laticornis). SILFVERBERG, 1992: 44 [#6723] (synonym of laticornis). CURLETTI, 1994: 155 [#658] (synonym of laticornis). BÍLÝ, 2002: 20 [#5877] (synonym of laticornis). BRECHTEL & KOSTENBADER, 2002: 273 [#5766] (synonym of laticornis). JENDEK, 2006: 392 [#7441] (synonym of laticornis; Palaearctic catalog). BELLAMY, 2008: 2160 [#8576] (synonym of laticornis; world catalog). asperrimus Marseul, 1866 (Agrilus) MARSEUL, 1866: 437, 492 493 [#676] (description). GEMMINGER & HAROLD, 1869: 1437 [#1057] (catalog). SAUNDERS, 1871: 121 [#2552] (catalog). STEIN & WEISE, 1877: 91 [#2660] (catalog; Europe). MARSEUL, 1887: 252 [#4707] (catalog). REITTER, 1891: 200 [#5278] (catalog). KERREMANS, 1892: 247 [#1397]

119

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA (catalog). ABEILLE DE PERRIN, 1897: 3, 11 [#19] (characters in key). HEYDEN, 1897: 194 [#5286] (Mulsant is cited as the author; notes; faunal records; Germany). KERREMANS, 1903: 275 [#1441] (catalog). REITTER, 1911: 195 [#2518] (characters in key; Germany). CAILLOL, 1913: 514 [#4053] (biology; faunal records; France). JAKOBSON, 1913: 796 [#4386] (catalog; Russia and Europe). BEDEL, 1921: 203, 215 [#170] (characters in key; references; France). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). PORTA, 1929: 402 [#2485] (characters in key; Italy). PORTEVIN, 1931: 330 [#3832] (characters in key; France). OBENBERGER, 1935: 52 [#2178] (catalog; summary of distribution; Bulgaria). OBENBERGER, 1936: 950 951 [#2191] (world catalog). THÉRY, 1942: 134, 162 163 [#2873] (subgenus Agrilus). SCHAEFER, 1945: 68 [#3583] (synonym of laticornis; lectotype designation). SCHAEFER, 1945: 149 [#3584] (synonym of laticornis). THÉRY, 1945: 68 [#2884] (taxonomic remarks). PORTA, 1949: 249 [#2487] (synonym of laticornis). HORION, 1955: 83 84 [#1285] (distribution overview; faunal records; biology; Deutschland; Österreich). RIKHTER & ALEXEEV, 1965: 296 [#2522] (synonym of laticornis). LOMPE, 1979: 232 [#3628] (synonym of laticornis). COBOS, 1986: 315 [#609] (synonym of laticornis). MERKL, 1991: 406 [#6564] (synonym of laticornis). CURLETTI, 1994: 155 [#658] (synonym of laticornis). JENDEK, 1998: 317 318 [#4481] (invalid lectotype designation). BÍLÝ, 2002: 20 [#5877] (SYNONYM OF LATICORNIS). BRECHTEL & KOSTENBADER, 2002: 273 [#5766] (synonym of laticornis). JENDEK, 2006: 392 [#7441] (synonym of laticornis; Palaearctic catalog). BELLAMY, 2008: 2160 [#8576] (synonym of laticornis; world catalog). sibiricola Obenberger, 1924 (Agrilus) syn. nov. OBENBERGER, 1924: 43 44 [#2040] (description). OBENBERGER, 1926: 656 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1041 [#2191] (world catalog). JENDEK, 1995: 147 [#3768] (lectotype designation; characters). ALEXEEV, 1998: 374 [#4322] (subgenus Quercuagrilus; subgeneric classification). JENDEK, 2006: 392 [#7441] (subgenus Quercuagrilus; Palaearctic catalog). BELLAMY, 2008: 2291 [#8576] (subgenus Quercuagrilus; erroneously cited as sibiricolus; world catalog).

Type material: Buprestis laticornis Illiger, 1803 Type locality: not given [Portugall is cited in the title of the publication]. Type specimens: Not located. Described from 1 specimen. Buprestis scaberrima Ratzeburg, 1837 Type locality: not given [Wäldern Preussens are cited in the title of the publication]. Type specimens: Not located. Described from unknown number of syntypes. Agrilus asperrimus Marseul, 1866 Type locality: Italie. Type specimens: Primary: Lectotype (Fig. 42AA), sex not examined, designated by SCHAEFER (1945: 68) [#3583], (MNHN): “Asperrimus (Laferté) type Marse[ul] Italie [h] [data from label on bottom of box]”. Described from unknown number of syntypes. Agrilus sibiricola Obenberger, 1924 Type locality: Sibiria: Kansk. Type specimens: Primary: Lectotype (Fig. 52V), ♀, designated by JENDEK (1995: 147) [#3768], (NMPC): “Sibiria: Kansk [h] \ Mus. Nat. Pragae Inv. [p] 24257 [h] [orange label] \ Agrilus sibiricola m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes.

Diagnosis: 4.3-5.9 mm; antennomeres in male markedly dilated; anterior pronotal lobe moderate; prosternal process distinctly enlarged. Species group: Sulcicollis. Relevant species: A. obscuricollis Kiesenwetter, 1857. Specimens examined: Unknown from East Asia (see Remarks). Host plants: Castanea: CURLETTI, 1994 [#658]; BÍLÝ, 2002 [#5877]; MUSKOVITS & HEGYESSY, 2002 [#5806]. – C. sativa: BRECHTEL & KOSTENBADER, 2002 [#5766]. – Corylus: CURLETTI, 1994 [#658]; BÍLÝ, 2002 [#5877]. – C. avellana: HASTIR & GASPAR, 2002 [#6211]. – Ostrya: CURLETTI, 1994 [#658]. – Quercus: RATZEBURG, 1844 [#8764]; MARSEUL, 1866 [#676]; LEDER, 1872 [#5596]; HUBENTHAL, 1902 (as coerulescens) [#8530]; BEDEL, 1921 [#170]; SAINTE CLAIRE, 1935 [#8551]; RIKHTER, 1945 [#3507]; TER MINASYAN, 1955 [#7893]; SCHAEFER, 1949 [#2565]; HORION, 1955 [#1285]; YABLOKOV KHNZORYAN, 1957 [#5612]; ZAGAIKEVICH, 1962 [#3249]; RIKHTER & ALEXEEV, 1965 [#2522]; NIEHUIS, 1988 [#1912]; CURLETTI, 1994 [#658]; BÍLÝ, 2002 [#5877]; HASTIR & GASPAR, 2002 [#6211]; MÜHLE, 2007 [#8787]; BUSSLER & SCHMIDL, 2009 [#8740]. – Q. cerris: GOBBI, 1973 [#1093]. – Q. ilex: GOBBI, 1973 [#1093]. – Q. petraea: BRECHTEL & KOSTENBADER, 2002 [#5766]; GUTOWSKI & MILKOWSKI, 2008 [#8244]. – Q. robur: BRECHTEL & KOSTENBADER, 2002 [#5766]; GUTOWSKI & MILKOWSKI, 2008 [#8244]. Dubious: Tilia: RIKHTER, 1945 [#3507]; TER MINASYAN, 1955 [#7893]; ZAGAIKEVICH, 1962 [#3249]. Erroneous: Rosa pimpinellifolia: CAILLOL, 1913 [#4053]. – R. rubiginosa: CAILLOL, 1913 [#4053]. – Salix: RIKHTER, 1945 [#3507]; TER MINASYAN, 1955 [#7893]. Distribution: AFRICA:ALGERIA. MOROCCO. TUNISIA. ASIA: ARMENIA. AZERBAIJAN. GEORGIA. ISRAEL. RUSSIA. TURKEY. EUROPE: ALBANIA. AUSTRIA. BELARUS. BELGIUM. BOSNIA & HERZEGOVINA. BULGARIA. CROATIA. CZECH REPUBLIC. DENMARK. ESTONIA. FINLAND. FRANCE. GERMANY. GREECE. HUNGARY. ITALY. LATVIA. LIECHTENSTEIN. LITHUANIA. LUXEMBOURG. MACEDONIA. MOLDOVA. NETHERLANDS. NORWAY. POLAND. PORTUGAL. ROMANIA. SERBIA & MONTENEGRO. SLOVAKIA. SLOVENIA. SPAIN. SWEDEN. SWITZERLAND. UKRAINE. UNITED KINGDOM. Remarks: The re-examination of the lectotype of A. sibiricola from Kansk confirmed its conspecifity with A. laticornis. This opinion is also supported by the recent record of A. laticornis from this region (NIKITSKY et al. 2008 [#8610]). Agrilus laticornis is not known from East Asia but due to the finding in Krasnodarskiy kray (Russia) its occurence in East Siberia is possible.

Agrilus latipalpis Jendek Fig. 24G (habitus of HT); Fig. 38N (aedeagus) latipalpis Jendek, 2007 (Agrilus) JENDEK, 2007: 39 40 [#8006] (description). [#8576] (world catalog).

BELLAMY, 2008: 2161

120 Type material: Agrilus latipalpis Jendek, 2007 Type locality: China, Yunnan, 100 km W of Kunming, Diaolin Nat. Reservation. Type specimens: Primary: Holotype (Fig. 24G), ♂, (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation”. Secondary: 1 PT (CBCS); 1 ♂ PT (SOCI); 37 PT (EJCB). Described from 40 specimens (HT, PT). Diagnosis: 3.6-4.8 mm; antennomeres in male markedly enlarged; ventral side in male with medial stripe of long pubescence on sterna and base of abdomen. Species group: Sulcicollis. Relevant species: A. kaluganus Obenberger, 1940. Specimens examined: CHINA: Yunnan: 9 PT (EJCB): “Yunnan 2000 m, 25.03N 101.55E, Yipinglang 8 10/6.1993” (JENDEK, 2007 [#8006]); 2 PT (EJCB): “Yunnan 8 10.Jun, Yipinglang, 1800 m, 25.05N 101.53E, 1993” (JENDEK, 2007 [#8006]); 28 PT (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation” (JENDEK, 2007 [#8006]).

Adult occurence: 5 - 6 (n=3). Altitude range: 1800-2000 m (n=2). Host plants: All specimens from Diaolin were found on Quercus foliage.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Truncation: transverse. Elytral pubescence: entire, Color: unicolored, Character: homogenous. Tomentum: absent. Prosternal lobe: moderate, Anterior margin: arcuate, Emargination (deepness): shallow, Emargination (width): narrow. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: rectangular, Angles (tips): sharp, Disk: flat. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Ovipositor: elongate. Diagnosis: body more elongate; head markedly convex; pronotum with conspicuous, arcuate anterior lobe; posterolateral pronotal impression shallow. Species group: Japanocarinatus. Relevant species: A. bocaki sp. nov. Type locality: China, Shaanxi province, Qing Ling Shan mts., road Baoji-Taibai village, pass 40 km south of Baoji. Type specimens: Holotype (Fig. 29D), ♀, (EJCB): “China, Shaanxi prov., 21-23.June 1998, Qing Ling Shan mts \ road Baoji-Taibai vill., pass 40 km S Baoji, Zd. Jindra lgt.”. Adult occurence: 6 (n=1). Host plants: Unknown.

Distribution: CHINA: Yunnan.

Distribution: CHINA: Shaanxi.

Agrilus lei Jendek, sp. nov.

Etymology: The specific name is the Chinese noun lei (rib) in apposition. It refers to the long humeral carina of this species.

Fig. 29D (habitus of HT)

Agrilus liscapia Jendek Description: Holotype 5.6 mm. BODY: Shape: subparallel, Build: slender, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Shape: markedly convex, Medial impression: present, Impression (extent): vertex and frons, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: punctures, Sculpture (aspect): irregular, Sculpture (density): sparse, Sculpture (intensity): superficial. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: visually elongate, Sides: arcuate, Maximal width: at middle, Anterior margin: narrower than posterior. Anterior lobe: obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: posterolateral. Prehumerus: carinal, Shape: bisinuate, Extent: beyond half of pronotal length, Posterior end: distant from pronotal angle or margin, Anterior end: close to pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate, Convergence: strongly convergent, Junction: present, Narrowest point: at posterior third of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: present. Apices: Arrangement: conjoint, Shape: arcuate or subtruncate,

Fig. 20K (habitus); Fig. 42T (type) liscapia Jendek, 2003 (Agrilus; replacement name for apicalis Bourgoin not Waterhouse) JENDEK, 2003: 182 [#5823] (replacement name proposal). BELLAMY, 2008: 2167 [#8576] (world catalog). JENDEK & GREBENNIKOV, 2009: 5, 13 [#8563] (characters in key; faunal record; distributional summary, remark on lectotype designation; note; North Vietnam; China (Fujian)). apicalis (Bourgoin, 1923) (Sambus; [Note: Preoccupied as secondary homonym]) DESCARPENTRIES BOURGOIN, 1923: 261 [#367] (description). & VILLIERS, 1967: 1008 [#712] (Therysambus; lectotype designation). JENDEK, 2000: 502 503 [#4851] (notes). JENDEK, 2003: 182 [#5823] (synonym of liscapia; remark on lectotype designation). BELLAMY, 2008: 2167 [#8576] (synonym of liscapia; world catalog). JENDEK & GREBENNIKOV, 2009: 13, 31 [#8563] (synonym of liscapia; taxonomic notes).

Type material: Agrilus liscapia Jendek, 2003 Type locality and specimens: See: Sambus apicalis Bourgoin, 1923. Sambus apicalis Bourgoin, 1923 Type locality: Tonkin: Pho Vi. Type specimens: Primary: Lectotype (Fig. 42T), ♀, designated by DESCARPENTRIES & VILLIERS (1967: 1008) [#712], (MNHN): “Tonkin Pho-Vi 6-07 Cap, Fouquet [h] Type [p] [red label] \ Sambus apicalis Bourgoin Type

121

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA [Bourgoin’s MS] \ Thérysambus apicalis n. gen. Bourgoin A. Descarpentries et [h] A. Villiers det. 19 [p] 66 [h]”. Described from unknown number of syntypes. Diagnosis: 5.0-5.2 mm; humeral carina absent; apical elytral pubescence more contrasting and more densely pubescent than on remaining parts. See also JENDEK & GREBENNIKOV, 2009 [#8563]. Species group: Muscarius. Relevant species: A. apicaureus Jendek, 2009; A. hunanus Jendek, 2009. Specimens examined: CHINA: Fujian: 1 ♀ (EJCB): “China prov. Fujian centr., Yongan env. 21.6., Fujian Shilin, R. Červenka lgt. 1991” (JENDEK & GREBENNIKOV, 2009 [#8563]).

Adult occurence: 6 (n=1). Host plants: Unknown. Distribution: CHINA: Fujian. VIETNAM.

Agrilus lisu Jendek, sp. nov. Fig. 30L (habitus of HT); Fig. 41N (aedeagus)

Description: 4.6-5.5 mm (Holotype 4.6 mm). BODY: Shape: subparallel, Build: slender, Color (dorsally): unicolored, Color (dominant): blue or green. HEAD: Shape: markedly convex, Medial impression: present, Impression (extent): vertex, Impression (depth): deep, Epistoma: in plane with frons. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): irregular, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: large, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: long, Sexual modification in male: not apparent. PRONOTUM: Shape: visually elongate, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles or projecting beyond anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: strongly convex, Disk impressions: posterolateral. Prehumerus: carinal, Shape: arcuate, Extent: to half of pronotal length, Posterior end: distant from pronotal angle or margin, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent or subparallel, Junction: absent or present, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate or conjoint, Shape: arcuate or subtruncate, Truncation: transverse. Elytral pubescence: absent. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): deep. Prosternal process: moderate, Shape: subparallel or dilated, Sides: arcuate, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual

modifications in male: medial sulcus on ventrite 1, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): medial lobe distinctly wider than parameres. Diagnosis: head markedly convex; eyes protruding; pronotum strongly convex, widest in middle; prehumerus present. Species group: Unassigned. Relevant species: A. hattorii Nakane, 1983. Type locality: China, Sichuan province, Jiulonggou near Dayi, cca 70 km west of Chengdu. Type specimens: Primary: Holotype (Fig. 30L), ♂, (EJCB): “China, Sichuan, 23-27 June, Jiulonggou near Dayi, 1993, cca 70 km W of Chengdu, Zd. Jindra et M. Trýzna lgt.”. Secondary: CHINA: Sichuan: 1 ♀ PT (EJCB): “China (Sichuan) 1995, Qingcheng-shan (Umg. Heavenly Old Village), 1000-1300m, 18/20.vii., Heinz leg.”. Adult occurence: 6-7 (n=2). Altitude range: 1000-1300 m (n=1). Host plants: Unknown. Distribution: CHINA: Sichuan. Etymology: Noun in apposition; the specific name refers to the ethnic group Lisu, inhabiting the mountainous regions of Myanmar, southwest China, Thailand and the Indian state of Arunachal Pradesh.

Agrilus lobatus Jendek, sp. nov. Fig. 28I (habitus of HT); Fig. 40M (aedeagus)

Description: Holotype 4.3 mm. BODY: Shape: subparallel, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Medial impression: absent. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): irregular. Eyes: Size: large, Shape: not protruding from head outline, Lower margin: in line with antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: visually elongate, Sides: arcuate, Maximal width: at middle or at posterior margin, Anterior margin: narrower than posterior. Anterior lobe: moderate, Shape: subangulate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: posterolateral. Prehumerus: carinal, Shape: bisinuate, Extent: to anterior pronotal angle, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina, Arc: obvious. Marginal and submarginal carinae: Interspace: wide, Convergence: strongly convergent, Junction: present, Narrowest point: at posterior third of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical third:

122 feebly attenuate. Apices: Arrangement: conjoint, Width: wide, Shape: arcuate. Elytral pubescence: entire, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: large, Anterior margin: subtruncate. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: acute or rectangular, Angles (tips): sharp, Disk: impressed. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): parameres with obvious membranous lobes. Diagnosis: eyes larger; pronotum more elongate; elytra without humeral carina; sides of aedeagus with conspicuous membraneous lobes. Species group: Viridiobscurus. Relevant species: A. pliculipennis Obenberger, 1940. Type locality: China, north Fujian province, Wuyishan city, southwest env., 27.7N, 117.95E. Type specimens: Holotype (Fig. 28I), ♂, (EJCB): “China, N Fujian, 6.V., Wuyishan city, SW env., 27.7N, 117.95E, Jaroslav Turna leg., 2005”. Adult occurence: 5 (n=1). Host plants: Unknown. Distribution: CHINA: Fujian. Etymology: The specific name is derived from the Latin noun lobus (lobe). It refers to the conspicuous lateral membranous lobes on the aedeagus of this species.

Agrilus loosdregti Baudon Fig. 15I (habitus); Fig. 34W (aedeagus); Fig. 48N (type) loosdregti Baudon, 1965 (Agrilus) BAUDON, 1965: 224 225 [#158] (description). BAUDON, 1963: 55 [#148] ([Note: Unavailable name, cited without characters]). DESCARPENTRIES & Villiers, 1967: 149 [#710] (sinensis species group). BAUDON, 1968: 106, 109 [#162]. BELLAMY, 2008: 2169 [#8576] (subgenus Sinagrilus; world catalog).

Type material: Agrilus loosdregti Baudon, 1965 Type locality: Laos: Paklay. Type specimens: Primary: Holotype (Fig. 48N), ♂, (MHNB): “Paklay 10.vi.[19]63 [h] Laos (Baudon) [p] Type [p][red label] \ Agrilus loosdregti mihi [h] A. Baudon det [p] [blue label]”. Described from 1 specimen (HT). Diagnosis: 9.1-10.6 mm; prehumerus obsoletely filamentary; paramerae of aedeagus with obvious sensoric zone. Species group: Smaragdinus. Relevant species: A. priamus Kerremans, 1912. Specimens examined: CHINA: Yunnan: 1 ♂ (NMPC): “Yun’nan’.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Ganlanba, 570 m, 16.iv.1957, Van Shu yun [in Russian]”. MYANMAR: Shan State: 1 ♂ (EJCB): “Burma, SW Shan State, Kalaw, 10 11.vi.1997 (70 km WSW Taunggyi), J. Kaláb”.

Adult occurence: 4-6 (n=2). Altitude range: ~570 m (n=1). Host plants: Unknown Distribution: CHINA: Yunnan. LAOS. MYANMAR.

Agrilus lubopetri Jendek Figs. 8E, 8F (habitus); Fig. 32C (aedeagus); Fig. 48P (type) lubopetri Jendek, 2000 (Agrilus) JENDEK, 2000: 188, 190 [#4852] (description). 2170 [#8576] (world catalog).

BELLAMY, 2008:

Type material: Agrilus lubopetri Jendek, 2000 Type locality: N Vietnam, 21°27N, 105°39E, 70 km NW of Hanoi, Tam Dao, 900-1200 m. Type specimens: Primary: Holotype (Fig. 48P), ♂, (EJCB): “N Vietnam, 21°27N, 105°39E, 70 km NW of Hanoi, Tam Dao, 9-19.v.1996, 900-1200 m, Dembický & Pacholátko leg.”. Secondary: 2 PT (EJCB). Described from 3 specimens (HT, PT). Diagnosis: 11-17 mm; body strikingly elongate; dorsally silky dark-green; elytra without pair of adsutural tomentose spots in apical half; female with obvious orange tomentum on pronotum. Species group: Cyaneoniger. Relevant species: A. auristernum Obenberger, 1924. Specimens examined: CHINA: Yunnan: 1 ♂ (EJCB): “Yunnan, Jing Gu County, Mt. Yingpantou, 7 14.vi.2009 local collector”. LAOS: Houaphan: 1 ♀ (EJCB): “NE Laos, Hua Phan prov., Ban Saluei, 1300 2000m, 20°13’N, 103°59’E, 6 18.iv., Phu Phan Mt. env. 2004, F. & L. Kantner leg.”; 8 (EJCB): “Lao NE, Hua Phan pr., ~20°12’N,104°01’E, Phu Phan Mt., 1500 1900m, 17.v. 3. vi.2007, Vít Kubáň leg.”; 27 (EJCB): “Laos NE, Houa Phan prov., 20°12 13.5’N, 103°59’.5 104°01’E, Ban Saluei → Phou Pane Mt., 1340 1870m, 15.iv. 15.v.2008, Lao collectors leg.”; 50 (NMPC): “Laos NE, Houa Phan prov., 20°12 13.5’N, 103°59’.5 104°01’E, Ban Saluei → Phou Pane Mt., 1340 1870m, 15.iv. 15.v.2008, Lao collectors leg.”. VIETNAM: 2 ♀ PT (EJCB): “N Vietnam, 21°27N, 105°39E, 70 km NW of Hanoi, Tam Dao, 1 8.vi.1996, 900 1200 m, Dembický & Pacholátko leg.” (JENDEK, 2000 [#4852]); 1 ♀ (EJCB): “N Vietnam, 55 km NNW Hanoi, Tam Dao env., 4 8.1998, local collector”.

Adult occurence: 4-5-6 (n=6). Altitude range: 900-2000 m (n=5). Host plants: Unknown. Distribution: CHINA: Yunnan. LAOS. VIETNAM.

Agrilus lugu Jendek, sp. nov. Fig. 22B (habitus of HT); Fig. 37K (aedeagus)

Description: 4.5-6.6 mm (Holotype 6.2 mm). BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Medial impression: present, Impression

123

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA (extent): vertex or vertex and frons, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex or markedly convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, rarely punctures, Sculpture (aspect): irregular, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: small, Shape: not protruding from head outline or protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short, rarely moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: vague or moderate, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: flat, Disk impressions: medial and lateral or lateral only, Lateral impressions (deepness): shallow, Lateral impressions (width): wide. Prehumerus: obsolete, filamentary or carinal, Shape: arcuate, Extent: to third of pronotal length, Posterior end: joining posterior pronotal angle or joining posterior pronotal margin, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior third of pronotal length, Modifications: rarely submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Shape: arcuate. Elytral pubescence: perisutural stripes, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate or large, Anterior margin: arcuate. Prosternal process: narrow or moderate, Shape: narrowed or subparallel, Sides: straight or arcuate, Angles: absent or obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): weakly sclerotised, parameres with obvious membranous lobes and apex of medial lobe markedly obtuse, Ovipositor: uritiform. Diagnosis: body dorsally carmineous-brown; pronotum widest in middle; medial lobe of aedeagus subtruncate on apex. Species group: Hyperici. Relevant species: A. klapperichianus Cobos, 1966. Type locality: China, Yunnan province, Lugu Lake-Luo Shui, 27.45N 100.45E. Type specimens: Primary: Holotype (Fig. 22B), ♂, (EJCB): “China, Yunnan, 8-9.VII., Lugu Lake-Luo Shui, 27.45N 100.45E, E. Jendek leg. 1992”. Secondary: CHINA: Sichuan: 1 ♂ PT (MNHN): “Chasseurs Indigènes des Missionnaire de Ta-tsien-Lou, 1906”; 1 ♂ PT (EJCB): “China, W Sichuan, Sabde env., J. Kaláb leg., 20.vi.1994, 2000 m alt.”; 1 ♂ PT

(EJCB): “China, Sichuan, Moxi, 29°13N, 102°10E, 1600 m, 2.VII.1998, D. Král \ 1998 China Expedition J. Farkač, D. Král, J. Schneider & A. Smetana”; Yunnan: 5 PT (EJCB): “China, Yunnan prov., 27°08’N, 100°14’E, 2900-3500 m, Yulongshan mts., Baishui, 7-12.vii.1990, Vít Kubáň leg.”; 1 ♂ PT (EJCB): “China, N-Yunnan, 27°08’N, 100°14’E, 29003500 m, Baishui vill., lgt. D. Král, 7-12/7’90”; 2 ♂ PT (EJCB): “China, Yunnan, 3500 m, 30 km N of Lijiang, Ganhaizi pass, 5.VII., M. Bocák leg. 1990”; 7 PT (EJCB): “China Yunnan, 8-9.VII., Lugu Lake-Luo Shui, 27.45N 100.45E, E. Jendek leg. 1992”; 1 ♂ PT (EJCB): “Yunnan, 2600-3100 m, 25.38N 100.09E, Cangshan Mts., 5-6/6, Vít Kubáň leg. 1993”; 1 PT (ZIN): “China, Yunnan, Loc. 26, Road #214, Dali-Lijiang, 72nd km NNW of Xiaguan, 26°09’41.0’’N, 100°10’14.1’’E, 2500m, 5.VI.2002, Konstantinov & Volkovitsh leg.”. Adult occurence: 6-7 (n=8). Altitude range: 1600-3500 m (n=7). Host plants: Unknown. Distribution: CHINA: Sichuan, Yunnan. Etymology: The specific name is the noun in apposition. It refers to the Lugu lake, the type locality of the species.

Agrilus macrotatopus Obenberger Fig. 21B (habitus of LT); Fig. 36X (aedeagus); Figs. 21B, 42V (types) macrotatopus Obenberger, 1930 (Agrilus) OBENBERGER, 1930: 110 [#2088] (description). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2174 [#8576] (world catalog). aridosus Obenberger, 1936 (Agrilus) syn. nov. OBENBERGER, 1936: 111 112 [#2205] (description). OBENBERGER, 1936: 949 [#2191] (Agrilus; world catalog). ALEXEEV, 1998: 375 [#4322] (Agrilus; subgenus Xeragrilus; subgeneric classification). JENDEK, 2006: 396 [#7441] (Agrilus; subgenus Xeragrilus; Palaearctic catalog). BELLAMY, 2008: 1975 1976 [#8576] (Agrilus; world catalog).

Type material: Agrilus macrotatopus Obenberger, 1930 Type locality: Asia centralis: Ur Maral. Type specimens: Primary: Lectotype (Fig. 21B) by present designation, ♂, (NMPC): “Asia centr. Ur Maral [h] \ Typus [p] [red label] \ Agrilus macrotatopus m. Type [h] Det. Dr Obenberger [p]”. Secondary: 2 PLT (AAOR). Described from unknown number of syntypes. Agrilus aridosus Obenberger, 1936 Type locality: Asia centralis: Deserta Ak-Kum. Type specimens: Primary: Lectotype (Fig. 42V) by present designation, ♂, (NMPC): “Ur Maral [h] \ Typus [p] [red label] \ Agrilus aridosus m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24322 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 4.8-5.9 mm; very similar to A. ecarinatus but aedeagus distinctly more robust. Species group: Albogularis. Relevant species: A. ecarinatus Marseul, 1866, A. zigzag Marseul, 1866.

124 Specimens examined: Known only from the type specimens.

Host plants: Unknown. Distribution: Ur Maral. See Remarks. Remarks: Lectotypes of both species, A. macrotatopus and A. aridosus, come from the same locality “Ur Maral”, though published locality of A. aridosus is “Asia centralis: Deserta Ak-Kum”. According to Volkovitsh (pers. com.), there are two sites with the similar name: 1) Ur Maral in Talasskii Alatau, Kyrgyzstan; 2) Urochishche Maral-Sai (tourist place) in Zailiiskii Alatau, near Talgar, Almaty, Kazakhstan. Deserta Ak-Kum [= Akkum sands] are in Kazakhstan and Turkmenistan, but it may be a local name for many places in Central Asia. This species has not yet been recorded from East Asia but it is possible in Western China or Mongolia.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA [h] det. T. Nakane [p] [pink label] \ Nakane Coll. Sehu Japan 1999 [p] [green label]”. Described from 1 specimen (HT). Diagnosis: 5.1-5.7 mm; body with distinct carmineous tinge; pronotum without anteromedial impression; elytra with ornamental pubescence. Species group: Maculifer. Relevant species: A. daimio Obenberger, 1936. Specimens examined: CHINA: Guangxi: 1 ♀ (EJCB): “Guangxi, Linchuan, 1984.vi.6”. LAOS: Houaphan: 1 ♀ (EJCB): “Laos, Hua Phan prov. 2004, Ban Saluei env., alt. 1500m, 30 km S of Xam Neua, Phu Phan Mt., 6 17.5.2004, 103°59’33’’E, 20°13’39’’N, P. Kresl leg.”.

Adult occurence: 5-6 (n=2). Altitude range: 1500 m (n=1). Host plants: Unknown.

Agrilus maculifer Saunders

Distribution: CHINA: Guangxi. JAPAN: Kyushu, Ryukyu. LAOS.

Fig. 11H (habitus of LT); Fig. 33H (aedeagus); Figs. 11H, 49S (types)

Agrilus madeci Baudon

maculifer Saunders, 1873 (Agrilus) SAUNDERS, 1873: 514 [#2554] (description). LEWIS, 1879: 15 [#3345] (catalog; Japan). KERREMANS, 1885: 153 [#1381] (catalog). SCHÖNFELDT, 1887: 113 [#8901] (Anambus; catalog; Japan). KERREMANS, 1892: 263 [#1397] (catalog). KERREMANS, 1903: 287 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 18 [#1809] (catalog; Japan). OBENBERGER, 1936: 1012 [#2191] (world catalog). KUROSAWA, 1963: 153 [#1618] (characters; Japan). KUROSAWA, 1974: 2 [#4577] (note). TÔYAMA, 1981: 32 [#4590] ([Note: Misidentification of new species described later as naomii by Tôyama]). TÔYAMA, 1985: 21 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 35 [#4091] (checklist; Japan: Kyushu). JENDEK, 2005: 13 [#7217] (lectotype designation). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2174 [#8576] (world catalog). nagaoi Nakane, 1983 (Agrilus) syn. nov. HIRASHIMA, NAKANE, 1983: 110 111 [#4589] (description). 1989: 323 [#8078] (subspecies of maculifer; checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (subspecies of maculifer; checklist; Japan). AKIYAMA & OHMOMO, 1997: 36 [#4091] (checklist; Japan: Ryukyus (Amami oshima I.)). JENDEK, 2006: 400 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2197 [#8576] (world catalog).

Type material: Agrilus maculifer Saunders, 1873 Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 11H), ♂, designated by JENDEK (2005: 13) [#7217], (BMNH): “Type H.T. [p] [round label with red border] \ Japan. G. Lewis. 1910 - 320. [p] \ Agr. maculifera ES. Type [h] [by Saunders hand] \ Agrilus maculifera E. S. Type [h]”. Described from unknown number of syntypes. Agrilus nagaoi Nakane, 1983 Type locality: Hatsuno, Amami-Oshima Is., Japan. Type specimens: Primary: Holotype (Fig. 49S), ♀, (EIHU): “Hatsuno, Amami-Oshima [p] 16.VII. [h] 19 [h] 62 [h] Coll. J. Nagao [p] \ Holotype [p] [red label] \ Agrilus nagaoi Nakane

Fig. 10J (habitus); Fig. 32Y (aedeagus); Figs. 48T, 49R, 52B (types) madeci Baudon, 1968 (Agrilus) BELLAMY, BAUDON, 1968: 131, 145 147 [#162] (description). 2008: 2175 [#8576] (world catalog). rokuyai Kurosawa, 1976 (Agrilus) syn. nov. TÔYAMA, KUROSAWA, 1976: 134 136 [#1628] (description). 1985: 24 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 185 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 37 [#4091] (checklist; Japan: Honshu). ALEXEEV, 1998: 372 [#4322] (subgenus Uragrilus; subgeneric classification). AKIYAMA & OHMOMO, 2000: 88 [#6240] (faunal records; Japan (Hiroshima)). HOU TAOQIAN & ZENG RUI, 2002: 357 [#6626] (cited as rokyai; checklist; China: Hainan). MÜHLE, 2003: 47 [#6155] (checklist). JENDEK, 2006: 395 [#7441] (subgenus Uragrilus; Palaearctic catalog). BELLAMY, 2008: 2267 [#8576] (subgenus Uragrilus; world catalog). LEE, KIM & AHN, 2010: 264 266 [#8939] (characters; faunal records; South Korea). mushanus Kurosawa, 1976 (subspecies of rokuyai) syn. nov. KUROSAWA, 1976: 136 [#1628] (description). HUA LI ZHONG, 2002: 90 [#6696] (subspecies of rokuyai; checklist; China: Taiwan). MÜHLE, 2003: 47 [#6155] (subspecies of rokuyai; checklist; Taiwan). JENDEK, 2006: 395 [#7441] (subgenus Uragrilus; subspecies of rokuyai; Palaearctic catalog). BELLAMY, 2008: 2267 [#8576] (subspecies of rokuyai; world catalog).

Type material: Agrilus madeci Baudon, 1968 Type locality: Laos: Muong Panh, alt. circa 1100 m. Type specimens: Primary: Holotype (Fig. 48T), ♂, (MHNB): “Muong Panh 1.iv.64 [h] Laos (Baudon) [p] \ Type [p] [red label] \ Agrilus madeci Type mihi [h] A, Baudon det. [p]”. Secondary: 1 PT (MHNB). Described from unknown number of specimens (HT, AT, PT). Agrilus rokuyai Kurosawa, 1976 Type locality: Ikada, Hyôgo Pref. Type specimens: Primary: Holotype (Fig. 52B), ♂, (NSMT): “IKADA HYOGO PREF. 6. VII. 1951 R. MORIMOTO [leg.]

125

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA [h] \ Holotype [p] Agrilus rokuyai Y. Kurosawa, 1976 [h] [red label with black border]”. Secondary: 3 PT (NSMT). Described from 10 specimens (HT, AT, PT). Agrilus rokuyai mushanus Kurosawa, 1976 Type locality: Musha, C. Formosa. Type specimens: Primary: Holotype (Fig. 49R), ♂, (NSMT): “Formosa [p] Musha [h] 19 [p] 62 [h] \ Holotype [p] Agrilus rokuyai mushanus Y. Kurosawa, 1976 [h] [red label]”. Secondary: 2 PT (NSMT); 1 PT (MNHN). Described from 11 specimens (HT, AT, PT). Diagnosis: 6.6-8.1 mm; antennae slender, elytral tomentose spots larger and elongate, elytral apices less sharply acuminate. Species group: Niveoguttatus. Relevant species: A. jaminae Baudon, 1968. Specimens examined: CHINA: Shaanxi: 1 ♂, 1 ♀ (EJCB): “China, S Shaanxi, Qinling Mts. S slope, Xunyangba S+W env., 33º28 37’ / 108º23 33’, 1400 2100m, 5 9.6.[19]95, L.+R. Businský lgt.”; 2 ♂, 2 ♀ (EJCB): “China, Shaanxi Henan border, 900 1500m, 29 31.5.[19]95, 33º48’ 53’ / 110º40’ 46’, L.+R. Businský lgt.”; 3 (EJCB): “China Shaanxi, Lüeang 33°07’N 106°05’E, 8.6. 14.6.1996, lgt. E. Kučera”; 2 ♀ (EJCB): “China, 1000 1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v. 13.vi.1998, I. H. Marshal leg.”; 8 (EJCB): “China, 1150 1300 m, Shaanxi, Qinling mts., Foping (6 km N), 20 21.VI.1998, I. H. Marshal leg.”; 1 ♂, 3 ♀ (EJCB): “C China, Shanxi [sic! should be Shaanxi], Qinling Shan, 6km E of Xunyangba, 1000 1300 m, 23.v. 13.vi., leg. C. Holzschuh, 2000”; 5 (EJCB): “China Shaanxi, Lüeang, 22.5. 25.5.2000, leg. E. Kučera”; 1 ♂ (EJCB): “C China, Shanxi [sic! should be Shaanxi], Qinling, Shan, 6 km N of Foping, 1150 1300m, 20 21.VI.2000, leg. C. Holzschuh”; 6 (EKCS): “China Shaanxi, Lueyang, 29.5. 2.6.2005, lgt. E. Kučera”. JAPAN: Honshu: 1 ♂ (EJCB): “Kameoka City, Kyoto pref., 26.vi.1982, J. Nakata”; 23 (EJCB): “Japan, Hyogo Pref., Komo, 20 km NW, Takarazuka, 400 m, 29.Jun 2003, Bolm lgt.”. LAOS: Houaphan: 12 (EJCB): “Laos NE, Houa Phan prov., 20°12 13.5’N, 103°59’.5 104°01’E, Ban Saluei → Phou Pane Mt., 1340 1870m, 15.iv. 15.v.2008, Lao collectors leg.”.

Adult occurence: 4-5-6-7 (n=16). Faunal records published by KUROSAWA, 1976 [#1627] are also included. Altitude range: 400-2100 m (n=9). Host plants: Quercus: AKIYAMA & OHMOMO, 1997 (as rokuyai) [#4091]. Distribution: CHINA: Henan, Shaanxi, Taiwan. JAPAN: Honshu. KOREA SOUTH. LAOS. Remarks: Distribution in South Korea was complemented from the recent publication of LEE, KIM & AHN, 2010 [#8939] and is not included in the Analysis.

Eyes: Size: small, Shape: not protruding from head outline or protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short. PRONOTUM: Shape: transverse, Sides: arcuate or subangulate, Maximal width: at middle, Anterior margin: subequal to posterior or narrower than posterior. Anterior lobe: absent or vague, Shape: subangulate, Position to anterior angles: not reaching level of anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: flat, Disk impressions: medial and lateral or lateral only, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow. Prehumerus: obsolete, tubercular, rarely carinal, Shape: arcuate, Extent: to half of pronotal length, Posterior end: distant from pronotal angle or margin, Anterior end: distant from pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate or wide, Convergence: strongly convergent, Junction: absent or present, Narrowest point: at posterior third of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate, Width: wide, Shape: arcuate. Elytral pubescence: absent. Tomentum: spots, Spots (locations): postmedial. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: large, Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): wide. Prosternal process: moderate, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: subtruncate. Sternal groove: Shape on apex of last ventrite: subtruncate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): shorter or subequal to 2-3 combined. GENITALIA: Aedeagus (symmetry): symmetrical. Diagnosis: body subparallel; elytra vaguelly attenuate in apical third; elytral apices wide; prehumerus obsolete; elytral disk with smooth sculpure; pubescent spots on apical third of elytra wide and distant from sutural margin. Species group: Biguttatus. Relevant species: A. biguttatus (Fabricius, 1777); A. decoloratus Kerremans, 1892; A. ventricosus Fairmaire, 1888. Type locality: North Vietnam, Tonkin, Vinphu province, Tamdao.

Agrilus magnus Jendek, sp. nov.

Type specimens: Primary: Holotype (Fig. 8L), ♂, (EJCB): “N Vietnam (Tonkin), Tamdao, (pr. Vinphu), 2.-11.6.1985, Vít Kubáň leg.”. Secondary: CHINA: Guangxi: 1 ♂ PT (EJCB): “Guang Xi, [19]80.5.31”.

Fig. 8L (habitus of HT); Fig. 32H (aedeagus)

Adult occurence: 5-6 (n=2).

Description: 12-14.8 mm (Holotype 12 mm). BODY: Shape: subparallel, Build: robust, Color (dorsally): unicolored, Color (dominant): blue or brown. HEAD: Medial impression: present, Impression (extent): vertex and frons, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: protruding from head outline. Vertex: Sculpture on vertex: punctures, Sculpture (density): sparse.

Host plants: Unknown. Distribution: CHINA: Guangxi. VIETNAM: Vinh Phu. Etymology: The specific name is the Latin adjective magnus (large). It refers to the large size of the species.

126

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Agrilus majzlani Jendek Fig. 13I (habitus of HT) majzlani Jendek, 2007 (Agrilus) JENDEK, 2007: 42 43 [#8006] (description). [#8576] (world catalog).

BELLAMY, 2008: 2176

Type material: Agrilus majzlani Jendek, 2007 Type locality: China, Yunnan, Ninglang env., 27º19’N, 100º55’E. Type specimens: Primary: Holotype (Fig. 13I), ♀, (EJCB): “China, Yunnan, 6-10.VII., Ninglang env., 27º19’N, 100º55’E, 1992”. Described from 1 specimen (HT). Diagnosis: 7.9 mm; dorsally dark violet; elytral apices longer and subangulate. Species group: Plagiatus. Relevant species: A. kostali Jendek, 2007. Specimens examined: Known only from the holotype.

Host plants: Unknown. Distribution: CHINA: Yunnan.

Agrilus mali Matsumura Fig. 14D (habitus); Fig. 34I (aedeagus), Fig. 47Q (type) mali Matsumura, 1924 (Agrilus) Matsumura in MURAMATSU, 1924: 4 6 [#3482] (description). OGLOBLIN & REICHARDT, 1932: 277 [#8654] (pest). YUASA, 1933: 281 [#3132] (pest of apple tree in Korea). MIWA & CHÛJÔ, 1936: 18 [#1809] (catalog; Japan). OBENBERGER, 1936: 1012 [#2191] (world catalog). MÜHLMANN, 1954: 83 [#8152] (notes on biology). TER MINASYAN, 1955: 452 [#7893] (host plants; distribution; references). BALACHOWSKI et al., 1962: 286 [#96] (economic importance). GURYEVA, 1974: 99 [#5428] (pest of agriculture cultures; references; no data from USSR). KUROSAWA, 1974: 2 [#4575] (notes). KUROSAWA, 1974: 2 [#4577] (note). ALEXEEV, 1979: 128 [#57] (subgenus Sinuatiagrilus; characters; from Transbaicalia to Far East; North China; Korea). NIKRITIN & SHUTOVA, 1985: 46 [#4249] (biology; Russia: Amurskaya oblast’, Primorskii i Khabarovskii krai). ANONYMOUS, 1986: 177 [#8848] (insect pest; Korea). PENG ZHONGLIANG, 1987: 356 [#2429] (cited as malyi; checklist; China). ALEXEEV, 1989: 486 [#64] (key to species; Khabarovsk; Amursk; Primorsk; Chita; Buryatia; Japan; Korean peninsula; northeastern China; north Mongolia). ALEXEEV & VOLKOVITSH, 1989: 320, 349, 357 [#3241] (characters in key; faunal records; Mongolia). LI JINGKE, 1992: 92 [#6755] (checklist; northeastern China). NIKRITIN, 1994: 46 [#8675] (characters; biology; harms; eradication). SOKOLOV et al., 1995: 40 [#6372] (biology; distribution; quarantine measures). ALEXEEV, 1998: 372 [#4322] (subgenus Sinuatiagrilus; subgeneric classification). NONNAIZAB et al., 1999: 113 [#8247] (checklist; China: Inner Mongolia). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Heilongjiang, Hebei, Gansu, Qinghai, Henan, Shandong, Hubei, Guangxi). LIN GUI RUI, 2002: 234 [#7926] (checklist; pest). JI YING, JI RONG & HUANG REN XIN, 2004: 31 33 [#7326] (invasive species; China: Xinjiang). JENDEK, 2006: 394 [#7441] (subgenus Sinuatiagrilus; Palaearctic catalog). BELLAMY, 2008: 2176 2177 [#8576] (subgenus Sinuatiagrilus; world catalog). VOLKOVITSH, 2009: 136 [#8850] (subgenus Sinuatiagrilus; faunal records; Russia: Primorskiy kray). jenissejensis Obenberger, 1924 (Agrilus) OBENBERGER, 1924: 56 57 [#2040] (description). OBENBERGER,

1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1003 [#2191] (cited as jenisejensis; world catalog). KUROSAWA, 1975: 1 [#4578] (cited as jenisseiensis). ALEXEEV, 1979: 128 [#57] (subgenus Sinuatiagrilus; variety of mali; characters; notes). ALEXEEV, 1989: 486 [#64] (cited as enisejensis; synonym of mali). ALEXEEV & VOLKOVITSH, 1989: 320 [#3241] (cited as jenisejensis; synonym of mali; type examined). NONNAIZAB et al., 1999: 113 [#8247] (synonym of mali). JENDEK, 2006: 394 [#7441] (synonym of mali; Palaearctic catalog). BELLAMY, 2008: 2176 2177 [#8576] (synonym of mali; world catalog).

Type material: Agrilus mali Matsumura, 1924 Type locality: Pyongyang, Taegu, Inchon. Type specimens: Not located. Described from unknown number of syntypes. Agrilus jenissejensis Obenberger, 1924 Type locality: Regionem fluvii Jenissej (Siberia) superioris. Type specimens: Primary: Lectotype (Fig. 47Q) by present designation, ♂, (NMPC): “Jenissej super [h] \ Typus [p] [red label] \ A. jenisejensis [sic!] m. Type [h] Det. Dr. Obenberger [p]”. Described from unknown number of syntypes. Note: There are two additional specimens in NMPC labeled as types and collected by Lukjanovitsh in 1928 in “Transbaicalia”. Diagnosis: 6.1-8.2 mm; elytra with pubescent spots; aedeagus widely enlarged distally. Species group: Sinuatus. Relevant species: A. sinuatus sachalinensis Obenberger, 1935. Specimens examined: CHINA: Gansu: 1 ♀ (EJCB): “China Kansu Ponggartang 30.Jun 1992 M. Bok lgt.”; 1 ♂ (EJCB): “China Gansu, Dingxi, 23.5. 30.5.1996, E. Kučera leg.”. Hubei: 1 ♂ (EJCB): “China, W Hubei, 8.+15 17.vi., Muyuping S env., 1100m, 31.45 N, 110.4 E, Jaroslav Turna leg. 2002”. Sichuan: 1 ♂ (EJCB): “China NW, Sichuan prov., Songpan, 2000m, 32°30’N, 103°40’E, 13 17.vii.1990, J. Kolibáč”; 1 ♀ (EJCB): “SE Tibet, 3000 3500 m, ca 20 km N of Yanjing, 23.vii. 8.viii.1998, O. Šafránek et M. Tryzna”; 1 ♀ (EJCB): “China Sechuan, Jiuzhaigou [sic!], 11.6. 16.6.2001, leg. E. Kučera”. MONGOLIA: 1 ♂ (ZIN): “Mogoliya, Vostochnyi aimak, r. Numrègin Gol, 32 km YuV g. Salkhit, Gur’eva, 8.viii.[1]976 [in Russian]”; 1 (EJCB): “MNR, Centr. aimak, Unchneul, 16.7.1976 [in Russian]”; 1 ♀ (ZIN): “Mongoliya, Vostochnyi aimak, Bayan Dun, Kerzhner, 10.vii. [1]976 [in Russian]”. RUSSIA: Amurskaya oblast’: 1 ♀ (ZIN): “g. Blagoveshchensk 19.vii.[19]27 [in Russian]”; 1 ♂ (ZIN): “g. Blagoveshchensk 5.ix.[19]27 [in Russian]”; 1 (ZIN): “Amursk. reg., rr. M. Pera B. Ergel’ mezhdur., Amur., Zinovjev 21.VII.[19]58 [in Russian]”. Primorskiy kray: 1 (ZIN): “Shmakovka 25/VI 1926 [in Russian]”; 1 ♂ (ZIN): “Primorskii kr., g. Ussuriisk, 7 viii 1958g, na yablone [in Russian]”; 1 ♀ (ZIN): “Primorskii kr., Slavyanskii r n., Gorno Taezhen, st. Uchkhoz., 21 vii 59g [in Russian]”.

Adult occurence: 5-6-7-8-9 (n=15). Altitude range: 11003500 m (n=3). Host plants: Malus: MATSUMURA in MURAMATSU, 1924 [#3482]; OGLOBLIN & REICHARDT, 1932 [#8654]; MÜHLMANN, 1954 [#8152]; TER MINASYAN, 1955 [#7893]; GURYEVA, 1974 [#5428]; ALEXEEV, 1979 [#57]; ALEXEEV, 1979 (as jenissejensis) [#57]; NIKRITIN & SHUTOVA, 1985 [#4249]; ALEXEEV, 1989 [#64]; ALEXEEV & VOLKOVITSH, 1989 [#3241]; NIKRITIN, 1994 [#8675]; SOKOLOV et al.,1995 [#6372]; JI YING, JI RONG & HUANG REN XIN, 2004 [#7326]. − M. pumila: HUA LI ZHONG, 2002 [#6696]. − M. spectabils: HUA LI ZHONG, 2002 [#6696]. − Prunus persica: HUA LI ZHONG, 2002

127

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA [#6696]. − P. armeniaca: HUA LI ZHONG, 2002 [#6696]. − Pyrus: ALEXEEV, 1979 [#57]; ALEXEEV, 1979 (as jenissejensis) [#57]; ALEXEEV, 1989 [#64]; ALEXEEV & VOLKOVITSH, 1989 [#3241]; HUA LI ZHONG, 2002 [#6696]. Erroneous: Juglans regia: HUA LI ZHONG, 2002 [#6696]. − Salix: HUA LI ZHONG, 2002 [#6696]. Distribution: CHINA: Gansu, Guangxi, Hebei, Heilongjiang, Henan, Hubei, Nei Mongol, Qinghai, Shandong, Sichuan, Xinjiang, Xizang. KOREA NORTH. KOREA SOUTH. MONGOLIA. RUSSIA: Amurskaya oblast’, Buryatiya Respublika, Khabarovskiy kray, Primorskiy kray, Zabaykalskiy kray. Remarks: Type specimens of A. mali Matsumura, 1924 have not been found. The species concept of this taxon used herein follows that of ALEXEEV 1979 [#57] and 1989 [#64]. There is still a possibility that A. mali Matsumura, 1924 might be conspecific with A. sinuatus sachalinensis Obenberger, 1935 partly because Malus, Prunus and Pyrus are known host plant also for A. sinuatus (Olivier, 1790).

Agrilus malloti Théry Fig. 25D (habitus); Fig. 38W (aedeagus); Figs. 45K, 48V (types) malloti Théry, 1930 (Agrilus) THÉRY, 1930: 168 170 [#2767] (description). OBENBERGER, 1936: 1090 [#2191] (world catalog). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2177 [#8576] (world catalog). diolaus Obenberger, 1958 (Agrilus) syn. nov. OBENBERGER, 1958: 238 [#2389] (description). YANG XINGKE et al., 1991: 31 [#6756] (type specimen in Institute of Zoology, Academica Sinica, Beijing). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Yunnan). JENDEK, 2005: 10 [#7217] (lectotype designation; faunal records; India). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2071 [#8576] (world catalog).

Type material: Agrilus malloti Théry, 1930 Type locality: Barkote, Dehra-Dun. Type specimens: Primary: Lectotype (Fig. 48V) by present designation, ♂, (MNHN): “Barkote Dehra Dun N. C. Chatterjee [p] II. V. [h] 192 [p] 6 [h] \ R.R.D. 529 B.C.R. 50 Cage 570 [p] \ Ex Mallotus philippinensis [p] \ malloti Thery ♂ [h] Type [p] [red ink]”. Described from unknown number of syntypes. Agrilus diolaus Obenberger, 1958 Type locality: Chine: Yunnan, Valley of Salveen, W. Paoshan. 100 m [sic!]. Type specimens: Primary: Lectotype (Fig. 45K), ♂, designated by JENDEK (2005: 10) [#7217], (NMPC): “Yünnan [h] \ Typus [p] [red label] \ A. Diolaus m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24904 [h] [orange label]”. Secondary: PLT (IZAS). Described from unknown number of syntypes. Diagnosis: 4.4-6.0 mm; body silky-green; elytral pubescence disrupted in apical third by upturned V-shaped glabrous spot; prosternal lobe and apex of last ventrite slightly emarginated; prosternal process glabrous in both sexes. Species group:

Tempestivus. Relevant species: A. fevreae Descarpentries & Villiers, 1963. Specimens examined: CHINA: Shaanxi: 1 ♂, 1 ♀ (EJCB): “China, 17 22.vi., Shaanxi prov., 1991, Hua Shan peak env., 100 km E of Xi’an, Z. Kejval lgt.”. INDIA: Assam: 50 (EJCB): “NE India, Assam, 1999, 5 km N of Umrongso, 700m, 25°27’N, 92°43’E, 17. 25.v., Dembický & Pacholátko leg.” (JENDEK, 2005 [#7217] as diolaus); 20 (EJCB): “NE India, Assam, 1999, 5 km N of Umrongso 700m, 25°27’N, 92°43’E, 17. 25.v., J. Rolčík leg.” (JENDEK, 2005 [#7217] as diolaus); 2 (EJCB): “NE India, Assam, 1999, 5 km N of Umrongso 700m, 25°27’N, 92°43’E, 17. 25.v., Zd. Košťál lgt.” (JENDEK, 2005 [#7217] as diolaus); 1 ♂, 1 ♀ (EJCB): “NE India, Assam prov. Umrongso env. 700m, 25°27’N, 92°43’E, 3 8.vi., Tryzna & Benda leg. 2002” (JENDEK, 2005 [#7217] as diolaus). Kerala: 1 ♂, 4 ♀ (EJCB): “S India, Kerala, Thekkady Periyar Lake, 9.34N, 77.10E, 900 1000 m, 19 27.IV.1997, Dembický & Pacholátko leg.”. Tamil Nadu: 2 ♂ (EJCB): “S India, Tamil Nadu, 17 22.V.1997, 15 km SE Kotagiri, 11°22’N, 76°56’E, Kunchappanai, Dembický & Pacholátko leg.”; 1 ♂ (EJCB): “India S, Tamil Nadu, Nilgiris, 15 km SE of Kotagiri, Kunchappanai, 900 m, 11.22N 76.56E, 7 22.v.2000, leg. P. Pacholátko”; 1 ♂ (EJCB): “India S, Tamil Nadu, Nilgiri Hills, 11 km SE of Kotagiri, 1100±100 m, 11.24N 76.56E, Kunchappanai, leg. L. Dembický, 3 15.v.2002”. Uttarakhand: 1 ♀ (MNHN): “Phanduwala, Dehra Dun, U.P., F. Ent., 21.v.1929 \ ex Mallotus philippinensis”. LAOS: Borikhamxai: 2 ♂ (EJCB): “Laos centr., Bolikhamsai prov., Ban Nape Kaew Nua Pass, 18.4. 1.5.1998, alt. 600±100 m, N 18º22.3, E 105º09.1 (GPS), E. Jendek & O. Šauša leg.”. Houaphan: 1 ♂ (EJCB): “Laos NE, Hua Phan prov., 20º10’N, 103º12’E, Ban Sakok, 23 26.v. (Phue Loei N.P.), J. Bezděk, 2001”. Louang Namtha: 1 ♂ (EJCB): “Laos north, 5 11.V.1997, 20 km NW Louang Namtha, N 21°09.2, E 101°18.7, alt. 900±100 m, E. Jendek & O. Šauša leg.”. THAILAND: Mae Hong Son: 1 ♂ (EJCB): “NW Thailand, 1 15.5., 1991, Mae Hong Son, Ban Huai Po, 800 1600 m, S. Bílý leg.”; 1 ♂, 1 ♀ (EJCB): “NW Thailand, 8 18.V., Mae Hong Son, 1992, Ban Si Lang, 1200 m, J. Horák leg.”. VIETNAM: 1 ♂ (EJCB): “N Vietnam, 21°35N, 106°30E, 52 km SW of Lang Son, 27.iv. 6.v.1996, 370 m, Dembický & Pacholátko leg.”.

Adult occurence: 4-5-6 (n=16). Altitude range: 370-1600 m (n=12). Host plants: Mallotus philippinensis: THÉRY, 1930 [#2767]. Distribution: CHINA: Shaanxi, Yunnan. INDIA: Assam, Kerala, Tamil Nadu, Uttarakhand. LAOS: Borikhamxai, Houaphan, Louang Namtha. THAILAND: Mae Hong Son. VIETNAM.

Agrilus mallotiellus Kurosawa Fig. 20I (habitus); Fig. 36T (aedeagus); Fig. 48U (type) mallotiellus Kurosawa, 1985 (Agrilus; replacement name for malloti Kurosawa not Théry) KUROSAWA, 1985: 162 [#1640] (replacement name proposal). TÔYAMA, 1985: 20 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 35 [#4091] (checklist; Japan: Honshu, Shikoku, Kyushu, Yakushima I., Ryukyus (Amami oshima I., Tokuno oshima I., Tokaranakanoshima I.)). ALEXEEV, 1998: 374 [#4322] (subgenus Austragrilus; cited as malotiellus; subgeneric classification). JENDEK, 2006: 390 [#7441] (subgenus Austragrilus; Palaearctic catalog). BELLAMY, 2008: 2177 [#8576] (world catalog). JENDEK & GREBENNIKOV, 2009: 4, 11 [#8563] (characters in key; faunal

128 record; distributional summary; Japan). HATTORI, 2010: 4 [#8899] (faunal records; Japan: Ryûkyû). HATTORI, 2010: 21 [#8921] (faunal record; Japan: Ryûkyû) malloti Kurosawa, 1957 (Agrilus; [preoccupied]) KUROSAWA, 1957: 189 190 [#1615] (description). KUROSAWA, 1963: 154 [#1618] (notes; Japan). KUROSAWA, 1974: 2 [#4577] (note). AKIYAMA, 1975: 4 [#3178] (faunal records; Ryûkyû Archipelago). KUROSAWA, 1985: 162 [#1640] (synonym of mallotiellus). AKIYAMA & OHMOMO, 1997: 35 [#4091] (synonym of mallotiellus). JENDEK, 2006: 390 [#7441] (synonym of malotiellus; Palaearctic catalog). BELLAMY, 2008: 2177 [#8576] (synonym of mallotiellus; world catalog). JENDEK & GREBENNIKOV, 2009: 11, 31 [#8563] (synonym of mallotiellus; HT examined).

Type material: Agrilus mallotiellus Kurosawa, 1985 Type locality and specimens: See: Agrilus malloti Kurosawa, 1957. Agrilus malloti Kurosawa, 1957 Type locality: Onoaida, Yakushima, S. Kyûshû, Japan. Type specimens: Primary: Holotype (Fig. 48U), ♂, (NSMT): “(Yakushima) Onoaida [p] 23. [h] VII, 1952 Y. Kurosawa [p] \ Holotype [p] Agrilus malloti Y. Kurosawa, 1957 [h] [red label]”. Described from 15 specimens (HT, AT, PT). Diagnosis: 3.9-4.7 mm; aedeagus as in Fig. 36T. For further characters see JENDEK & GREBENNIKOV, 2009 [#8563]. Species group: Muscarius. Relevant species: A. aurarius (Kerremans, 1892). Specimens examined: JAPAN: Ryukyu isl. (Okinawa incl.): 1 ♂ (EJCB): “Yakushima Is. 6.vii.1989 H. Akiyama // Funayuki” (JENDEK & GREBENNIKOV, 2009 [#8563]).

Adult occurence: 7-8 (n=5). Faunal records published by KUROSAWA, 1957 [#1615] are also included. Host plants: Mallotus japonicus: KUROSAWA, 1957 [#1615] (as malloti); AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: JAPAN: Honshu, Kyushu, Ryukyu Islands, Shikoku.

Agrilus manchu Jendek, sp. nov. Fig. 27M (habitus); Fig. 39S (aedeagus)

Description: 3.2-5.0 mm (Holotype 4.6 mm). BODY: Shape: cuneiform, Build: slender, Color (dorsally): unicolored, Color (dominant): green. HEAD: Medial impression: absent or present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: flat or moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: protruding from head outline, Sculpture on vertex: rugae, Sculpture (intensity): superficial. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square, Sides: arcuate, Maximal width: at anterior margin, rarely at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles:

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA at level with anterior pronotal angles or projecting beyond anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: flat, Disk impressions: medial and lateral, Medial impression (shape): posteromedial only, rarely anteromedial and posteromedial, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial. Prehumerus: carinal, Shape: bisinuate, Extent: beyond half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: narrow or moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior angles or at posterior 1/5-1/4 of pronotal length, Modifications: rarely submarginal carina divergent at posterior angles. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: bicolored, complementary with pubescence, Humeral carina: absent. Apices: Arrangement: separate, rarely conjoint, Shape: arcuate. Elytral pubescence: proximal and distal, Proximal: humeral, Distal: apical, Color: unicolored. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): narrow. Prosternal process: moderate or wide, Shape: dilated, Sides: straight, Angles: acute or rectangular, Angles (tips): sharp, Disk: impressed. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: angulately sinuate, Sinuosity (deepness): markedly deep. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: elongate. Diagnosis: apex of elytra entirely pubescent; apices jointly or separately arcuate. Species group: Cyaneovirens. Relevant species: A. gussakovskiji Alexeev, 1981. Type locality: China, north Henan province, Jiuligou, altitude 550 m, 35°12’N, 112°26’E. Type specimens: Primary: Holotype, ♂, (EJCB): “China, N. Henan, 11-12.VI., Jiuligou, 550 m, 35°12’N, 112°26’E, Jaroslav Turna leg. 2007”. Secondary: CHINA: Henan: 8 PT (EJCB): “China, N. Henan, 11-12.VI., Jiuligou, 550 m, 35°12’N, 112°26’E, Jaroslav Turna leg. 2007”; Shaanxi: 1 ♂ PT, 3 ♀ PT (EJCB): “China, Shaanxi pr., Hua Shan, 17-21. VI.1991, R. Dunda lgt.”; 2 ♀ PT (EJCB): “China, Shaanxi province, Hua Shan Mt., 17-21.VI.1991, leg. R. Sauer”; 2 ♂ PT, 1 ♀ PT (EJCB): “China, Shaanxi, 110 km E of Xi’An, HuaShan, ~500 m, 23-24.vi.1998, I. H. Marshal leg.”; 1 PT (EJCB): “China-Shaanxi, Hua-Shan, 30.6.1.7.2000, leg. E. Kučera”; Yunnan: 1 ♂ PT (EJCB): “China, Yunnan, 22.V.-2. VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”; 1 ♂ PT (EJCB): “China-Yunnan, 24-29.6.[19]94, Daju, lgt. E. Kučera”; 1 ♀ PT (EJCB): “China, Yunnan, Daju, 17.VI.1995, Šiška & Spevár & Pekarovič”; LAOS: Houaphan: 1 ♀ PT (EJCB): “Laos-NE, Houa Phan prov., 20°12-13.5’N, 103°59’.5-104°01’E, Ban

129

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Saluei → Phou Pane Mt., 1340-1870m, 15.iv.-15.v.2008, Lao collectors leg.”. Adult occurence: 4-5-6-7 (n=9). Altitude range: 500-1870 m (n=3). Host plants: Unknown. Distribution: CHINA: Henan, Shaanxi, Yunnan. LAOS: Houaphan.

margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): distinctly longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): markedly dorsoventrally bent, parameres with obvious membranous lobes and apex of medial lobe sharply pointed.

Etymology: Noun in apposition; the specific name refers to the ethnic group Manchu, a Tungusic people who originated in Manchuria.

Diagnosis: posterior third of pronotum with carinate prehumerus; aedeagus longer, gradually dilated from base to apex. Species group: Pekinensis. Relevant species: A. monogrammus Thomson, 1879.

Agrilus mandala Jendek, sp. nov.

Type locality: Southeastern Tibet, about 20 km north of Yanjing, altitude 3000-3500 m.

Fig. 22F (habitus of HT); Fig. 37O (aedeagus)

Description: Holotype 7.3 mm. BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): green or brown. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): subparallel, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: small or moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: long. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: vague or moderate, Shape: arcuate or subangulate, Position to anterior angles: not reaching level of anterior pronotal angles. Posterior angles: Shape: rectangular, Apex: sharp. Disk: Convexity: flat, Disk impressions: medial and lateral or lateral only, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): deep, Lateral impressions (width): wide. Prehumerus: carinal, Shape: arcuate, Extent: to third of pronotal length or to half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: absent, rarely present, Narrowest point: at posterior 1/51/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: bicolored, Alternative color (location): pleural portions, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate, Width: wide, Shape: subangulate or subtruncate. Elytral pubescence: absent. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: large, Anterior margin: arcuately emarginate, Emargination (width): moderate. Prosternal process: moderate, Shape: narrowed, Sides: straight or arcuate, Angles: absent or obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical

Type specimens: Holotype (Fig. 22F), ♂, (EJCB): “SE Tibet, 3000-3500 m, ca 20 km N of Yanjing, 23.vii.-8.viii.1998, O. Šafránek et M. Tryzna”. Adult occurence: 7-8 (n=1). Altitude range: 3000-3500 m (n=1). Host plants: Unknown. Distribution: CHINA: Sichuan. Etymology: Noun in apposition; the specific name is the Sanskrit noun mandala (circle). In common use, mandala has become a generic term for any plan, chart or geometric pattern that represent the cosmos.

Agrilus marginicollis Saunders Fig. 23L (habitus); Fig. 38G (aedeagus); Figs. 43M, 46X, 49B (types) marginicollis Saunders, 1873 (Agrilus) SAUNDERS, 1873: 516 [#2554] (description). LEWIS, 1879: 15 [#3345] (catalog; Japan). KERREMANS, 1885: 153 [#1381] (catalog). SCHÖNFELDT, 1887: 113 [#8901] (Anambus; catalog; Japan). KERREMANS, 1892: 263 [#1397] (catalog). KERREMANS, 1903: 287 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 18 [#1809] (catalog; Japan). OBENBERGER, 1936: 1013 [#2191] (world catalog). CHÛJÔ & KUROSAWA, 1950: 10 [#501] (catalog; faunal records; Japan (Shikoku)). KUROSAWA, 1963: 154 [#1618] (characters; Japan). KUROSAWA, 1974: 3 [#4577] (note). TÔYAMA, 1985: 30, 32 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). JENDEK, 1994: 22 [#1355] (lectotype designation; synonymy; characters; Japan; China). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 185 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 35 [#4091] (checklist; Japan: Hokkaido, Honshu, Shikoku, Kyushu). FUKUTOMI & HORI, 2004: 7 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2005: 13 [#7217] (synonymy). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). ANONYMOUS, 2007: 32 [#8250] (checklist; Japan: Honshu). BELLAMY, 2008: 2179 [#8576] (world catalog). LEE, KIM & AHN, 2010: 264 266 [#8939] (characters; faunal records; South Korea). hastuliferoides Gebhardt, 1929 (Agrilus) OBENBERGER, GEBHARDT, 1929: 32 33 [#1051] (description). 1936: 997 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 355 [#2429] (checklist; China). JENDEK, 1994: 22 [#1355] (synonym

130 of marginicollis; lectotype designation). AKIYAMA & OHMOMO, 1997: 35 [#4091] (synonym of marginicollis). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Zhejiang). JENDEK, 2006: 399 [#7441] (synonym of marginicollis; Palaearctic catalog). BELLAMY, 2008: 2179 [#8576] (synonym of marginicollis; world catalog). beppuensis Obenberger, 1936 (Agrilus) OBENBERGER, 1936: 140 [#2207] (description). MIWA & CHÛJÔ, 1936: ADDENDA P. 4 [#1809] (catalog; Japan). OBENBERGER, 1936: 959 [#2191] (world catalog). KUROSAWA, 1975: 1 [#4591] (synonym of subrobustus). AKIYAMA & OHMOMO, 1997: 40 [#4091] (synonym of subrobustus). JENDEK, 2005: 13 [#7217] (synonym of marginicollis; lectotype designation). JENDEK, 2006: 402 [#7441] (synonym of subrobustus; Palaearctic catalog). BELLAMY, 2008: 2179 [#8576] (synonym of marginicollis; world catalog).

Type material: Agrilus marginicollis Saunders, 1873 Type locality: Hiogo. Type specimens: Primary: Lectotype (Fig. 49B), ♂, designated by JENDEK (1994: 22) [#1355], (BMNH): “Type H. T. [p] [round label with red border] \ Hiogo 1871 [h] \ Japan. G. Lewis 1910-320 [p] \ Agr. marginicollis ES Type [by Saunders hand] \”. Described from 4 syntypes. Agrilus hastuliferoides Gebhardt, 1929 Type locality: China: Chekiang. Type specimens: Primary: Lectotype (Fig. 46X), ♀, designated by JENDEK (1994: 22) [#1355], (MNHN): “China Chekiang [p] \ Typus [p] [red label] \ Agrilus hastuliferoides Type [h] det. Dr. A.v. Gebhardt [p] \ Museum Paris 1936 coll. A. Thery [p]”. Secondary: 1 PLT (NMPC); 1 PLT (HNHM). Described from unknown number of syntypes. Agrilus beppuensis Obenberger, 1936 Type locality: Japan: Beppu. Type specimens: Primary: Lectotype (Fig. 43M), ♀, designated by JENDEK (2005: 13) [#7217], (NMPC): “Japan, VII. 37 Beppu [h] \ Typus [p] [red label] \ A. beppuensis m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24954 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 4.3-5.8 mm; elytral apices without spinate adsutural projection, basal ventrite in male without tubercles. Species group: Sulcicollis. Relevant species: A. suturicuspidatus Jendek, 2007. Specimens examined: CHINA: Henan: 1 ♀ (EJCB): “China, N. Henan, 11 12.VI., Jiuligou, 550 m, 35º12’N, 112º26’E, Jaroslav Turna leg. 2007”. Hubei: 1 ♀ (EJCB): “China, E Hubei, 17 18. vi., Dabie Shan, 31.1N, 115.8E, Wujiashan, forest park, Jaroslav Turna leg., 2003”; 1 (EJCB): “China, W Hubei, 17.vii. saddle 5 km N Gaucho, road Xingshan Badong, 31.2N, 110.5E, 1500m, Jaroslav Turna leg., 2003”. Shaanxi: 5 (EJCB): “China Shaanxi pr. Hua Shan 17 21.VI.1991 R. Dunda & R. Sauer lgt.” (JENDEK, 1994 [#1355]). JAPAN: Honshu: 1 ♀ (EJCB): “Sarushima Ja, Yokosuka City, Kanagawa Pref., 13.VII.[19]80, leg. M. Horikawa” (JENDEK, 1994 [#1355]); 1 (EJCB): “Kohoku Ku, Yokohama City, Kanagawa Pref., 5.VI.[19]81 leg. V. Kusakabe” (JENDEK, 1994 [#1355]); 1 ♂ (EJCB): “Japan, C Honshu is., Yamanashi pref., Misaka pass, 4.vii.1985”. Kyushu: 1 ♀ (EJCB): “Kuro dake, Mts. Kujyusan, Oita Pref., 27.VI.1982, N. Takei leg.” (JENDEK, 1994 [#1355]). KOREA SOUTH: 1 (JLCS): “Nari bunji, Ulreungdo Is., Korea,

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA 15.viii.1995, S. L. An leg.”; 1 (JLCS): “Mt. Odaesan, Gangwon do, Korea, ii.2003, Hae yong Oh”. RUSSIA: Primorskiy kray: 1 ♂ (ZIN): “Prim., kr. Partizansk r n, p. Partizansk, 20.09.1986 g [in Russian]”; 2 (ZIN): “ex pupo 1 5.XI.1997, VIII.1997, okr. Vladivostoka, bukhta Tavaiza, s vinograda [Vitis] leg. S. Y. Mukhanov [in Russian]”.

Adult occurence: 2-6-7-8-9 (n=11). Altitude range: 5501500 m (n=2). Host plants: Vitis coignetiae: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: CHINA: Henan, Hubei, Shaanxi, Zhejiang. JAPAN: Hokkaido, Honshu, Kyushu, Shikoku. KOREA NORTH. KOREA SOUTH. RUSSIA: Primorskiy kray.

Agrilus miao Jendek, sp. nov. Fig. 13J (habitus of HT); Fig. 34D (aedeagus)

Description: 6.5-8.3 mm (Holotype 6.7 mm). BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): green. HEAD: Medial impression: absent or present, Impression (extent): vertex and frons, Impression (depth): shallow, Epistoma: in plane with frons. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Sculpture on vertex: punctures or rugae, Sculpture (aspect): regular, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: small, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square, Sides: straight or arcuate, Maximal width: at middle, Anterior margin: subequal to posterior or narrower than posterior. Anterior lobe: vague or moderate, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles or at level with anterior pronotal angles. Posterior angles: Shape: acute or rectangular, Apex: sharp. Disk: Convexity: flat or moderately convex, Disk impressions: medial and lateral, Medial impression (shape): entire, rarely anteromedial and posteromedial, Medial impression (deepness): shallow. Prehumerus: filamentary or carinal, Shape: arcuate, Extent: to third of pronotal length, Modifications: with rudiment at anterior pronotal angle, Posterior end: joining posterior pronotal angle or joining posterior pronotal margin, Anterior end: distant from pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present, rarely obsolete. ELYTRA: Color: unicolored with darker hue in apical half, Humeral carina: absent, Apical part: markedly attenuate, rarely feebly attenuate. Apices: Arrangement: separate, Width: narrow, Shape: arcuate or subangulate. Elytral pubescence: entire except glabrous part at apical half, Color: unicolored, Character: with patches of denser pubescence. Tomentum: rarely spots, Spots (locations): rarely premedial or postmedial. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate, Anterior

131

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA margin: arcuately emarginate, Emargination (deepness): moderate or deep, Emargination (width): wide. Prosternal process: moderate, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: impressed with sides protruding, Modification: rarely sides crenulate. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): distinctly longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed. Diagnosis: prehumerus feebly carinal or filamentary, short and feebly arcuate; aedeagus subparallel. Species group: Plagiatus. Relevant species: A. pumi sp. nov. Type locality: China, Yunnan province, Sabe. Type specimens: Primary: Holotype (Fig. 13J), ♂, (EJCB): “China-W, Yunnan, Sabe, 28.6.-1.7.2002, lgt. E. Kučera”. Secondary: CHINA: Sichuan: 1 ♂ PT (MNHN): “Chasseurs Indigènes des Missionnaire de Ta-tsien-Lou, 1906”; Yunnan: 1 ♂ PT (EJCB): “China: N. Yunnan, Lijiang, 2600 m, 30.6.- 2.7.1990, L.& M. Bocák lgt.”; 1 ♂ PT (EJCB): “China, Yunnan prov., 27°08’N, 100°14’E, 2900-3500 m, Yulongshan mts., Baishui, 7-12.vii.1990, Vít Kubáň leg.”; 1 ♀ PT (EJCB): “China-Yunnan, Baishui, 27°N, 100°12E, 10.6.-17.6.1998, lgt. E. Kučera”. Adult occurence: 6-7 (n=4). Altitude range: 2600-3500 m (n=2). Host plants: Unknown. Distribution: CHINA: Sichuan, Yunnan.

to anterior angles: at level with anterior pronotal angles or projecting beyond anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: flat or moderately convex, Disk impressions: medial and lateral or lateral only, Medial impression (shape): anteromedial and posteromedial, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial, Lateral impressions (deepness): deep. Prehumerus: carinal, Shape: arcuate, Extent: to third of pronotal length, Posterior end: joining posterior pronotal angle or joining posterior pronotal margin, Anterior end: distant from pronotal marginal carina, Arc: moderate or weak. Marginal and submarginal carinae: Interspace: moderate, Convergence: subparallel, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: rudimentary, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Width: wide, Shape: arcuate. Elytral pubescence: absent. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): moderate. Prosternal process: moderate, Shape: subparallel or dilated, Sides: straight or arcuate, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: subangulate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: elongate.

Etymology: Noun in apposition; the specific name refers to the ethnic group Miao inhabiting mainly regions of Yunnan province of China.

Diagnosis: body ventrally less lustrous, apex of elytra more elongate; pronotum without postero-lateral impression; prosternal process subparallel or dilated. Species group: Betuleti. Relevant species: A. cupes Lewis, 1893.

Agrilus mio Jendek, sp. nov.

Type locality: China, Shaanxi province, Qinling mts., 6 km east Xunyangba, altitude 1000-1300 m.

Fig. 18D (habitus of HT); Fig. 35T (aedeagus)

Description: 8.1-9.0 mm (Holotype 8.2 mm). BODY: Shape: cuneiform, Build: robust, Color (dorsally): unicolored, Color (dominant): blue or green. HEAD: Medial impression: present, Impression (extent): vertex or vertex and frons, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: flat, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): regular, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: small or moderate, Shape: not protruding from head outline or protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short or moderate, Sexual modification in male: antennae longer. PRONOTUM: Shape: transverse, Sides: arcuate, rarely straight, Maximal width: at middle or at posterior margin, Anterior margin: subequal to posterior or narrower than posterior. Anterior lobe: vague, Shape: arcuate, Position

Type specimens: Primary: Holotype (Fig. 18D), ♂, (EJCB): “China, 1000-1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.-13.vi.1998, I. H. Marshal leg.”. Secondary: CHINA: Hubei: 1 ♂ PT (EJCB): “China, W-Hubei prov., Dashennongjia Nat. Res., Muyi, E-slope, 2000m, 12-16 Jun 1997, Bolm lgt.”; Shaanxi: 2 ♀ PT (EJCB): “China, 10001300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.13.vi.1998, I. H. Marshal leg.”. Adult occurence: 5-6 (n=2). Altitude range: 1000-2000 m (n=2). Host plants: Unknown. Distribution: CHINA: Hubei, Shaanxi. Etymology: The specific name is a random combination of letters.

132

Agrilus miwai Obenberger Fig. 23E (habitus); Fig. 37Y (aedeagus); Fig. 49K (type) miwai Obenberger, 1936 (Agrilus) OBENBERGER, 1936: 37 [#2200] (description). MIWA & CHÛJÔ, 1936: 18 [#1809] (catalog; Japan). OBENBERGER, 1936: 1092 [#2191] (world catalog). MIWA & CHÛJÔ, 1940: 54, 58 60 [#1810] (characters; faunal record; Formosa). PENG ZHONGLIANG, 1987: 356 [#2429] (checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Taiwan). MÜHLE, 2003: 47 [#6155] (checklist; Taiwan). JENDEK, 2007: 91 [#8008] (lectotype designation; type locality determined by lectotype). BELLAMY, 2008: 2189 [#8576] (world catalog).

Type material: Agrilus miwai Obenberger, 1936 Type locality: Formosa. Type specimens: Primary: Lectotype (Fig. 49K), ♀, designated by JENDEK (2007: 91) [#8008], (NMPC): “Formosa, Koshun [h] \ Typus [p] [red label] \ Agrilus Miwai m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 26858 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 4.5-6.4 mm; body wider; prosternal process in male glabrous; basal ventrite in male with pair of almost merging tubercles on distal margin. Species group: Sulcicollis. Relevant species: A. reichardti Obenberger, 1935; A. volkovitshi Jendek, 2007. Specimens examined: CHINA: Henan: 2 ♀ (EJCB): “China, N. Henan, 11 12.VI., Jiuligou, 550 m, 35º12’N, 112º26’E, Jaroslav Turna leg. 2007”. Shaanxi: 1 ♂, 1 ♀ (EJCB): “China, 1000 1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v. 13.vi.1998, I. H. Marshal leg.”; 1 ♀ (EJCB): “C China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000 1300m, 23.V. 13.VI. leg. C. Holzschuh 2000”. LAOS: Houaphan: 50 (EJCB): “Laos NE, Houa Phan prov., 20°12 13.5’N, 103°59’.5 104°01’E, Ban Saluei → Phou Pane Mt., 1340 1870m, 15.iv. 15.v.2008, Lao collectors leg.”.

Adult occurence: 4-5-6 (n=4). Altitude range: 550-1870 m (n=4). Host plants: Unknown. Distribution: CHINA: Henan, Shaanxi, Taiwan. LAOS.

Agrilus moerens Saunders Fig. 29I (habitus); Fig. 40X (aedeagus); Figs. 42U, 48X, 49C, 49L, 50N, 52D, (types) moerens Saunders, 1873 (Agrilus) SAUNDERS, 1873: 517 [#2554] (description). KERREMANS, 1885: 153 [#1381] (catalog). SCHÖNFELDT, 1887: 113 [#8901] (Anambus; catalog; Japan). KERREMANS, 1892: 264 [#1397] (catalog). KERREMANS, 1903: 287 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 18 [#1809] (catalog; Japan). OBENBERGER, 1936: 1014 [#2191] (world catalog). KUROSAWA, 1974: 3 [#4577] (note). TÔYAMA, 1985: 32 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). JENDEK, 1995: 138 [#3768] (lectotype designation; synonymy; notes). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 185 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO,

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA 1997: 35 36 [#4091] (checklist; Japan: Hokkaido, Honshu, Shikoku, Kyushu, Tsushima; Korea; North China; Russia: East Siberia). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). NONNAIZAB et al., 1999: 113 [#8247] (checklist; China: Inner Mongolia). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Taiwan). FUKUTOMI & HORI, 2004: 12 13 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2005: 14 [#7217] (synonymy). JENDEK, 2006: 393 [#7441] (subgenus Robertius; Palaearctic catalog). BELLAMY, 2008: 2190 2191 [#8576] (subgenus Robertius; world catalog). VOLKOVITSH, 2009: 136 [#8850] (subgenus Robertius; faunal records; Russia: Primorskiy kray). rotundicollis Saunders, 1873 (Agrilus) SAUNDERS, 1873: 517 518 [#2554] (description). LEWIS, 1879: 15 [#3345] (catalog; Japan). KERREMANS, 1885: 154 [#1381] (catalog). SCHÖNFELDT, 1887: 113 [#8901] (Anambus; catalog; Japan). KERREMANS, 1892: 270 [#1397] (catalog). KERREMANS, 1903: 290 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 20 [#1809] (catalog; Japan). OBENBERGER, 1936: 1032 [#2191] (world catalog). MIWA & CHÛJÔ, 1940: 57, 70 [#1810] (cited as vladivostokanus; [NOTE: MISSIDENTIFIED AS VLADIVOSTOKANUS ACCORDING TO CHÛJÔ & KUROSAWA, 1950: 10 [#501]]; notes; Japan: Hokkaido). KUROSAWA, 1949: 14 [#5028] (characters). CHÛJÔ & KUROSAWA, 1950: 10 [#501] (catalog; faunal records; Japan (Shikoku)). IGA, 1955: 79 [#3828] (iconography; Japan). IGA, 1962: 79 [#3351] (iconography; Japan). KUROSAWA, 1963: 154 [#1618] (cited as ratundicollis; characters; Japan). KUROSAWA, 1974: 4 [#4577] (note). TÔYAMA, 1985: 30 [#3678] (iconography; Japan). ANONYMOUS, 1986: 177 [#8848] (cited as ratundicollis; insect pest; Korea). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). CHEN, 1992: 401 402 [#6192] (cited as ratundicollis; characters; bionomy). LI JINGKE, 1992: 93 [#6755] (checklist; northeastern China). JENDEK, 1994: 15 16 [#1355] (lectotype designation; synonymy; characters; distribution; Russian Far East; Korea; China; Japan). JENDEK, 1995: 138 [#3768] (synonym of moerens; notes). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 186 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 36 [#4091] (synonym of moerens). NONNAIZAB et al., 1999: 113 [#8247] (checklist; China: Inner Mongolia). HUA LI ZHONG, 2002: 90 [#6696] (subspecies of rokuyai; checklist; northeastern China). JENDEK, 2006: 393 [#7441] (synonym of moerens; Palaearctic catalog). BELLAMY, 2008: 2190 [#8576] (synonym of moerens; world catalog). araxicola Abeille de Perrin, 1897 (Agrilus) ABEILLE DE PERRIN, 1897: 3, 11 12 [#19] (description). KERREMANS, 1903: 282 [#1441] (catalog). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 949 [#2191] (world catalog). JENDEK, 2005: 14 [#7217] (synonym of moerens; lectotype designation; type locality confused). JENDEK, 2006: 393 [#7441] (synonym of moerens; Palaearctic catalog). BELLAMY, 2008: 2190 [#8576] (synonym of moerens; world catalog). mandjuricus Obenberger, 1922 (Agrilus) OBENBERGER, 1922: 23 24 [#2021] (description). OBENBERGER, 1924: 47 [#2040] (note). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1012 1013 [#2191] (world catalog). KUROSAWA, 1974: 4 [#4577] (related to A. rotundicollis). JENDEK, 1994: 15, 16 [#1355] (synonym of rotundicollis; lectotype designation). JENDEK, 1995: 138 [#3768] (synonym of moerens). AKIYAMA & OHMOMO, 1997: 36 [#4091] (synonym of moerens). JENDEK, 2006: 393 [#7441] (synonym

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA of moerens; Palaearctic catalog). BELLAMY, 2008: 2190 [#8576] (synonym of moerens; world catalog). marinus Obenberger, 1922 (Agrilus) OBENBERGER, 1922: 24 25 [#2021] (description). OBENBERGER, 1924: 47 [#2040] (note). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1013 [#2191] (world catalog). ALEXEEV, 1979: 129 [#57] (synonym of vladivostokanus). JENDEK, 1994: 15, 16 [#1355] (synonym of rotundicollis; lectotype designation). JENDEK, 1995: 138 [#3768] (synonym of moerens). AKIYAMA & OHMOMO, 1997: 36 [#4091] (synonym of moerens). JENDEK, 2006: 393 [#7441] (synonym of moerens; Palaearctic catalog). BELLAMY, 2008: 2191 [#8576] (synonym of moerens; world catalog). paphius Théry, 1942 (Agrilus) THÉRY, 1942: 268 270 [#2875] (description). OBENBERGER, 1943: 10 [#2333] (synonym of vladivostokanus). JENDEK, 1994: 15 [#1355] (synonym of rotundicollis). JENDEK, 1995: 138 [#3768] (synonym of moerens). AKIYAMA & OHMOMO, 1997: 36 [#4091] (synonym of moerens). JENDEK, 2006: 393 [#7441] (synonym of moerens; Palaearctic catalog). BELLAMY, 2008: 2191 [#8576] (synonym of moerens; world catalog).

Type material: Agrilus moerens Saunders, 1873 Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 49L), ♂, designated by JENDEK (1995: 138) [#3768], (BMNH): “Type H.T. [p] [round label with red border] \ Japan. G. Lewis 1910320 [p] \ Moerens ES [by Saunders hand] \ Agrilus moerens E.S. Type [by Lewis hand]”. Described from unknown number of syntypes. Agrilus rotundicollis Saunders, 1873 Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 52D), ♀, designated by JENDEK (1994: 15-16) [#1355], (BMNH): “Type H. T. [p] [round label with red border] \ Japan. G. Lewis 1910-320 [p] \ Agr. rotundicollis ES Type [by Saunders hand] \ Agrilus rotundicollis ES Type [by Lewis hand] \”. Described from unknown number of syntypes. Agrilus araxicola Abeille de Perrin, 1897 Type locality: Caucase, vallée de l’Araxe [Note: locality confused, see JENDEK, 2005 [#7217]]. Type specimens: Primary: Lectotype (Fig. 42U), ♂, designated by JENDEK (2005: 14) [#7217], (MNHN): “Araxe [h] \ araxicola Ab. Type [h] \ Muséum Paris Coll. Abeille De Perrin 1919 [p] \ vid Niehuis 1992 [p]”. Described from unknown number of syntypes. Agrilus mandjuricus Obenberger, 1922 Type locality: Siberia orientalis: Vladivostok. Type specimens: Primary: Lectotype (Fig. 48X), ♂, designated by JENDEK (1994: 15, 16) [#1355], (NMPC): “COLLECTIO DR. OBENBERGER MUS. PRAGENSE [p] \ Vladivostok [h] \ Typus [p] [red label] \ Agrilus mandjuricus m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24879[h] [orange label]”. Described from unknown number of syntypes. Agrilus marinus Obenberger, 1922 Type locality: Siberia orientalis: Vladivostok. Type specimens: Primary: Lectotype (Fig. 49C), ♂, designated by JENDEK (1994: 15, 16) [#1355], (NMPC): “COLLECTIO DR. OBENBERGER MUS. PRAGENSE [p]

133 \ Vladivostok [h] \ Typus [p] [red label] \ marinus m. Type male symbol [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Agrilus paphius Théry, 1942 Type locality: Erzendjanzcy, Mandchourie. Type specimens: Holotype by monotypy (Fig. 50N), (NHMB): “Erzendjanzcy [p] 16.6.40 [h] 19 [p] 39 [p] [scratched] Mandschurei [p] \ Type [p] [red label] \ Sammlung Georg Frey [p] \ Agrilus paphius Thery [h] Type [p] [red ink] \ Agrilus 1943 vladivostokanus m. comp. type [h] Det. Dr. Obenberger [p]”. Described from 1 specimen. Diagnosis: 4.2-5.5 mm, dorsally glabrous; elytral apices broader; basal ventrite in male with deep medial impression. Species group: Roscidus. Relevant species: A. roscidus Kiesenwetter, 1857. Specimens examined: CHINA: Beijing: 1 (EJCB): “CHINA Beijing pr. Qinglongqiao 8 9.6.1991” (JENDEK, 1994 [#1355] as rotundicollis); 1 (EJCB): “China: Beijing Region, Xiaolongmen, ca 1000m, N 39°58.074’, E 115°25.882’, 9 13.VI.2004, Leg. J. Cooter”. Gansu: 1 ♀ (EJCB): “China, S Gansu, Dengkagoin, 5 9.vi.1996, Beneš & Štepař leg.”. Hebei: 1 ♀ (EJCB): “China, N Hebei, 3.vi.2000, Fengning env. NW slope of Yunwu Shan, 41.2, 116.6E, 1200m, Jaroslav Turna leg.”; 7 (EJCB): “China, Hebei/Nei Mongol, pass Chengde Chifeng, 41.6N, 118.2E, 14 16.VI., Jaroslav Turna leg. 2001”; 10 (EJCB): “China, Hebei/Nei Mongol, pass Chengde Chifeng, 41.6N, 118.2E, 14 16.VI.2001, O. Šafránek leg.”. Shaanxi: 5 (EKCS): “China Shaanxi, Lüeyang, 8.6. 14.6.1996, lgt. E. Kučera”. Shanxi: 1 ♂, 2 ♀ (EJCB): “China, W Shanxi, 9.VI.2000, 37.7N, 111.6E, Lüliang Shan, road Fangshan Jiaocheng, Hengjian env., ~1000m, Jaroslav Turna leg.”. Sichuan: 5 (EJCB): “China, Sichuan 8 13.VI.91 Abazhou (102) 2000m Nanping Jiuzhaigou 33.15 N / 104.15 E” (JENDEK, 1994 [#1355] as rotundicollis); 1 ♀ (EJCB): “China, N Sichuan 30 km W Nanping 11.,16.VI.1992 Jiuzhaigou, 2100 m, Jaroslav Turna leg.” (JENDEK, 1994 [#1355] as rotundicollis). JAPAN: Honshu: 1 ♀ (NMPC): “Tokyo Japan” (JENDEK, 1994 [#1355] as rotundicollis); 1 ♂ (EJCB): “Japan Kumanotaira pr. Kuruizawa 12.7.[19]08” (JENDEK, 1994 [#1355] as rotundicollis); 1 ♂ (EJCB): “Mt. Tsukubasan Ibaraki Pref. 16.V.1982 S. Ohmomo” (JENDEK, 1994 [#1355] as rotundicollis); 1 ♂ (EJCB): “Yoshise Tsuchiura C. Ibaraki Pref. 13.V.1982 S. Ohmomo” (JENDEK, 1994 [#1355] as rotundicollis). KOREA NORTH: 1 (VSCS): “Korea, Sujang san bei Hedzu, 4.6.1975, Josifov”. 1 (EJCB): “Korea prov. Phondjang Si, leg. M. Mroczkowski et A. Riedel Songmun ri, distr. Samsok 22.V.1965” (JENDEK, 1994 [#1355] as rotundicollis); 1 (VSCS): “Korea, Kymgang san, Samilpo, 25.5.1975, Josifov leg.”; 1 (VSCS): “Korea, Camo san, Pjômgsang si, 18.5.1975, Josifov leg.”; 5 (EJCB): “D.P.R. Korea Ryongaksan Mts. 10 km W of Pyongyang lgt. M. Slovák, M. Kozánek, 14 16.V. 1988” (JENDEK, 1994 [#1355] as rotundicollis); 1 (EJCB): “D.P.R. Korea Okryu Valley Kumgangsan Mts. lgt. M. Slovák 19.V.1988” (JENDEK, 1994 [#1355] as rotundicollis). RUSSIA: Primorskiy kray: 5 (MNHN): “Ile Askold, Mantschourie, M. Jankowski, 1880”; 1 (NMPC): “Sibir. or. Ussuri Vladivostok Dr. Jurecek 1919” (JENDEK, 1994 [#1355] as rotundicollis); 1 (ZIN): “Primorsk. krai 15 17 km NNO st. Khasan Loginova 20.VI.[19]74 [in Russian]” (JENDEK, 1994 [#1355] as rotundicollis); 1 (IMCM): “Primor’e, g. Barabash Levada 6.vi.1989, Nikireev, S. [in Russian]”; 1 (EJCB): “Primorje Novickoje 15 km IU Partizanska les 19.06.1990 Belokobylskij” (JENDEK, 1994 [#1355] as rotundicollis); 1 (EJCB): “USSR, Primorskij kraj Arseneev env. 300 400 m 11. 19.VII.1990 B. Šiška & R. Čermák leg.” (JENDEK, 1994 [#1355] as rotundicollis); 2 (ZIN): “Primorskij kraj Lyalitchi r.Ilistaya

134 4.06.1990 Belokobylskij [in Russian]” (JENDEK, 1994 [#1355] as rotundicollis); 3 (ZIN): “Primorskij kraj Spassk forest 13 15. VI.1990 Belokobylskij [in Russian]” (JENDEK, 1994 [#1355] as rotundicollis); 3 (ZIN): “Vladivostok, Morskoe Kladbishsche, les, 17.06.1990 Belokobylskij [in Russian]” (JENDEK, 1994 [#1355] as rotundicollis); 1 ♀ (EJCB): “Sib.or. m., Primorje Sichote Alin Mts. Sokolči 1. 15.7.1990 Kadlec + Voříšek lg.” (JENDEK, 1994 [#1355] as rotundicollis); 1 (EJCB): “USSR, Primorsk. kraj, Arsenev env., VI.1991, 44.11N, 133.19E, O. Šausa leg.” (JENDEK, 1994 [#1355] as rotundicollis); 6 (EJCB): “USSR, Primorskij kraj Arseneev env., VI.1991, 44.11 N, 133.19 E, M. Štrba leg.” (JENDEK, 1994 [#1355] as rotundicollis); 1 (MNCA): “Russia: Primorskiy Kray, Kremovo, 25 km SW Sibirtsevo, 43.59N, 132.18E, 23.V.1993, 120 m, leg. L. Zerche et al.”; 1 ♀ (EJCB): “Russia, Primorskij kraj, Černigovka env., Sinnij chrebet Mts., 14 16.vi.1997, leg. L. Roller, L. Vidlička”; 1 (MSCI): “SE Primorskii kr., Lazovskii zap. 10 km SW pos. Sokolchi, okr. kordona Korpad’, 43°15’N, 134°07’E, 23 24.6.2006, leg. M. & L. Smirnov [in Russian]”; 1 ♂ (ZIN): “Primorskii kr., s. Lazo 8 12.vi.2007 V. Shokhrin”.

Adult occurence: 5-6-7 (n=33). Altitude range: 120-2100 m (n=7). Host plants: Quercus: ALEXEEV, 1979 (as vladivostokanus) [#57]. − Q. acutissima: AKIYAMA & OHMOMO, 1997 [#4091]. − Q. serrata AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: CHINA: Beijing, Gansu, Hebei, Heilongjiang, Nei Mongol, Shaanxi, Shanxi, Sichuan. JAPAN: Hokkaido, Honshu, Kyushu, Shikoku, Tsushima. KOREA NORTH. KOREA SOUTH. RUSSIA: Primorskiy kray.

Agrilus mongolorum Obenberger Fig. 21D (habitus of LT); Fig. 37A (aedeagus) mongolorum Obenberger, 1936 (Agrilus) OBENBERGER, 1936: 104 [#2205] (description). OBENBERGER, 1936: 1015 [#2191] (world catalog). COBOS, 1972: 56 [#591] (subgenus Agrilus; checklist; Mongolia). ALEXEEV, 1975: 156 157 [#52] (presumable synonym of aerosus). PENG ZHONGLIANG, 1987: 356 [#2429] (checklist; China). ALEXEEV, 1998: 375 [#4322] (subgenus Xeragrilus; subgeneric classification). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Inner Mongolia). JENDEK, 2006: 396 [#7441] (subgenus Xeragrilus; Palaearctic catalog). BELLAMY, 2008: 2192 [#8576] (subgenus Xeragrilus; world catalog).

Type material: Agrilus mongolorum Obenberger, 1936 Type locality: Mongolia mer. or. Type specimens: Primary: Lectotype (Fig. 21D) by present designation, ♂, (NMPC): “Mongolia mer. or. [h] \ Typus [p] [red label] \ Agrilus Mongolorum m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24326 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: (♂) 6.4-6.6 mm; vertex with fine medial impression; antennae very short, not reaching to half of pronotal length; metatarsi distinctly shorter than metatibiae; pronotum markedly convex with distinctly pubescent and finely impressed sides; prehumerus absent; prosternal lobe large and deeply, arcuately emarginated. Species group: Albogularis. Relevant species: A. morawitzi Obenberger, 1936.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Specimens examined: Mongolia: 1 ♂ (ZIN): “Mongoliya, Yu. Gob. aim., uroch. Udzur Dzag, 40 km VYUV g. Khan Bogdo, Kozlov, 24.VI.[1]971 [in Russian]”.

Adult occurence: 6 (n=1). Host plants: Unknown. Distribution: MONGOLIA. Remarks: The distributional record from Nei Mongol (China) cited without supporting data by PENG ZHONGLIANG, 1987 [#2429] and HUA LI ZHONG, 2002 [#6696] was omitted.

Agrilus morawitzi Obenberger Fig. 21F (habitus of LT); Fig. 37C (aedeagus) morawitzi Obenberger, 1936 (Agrilus) OBENBERGER, 1936: 106 107 [#2205] (description). OBENBERGER, 1936: 1015 [#2191] (world catalog). VOLKOVITSH & ALEXEEV, 1994: 437 (as pseudoalbogularis) [#2974]. JENDEK, 2006: 400 [#7441] (Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2193 [#8576] (world catalog). richteri Alexeev, 1975 (Agrilus; ssp. of albogularis) syn. nov. ALEXEEV, 1975: 156 [#52] (description). KRIVOSHEINA, 1976: 23 [#8641] (subspecies of albogularis; rhizobiont; Turkmenistan). KAPLIN, 1981: 48 [#5853] (subspecies of albogularis; notes). ALEXEEV & VOLKOVITSH, 1989: 345, 356 [#3241] (subspecies of albogularis; characters in key; faunal records; Mongolia). BÍLÝ, 2002: 10 [#5877] (subspecies of albogularis; DISTRIBUTION; MONGOLIA). TLEPPAEVA & ISHKOV, 2004: 84 [#7211] (subgenus Xeragrilus; subspecies of albogularis; checklist; Kazakhstan). JENDEK, 2006: 396 [#7441] (subgenus Xeragrilus; subspecies of albogularis; Palaearctic catalog). BELLAMY, 2008: 1958 [#8576] (subspecies of albogularis; world catalog).

Type material: Agrilus morawitzi Obenberger, 1936 Type locality: Semirječia: Djarkent. Type specimens: Primary: Lectotype (Fig. 21F) by present designation, ♂, (NMPC): “Семиръченская обл. р. Или, Джаркентскій уъзд, 1906, В. Рюкбейль [p] \ Typus [p] [red label] \ Agrilus Morawitzi m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24341 [h] [orange label]”. Secondary: 1 PLT (NMPC). Described from unknown number of syntypes. Agrilus albogularis richteri Alexeev, 1975 Type locality: Mongolia, Vostochno-Gobiiskii aimak: NomtUla, 30 km YUYUV [south-southeastern] Schokhoi-Nur [in Russian]. Type specimens: Primary: Holotype (ZIN, temporarily in AAOR). Secondary: 2 PT (AAOR), 1 PT (ZIN). Described from 4 specimens (HT, PT). Diagnosis: 3.9-5.3 mm; head conspicuously large; vertex with deep medial carina; antennae long, reaching beyond half of pronotal length, prehumerus absent; whole ventral side of body distinctly pubescent; ventral side with tomentose patterns; prosternal lobe subtruncate or feebly emarginate. Species group: Albogularis. Relevant species: A. mongolorum Obenberger, 1936.

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Specimens examined: KAZAKHSTAN: 2 ♀ (ZIN): “Kazakhstan, Almaty reg., Qumbasy sandy desert, 69 km NNW of Qapchaghay, 16.V.2005, Volkovitsh”. 1 ♂ (ZIN): “ur. Kurtogai. r. Charyn, 35 km SZ pos. Kegen, Kazakhstan, Volkovitsh, 15 16.VII.1986 [in Russian]”; 1 ♂, 1 ♀ (ZIN): “ur. Kurtogai. r. Charyn, 35 km SZ Kegena, Kazakhstan, Volkovitsh, 2 4.VII.1987 [in Russian]”; 1 ♂ (EJCB): “Alma Ata, 8.V.1992”. KYRGYZSTAN: 2 ♂ (ZIN): “Kirg. CCP, r. Chu, Zoloty Zamki, 20.07.1978 [in Russian]”; 1 ♂ (ZIN): “r. Chu, 10 km VSV pos. Kochkorka, Kirgiziya, Volkovich 10.vii.1986 [in Russian]”; 1 ♂ (ZIN): “okr. pos. Kyzyl Oi, dolina reki Susamir, Kirgiziya, 1986, Dolgovskaya 5 6.vii. [in Russian]”; 1 ♀ (ZIN): “okr. pos. Kyzyl Oi, dolina reki Susamir, Kirgiziya, 1986, Emel’yanov 5 6.vii. [in Russian]”; 1 (ZIN): “r. Kasansai, 17 km YuV Tereksaya, Chatkal’sk khr., Ferg., Volkovitsh, 31.5. 1.6.1989 [in Russian]”. MONGOLIA: 1 ♀ (EJCB): “Mongolia 16.7.1975, Gobi Altaj ajm., 10 km SEE Altaj, exp. USSR”. RUSSIA: Tyva Respublika: 1 (ZIN): “Tuva, g. Kyzyl, B. Korotyaev, 3.7.[19]73 [in Russian]”; 2 ♂, 1 ♀ (EJCB): “Tuva, Kyzyl, right bank, 6 10. vi.1979, on Caragana”. TAJIKISTAN: 1 ♂ (ZIN): “vost. bereg. oz. Yashil’ kul’, 3900 4000 m, Pamir, GBAO, Volkovitsh, 22.vi.1986 [in Russian]”.

Adult occurence: 5-6-7 (n=15). Faunal records published by ALEXEEV, 1975 [#52] (as albogularis richteri) are also included. Altitude range: 3900-4000 m (n=1). Host plants: Ceratoides papposa [= Eurotia ceratoides]: ALEXEEV & VOLKOVITSH, 1989 (as albogularis richteri) [#3241]. − Eurotia ceratoides: ALEXEEV, 1975 (as albogularis richteri) [#52]; VOLKOVITSH & ALEXEEV, 1994 (as pseudoalbogularis) [#2974]; TLEPPAEVA & ISHKOV, 2004 (as albogularis richteri) [#7211]. Dubious: Caragana pygmaea: ALEXEEV & VOLKOVITSH, 1989 (as albogularis richteri) [#3241]. Erroneous: Artemisia: KRIVOSHEINA, 1976 (as albogularis richteri) [#8641]. − A. dimoana: KAPLIN, 1981 (as albogularis richteri) [#5853]. Distribution: KAZAKHSTAN. KYRGYZSTAN. MONGOLIA. RUSSIA: Tyva Respublika. TAJIKISTAN. Remarks: In ZIN, this species was frequently labeled by Alexeev with the unavailable name “pseudoalbogularis”.

Agrilus muongoides Jendek, sp. nov. Fig. 31A (habitus of HT); Fig. 41O (aedeagus)

Description: 3.6-5.1 mm (Holotype 4.3 mm). BODY: Shape: cuneiform, Build: slender, Color (dominant): black. HEAD: Shape: obviously flat, Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: flat or moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture (intensity): superficial. Eyes: Size: small or moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square, Sides: straight, arcuate or subangulate, Maximal width: at anterior margin, Anterior margin: wider than posterior or subequal to posterior. Anterior lobe: moderate, obvious, rarely vague, Shape: arcuate, Position to anterior angles:

135 at level with anterior pronotal angles, projecting beyond anterior pronotal angles, rarely not reaching level of anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: posterolateral. Prehumerus: carinal, Shape: bisinuate, Extent: beyond half of pronotal length or to anterior pronotal angle, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina or joining anterior pronotal angle, Arc: obvious. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior angles or at posterior 1/51/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate or conjoint, Shape: arcuate or subtruncate, Truncation: transverse. Elytral pubescence: absent or proximal only, Proximal: quadrangular, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: large, Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): narrow. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: rectangular, Angles (tips): sharp, Disk: impressed with sides protruding. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Male sexual modifications: metafemur markedly incrassate. Metatarsus (size to mesotarsus): somewhat longer than mesotarsus or distinctly longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): markedly dorsoventrally bent, Ovipositor: elongate. Diagnosis: frons flat; pronotum more elongate; prosternal process with impressed disk; male glabrous, female with white pubescence in basal half of elytra. Species group: Unassigned. Relevant species: A. muong Descarpentries & Villiers, 1967. Type locality: China, southeastern Hunan province, Guidong env., 26.04N 113.56E. Type specimens: Primary: Holotype (Fig. 31A), ♂, (EJCB): “China, Hunan SE, 1994, Guidong env. 26-31.V., 26.04N 113.56E, E. Jendek & O. Šauša leg.”. Secondary: CHINA: Hubei: 1 ♀ PT (EJCB): “China, W-Hubei prov., Dashennongjia Nat. Res., Muyi, E-slope, 2000m, 12-15 June 1997, Bolm lgt.”; 1 PT (EJCB): “China, W Hubei, 8.vii., Jiangshi SE env., 30.6N, 109.7E, 800 m, Jaroslav Turna leg., 2003”; Hunan: 2 ♂ PT (EJCB): “China, Hunan SE, 1994, Guidong env. 26-31.V., 26.04N 113.56E, E. Jendek & O. Šauša leg.”; 1 ♀ PT (EJCB): “China, N Hunan, 4-7.vii., Wuling Shan, 29.4N, 110.4E, Zhangjiajie, 700 m, Jaroslav Turna leg. 2003”; Jiangxi: 1 ♂ PT (EJCB): “China, Jiangxi W, Jinggang Shan, Ciping, 2-14.VI.1994, E. Jendek & O. Šauša leg.”; Yunnan: 1 ♂ PT (EJCB): “Yunnan, 12501500m, 23.14N 104.34E, Xinjie, 24.6.1994, Vít Kubáň leg.”; 1 ♂ PT (EJCB): “Yunnan, 1500-2500 m, 25°22’N, 98°49’E,

136 17-24 / 5 Gaoligong mts, Vít Kubáň leg., 1995”; LAOS: Borikhamxai: 1 ♂ PT (EJCB): “Laos centr., Bolikhamsai prov., Ban Nape - Kaew Nua Pass, 18.4.-1.5.1998, alt. 600±100 m, N 18°22.3, E 105°09.1 (GPS), E. Jendek & O. Šauša leg.”; Houaphan: 19 PT (EJCB): “Laos-NE, Houa Phan prov., 20°12-13.5’N, 103°59’.5-104°01’E, Ban Saluei → Phou Pane Mt., 1340-1870m, 15.iv.-15.v.2008, Lao collectors leg.”; Louang Namtha: 3 ♂ PT (EJCB): “Laos, Louang Namtha pr., 21°09’N, 101°19’E, Namtha - Muang Sing, 5-31.v.1997, 900-1200 m, Vit Kubáň leg.”; Louangphrabang: 1 ♀ PT (EJCB): “Laos-N, 23.iv.1999, Louangphrabang prov. 20°42’ N 102°54’E, 25 km E Muang Ngoy, 1000 m, Vít Kubáň leg.”; Phongsali: 2 ♂ PT (EJCB): “Lao-N, Phongsaly prov., 21°41’-2’N, 102°06’-08’E, 28.v.20.vi.2003 Phongsaly env., ~1500m, Vít Kubáň leg.”; 1 ♀ PT (EJCB): “Lao-N, Phongsaly prov., 21°41’-2’N, 102°06’08’E, Phongsaly env., 28.v.-20.vi.2003, 1500m, P. Pacholátko leg.”; VIETNAM: 1 ♂ PT (EJCB): “N Viet Nam (Tonkin) pr. Vin Phu, Tam Dao, 2-11.6.1985, Vít Kubáň leg.”; 1 ♂ PT (EJCB): “N Vietnam, 21°27N, 105°39E, 70 km NW of Hanoi, Tam Dao, 1-8.vi.1996, 900-1200 m, Dembický & Pacholátko leg.”. Adult occurence: 4-5-6-7 (n=15). Altitude range: 5002500 m (n=12). Host plants: Unknown. Distribution: CHINA: Hubei, Hunan, Jiangxi, Yunnan. LAOS: Borikhamxai, Houaphan, Louang Namtha, Louangphrabang, Phongsali. VIETNAM. Etymology: The specific name is a combination of the muong (specific name of another Agrilus) and -oides (Greek suffix denoting likeness) to stress the species similarity to A. muong Descarpentries and Villiers, 1967.

Agrilus muscarius Kerremans Figs. 20F 20H (habitus); Fig. 36S (aedeagus); Figs. 47Z, 49Q, 52Q (types) muscarius Kerremans, 1895 (Agrilus) KERREMANS, 1895: 224 [#1416] (description). KERREMANS, 1903: 279 [#1441] (catalog). OBENBERGER, 1936: 1092 [#2191] (world catalog). BAUDON, 1961: 73 [#143] (faunal records; Laos). DESCARPENTRIES & VILLIERS, 1963: 105, 110 [#700] ([Note: An inference of type: “Tonkin: Ha Lang (Lamey, type, British Museum)” is not valid lectotype designation because there were two syntypes in the Natural History Museum London (ICZN, Article 74.3)]; characters; faunal records; Tonkin; Laos). BAUDON, 1968: 132, 149 [#162] (catalog; Laos). BELLAMY, 2008: 2196 [#8576] (world catalog). JENDEK & GREBENNIKOV, 2009: 5, 17 19 [#8563] (characters in key; synonymy; lectotype designation; faunal records; distributional summary; China (Shaanxi, Sichuan, Yunnan); India (Assam, Meghalaya, Tamil Nadu, Uttar Pradesh, West Bengal (paratype of komiyai)); Laos (Borikhamxai, Louang Namtha, Louangphrabang, Phongsali, Vientiane); Myanmar (Shan State); Nepal; Sri Lanka; Thailand (Chiang Mai, Kanchanaburi, Loei, Mae Hong Son, Nan, Tak); Vietnam). seladon Obenberger, 1940 (Agrilus) PENG OBENBERGER, 1940: 174 175 [#2273] (description). ZHONGLIANG, 1987: 358 [#2429] (cited as seladom; checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA JENDEK, 2005: 18 [#7217] (lectotype designation). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2285 [#8576] (world catalog). JENDEK & GREBENNIKOV, 2009: 18, 31 [#8563] (synonym of muscarius). komiyai Tôyama, 1987 (Agrilus) TÔYAMA, 1987: 305 306 [#2931] (description). JENDEK, 2005: 18 [#7217] (synonym of seladon). JENDEK, 2006: 401 [#7441] (synonym of seladon; Palaearctic catalog). BELLAMY, 2008: 2285 [#8576] (synonym of seladon; world catalog). JENDEK & GREBENNIKOV, 2009: 18, 31 [#8563] (synonym of muscarius; HT examined).

Type material: Agrilus muscarius Kerremans, 1895 Type locality: Tonkin. Type specimens: Primary: Lectotype (Fig. 49Q), ♂, designated by JENDEK & GREBENNIKOV (2009: 5, 17-19) [#8563], (MNHN): “Meliboeus muscarus nov. s. Fairm. Ht. Tonkin [h] \ Ha-Lang Tonkin N. Lamey \ muscarius Kerr. [h] PARAType [p] [red label] \ Muséum Paris 1935 Coll. A. Théry [p] \ Muséum Paris Coll. Générale [p] [yellow label]”. Secondary: 2 PLT (BMNH); 1 PLT (MNHN). Described from unknown number of syntypes. Agrilus seladon Obenberger, 1940 Type locality: China: Yunnan, vallis fluminis Soling-Ho. Type specimens: Primary: Lectotype (Fig. 52Q), sex not examined, designated by JENDEK (2005: 18) [#7217], (NMPC): “Vallis flumin. Soling-ho, Yun. [p] \ Typus [p] [red label with black border] \ Mus. Nat. Pragae Inv. [p] 24926 [h] [orange label] \ Agrilus Seladon m. Type [h] Det. Dr. Obenberger [p]”. Described from unknown number of syntypes. Agrilus komiyai Tôyama, 1987 Type locality: Doi Pui near Chiang Mai, NW Thailand. Type specimens: Primary: Holotype (Fig. 47Z), sex not examined, (NSMT): “Doi Pui N. Thailand 4-7. v. 1983 M. Takakuwa [p] \ Holotype Agrilus komiyai Tôyama, 1983 [h]”. Described from 7 specimens (HT, PT). Diagnosis: 3.5-4.9 mm; aedeagus as in Fig. 36S; for further characters see JENDEK & GREBENNIKOV, 2009 [#8563]. Species group: Muscarius. Relevant species: A. semicaducus Jendek, 2009. Specimens examined: CHINA: Shaanxi: 1 (EJCB): “China Shaanxi, Lüeyang, 8.6. 14.6.1996, lgt. E. Kučera” (JENDEK & GREBENNIKOV, 2009 [#8563]). Sichuan: 1 ♂ (EJCB): “China N, Sechuan, Nanjiang, 21 23.5.2002, lgt. E. Kučera” (JENDEK & GREBENNIKOV, 2009 [#8563]). Yunnan: 1 (EJCB): “Yun’nan’. Syaomon’yan. 810 m, 27.iii.1957, Pu Fu di leg. [in Russian]” (JENDEK & GREBENNIKOV, 2009 [#8563]); 2 ♂ 1 ♀ (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 (EJCB): “China, Yunnan, Loc. 9., Road 214, Dali Lijiang, 65th 71st km NNW of Xiaguan, 26°03’N, 100°01’ 09’E, 2350 2528 m, 24.05.2002, Konstantinov & Volkovitsh leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). INDIA: Assam: 6 (EJCB): “NE India, Assam, 1999, 5 km N of Umrongso 700m, 25°27’N, 92°43’E, 17. 25.v., Dembický & Pacholátko leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). Meghalaya: 10 (EJCB): “NE India, Meghalaya state, West Garo Hills, Nokrek Nat.Park, 9 17.V.1996 alt.1100±150m, GPS N25°29.6’, E90°19.5’ (WGS 84), E. Jendek & O. Šauša leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 ♂ (EJCB):

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA “NE India, Meghalaya, 1400 m, Nokrek N.P., 3 km S Daribokgiri, 25°27’N, 90°19’E, 26.iv.1999, Dembický & Pacholátko leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 ♂ (EJCB): “NE India, Meghalaya, 1999, 9 km NW of Jowai, 1400 m, 25°30’N, 92°10’E, 11 15.v., Dembický & Pacholátko leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). Tamil Nadu: 1 ♂ (EJCB): “S India, Tamil Nadu, 17 22.V.1997, 15 km SE Kotagiri, 11°22’N, 76°56’E, Kunchappanai, Dembický & Pacholátko leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). Uttar Pradesh: 1 (EJCB): “India, Kumaon Himalaya, U.P., Nainital, Bhimtal, 20 23.5.1981, 1500 m leg. C. Holzschuh” (JENDEK & GREBENNIKOV, 2009 [#8563]). Uttarakhand: 1 ♀ (EJCB): “N India, Uttaranchal state, ca 13 km NW of Nainital, Khairna bridge env., 900 1000 m, 13 17.vii.2003, Z. Kejval & M. Trýzna leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). LAOS: Borikhamxai: 7 (EJCB): “Laos centr., Bolikhamsai prov., Ban Nape Kaew Nua Pass, 18.4. 1.5.1998, alt. 600±100 m, N 18º22.3, E 105º09.1 (GPS), E. Jendek & O. Šauša leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). Louang Namtha: 2 ♂ (EJCB): “Laos, Louang Namtha pr., 21°09’N, 101°19’E, Namtha Muang Sing, 5 31.v.1997, 900 1200 m, Vit Kubáň leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). Louangphrabang: 2 ♂ (EJCB): “Laos N, 23.iv.1999, Louangphrabang prov. 20°42’N 102°54’E, 25 km E Muang Ngoy, 1000 m, Vít Kubáň leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). Phongsali: 1 ♂ (EJCB): “Lao N, Phongsaly prov., 21°41’ 2’N, 102°06’ 08’E, 28.v. 20.vi.2003 Phongsaly env., ~1500m, Vít Kubáň leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). Vientiane: 1 ♀ (EJCB): “Laos centr., 27.IV. 1.V.1997, 70 km NE Vientiane, Ban Phabat env., 150 m, N 18°16.1, E 103°10.9, E. Jendek & O. Šauša leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). Xiangkhoang: 1 ♀ (EJCB): “Laos N, Xieng Khuang prov., 19º30’N, 104º03’E, Nong Haet, 29 31.v.2001, J. Bezděk leg.”. MYANMAR: Shan State: 1 ♂ (EJCB): “Burma (Myanmar), SW Shan State, Taunggyi, J. Rejsek 1 18.6.1997” (JENDEK & GREBENNIKOV, 2009 [#8563]). NEPAL: 1 (EJCB): “E Nepal, Dhankuta, Arun Valley, Lamobagar Gola, 27.5. 3.6.1980, 1000 1400 m, leg. C. Holzschuh” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 ♂ (EJCB): “W Nepal, NW Pokhara, Khali Gandaki Khola, 1100 1300 m, Tatopani, 12 14.V.1994, leg. C. Holzschuh” (JENDEK & GREBENNIKOV, 2009 [#8563]). SRI LANKA: 6 (EJCB): “SRI LANKA 300m, DAMBULLA env., 19.IV. 9.V.1991, leg. J. Kolibáč” (JENDEK & GREBENNIKOV, 2009 [#8563]). THAILAND: Chiang Mai: 1 ♂ (EJCB): “NW Thailand, 19 23.iv.1991, Doi Suthep to Doi Pui, J. Horák leg.”; 1 ♂, 1 ♀ (EJCB): “NW Thailand, 19 23.iv.1991, Doi Suthep, P.Pacholátko leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 ♂ (EJCB): “Thai, 17 24.V.1991, Chiang Dao, 1000 m, 19°25’N, 98°52’E, Vít Kubáň leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 2 ♂ (EJCB): “NW THAI, 7 14.V.1992, Chiang Mai, Doi Suthep Pui, P.Pacholátko leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 (EJCB): “THAI 15/5.1993, 19°12’N, 98°42’E, Mae Sae 750 m, Vít Kubáň leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 ♂ (EJCB): “Thailand 1000 1600 m, Chiang Mai prov., 20 km NW from Fang 2 5.v.1996, lgt. S. & E. Becvar” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 ♂ (EJCB): “NW Thailand 25.iv., Chiang Mai prov. 7.v., Ban San Pakia 1996, Sv. Bílý leg. 1700 m” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 (EJCB): “Thailand north, Chiang Mai env., 16.V.1997, M. Snížek leg.”; 1 ♂, 2 ♀ (EJCB): “Thailand, Chiang Mai prov., 19°19’N, 98°50’E, 1400 m, Sanpakia vill., 1 15.V.1998, Vít Kubáň leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). Kanchanaburi: 3 ♂ (EJCB): “THAI, 4 7.iv.1991, Erawan n.p., 300 m, 14°16’N, 99°12’E, Vít Kubáň leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 ♀ (EJCB): “Thai, 9 13.IV.1991, Thimonghta, 350 m, 15°02’N, 98°35’E, Vít Kubáň leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). Loei: 1 ♀ (EJCB): “Thailand NE, Loei prov., Phu Rua N. P., 17°30’N, 101°21’E, 6 9.v.1999, 1100m, D. Hauck leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). Mae Hong Son: 2 ♂, 3 ♀ (EJCB):

137 “NW Thailand, 9 16.5.1991, Mae Hong Son, Ban Huai Po, 1600 m, leg. Pacholátko” (JENDEK & GREBENNIKOV, 2009 [#8563]); 2 (EJCB): “NW Thailand, 9 16.5., Mae Hong Son, 1991, Ban Huai Po, 1600 2000 m, J. Horák leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 (EJCB): “NW Thailand, 19°19’N, 97°59’E, Mae Hong Son, 1991, Ban Huai Po, 1600 2000 m, 30.4 4.5., L. Dembický leg.”; 1 ♀ (EJCB): “NW Thailand, 1 6.5.1991, Soppong Pai, 1800 m, Leg. Pacholátko” (JENDEK & GREBENNIKOV, 2009 [#8563]); 2 (EJCB): “NW Thailand, 17 23.5., Mae Hong Son, 1991, Ban Huai Po, 1600 2000 m, J. Horák leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 ♂ (EJCB): “NW Thai, 30.iv. 4.v., Mae Hong Son, 1991, Ban Huai Po, 1600 2000, L. Dembický leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 2 (EJCB): “NW Thailand, 19°19’N, 97°59’E, Mae Hong Son, 1991, Ban Huai Po, 1600 2000 m, 9 16.5., L. Dembický leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 16 (EJCB): “NW Thai, 25.4. 5.5.1992, Soppong Pai pass, P. Pacholátko leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 ♀ (EJCB): “NW Thailand, 1 8.V., Mae Hong Son, 1992, Ban Si Lang, 1200 m, J. Horák leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 8 (EJCB): “NW Thailand, 8 18.V., Mae Hong Son, 1992, Ban Si Lang, 1200 m, J. Horák leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 ♂, 1 ♀ (EJCB): “Thailand 7 8.v.1992, Suan Pu, L. Dembický leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 ♀ (EJCB): “THAI, 1 5.V.1992, Ban Huai Po, L. Dembický leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 20 (EJCB): “NW Thailand 28.iv. 3.v.1992, Mae Hong Son, P. Pacholátko leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 4 ♂ (EJCB): “Thailand 7 8.v.1992, Nupa ah, L. Dembický leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 ♀ (EJCB): “Thailand, 1 8.5.1993, Soppong Pai, 1800 m, Pacholátko & Dembický leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 1 (EJCB): “Thailand 13.5.1993, 19°29’N, 98°18’E, Soppong, 750 m” (JENDEK & GREBENNIKOV, 2009 [#8563]); 5 (NMPC): “THAI 10 13/5.1993, 19°27’N, 98°20’E, SOPPONG 1550 m, Vít Kubáň leg.”; 9 (EJCB): “Thai 10 13/5.1993, 19°27’N, 98°20’E, SOPPONG 1550 m, Vít Kubáň leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 2 (EJCB): “Thailand, 7 12.5.1996, Mae Hong Son prov., SOPPONG, 1500 m 19°27’N, 98°20’E, S. Bečvář leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 5 (EJCB): “Thai, Mae Hong Son pr., 19°27’N, 98°20’E, 1500 m, Soppong 7 12.V., Vít Kubáň leg., 1996” (JENDEK & GREBENNIKOV, 2009 [#8563]). Nan: 1 ♂ (EJCB): “Thailand N, Nan prov., Bo Klua 700 m, 19°03’N, 101°10’E, 28.iv. 1.v.1999, D. Hauck leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). Tak: 1 ♀ (EJCB): “THAI, 28.IV. 6.V.1991, Umphang riv., 1000 m, 16°07’N, 99°00’E, Vít Kubáň leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 2 ♂ (EJCB): “Thai, 18 24.IV.1991, Lansang n. p. 500 m, 16°48’N, 98°57’E, Vít Kubáň leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]). VIETNAM: 2 ♂ 1 ♀ (EJCB): “S Vietnam, 14.10N 108.30E, 40 km NW Ankhe, Buon Luoi, 620 750 m, 28.3. 12.4.1995, Pacholátko & Dembický leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]); 9 (EJCB): “S Vietnam, 11.54N 108.27E, 10 km S of Dalat, 1500 m, 16.IV.1995 Pacholátko & Dembický leg.” (JENDEK & GREBENNIKOV, 2009 [#8563]).

Adult occurence: 2-3-4-5-6-7 (n=66). Faunal records published by BAUDON, 1960 [#157] and TÔYAMA, 1987 [#2931] (as komiyai) are also included. Altitude range: 1502528 m (n=44). Host plants: Indigofera galegoides: BAUDON, 1968 [#162]. Distribution: CHINA: Shaanxi, Sichuan, Yunnan. INDIA. LAOS. MYANMAR. NEPAL. SRI LANKA. THAILAND. VIETNAM.

138

Agrilus myopic Jendek Fig. 22C (habitus of HT); Fig. 37L (aedeagus) myopic Jendek, 2000 (Agrilus) JENDEK, 2000: 150 [#4853] (description). JENDEK, 2006: 400 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2197 [#8576] (world catalog).

Type material: Agrilus myopic Jendek, 2000 Type locality: China-Sichuan: Paß zw.[ischen] Zhangla u.[nd] Huanglong, 4000 m. Type specimens: Primary: Holotype (Fig. 22C), ♂, (EJCB): “China – Sichuan: Paß zw[ischen]. Zhangla [32°48’N, 103°40’E] u[nd]. Huanglong, 4000 m, 27. 6. [19] 96, Erber [leg.]”. Secondary: 6 PT (EJCB). Described from 7 specimens (HT, PT). Diagnosis: 5.4-6.0 mm; pronotum flat; prehumerus short, carinal or tubercular; elytra almost glabrous, with very sparse, whitish, obsolete, perisutural pubescence; elytral apices markedly broad. Species group: Hyperici. Relevant species: A. hyperici (Creutzer, 1799). Specimens examined: Known only from the type specimens.

Host plants: Unknown. Distribution: CHINA: Sichuan.

Agrilus naxi Jendek, sp. nov. Fig. 27K (habitus of HT); Fig. 40D (aedeagus)

Description: 3.6-5.7 mm (Holotype 4.6 mm). BODY: Shape: subparallel, Color (dorsally): unicolored, Color (dominant): green or brown. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture on vertex: punctures, Sculpture (aspect): irregular, Sculpture (density): sparse. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: in line with antennal socket. Antennae: Length: moderate, Shape: slender, Sexual modification in male: antennae longer. PRONOTUM: Shape: visually square, Sides: straight or arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate or obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Lateral impressions (deepness): shallow. Prehumerus: carinal, Shape: bisinuate, Extent: to anterior pronotal angle, Posterior end: joining posterior pronotal angle, Anterior end: joining anterior pronotal angle, Arc: obvious. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: absent or present, Narrowest point: at posterior angles. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color:

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA unicolored with darker hue in apical half, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate, Width: wide, Shape: arcuate. Elytral pubescence: entire except glabrous part at apical half, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate or large, Anterior margin: arcuately emarginate, Emargination (deepness): shallow or moderate. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: acute or rectangular, Angles (tips): sharp, Disk: impressed. ABDOMEN: Sexual modifications in male: pair of medial tubercles delimited with sulcus on ventrite 1, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Male sexual modifications: metafemur markedly incrassate. Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe markedly obtuse, Ovipositor: elongate. Diagnosis: prosternum longly, erectly pubescent; prosternal process dilated between coxae; basal ventrite in male with medial impression. Species group: Cyaneovirens. Relevant species: A. coloratus Kerremans, 1894. Type locality: China, Yunnan province, 50 km north of Lijiang, Daju, 27.21N, 100.19E. Type specimens: Primary: Holotype (Fig. 27K), ♂, (EJCB). Secondary: CHINA: Yunnan: 41 PT (EJCB): “China, Yunnan, 27-28.VI., Daju (50 km N of Lijiang), 27.21N, 100.19E, E. Jendek, 1992”; 5 PT (EJCB): “China-Yunnan, 24-29.6.[19]94, Daju, lgt. E. Kučera”; 2 PT (EJCB): “China, Yunnan, 25-27.VI.1994, Daju, B. Šiška & T. Spevár leg.”; 27 PT (EJCB): “China, Yunnan, Daju, 17.VI.1995, Šiška & Spevár & Pekarovič”; 11 PT (EJCB): “China, Yunnan prov., 16-17.VII.1996, Daju, 27.15N, 100.15E, B. Šiška leg.”. Adult occurence: 6-7 (n=5). Host plants: Unknown. Distribution: CHINA: Yunnan. Etymology: Noun in apposition; the specific name refers to the ethnic group Naxi living in Yunnan and Sichuan provinces of China.

Agrilus nelsonodaimio Ohmomo Fig. 11I (habitus of PT); Fig. 33I (aedeagus) nelsonodaimio Ohmomo, 2006 (Agrilus) OHMOMO, 2006: 245 247 [#7546] (description). 2199 [#8576] (world catalog).

BELLAMY, 2008:

Type material: Agrilus nelsonodaimio Ohmomo, 2006 Type locality: Mt. Akaguna-yama, Fujioka, Gunma Pref. Type specimens: Primary: Holotype, ♂, (NSMT). Secondary: 1 PT (EJCB). Described from 31 specimens (HT, PT). Paratype (Fig. 11I) from EJCB is illustrated.

139

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Diagnosis: 5.3-6.35 mm; pronotum widest in middle; aedeagus expanded apically. Species group: Maculifer. Relevant species: A. shashamboe Kurosawa, 1963. Specimens examined: KOREA SOUTH: 1 (EJCB): “Korea: Kyenggi prov., Yangpyangun, Yongmusan, 1 13.VII.1998, K L. You, H J. Lim, ex FIT”.

Adult occurence: 7 (n=1). Host plants: Rhododendron: OHMOMO, 2006 [#7546] Distribution: JAPAN: Honshu, Kyushu, Ryukyu Islands, Shikoku, Tsushima. KOREA NORTH.

Agrilus nepalensis Tôyama Fig. 12K (habitus); Fig. 33T (aedeagus); Fig. 49U (type).

nepalensis Tôyama, 1988 (Agrilus) TÔYAMA, 1988: 759 761 [#2932] (description). JENDEK, 2006: 400 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2200 [#8576] (world catalog).

Type material: Agrilus nepalensis Tôyama, 1988 Type locality: Nagarjun R. F., C. Nepal. Type specimens: Holotype by monotypy (Fig. 49U), ♂, (NSMT): “Nagarjun R. For. C. [h] Nepal [p] 29. IV. [h] 19 [p] 83 T. Shimomura [leg.] [h] \ Holotype [p] Agrilus nepalensis Tôyama, 1987 [h] [red label]”. Described from 1 specimen (HT). Diagnosis: 8-13 mm; elytra silky black, with narrow perisutural pubescent stripe not reaching sutural margin; prosternum with bunch of setae in middle and row of erect pubescence on prosternal process. Species group: Imitans. Relevant species: A. blatteicollis Bourgoin, 1922; A. sichuanus sp. nov. Specimens examined: INDIA: Arunachal Pradesh: 1 ♂, 1 ♀ (EJCB): “India, W Arunachal Pr., Dirang vicinity, 1500 1800m, 27°21’ 23’N 92°13’ 16’E, L. Dembický leg.; 1. 10.vi.2004”. Assam: 2 (EJCB): “I Assam, 18 25.5., Haflong Hills, St. Jákl lgt., 1991”; 20 (EJCB): “NE India, Assam, 1999, 5 km N of Umrongso, 700m, 25°27’N, 92°43’E, 17. 25.v., Dembický & Pacholátko leg.”; 3 (EJCB): “NE India, Assam, 1999, 5 km N of Umrongso 700m, 25°27’N, 92°43’E, 17. 25.v., J. Rolčík leg.”. West Bengal: 1 ♂ (EJCB): “Pedong, 27 30.iv.1987, Indien, Darjeeling D., Bhakta B.”. LAOS: 1 (EJCB): “Houei Khinin, 1500m, vi.1968, Laos (Baudon)”. Phongsali: 2 ♀ (EJCB): “Lao N, Phongsaly prov., 21°41’ 2’N, 102°06’ 08’E, Phongsaly env., 28.v. 20.vi.2003, 1500m, P. Pacholátko leg.”. NEPAL: 1 (EJCB): “Lamobagar Gola, 1400m, 8 14.vi.1983, E Nepal, Arun V., M. Brancucci”.

Adult occurence: 4-5-6 (n=8). Altitude range: 700-1800 m (n=6). Host plants: Unknown. Distribution: BHUTAN. INDIA. LAOS. NEPAL. Remarks: Not recorded from East Asia but possible in southwestern China.

Agrilus nicolanus Obenberger Fig. 17K (habitus); Fig. 35O (aedeagus); Figs. 49V, 50T (types) nicolanus Obenberger, 1924 (Agrilus; subspecies of foveicollis) OBENBERGER, 1924: 42 [#2040] (description). OBENBERGER, 1926: 656 [#2057] (subspecies of foveicollis; Palaearctic catalog). MIWA & CHÛJÔ, 1936: 20 [#1809] ([Note: Misidentified as ronino according to Jendek, 1994: 13 [#1355]]; catalog; Japan). OBENBERGER, 1936: 990 [#2191] (subspecies of foveicollis; world catalog). KUROSAWA, 1963: 153 [#1618] ([Note: Misidentified as ronino according to JENDEK, 1994: 13 [#1355]]; characters; Japan). KUROSAWA, 1974: 4 [#4577] ([Note: Misidentified as ronino according to JENDEK, 1994: 13 [#1355]]). TÔYAMA, 1985: 30 [#3678] ([Note: Misidentified as ronino according to JENDEK, 1994: 13 [#1355]]; iconography; Japan). JENDEK, 1994: 13 14 [#1355] (lectotype designation; synonymy; characters; Altay; Russian Far East; Japan). AKIYAMA & OHMOMO, 1997: 36 [#4091] (checklist; Japan: Honshu, Shikoku, Kyushu; Korea; Russia: Ussuri). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). AKIYAMA & OHMOMO, 2000: 88 [#6240] (faunal records; Japan (Hiroshima)). JENDEK, 2006: 393 [#7441] (subgenus Robertius; Palaearctic catalog). BELLAMY, 2008: 2201 [#8576] (subgenus Robertius; cited as nikolanus; world catalog). VOLKOVITSH, 2009: 136 [#8850] (subgenus Robertius; faunal records; Russia: Primorskiy kray). piscatorius Obenberger, 1935 (Agrilus) OBENBERGER, 1935: 169 170 [#2158] (description). OBENBERGER, 1936: 1022 [#2191] (world catalog). JENDEK, 1994: 13 [#1355] (synonym of nicolanus; lectotype designation). AKIYAMA & OHMOMO, 1997: 36 [#4091] (synonym of nicolanus). JENDEK, 2006: 393 [#7441] (synonym of nicolanus; Palaearctic catalog). BELLAMY, 2008: 2201 [#8576] (synonym of nicolanus cited as nikolanus; world catalog).

Type material: Agrilus foveicollis nicolanus Obenberger, 1924 Type locality: Ussuri (Nikolsk Ussurijsk). Type specimens: Primary: Lectotype (Fig. 49V), ♂, designated by JENDEK (1994: 13-14) [#1355], (NMPC): “Nikolsk - Ussuri [h] \ Typus [p] [red label] \ foveicollis ssp. nicolanus m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24267 [h] [orange label]”. Secondary: 2 PLT (NMPC). Described from unknown number of syntypes. Agrilus piscatorius Obenberger, 1935 Type locality: Ussuri mer.: Kamen Rybolov, lacus Chanka. Type specimens: Primary: Lectotype (Fig. 50T), ♀, designated by JENDEK (1994: 13) [#1355], (NMPC): “KamenRybolov oz. Chanka, Juzhnouss. A. Tscherskij [p] 30.V. [h] [19] 08 [p] [in Russian] \ Typus [p] [red label] \ Mus. Nat. Pragae Inv. [p] 24269 [h] [orange label] \ Agrilus piscatorius m. Type [h] Det. Dr Obenberger [p]”. Secondary: 1 PLT (ZIN). Described from unknown number of syntypes. Diagnosis: 3.7-5.5 mm; dorsally dark-brown or darkgreen; sides of pronotum almost subparallel; elytral apices diagonally subtruncate. Species group: Betuleti. Relevant species: A. delphinensis Abeille de Perrin, 1897. Specimens examined: CHINA: Beijing: 1 ♂ (EJCB): “China, 12 15 June 2000, Dongling Shan, 1500 m, 100 km W of Beijing, Zd. Jindra lgt.”; 2 ♂, 1 ♀ (EJCB): “China, Beijing distr., Yan Shan, Dongling Mts., 39º58’N, 115º26’E, 1400 m, Xiaolongmen, 15 16.VI.2001”. Hebei: 1 ♂ (EJCB): “China, Hebei (Nei Mongol) pass Chengde Chifeng 41.6N, 118.2E, 30 31.V., J.Turna leg. 2002”; 6 (EJCB): “China, Hebei/Nei Mongol, pass Chengde Chifeng, 41.6N, 118.2E,

140 14 16.VI., Jaroslav Turna leg. 2001”; 9 (EJCB): “China, Hebei/Nei Mongol, pass Chengde Chifeng, 41.6N, 118.2E, 14 16.VI.2001, O. Šafránek leg.”. Hubei: 1 ♂ (EJCB): “China, W Hubei prov., Shennongjia Co, Yannzi Pass, 31º43’/110º28’, 2200m, 23 26.6.1995, L.+R. Businský lgt.”. JAPAN: Honshu: 1 ♂ (EJCB): “Japan, Alp’s de Sasaggo prés Kôfu, 27.7.[19]08” (JENDEK, 1994 [#1355]); 1 ♂ (EJCB): “Japan, Kumanotaira, pr. Kuruizawa, 12.7.[19]08” (JENDEK, 1994 [#1355]); 1 (EJCB): “Tobira spa., Nagano Pref. 19.VI.1974 H. Hayakawa” (JENDEK, 1994 [#1355]); 1 (EJCB): “Tobira spa., Nagano Pref. 17.VII.1974 H. Hayakawa” (JENDEK, 1994 [#1355]); 2 ♂, 1 ♀ (EJCB): “Mts. Daibosatsu Yamanashi Pref. 21.VI.1985 S.Ohmomo” (JENDEK, 1994 [#1355]). Kyushu: 1 ♀ (SOCI): “2005.5.9, Kusui, Kokonoe, Oita, Kyushu, S Jpn, Y. Tsutsumiuchi leg.”. POLAND: Bialystok: 1 ♂ (RKCK): “Poland FD94, BPN, odd. 398 399, distr. Bialystok, 01.06.1998 r., coll. Jacek Kalisiak”. RUSSIA: Kemerovskaya oblast’: 1 (NMPC): “Altai m. Kuzneck” (JENDEK, 1994 [#1355]). Primorskiy kray: 1 (NMPC): “Kamen Rybolov, oz Chanka, Yuzhnouss, Tscherskij 18.VI.08” (JENDEK, 1994 [#1355]); 1 (ZIN): “Primorsk.obl., 31.V.1915, Sedanka, Rimskij Korsak leg” (JENDEK, 1994 [#1355]); 1 ♂ (ZIN): “Ussur., Grigorievka, 12.VII.1937 Stepanov leg.” (JENDEK, 1994 [#1355]); 4 ♂ (ZIN): “Primorskij kraj, Krasnoarmijsk, 18.VI.1951, V. N. Stepanov leg.” (JENDEK, 1994 [#1355]); 1 (ZIN): “Yu Primor’e Sr. Ter. r. Suchan 10 20.vii.1952 Kabakov [in Russian]”; 1 (ZIN): “Yu Primor’e Uglovoe 5.vi.1958 Solyanikov [in Russian]”; 5 (EJCB): “Sib.or. m., Primorje Sichote Alin Mts. Sokolči 1. 15.7.1990 Kadlec + Voříšek lg.” (JENDEK, 1994 [#1355]); 1 ♂, 1 ♀ (ZIN): “Primorskij kraj Spassk forest 13 15.VI.1990 Belokobylskij leg. [in Russian]” (JENDEK, 1994 [#1355]); 1 (EJCB): “Primorie, Novickoe 15km S of Partizansk forest 24.VI.1990 Belokobylskij leg.” (JENDEK, 1994 [#1355]); 2 ♀ (EJCB): “USSR, Primorsk. kraj, Arsenev env., VI.1991, 44.11N, 133.19E, O. Šausa leg.” (JENDEK, 1994 [#1355]); 1 ♀ (EJCB): “USSR, Primorskij kraj Arseneev env., VI.1991, 44.11N, 133.19E, M. Štrba leg.” (JENDEK, 1994 [#1355]); 1 (EJCB): “Prim. Chasan Rjazanovka, 5.6.1991, Filimonov [in Russian]” (JENDEK, 1994 [#1355]); 1 ♀ (EJCB): “Sib. or., Primorje, Komarovka flum., Kamenushka env., 300 m, Vořišek lgt., VI.1992” (JENDEK, 1994 [#1355]); 1 ♀ (EJCB): “Rossia, Primorskij k., Rjazanovka 4 8. vii.1993, Z. Kletečka leg.”; 1 ♀ (JMCZ): “Russia Far East, 19 20. VII.1993, Primorsk region, Lazo, lgt. M. Trýzna”; 1 ♂ (EJCB): “Russia, Primorski kray, Khankajski distr. 1995, lake Khanka, 17 19.vii., Kamen’ Rybolov”; 1 (FACF): “Russia Far East, S Primorie, Hanka lake, 30.vii.2002”; 1 ♀ (EJCB): “Russia, Primorski reg., Khankayski distr., Novokachalinsk, 04.07.2003, Filimonov leg.”; 2 ♂ (EJCB): “Russia Far East, 20 km SE Ussuriysk, Gornotaezhnoje vill. env., 28.06.2003, Lg. A. Napolov”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Preobrazhenie env. 14.07.04 leg. М.& L. Smirnov’s”.

Adult occurence: 5-6-7 (n=37). Altitude range: 300-2200 m (n=4). Host plants: Unknown. Distribution: CHINA: Beijing, Hebei, Hubei, Nei Mongol. JAPAN: Honshu, Kyushu, Shikoku. KOREA NORTH. KOREA SOUTH. POLAND: Bialystok. RUSSIA: Kemerovskaya oblast’, Primorskiy kray.

Agrilus ningpoensis Obenberger Fig. 15C (habitus of LT) ningpoensis Obenberger, 1927 (Agrilus) OBENBERGER, 1927: 85 [#2059] (description). OBENBERGER, 1936: 1016 [#2191] (world catalog). HUA LI ZHONG, 2002: 90

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA [#6696] (checklist; China: Zhejiang). JENDEK, 2006: 400 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2203 [#8576] (world catalog).

Type material: Agrilus ningpoensis Obenberger, 1927 Type locality: China: Ning-Po. Type specimens: Primary: Lectotype (Fig. 15C) by present designation, ♀, (NMPC): “China: Ning Po [h] \ Typus [p] [red label] \ Agrilus ningpoënsis m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24945 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 6.4 mm; very similar to A. caligans from which it differs mainly by deeper medial impression on vertex. Species group: Subrobustus. Relevant species: A. caligans Bourgoin, 1925. Specimens examined: Known only from the lectotype.

Host plants: Unknown. Distribution: CHINA: Zhejiang.

Agrilus nipponigena Obenberger Fig. 11G (habitus); Fig. 33G (aedeagus); Figs. 49E. 49X (types) nipponigena Obenberger, 1935 (Agrilus) OBENBERGER, 1935: 167 168 [#2158] (description). LEWIS, 1879: 15 [#3345] ([Note: Misidentified as moerens according to JENDEK, 1995:138 [#3768]]; catalog; Japan). MIWA & CHÛJÔ, 1936: 19 [#1809] (cited as nipponigera; catalog; Japan). OBENBERGER, 1936: 1016 [#2191] (world catalog). CHÛJÔ & KUROSAWA, 1950: 10 [#501] (catalog; faunal records; Japan (Shikoku)). IGA, 1955: 79 [#3828] (iconography; Japan). IGA, 1962: 79 [#3351] (iconography; Japan). KUROSAWA, 1963: 154 [#1618] (characters; Japan). KUROSAWA, 1974: 3 [#4577] (synonym of moerens; notes). PENG ZHONGLIANG, 1987: 356 [#2429] (cited as nigponigena; checklist; China). LI JINGKE, 1992: 92 [#6755] (checklist; China: Liaoning). JENDEK, 1995: 138 [#3768] (lectotype designation; synonymy; characters; Japan; China: Shaanxi). AKIYAMA & OHMOMO, 1997: 36 [#4091] (checklist; Japan: Honshu, Shikoku, Kyushu, Tsushima; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Liaoning, Hebei, Shandong, Henan, Zhejiang). JENDEK, 2006: 400 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2203 [#8576] (world catalog). melanolucidus Ohmomo, 2005 (Agrilus) syn. nov. OHMOMO, 2005: 236 238 [#7253] (description). BELLAMY, 2008: 2183 [#8576] (world catalog).

Type material: Agrilus nipponigena Obenberger, 1935 Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 49X), ♀, designated by JENDEK (1995: 138) [#3768], (NMPC): “Japan [h] \ Typus [p] [red label] \ Mus. Nat. Pragae Inv. [p] 24292 [h] [orange label] \ Agrilus nipponigena m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Agrilus melanolucidus Ohmomo, 2005 Type locality: Nobeyama-kogen, Nagano Pref. Type specimens: Primary: Holotype (NSMT). Secondary: 4 PT (SOCI), 1 PT (EJCB). Described from 6 specimens (HT, PT). Paratype (Fig. 49E) from EJCB is illustrated.

141

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Diagnosis: 5.0-6.1 mm; Body cuneiform with distinctly attenuate apical third of elytra; pronotal disk with very fine sculpture. Species group: Quadrisignatus. Relevant species: A. ornatoides sp. nov. Specimens examined: CHINA: Shaanxi: 4 ♂, 2 ♀ (EJCB): “China, Shaanxi province, Hua Shan Mt., 17 21.VI.1991, leg. R. Sauer” (JENDEK, 1995 [#3768]); 1 (EJCB): “China Shaanxi, Lueang env., 15 km NW, 26.5. 31.5.2001, lgt. E. Kučera”; 1 (EKCS): “China Shaanxi, Lueyang, 23 30.5.2009, lgt. E. Kučera”. JAPAN: Honshu: 1 (EJCB): “Japan, Todoromi, Osaka Prefecture, 9.VI.1968, leg. M. Yagi” (JENDEK, 1995 [#3768]); 1 (MNCA): “Japan, Hattoji, Okayma Prefecture, 28.V. 1979, leg. H. Jamazaki” (JENDEK, 1995 [#3768]); 2 ♀ (EJCB): “Japan, Nishinasuno, Tochigi Prefecture, 15.VI.1991, leg. S. Ohmomo” (JENDEK, 1995 [#3768]); 1 ♀ (EJCB): “Japan, Honshu, Osaka fu, Kongosan, 300 m, 2 km E of Amami, 12. May 2002, Bolm Leg.”.

Adult occurence: 5-6 (n=7). Altitude range: ~300 m (n=1). Host plants: Diospyros kaki AKIYAMA & OHMOMO, 1997 [#4091]. − D. lotus: AKIYAMA & OHMOMO, 1997 [#4091]. Erroneous: Salix?: OHMOMO, 2005 (as melanolucidus) [#7253]. – Salix matsudana: HUA LI ZHONG, 2002 [#6696]. Distribution: CHINA: Hebei, Henan, Liaoning, Shaanxi, Shandong, Zhejiang. JAPAN: Honshu, Kyushu, Shikoku, Tsushima. Remarks: Kurosawa (1974: 3 [#4577]) due to the misinterpretation of A. moerens, considered A. nipponigena to be A. moerens. All specimens of A. nipponigena examined from Japan were misidentified as A. moerens while “true” A. moerens was misdetermined as A. rotundicollis Saunders, 1873. This confusions might be the cause that Ohmomo, 2005 [#7253] had overlooked A. nipponigena and described this species again as A. melanolucidus. For the same reason, the host plants cited by AKIYAMA & OHMOMO, (1997 [#4091]) need reconfirmation.

attenuate; elytral apices sharply spinose. Species group: Niveoguttatus. Relevant species: A. satoi Kurosawa, 1954 Specimens examined: CHINA: Yunnan: 1 (EJCB): “Yunnan, 2000 2800m, 25.11N, 100.24E, Weibaoshan mts., W slope, 25 28/6.1992, Vít Kubáň”; 1 ♀ (EJCB): “CH, Yunnan, vii.1993, Weishan (County), Weibaoshan, 2600 2900 m”; 1 ♀ (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”. INDIA: Arunachal Pradesh: 1 ♂ (EJCB): “India, W Arunachal Pr., betw. Dirang & Bomdila Pass, 27°19’N, 92°22’E; 1900±300m L. Dembický leg. 12. 16.vi.2004”. Uttarakhand: 1 ♂, 1 ♀ (EJCB): “N India, Uttaranchal state, ca 30 km N of Bageshwar, SE of Dhakuri vill., 2600 2800, 25 26.vi.2003, Z. Kejval & M. Trýzna lgt.”. West Bengal: 1 (EJCB): “Chivo Busty, 9.iv.1985, 1100m, India, Darjeeling, Bhakta, B.”. LAOS: Houaphan: 1 ♀ (EJCB): “NE Laos, Hua Phan prov., Ban Saluei, 1300 2000m, 20°13’N, 103°59’E, 6 18.iv., Phu Phan Mt. env. 2004, F. & L. Kantner leg.”; 3 (EJCB): “Lao NE, Hua Phan pr., ~20°12’N,104°01’E, Phu Phan Mt., 1500 1900m, 17.v. 3.vi.2007, Vít Kubáň leg.”; 1 ♀ (EJCB): “Laos NE, Houa Phan prov., 20°12 13.5’N, 103°59’.5 104°01’E, Ban Saluei → Phou Pane Mt., 1340 1870m, 15.iv. 15.v.2008, Lao collectors leg.”; 6 (EJCB): “Laos NE, Houa Phan prov., 20°12 13.5’N, 103°59’.5 104°01’E, Ban Saluei → Phou Pane Mt., 1340 1870m, 15.iv. 15.v.2008, Lao collectors leg.”. Phongsali: 1 ♂ (EJCB): “Lao N, Phongsaly prov., 21°41’ 2’N, 102°06’ 08’E, Phongsaly env., 28.v. 20.vi.2003, 1500m, P. Pacholátko leg.”. NEPAL: 1 ♂ (MNCA): “Nepal, prov. Karnali, Distr. Humla, 5 km SE Simikot, NE Chipra Chuwa Khola valley, 2200 m, NN, 09.VII.2001, riverside, 29°56’33’’N, 81°51’24’’E, HF, leg. M. Hartmann”. 1 (EJCB): “E Nepal, Koshi, Basantapur, 2300 m, 30.V. 2.VI.1985, leg. C. Holzschuh”. 2 (EJCB): “Nepal, Kathmandu, Jamajok, 18 19.v.1993, 1800m, Schimmel”; 1 ♀ (EJCB): “E Nepal, Dhankuta, Arun Valley, 19.6.1983, Chichila, 2000 m, leg. C. Holzschuh”. THAILAND: Chiang Mai: 1 ♀ (EJCB): “Thai, Chiang Mai prov. 18°49’N, 98°54’E, 1600 m, Doi Pui mt., 2 6.V.1996, Vít Kubáň leg.”. Mae Hong Son: 1 ♂ (EJCB): “Thai, Mae Hong Son pr., 19°27’N, 98°20’E, 1500 m, Soppong 7 12.V., Vít Kubáň leg., 1996”.

Adult occurence: 4-5-6-7 (n=17). Altitude range: 11002900 m (n=16). Host plants: Unknown.

Agrilus niveoguttatus Kerremans Fig. 10K (habitus); Fig. 32Z (aedeagus); Fig. 49Y (type) niveoguttatus Kerremans, 1892 (Agrilus) KERREMANS, 1892: 210 211 [#1402] (description). KERREMANS, 1903: 288 [#1441] (catalog). OBENBERGER, 1936: 1093 [#2191] (world catalog). PENG ZHONGLIANG, 1992: 399 400 [#2431] (characters; notes; Hunan). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Jiangxi, Hunan). JENDEK, 2006: 400 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2203 [#8576] (world catalog).

Type material: Agrilus niveoguttatus Kerremans, 1892 Type locality: Inde. Type specimens: Holotype by monotypy (Fig. 49Y), ♂, (MNHN): “niveoguttatus (Laferté) [h] Inde \ Ex-musaeo Mniszech [p] \ niveoguttatus Kerr. Type [h] \ Kerremans vidit 1892 [p] \ collectio Oberthür [p]”. Described from 1 specimen. Diagnosis: 6.6-10.8 mm; apical third of elytra markedly

Distribution: CHINA: Yunnan. INDIA: Arunachal Pradesh, Uttarakhand, West Bengal. LAOS: Houaphan, Phongsali. NEPAL. THAILAND: Chiang Mai, Mae Hong Son. Remarks: Distributional records cited without supporting data from Hunan and Jiangxi (China) by PENG ZHONGLIANG, 1992 [#2431] and by HUA LI ZHONG, 2002 [#6696] were omitted.

Agrilus nivosus Abeille de Perrin Fig. 21C (habitus); Fig. 36Y (aedeagus); Figs. 44B, 48C, 50A (types) nivosus Abeille de Perrin, 1900 (Agrilus) ABEILLE DE PERRIN, 1900: 13 [#21] (description). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 660 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1016 [#2191] (world catalog). TER MINASYAN, 1955: 452 [#7893] ([Note: Misidentification of pistaciophagus]; host plants; distribution; references). GURYEVA, 1974: 99 [#5428] ([Note: Probably misidentification according to host plant cited]; pest of agriculture

142 cultures; references; Uzbekistan). JENDEK, 2003: 183 [#5823] (lectotype designation). JENDEK, 2006: 400 [#7441] (Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2204 [#8576] (world catalog). kuldjensis Obenberger, 1936 (Agrilus) syn. nov. OBENBERGER, OBENBERGER, 1936: 111 [#2205] (description). 1936: 1004 [#2191] (world catalog). COBOS, 1972: 56 [#591] (subgenus Agrilus; checklist; Mongolia). ALEXEEV, 1975: 156 [#52] (subspecies of sericans; notes; faunal records; distributional overview; Mongolia). PENG ZHONGLIANG, 1987: 356 [#2429] (cited as kurdjensis; checklist; China). ALEXEEV, 1989: 481 [#64] (subspecies of sericans; key to species; northwestern China; Mongolia). ALEXEEV & VOLKOVITSH, 1989: 345, 356 [#3241] (subspecies of sericans; characters in key; Mongolia). OVTCHINNIKOV, 1996: 132 [#8621] (subspecies of sericans; checklist; Kyrgyzstan). NONNAIZAB et al., 1999: 113 [#8247] (checklist; China: Inner Mongolia). HUA LI ZHONG, 2002: 90 [#6696] (cited as kurdjensis; checklist; China: Inner Mongolia). JENDEK, 2006: 396 [#7441] (subgenus Xeragrilus; subspecies of sericans; Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (subspecies of sericans; complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2288 [#8576] (subspecies of sericans; world catalog). candidiventris Obenberger, 1936 (Agrilus) syn. nov. OBENBERGER, OBENBERGER, 1936: 113 [#2205] (description). 1936: 969 [#2191] (world catalog). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2013 [#8576] (world catalog).

Type material: Agrilus nivosus Abeille de Perrin, 1900 Type locality: Turkestan: Aoulié-Ata. Type specimens: Primary: Lectotype (Fig. 50A), ♀, designated by JENDEK (2003: 183) [#5823], (ZSMC): “Type ! [h] \ Aulie Ata [h] \ Turkestan (Stdgr. 1901) [h] \ nivosus [h] \ Sammlung Dr. K. Daniel [p] \ Holotypus [p] Agrilus nivosus Abeille [h] Zool. Staatssammlg. München [p] [red label] \ Zool. Staatsslg. München [p] [blue label]”. Secondary: 5 PLT (MNHN). Described from unknown number of syntypes. Agrilus kuldjensis Obenberger, 1936 Type locality: Kuldja. Type specimens: Primary: Lectotype (Fig. 48C) by present designation, ♀, (NMPC): “Kuldja [h] \ Typus [p] [red label] \ Agrilus kuldjensis m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24953 [h] [orange label]”. Described from unknown number of syntypes. Agrilus candidiventris Obenberger, 1936 Type locality: Deserta Mujun-Kum (Asia centr.). Type specimens: Primary: Lectotype (Fig. 44B) by present designation, ♀, (NMPC): “Mujun Kum, Asia centr. [h] \ Typus [p] [red label] \ Agrilus candidiventris m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24990 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 5-7.3 mm; very similar to A. sericans from which it differs by larger size; body more elongate and more pubescent; elytra often with perisutural strips of denser pubescence; lateral pronotal impressions and humeral part of elytra subtomentose; pronotum widest in anterior third;

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA posterior pronotal angles sharp. Species group: Albogularis. Relevant species: A. sericans Kiesenwetter, 1857, A. albogularis Gory, 1841. Specimens examined: CHINA: Xinjiang: 1 ♀ (JMCZ): “China occ., Jining, Boro Horo Shan, 26 31.VII.1991, 44º06ŠS, 81º56’VD, Ining H Sien, Snížek”. KAZAKHSTAN: 1 (EJCB): “Aulie Ata”; 2 (EJCB): “Aulie Ata, Syr Darya Gebiet”; 3 ♂ (EJCB): “USSR, Kazakhstan, Aksaj, 30 5.1974 (Alma Ata env.), Kr. Pospíšil lgt.”; 1 (ZIN): “g. Toraigyr, 28 km YuV Kokpeka, Kazakhstan, Volkovich, 1.vii.1987 [in Russian]”; 2 (ZIN): “YuV Kazakhstan, KZOKh Myndulak, khr. Malyi Kalkan, 19.v.1990, Prasolov [in Russian]”; 2 (ZIN): “YuV Kazakhstan, KZOKh Myndulak, 20.v.1990, Prasolov [in Russian]”; 25 (EJCB): “Kazakhstan m., Mikhaylovka, Djambul, 16 17.vii.1991, S. Bečvář leg.”; 1 ♂, 2 ♀ (EJCB): “SU Kazakhstan, 1991, 15 km NE Jambul, Moiynkum Desert, Mikhailovka, 16.17.7., Jaroslav Turna leg.”; 3 (EJCB): “SU Kazakhstan, 1991, 50 km NE Jambul, Moiynkum Desert, Kospan, 18 20.vii., Jaroslav Turna leg.”; 1 ♂ (EJCB): “Kazakhstan cent., Lake, 50 km E Balkash, 26 27.6.1992, leg. J. Halada”; 1 ♂ (EJCB): “W Kazkhstan, Aktyubinsk, Mugodzhany, Jurinllinoe vill., 14.6.1995, 6.6.1995, leg. Dolin”; 4 (ZIN): ““Yasenevaya Dacha”, r. Charyn, 10 km SZ Chundzhi, Kazakhstan, Volkovich, 22 23.06.1998 [in Russian]”; 3 (EJCB): “Kazakhstan, Uralsk reg., 10 m, Chapaev, 12.6.1999, M. Danilevskiy leg.”. KYRGYZSTAN: 6 (EJCB): “SSSR Kirgizia or., Čon Aryk (prope Frunze), 2.7.1980, Ing. K. Schön lgt.”; 9 (EJCB): “USSR, Kirgizia, Frunze env., Tšon Aryk, 1.7.1980, S. Kadlec lgt.”; 1 (ZIN): “5 km YuV Kulanaka, khr. Karatau, Naryn, Volkovich 19.06.1989 [in Russian]”; 1 (EJCB): “Kirg., Issyk Kul. reg., Barskau, 10 17.7.1991, I. Jeniš leg.”; 1 ♂ (ZIN): “Kirgiz., Sosnovka, 12.7.1991, Napolov [in Russian]”; 1 ♂ (EJCB): “Kyrgyzstan, Issyk Kul, S. Ufer, env. Kadzhidaj, 16.07.1998, DW &RA”; 1 ♂ (EJCB): “Kirgizstan, S slope, Turkestan range, Sokh riv., upper, 3500 m, 5 9.7.1998, Petrov leg.”; 1 ♀ (EJCB): “Kirgizstan, 10 km S Bishkek, 20.6.1999, Klimenko leg.”. MONGOLIA: Hovd: 1 ♂ (NMPC): “Mongolia, Khovd ajmak, Bajtag Bogd Nuru, 25.vii.1975”. UZBEKISTAN: 1 ♀ (EJCB): “USSR, Uzbekistan, Taškent env., Galvasai, 25.6.1980, S. Kadlec lgt.”; 4 (EJCB): “Uzbekistan, Iskander, 28.6. Kadlec lgt., 1980”.

Adult occurence: 5-6-7 (n=23). Altitude range: 10-3500 m (n=2). Host plants: Artemisia: ALEXEEV & VOLKOVITSH, 1989 [#3241] (as sericans kuldjensis). Erroneous: Pistacia: TER MINASYAN, 1955 [#7893]; GURYEVA, 1974 [#5428]. Distribution: CHINA: Nei Mongol, Xinjiang. KAZAKHSTAN. KYRGYZSTAN. MONGOLIA. RUSSIA: Buryatiya Respublika, Tyva Respublika, Zabaykalskiy kray. TAJIKISTAN. TURKMENISTAN. UZBEKISTAN.

Agrilus notoclavus Jendek Fig. 15J (habitus of HT); Fig. 34X (aedeagus) notoclavus Jendek, 2000 (Agrilus) JENDEK, 2006: 400 JENDEK, 2000: 153 [#4853] (description). [#7441] (Palaearctic catalog). BELLAMY, 2008: 2205 [#8576] (world catalog).

Type material: Agrilus notoclavus Jendek, 2000 Type locality: China, 1000-1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E). Type specimens: Primary: Holotype (Fig. 15J), ♂, (EJCB):

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA “China, 1000-1300 m, Shaanxi [prov.], Qinling mts., Xunyangba (6 km E), 23. v.- 13.vi.1998, I. H. Marhshal leg.”. Secondary: 14 PT (EJCB). Described from 15 specimens (HT, PT). Diagnosis: 5.9-9.1 mm; pronotum medially with obvious black stripe. Species group: Smaragdinus. Relevant species: A. smaragdinus habros ssp. nov. Specimens examined: CHINA: Henan: 1 ♂ (EJCB): “China, W Henan, 1.vii.2001, 20 km SE Luanchan “tunel”, 33.7ºN 111.8ºE, O. Šafránek leg.”. Shaanxi: 1 ♂ PT (EJCB): “China S Shaanxi, Qinling Mts. S slope, Xunyangba S + W env. 33º28 37’ [N] / 108º23 33’ [E] 1400 2100m, 5. 9.6.[19]95” (JENDEK, 2000 [#4853]); 1 PT (EJCB): “C. China SW Shaanxi, Houzhenzi [33°51’N, 107°50’E] env. 1600m, Qinling mts. VI 1996, Lgt. Dr. M. Häckel” (JENDEK, 2000 [#4853]); 3 PT (EJCB): “China Shaanxi, Lüeang 33º07’N 106º05’E, 18.6 24.6 [19]97 lgt. E. Kučera” (JENDEK, 2000 [#4853]); 4 PT (EKCS): “China Shaanxi, Lüeang 33º07’N 106º05’E, 18.6 24.6 [19]97 lgt. E. Kučera” (JENDEK, 2000 [#4853]); 1 ♀ PT (EJCB): “China, 1000 1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v. 13.vi.1998, I. H. Marshal leg.” (JENDEK, 2000 [#4853]); 1 ♂ PT, 3 ♀ PT (EJCB): “China, 1150 1300 m, Shaanxi, Qinling mts., Foping (6 km N), 20 21.VI.1998, I. H. Marshal leg.” (JENDEK, 2000 [#4853]); 5 ♀ (EJCB): “C China, Shanxi [sic! should be Shaanxi], Qinling, Shan, 6 km N of Foping, 1150 1300m, 20 21.VI.2000, leg. C. Holzschuh”; 2 ♀ (EJCB): “C China, Shanxi [sic! should be Shaanxi], Qinling Shan, 6km E of Xunyangba, 1000 1300 m, 23.v. 13.vi., leg. C. Holzschuh, 2000”.

Adult occurence: 5-6-7 (n=9). Altitude range: 1000-2100 m (n=6). Host plants: Unknown. Distribution: CHINA: Henan, Shaanxi.

Agrilus obscuricollis Kiesenwetter Fig. 23J (habitus); Fig. 38E (aedeagus); Figs. 50D, 51U (types) obscuricollis Kiesenwetter, 1857 (Agrilus) KIESENWETTER, 1857: 125, 144 145 [#3015] (description). MARSEUL, 1866: 438, 491 492 [#676] (characters in key; redescription; faunal record; distribution overview; France). GEMMINGER & HAROLD, 1869: 1443 [#1057] (catalog). SAUNDERS, 1871: 121 [#2552] (catalog). MARSEUL, 1887: 252 [#4707] (catalog). SCHILSKY, 1889: 360 [#8431] (checklist; Germany). KERREMANS, 1892: 265 [#1397] (catalog). ABEILLE DE PERRIN, 1897: 3 [#19] (characters in key). KERREMANS, 1903: 275 [#1441] (catalog). JAKOBSON, 1913: 796 [#4386] (catalog; Russia and Europe). BEDEL, 1921: 203, 215 [#170] (characters in key; references; France). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1016 1018 [#2191] (world catalog). RIKHTER, 1945: 167 [#3507] (characters in key; Kharkovsk; Kursk). SCHAEFER, 1945: 149 150 [#3584] (Agrilus). ILINSKIJ, 1948: 293 [#5670] (larval characters; notes). RIKHTER, 1948: 394 [#3879] (characters in key; Kharkovsk; Kursk). SCHAEFER, 1949: 353, 366 368, 373, 490 [#2565] (characters; references; faunal records; biology; notes; ditributional overview; notes; France). RIKHTER, 1950: 159 [#3881] (characters in key). HORION, 1955: 84 85 [#1285] (distribution overview; faunal records; biology; Deutschland; Österreich). TER MINASYAN, 1955: 452 [#7893] (host plants; distribution; references). ALEXEEV, 1959: 19 [#3229] (characters in key; Kharkovsk; Kursk; Voronezhsk; Kamensk; Voroshilovgradsk; Stalingradsk; Stalinsk; Rostovsk; Dagestan). RIKHTER & ALEXEEV, 1965: 297 [#2522] (characters in key; European part of USSR). VOLKOVITSH, 1977: 25 [#2985]

143 (faunal record; notes; USSR: Belgorod, Kursk, Kharkov, Lugansk; Voronezh). LOMPE, 1979: 236 [#3628] (characters in key; Central Europe). ALEXEEV, 1981: 73 [#61] (larval characters in key). COBOS, 1986: 228 229, 315 [#609] (subgenus Anambus). NIEHUIS, 1988: 66 67 [#1912] (faunal records; biology; Germany). NIEHUIS & TEZCAN, 1993: 11 12, 48 [#1910] (subgenus Anambus; faunal records from Turkey; distribution and host plants summary; Turkey). CURLETTI, 1994: 22, 154 [#658] (subgenus Agrilus; catalog; faunal records; Italia; Sicilia; Malta; Corsica). ALEXEEV, 1998: 374 [#4322] (subgenus Quercuagrilus; subgeneric classification). BÍLÝ, 2002: 22 [#5877] (bionomy; host plants; distribution). BRECHTEL & KOSTENBADER, 2002: 275 277 [#5766] (biology; ecology; distribution; Germany). FARRUGIA, 2002: 18, 63 [#6511] (characters in key; France). JENDEK, 2003: 184 [#5823] (lectotype designation). VERDUGO PAÉZ, 2003: 200 [#6522] (subgenus Anambus; faunal record; Spain). KARAGYAN, KUZNETSOVA & LACHOWSKA, 2004: 155 [#7272] (subgenus Quercuagrilus; karyotype; Armenia). JENDEK, 2006: 392 [#7441] (subgenus Quercuagrilus; Palaearctic catalog). BELLAMY, 2008: 2208 2209 [#8576] (subgenus Quercuagrilus; world catalog). reyi Bauduer, 1878 (Agrilus) BAUDUER, 1878: 75, 79, 83 [#165] (description). BAUDUER, 1883: 21, 22 23 [#5143] (characters in key; notes; France). DELHERM DE LARCENNE, 1885: 184 [#3924] (checklist; France). KERREMANS, 1885: 154 [#1381] (catalog). MARSEUL, 1887: 252 [#4707] (catalog). REITTER, 1891: 200 [#5278] (catalog). REY, 1891: 19 [#3666] (Agrilus). FAUCONNET, 1892: 255 [#4467] (characters in key; France). KERREMANS, 1892: 270 [#1397] (catalog). ABEILLE DE PERRIN, 1897: 11 [#19] (synonym of asperrimus). HEYDEN, 1897: 194 [#5286] (synonym of asperrimus). KUTHY, 1897: 113 [#4040] (faunal records; kingdom of Hungary). KUTHY, 1900: 113 [#4060] (faunal records; kingdom of Hungary). KERREMANS, 1903: 275 [#1441] (synonym of asperrimus). PIC, 1909: 167, 189 [#5295] (as a synonym of aperrimus on page 167 and as a valid species on page 189). REITTER, 1911: 195 [#2518] (synonym of asperrimus). CAILLOL, 1913: 514 [#4053] (synonym of angustulus). JAKOBSON, 1913: 796 [#4386] (synonym of asperrimus; catalog). KAUFMANN, 1914: 56 [#5970] (checklist; Hungary). CSIKI, 1915: 96 [#3914] (synonym of aperrimus). BEDEL, 1921: 215 [#170] (synonym of asperrimus). FUENTE, 1930: 152 [#1036] (synonym of asperrimus). PORTEVIN, 1931: 330 [#3832] (synonym of asperrimus). OBENBERGER, 1935: 52 [#2178] (synonym of asperrimus). SAINTE CLAIRE, 1935: 232 [#8551] (synonym of asperrimus). OBENBERGER, 1936: 951 [#2191] (synonym of asperrimus). SCHAEFER, 1945: 148 149 [#3584] (synonym of obscuricollis). COBOS, 1986: 315 [#609] (synonym of obsuricollis). CURLETTI, 1994: 154 [#658] (synonym of obscuricollis). BÍLÝ, 2002: 22 [#5877] (SYNONYM OF OBSCURICOLLIS). BRECHTEL & KOSTENBADER, 2002: 275 [#5766] (synonym of obscuricollis). FARRUGIA, 2002: 107 [#6511] (synonym of obscuricollis). MUSKOVITS & HEGYESSY, 2002: 128 [#5806] (synonym of obscuricollis). JENDEK, 2006: 392 [#7441] (synonym of obscuricollis; Palaearctic catalog). BELLAMY, 2008: 2208 [#8576] (synonym of obscuricollis; world catalog). obscurus Delherm de Larcenne, 1885 (Agrilus; variety of obscuricollis) DELHERM DE LARCENNE, 1885: 83 [#3924] ([Note: Bauduer is cited as the author but he never described this taxon and Delherm de Larcenne also presents characters]; description). OBENBERGER, 1936: 1018 [#2191] (synonym of obscuricollis). ALEXEEV, 1959: 19 [#3229] (synonym of obscuricollis). CURLETTI, 1994: 154 [#658] (synonym of obscuricollis). BÍLÝ, 2002: 22 [#5877] (synonym of obscuricollis). BRECHTEL & KOSTENBADER, 2002: 275 [#5766] (synonym of obscuricollis). FARRUGIA, 2002: 107 [#6511] (synonym of obscuricollis). BELLAMY, 2008: 2208 [#8576] (unavailable synonym of obscuricollis; world catalog).

144 foveola Rey, 1891 (Agrilus; variety of obscuricollis) REY, 1891: 19 [#3666] ([Note: This name was treated as a senior homonym and thus is an available name (Article 45.6.4.1)]; description). JAKOBSON, 1913: 796 [#4386] (variety of obscuricollis; catalog; Russia and Europe). KUHNT, 1913: 660 [#1601] (variety of obscuricollis). OBENBERGER, 1926: 655 [#2057] (aberration of obscuricollis; Palaearctic catalog). PORTA, 1929: 403 [#2485] (aberration of obscuricollis; characters; Italy). SAINTE CLAIRE, 1935: 232 [#8551] (synonym of obscuricollis). OBENBERGER, 1936: 1018 [#2191] (variety of obscuricollis; world catalog). THÉRY, 1942: 160, 161 [#2873] (subgenus Agrilus; aberration of obscuricollis). COBOS, 1986: 315 [#609] (synonym of laticornis). CURLETTI, 1994: 155 [#658] (synonym of laticornis). BÍLÝ, 2002: 20 [#5877] (synonym of laticornis). BRECHTEL & KOSTENBADER, 2002: 273 [#5766] (synonym of laticornis). FARRUGIA, 2002: 107 [#6511] (synonym of laticornis). JENDEK, 2006: 40 [#7359] (available name). JENDEK, 2006: 392 [#7441] (synonym of obscuricollis; Palaearctic catalog). BELLAMY, 2008: 2208 [#8576] (synonym of obscuricollis; world catalog). savalae Stark, 1953 (Agrilus) STARK, 1953: 300 301 [#3534] (description). STARK, 1955: 775 [#3533] (faunal record; USSR: Saratov). TER MINASYAN, 1955: 453 [#7893] (host plant; distribution; references; Voronezh). ALEXEEV, 1959: 26 [#3230] (synonym of obscuricollis). ALEXEEV, 1959: 19 [#3229] (synonym of obscuricollis). ZAGAIKEVICH, 1962: 108 [#3249] (synonym of obscuricollis). RIKHTER & ALEXEEV, 1965: 297 [#2522] (synonym of obscuricollis). JENDEK, 2006: 392 [#7441] (synonym of obscuricollis; Palaearctic catalog). BELLAMY, 2008: 2208 [#8576] (synonym of obscuricollis; world catalog).

Type material: Agrilus obscuricollis Kiesenwetter, 1857 Type locality: In Deutschland . . in Steyermark . . in Lyon. Type specimens: Primary: Lectotype (Fig. 50D), ♀, designated by JENDEK (2003: 184) [#5823], (ZSMC): “Lyon [h] \ Kiesenwetter [p] \ Sammlung C. Müller [p] \ Type von [p] Agrilus obscuricollis Kiesw. [h] [red label] \ Holotypus [p] Agrilus obscuricollis Kiesenw. [h] Zool. Staatssammlg. München [p] Secondary: Described from unknown number of syntypes. Agrilus reyi Bauduer, 1878 Type locality: Lyon, Pyrénées, Allier, Paris, Aube, Landes, Sos. Type specimens: 1 ST (Fig. 51U) (BMNH); 1 ST (ISNB). Described from unknown number of syntypes. Agrilus obscuricollis variety obscurus Delherm de Larcenne, 1885 Type locality: Sos. Type specimens: Not located. Described from unknown number of syntypes. Agrilus obscuricollis variety foveola Rey, 1891 Type locality: not given. Type specimens: Not located. Described from unknown number of syntypes. Agrilus savalae Stark, 1953 Type locality: v Saval’skom lesnichestve Voronezhskoi oblasti (levyi bereg Savaly) [in Russian]. Type specimens: 1 ST (AAOR) labeled as paratype. Described from unknown number of syntypes. Diagnosis: 3.6-4.7 mm; antennomeres in male not dilated;

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA anterior pronotal lobe prominent; prosternal process subparallel or feebly enlarged. Species group: Sulcicollis. Relevant species: A. laticornis (Illiger, 1803). Specimens examined: RUSSIA: Kaluzhskaya oblast’: 1 (MSCI): “C. Russia, Kaluzhskaja ar., Malojaroslavetsky dist., Afanasovo vill. 11 19.06.2005 leg. М. Smirnov”. Lipetskaya oblast’: 1 (MCCD): “30км В г. Елец, ур. Морозова Гора, 5.7.1998 Цуриков”; 1 (MCCD): “30км В г. Елец, ур. Морозова Гора, 27.7.2000 Пантелеева”. Saratovskaya oblast’: 1 (MSCI): “Saratovskaya obl. Novoburasskii r n, okr. st. Burasy, 52°12’N, 46°08’E, 21.06.2008, M. Smirnov”. Tyva Respublika: 1 ♂ (EJCB): “Siberia, NC Tuva, 5.vi.2002, Zapadnii Sayan, Uyukskiy Rdg., Uyuk vill. env., Uyuk riv., 52°05’N, 94°00’E, 800m, leg. Jan Hron”.

Adult occurence: 5-6-7 (n=11). Altitude range: ~800 m (n=1). Host plants: Carpinus betulus: BÍLÝ, 2002 [#5877]. – Quercus: BAUDUER, 1878 [#165] (as reyi); DELHERM DE LARCENNE, 1885 (as variety obscurus) [#3924]; CAILLOL, 1913 [#4053]; SAINTE CLAIRE, 1935 [#8551]; ILINSKIJ, 1948 [#5670]; STARK, 1953 (as savalae) [#3534]; HORION, 1955 [#1285]; TER MINASYAN, 1955 [#7893]; ZAGAIKEVICH, 1962 [#3249]; RIKHTER & ALEXEEV, 1965 [#2522]; VOLKOVITSH, 1977 [#2985]; ALEXEEV, 1981 [#61]; NIEHUIS, 1988 [#1912]; CURLETTI, 1994 [#658]; VOLKOVITSH, 2004 [#7001]; MÜHLE, 2007 [#8787]. – Q. cerris: GOBBI, 1973 [#1093]. – Q. ilex: NOVAK, 1964 (as jirnensis) [#6567]; NOVAK, 1964 [#6567]. – Q. petraea: BÍLÝ, 2002 [#5877]; BRECHTEL & KOSTENBADER, 2002 [#5766]. – Q. pubescens: SCHAEFER, 1949 [#2565]; BÍLÝ, 2002 [#5877]. – Q. robur: BEDEL, 1921 [#170]; SCHAEFER, 1949 [#2565]; BÍLÝ, 2002 [#5877]; BRECHTEL & KOSTENBADER, 2002 [#5766]. Dubious: Acer: RIKHTER & ALEXEEV, 1965 [#2522]. – Betula: SLIVKINA, 1958 [#8660]; RIKHTER & ALEXEEV, 1965 [#2522]. – B. alba: BEDEL, 1921 [#170]. – Ficus: CURLETTI, 1994 [#658]. Distribution: Local: RUSSIA: Respublika Tyva. Global: AFRICA: ALGERIA. ASIA: ARMENIA. AZERBAIJAN. IRAN. ISRAEL. KAZAKHSTAN. RUSSIA. TURKEY. EUROPE: ALBANIA. AUSTRIA. BELARUS. BOSNIA & HERZEGOVINA. BULGARIA. CROATIA. CZECH REPUBLIC. FRANCE. GERMANY. GREECE. HUNGARY. ITALY. MACEDONIA. MALTA. MOLDOVA. NETHERLANDS. POLAND. ROMANIA. SERBIA & MONTENEGRO. SLOVAKIA. SLOVENIA. SPAIN. SWITZERLAND. UKRAINE.

Agrilus obscurosericeus Obenberger Fig. 24J (habitus of LT); Fig. 38Q (aedeagus) obscurosericeus Obenberger, 1940 (Agrilus) OBENBERGER, 1940: 185 [#2273] (description). PENG ZHONGLIANG, 1987: 357 [#2429] (checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (cited as obscurisericeus; checklist; China). PENG ZHONGLIANG, 2002: 270 [#6628] (notes). JENDEK, 2006: 400 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2210 [#8576] (world catalog).

Type material: Agrilus obscurosericeus Obenberger, 1940 Type locality: China: Fokien: Taining-li.

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TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Type specimens: Primary: Lectotype (Fig. 24J) by present designation, ♀, (NMPC): “Fokien: Tainingli [h] \ Typus [p] [red label] \ A. obscurosericeus m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24944 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 4.8-5 mm; pronotum widest in middle; elytral pubescence in form of sutural wedge; aedeagus asymmetric. Species group: Sulcicollis. Relevant species: A. friebi Obenberger, 1922. Specimens examined: CHINA: Anhui: 1 ♂ (EJCB): “China, Anhui, 1998, Dabieshan, 65 km SW Huoshan, 1400 m, Bolm lgt. 21 24 Jun”.

Adult occurence: 6 (n=1). Altitude range: ~1400 m (n=1). Host plants: Unknown. Distribution: CHINA: Anhui, Fujian.

Agrilus odetteae Baudon Fig. 15L (habitus); Fig. 35A (aedeagus), Fig. 50G (type) odetteae Baudon, 1968 (Agrilus) BAUDON, 1968: 90, 96 97 [#162] (description). 2214 [#8576] (world catalog).

BELLAMY, 2008:

Type material: Agrilus odetteae Baudon, 1968 Type locality: Laos: Muong-Panh. Type specimens: Primary: Holotype (Fig. 50G), ♀, (MHNB): “Muong Panh 1.iv.64 [h] Laos (Baudon) [p] \ Type [p] [red label] \ Agrilus odetteae Type mihi [h] A, Baudon det. [p]”. Described from 1 specimen (HT). Diagnosis: 7-8.6 mm; dorsal side golden green except for vertex, medial pronotal stripe and tear-like spot on elytra; pronotum visually square; prehumerus absent. Species group: Smaragdinus. Relevant species: A. aurinotus sp. nov. Specimens examined: CHINA: Fujian: 1 ♀ (EJCB): “China, Fujian c, 700 1100m, Ziyungdogshan, NW slope, 25º46’N, 117º20’E, 31.V., Jaroslav Turna leg., 2008”. Hunan: 1 ♂ (EJCB): “China, Hunan SE, 1994, Guidong env. 26 31.V., 26.04N 113.56E, E. Jendek & O. Šauša leg.”. Jiangxi: 1 ♂ (EJCB): “China, Jiangxi W, Jingggang Shan Ciping, 2 14.VI.1994, E. Jendek & O. Šauša leg.”. Taiwan: 1 ♂, 2 ♀ (EJCB): “Mt. Kuantoushan, Taiwan, 2.vii.1981”; 1 ♂ (EJCB): “Rep. of China, Formosa (Taiwan), Sun Moon Lake, 7 9.6.2002, Moravec Petr”. INDIA: Arunachal Pradesh: 1 ♀ (EJCB): “India, W Arunachal Pr., Dirang vicinity, 1500 1800m, 27°21’ 23’N 92°13’ 16’E, L. Dembický leg.; 1. 10.vi.2004”. LAOS: Borikhamxai: 1 ♀ (EJCB): “Laos CE., 1 18.v.2001, Boli Kham Xai prov., 18º21’N, 105º08’E, Ban Nape (8 km NE), ~600m, Pacholátko leg.”; 1 ♀ (EJCB): “Laos CE., 1 18.v.2001, Boli Kham Xai prov., 18º21’N, 105º08’E, Ban Nape (8 km NE), ~600m, Dembický leg.”; 1 ♂ (EJCB): “Laos C., Bolikhamsai pr., Ban Nape env., 7 16.V.2004, alt., 400±100 m, 18º20’N, 105º08’E, E. Jendek & O. Šauša leg.”. Houaphan: 2 ♀ (EJCB): “Laos NE, Hua Phan prov., 20°15’N, 104°02’E, Phu Pan mt. (Ban Saluei), 1500 2000 m, 26.iv. 11.v.2001, J. Bezděk leg.”; 1 ♂, 1 ♀ (EJCB): “Laos NE, Houa Phan prov., 20°12’ 13.5’N, 103°59’.5 104°01’E, Ban Saluei → Phou Pane Mt., 1340 1870m, 15.iv. 15.v.2008, Lao collectors leg.”. VIETNAM: 1 ♀ (ZIN): “V’etnam, khr. Fan ci pan, 2200 2900m, 25.5.1963, Kabakov [in Russian]”; 1 ♀ (EJCB): “N. Vietnam (Tonkin), Tamdao (pr. (Vinphu), 2 11.6.1985, A. Olexa

leg.”; 1 ♀ (EJCB): “N. Vietnam, 1985, Tam dao, 3 11.6.1985, 900 1400m, J. Jelínek lgt.”.

Adult occurence: 4-5-6-7 (n=14). Altitude range: 3002900 m (n=9). Host plants: Unknown. Distribution: CHINA: Fujian, Hunan, Jiangxi, Taiwan. INDIA: Arunachal Pradesh. LAOS: Borikhamxai, Houaphan, Xiangkhoang. VIETNAM. Remarks: The specimens from Taiwan - Mt. Kuantoushan (see Specimens examined) were misidentified by K. Akiyama as A. obscurecinctus Obenberger, 1935. The latter species is larger (8.5-9 mm), has a single black medial strip on the pronotum and a longer aedeagus. Agrilus obscurecinctus is known to us only from Taiwan. Records for this species from mainland China (Hunan, Jiangxi) cited by JENDEK (2006: 400 [#7441]) should refer to A. odetteae.

Agrilus ogatai Ohmomo Fig. 18E (habitus of PT); Fig. 35U (aedeagus) ogatai Ohmomo, 2002 (Agrilus) OHMOMO, 2002: 160 162 [#5750] (description). FUKUTOMI & HORI, 2004: 10 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 400 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2214 [#8576] (world catalog).

Type material: Agrilus ogatai Ohmomo, 2002 Type locality: Mt. Kurodake, Kuju, Oita Pref., Kyushu. Type specimens: Primary: Holotype, ♂, NSMT (OHMOMO, 2002 [#5750]). Secondary: 1 PT (Fig. 18E)(EJCB). Described from 38 specimens (HT, PT). Diagnosis: 5.2-7.2 mm; for distinctive characters see OHMOMO, 2002: 162 [#5750]. Species group: Betuleti. Relevant species: A. sibiricus Obenberger, 1912. Specimens examined: Known to us only from the male paratype.

Host plants: Unknown. Distribution: JAPAN: Hokkaido, Honshu, Kyushu. Remarks: The diagnostic characters (shape of prosternal process and submarginal pronotal carina) given by OHMOMO, 2002: 162 [#5750] to distinguish A. ogatai from A. sibiricus are not adequate. The form of prosternal process of A. sibiricus fukushimensis is not enlarged but it is similar to that of A. ogatai. More specimens are needed to resolve status of A. ogatai.

Agrilus olivaceidorsis Obenberger nom. rev. Fig. 20E (habitus); Fig. 36R (aedeagus); Figs. 45P, 50I, 50K (types) olivaceidorsis Obenberger, 1917 (Agrilus) nom. rev. OBENBERGER, 1917: 211 212 [#1997] (description). OBENBERGER, 1926: 660 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 958 [#2191] (variety of auriventris; world catalog). HUA LI ZHONG,

146 2002: 89, 90 [#6696] (cited as a synonym of auriventris on page 89 and as a species on page 90; checklist; China). JENDEK, 2006: 396 [#7441] (synonym of auriventris; Palaearctic catalog). BELLAMY, 2008: 1985 [#8576] (synonym of auriventris; world catalog). duporti Bourgoin, 1922 (Agrilus) syn. nov. BOURGOIN, 1922: 23 [#365] (description). OBENBERGER, 1936: 1082 [#2191] (world catalog). DESCARPENTRIES & VILLIERS, 1963: 105, 112 [#700] ([Note: An inference of type: “Tonkin: Cho Ganh (Duport). Holotype au Muséum de Paris” is not valid lectotype designation because of more syntypes (ICZN, Article 74.3)]; characters; faunal records; Tonkin). BAUDON, 1968: 130, 141 [#162] (Laos). BELLAMY, 2008: 2078 [#8576] (world catalog). ornamentifer Obenberger, 1940 (Agrilus) syn. nov. PENG OBENBERGER, 1940: 181 182 [#2273] (description). ZHONGLIANG, 1987: 357 [#2429] (checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China). PENG ZHONGLIANG, 2002: 269 [#6628] (characters; Fujian). JENDEK, 2006: 400 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2220 [#8576] (world catalog).

Type material: Agrilus olivaceidorsis Obenberger, 1917 Type locality: China. Type specimens: Primary: Lectotype (Fig. 50I) by present designation, ♂, (NMPC): “Lilong. Chine [h] \ Typus [p] [red label] \ olivaceidorsis m. Type [h] Det. Dr Obenberger [p]”. Secondary: 1 PLT (NMPC). Described from unknown number of syntypes Agrilus duporti Bourgoin, 1922 Type locality: not given [Note: Indo-Chine française is cited in the title of the publication. Stipulated to “Cho-Ganh” by DESCARPENTRIES & VILLIERS, 1963:112 [# 700]]. Type specimens: Primary: Lectotype (Fig. 45P) by present designation, ♂, (MNHN): “Cho Ganh L. Duport [p] \ Agrilus Duporti A. Bourgoin [h] Type [h] [red ink] \ Type [p] [red label] \ Museum Paris Coll. Générale [p] [yellow label]”. Secondary: 7 PLT (MNHN); PLT (NMBS); 2 PLT (MCSN); 4 PLT (NMPC); 1 PLT; (BMNH). Described from unknown number of syntypes. Agrilus ornamentifer Obenberger, 1940 Type locality: China: Fokien: Taining-Li. Type specimens: Lectotype (Fig. 50K) by present designation, ♂, (NMPC): “Fokien: Tainingli [h] \ Typus [p] [red label] \ Agrilus ornamentifer m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24301 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 5.1-9 mm; body generally smaller and slimmer; pubescent elytral fasciae wider with narrower interspaces. Species group: Angulatus. Relevant species: A. grisator Kerremans, 1893. Specimens examined: CHINA: Yunnan: 1 (NMPC): “Yun’nan, Czindun, 1170 m, 20.v.1956, Kryzhanovskii [in Russian]”; 1 ♂ (EJCB): “S. China, Jinghong, 10 14.7.1990, pr. Yunnan, lgt. S. Bečvář”; 1 ♂ (EJCB): “China: S Yunnan (Xishuangbanna), 23 km NW Jinghong vic. Na Ban (NNNR) \ N22 09.49, E100 39.92, 730 m, 6.vi.2008, leg. A. Weigel traps, transition zone MF1”. THAILAND: 1 (MNHE): “Thailand, 13.05.2005, Huai Pong, 600 m, 18º50’50N/97º49’23E, leg. U. Roosileht”.

Adult occurence: 5-6-7 (n=5). Altitude range: 600-1170 m (n=3).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Host plants: Citrus: BAUDON, 1968 (as duporti) [#162]. Distribution (see Remarks): CHINA: Fujian, Guizhou, Yunnan. THAILAND. VIETNAM. Remarks. Distribution of this species given herein is incomplete due to the pending revision of the whole species group. Only records from China are given among examined specimens.

Agrilus ornatoides Jendek, sp. nov. Fig. 11F (habitus of HT); Fig. 33F (aedeagus)

Description: 5.0-5.5 mm (Holotype 5.5 mm). BODY: Shape: subparallel, Build: robust, Color (dorsally): unicolored, Color (dominant): black or brown. HEAD: Shape: markedly convex, Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): irregular or subparallel, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: small, Shape: not protruding from head outline, Lower margin: in line with antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp, rarely blunt. Disk: Convexity: flat or moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow. Prehumerus: filamentary or carinal, Shape: bisinuate, Extent: to anterior pronotal angle, Modifications: obsolete in anterior part, Posterior end: joining posterior pronotal angle, Anterior end: joining anterior pronotal angle, Arc: moderate. Marginal and submarginal carinae: Interspace: narrow or moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior angles or at posterior 1/51/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate, Width: wide, Shape: arcuate. Elytral pubescence: proximal and distal, Proximal: like V or upturned T, Distal: apical, Color: unicolored, Character: with spots of denser pubescence. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): moderate. Prosternal process: moderate, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate.

147

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): distinctly longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed, Ovipositor: elongate. Diagnosis: body more robust; suparallel; pronotal disk with rough transverse sculpture. Species group: Quadrisignatus. Relevant species: A. nipponigena Obenberger, 1935. Type locality: China, Hebei province, Fengning env., northwest slope of Yunwu Shan, altitude 1200 m. Type specimens: Primary: Holotype (Fig. 11F), ♂, (EJCB): “China, 3 June 2000, Fengning env., 1200m, NW slope of Yunwu Shan, Zd. Jindra lgt.”. Secondary: CHINA: Shaanxi: 1 ♀ PT (EJCB): “China c., 17-18.V., Ganguyi 110,3’-36,8’, 35 km NE of Yanan, leg. J. Halada 1996”. Adult occurence: 5-6 (n=2). Altitude range: 1200 m (n=1). Host plants: Unknown. Distribution: CHINA: Hebei, Shaanxi. Etymology: The specific name is a combination of the Latin adjective ornatus (decorate) and -oides (Greek suffix denoting likeness). It refers to the elytral ornamental pubescence of the species.

Agrilus paralleloides Jendek, sp. nov. Fig. 19A (habitus of HT); Fig. 36B (aedeagus)

Description: 7.7-8.4 mm (Holotype 7.8 mm). BODY: Shape: subparallel, Build: robust, Color (dorsally): unicolored, Color (dominant): red. HEAD: Medial impression: present, Impression (extent): vertex, Epistoma: in plane with frons. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): semispherical, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: small, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles or at level with anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: flat, Disk impressions: lateral only, rarely medial and lateral. Prehumerus: tubercular, Extent: to third of pronotal length. Marginal and submarginal carinae: Interspace: moderate, Convergence: strongly convergent, Junction: present, Narrowest point: at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices:Arrangement: separate, Width: wide, Shape: arcuate. Elytral pubescence: distal only, Distal: perisutural stripe, Color: unicolored. Tomentum: absent. STERNUM:

Metasternal projection: impressed, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): narrow. Prosternal process: moderate, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): sharp, Disk: flat. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate or angulately sinuate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed. Diagnosis: body dark carmineous; apical third of elytra with adsutural strip of pubescence. Species group: Laetecyanescens. Relevant species: A. laetecyanescens Obenberger, 1940. Type locality: China, Yunnan province, Julongxue Shan, 40 km north of Lijiang, 27°12’30.8’’N, 100°16’01.0’’E, altitude 3082 m. Type specimens: Primary: Holotype (Fig. 19A), ♂, (EJCB): “China, Yunnan, Loc.15, Julongxue Shan, 40 km N of Lijiang, 27°12’30.8’’N, 100°16’01.0’’E, 3082 m, 28.05.2002, Konstantinov & Volkovitsh leg.”. Secondary: CHINA: Yunnan: 1 ♂ PT, 1 ♀ PT (EJCB): “China, Yunnan, Loc.13, Julongxue Shan, 30 km N of Lijiang, 27°08’46.9’’N, 100°15’03.3’’E, 2802 m, 02.06.2002, Konstantinov & Volkovitsh leg.”. Adult occurence: 6 (n=2). Altitude range: 2802-3082 m (n=2). Host plants: Unknown. Distribution: CHINA: Yunnan. Etymology: The specific name is a combination of the Greek adjective parallelos (side by side) and -oides (Greek sufix denoting likeness). It refers to the subparallel body of this species.

Agrilus pecirkai Obenberger Fig. 17C (habitus); Fig. 35H (aedeagus); Fig. 50O (type) pecirkai Obenberger, 1916 (Agrilus) OBENBERGER, 1916: 273 274 [#1983] (description). OBENBERGER, 1926: 656 [#2057] (Palaearctic catalog). ROUBAL, 1930: 115 [#5172] (notes). OBENBERGER, 1936: 1021 [#2191] (world catalog). MAKHNOVSKIJ, 1958: 127 [#6090] ([Note: Misidentified as chrysoderes]; pest; Uzbekistan). MAKHNOVSKIJ, 1959: 32 33 [#5829] ([Note: Misidentified as chrysoders]; characters; biology; Uzbekistan). GURYEVA, 1974: 99 [#5428] (pest of agriculture cultures; references; Uzbekistan; Tadjikistan). KADYROV, 1976: 52 [#3461] (cited as pecsikoi; biology; Tajikistan). ALEXEEV, 1998: 381 [#4322] (subgenus Agrilus; subgeneric classification). JENDEK, 2003: 11, 19 [#5878] (notes; characters; faunal records; distribution). JENDEK, 2006: 389 [#7441] (subgenus Agrilus; Palaearctic catalog). BELLAMY, 2008: 2228 [#8576] (subgenus Agrilus; world catalog). GHAHARI et al., 2008: 638 [#8160] (faunal record; Iran).

148

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Type material: Agrilus pecirkai Obenberger, 1916 Type locality: Turkestan: Tschamkal. Type specimens: Holotype by monotypy (Fig. 50O), ♀, (NMPC): “Tchamkal Turkestan 198 / VII. 06 [h] \ Typus [p] [red label with black border] \ Agrilus Pečírkai m. Type [Obenberger’s MS] Det. Dr. Obenberger [p]”. Described from 1 specimen.

Type material: Agrilus pekinensis accolus Jendek, 1995 Type locality: Yunnan, 1950 - 2050 m, 27.18N, 100.14E, Daju, Jinsha r. Type specimens: Primary: Holotype (Fig. 42A), ♂, (EJCB): “YUNNAN, 1950 - 2050 m, 27.18N, 100.14E, DAJU, Jinsha r., 7-10 / 7. 92”. Secondary: 1 PT (TSCB); 2 PT (MNHN); 60 PT (EJCB). Described from 91 specimens (HT, PT).

Diagnosis: 4.4-6.6 mm; prehumerus carinal, strongly arcuate; pronotal disk with deep cavity between prehumerus and pronotal side. Species group: Viridis. Relevant species: A. cuprescens (Ménétriés, 1832).

Diagnosis: 6-9.1 mm. It differs from nominotypical taxon by having body more elongate and dorsally more flattened. Species group: Pekinensis. Relevant species: A. pekinensis Obenberger, 1924.

Specimens examined: KAZAKHSTAN: 2 (ZIN): “25 km YUV [ southeastern] Suzaka, khr. Karatau, c v [ northeastern] sklon, Kazakhstan, Volkovitsh, 23 VI 1987 [in Russian]”. KYRGYZSTAN: 1 (ZIN): “ur. r. Alamedin, 50 km Yu Frunze Kirgiziya Volkovitsh 7 8.vii.1984 [in Russian]”; 2 ♀ (EJCB): “Ferganskii khr., 15 km SV [ northeastern] Arkhangel’skogo, Kirgiziya Volkovich 28 29.VI.1986 [in Russian]” (JENDEK, 2003 [#5878]); 1 ♂ (AGCM): “Kirgizstan, Kirgizski range, Ala Archa river, h 2500 m, 25.05.1997”. TAJIKISTAN: 1 ♀ (ZIN): “y. Kondara, 1100 m, d. Varzoba, Tadzh., Gussakovskii, 17/VI [19]37 [in Russian]”; 1 ♀ (ZIN): “y. Kondara, 1100 m, d. Varzoba, Tadzh., Gussakovskii, 26/VI [19]37 [in Russian]”; 1 (ZIN): “NW bereg ozera Iskander kul’, Giss. khr. Kirichenko 19.vii.1947 [in Russian]”; 1 ♂ (EJCB): “Tadzik SSR, 25.6.1976 Hissar Mts. Ziddi J. Jelinek lgt.” (JENDEK, 2003 [#5878]); 1 (ZIN): “Tadzhikistan Ergeng 1300 m 8.vi.1979 Alekseev”; 1 ♀ (EJCB): “USSR, Tardzjekistan 1981, Hissar Mts, 1300 1600 m, 50 km N of Dushanbe, 19.6., Gushara, Varzob river, Vít Kubáň leg.” (JENDEK, 2003 [#5878]); 1 ♂, 1 ♀ (EJCB): “Tadjik, 17.V.1982 Hissar, Romit, P. Štys leg.” (JENDEK, 2003 [#5878]); 1 (MDCM): “Tadzhikistan Varzob 27.6.1986, S. Nikireev” (JENDEK, 2003 [#5878]); 1 (ZIN): “15 km SV [ Northeast] pos. Vanch, Darvazkii khr., GBAO, Dolgovskaya 16 17 VI.[1]986 [in Russian]”; 1 (MDCM): “Tadz. SSR, Romit, 8.VI.1988, M. Danilevski leg.” (JENDEK, 2003 [#5878]). TURKMENISTAN: 1 ♂ (ZIN): “Turkmeniya, khr. Kugitang tau, Svinc. Rudnik, 1500 2000m, 28.V.1977, G. S. Medvedev [in Russian]”.

Specimens examined: CHINA: Anhui: 1 ♂, 1 ♀ (EJCB): “China, Anhui, 1998, Dabieshan, 65 km SW Huoshan, 1400 m, Bolm lgt. 21 24 Jun”. Hunan: 1 ♀ (EJCB): “China, N Hunan, 4 7.vii., Wuling Shan, 29.4N, 110.4E, Zhangjiajie, 700 m, Jaroslav Turna leg. 2003”. Sichuan: 1 PT (MNHN): “Su Tchuen [ Sichuan province], Chass. Thibétains, 1897” (JENDEK, 1995 [#3768]); 1 ♂ (EJCB): “China, NW Sichuan prov., Maowen, 1000 m, 31°30’N 103°50’E, 10 18. vii.1990, J. Kolibáč leg.”; 1 PT (EJCB): “China: Sichuan Mt. Emei, 1050 m, 10.VII.1990, L.& M. Bocák lgt.” (JENDEK, 1995 [#3768]); 1 PT (EJCB): “China, N Sichuan prov., Jianchangba, ca 2000m, 8.7.1991, J. Kaláb leg.” (JENDEK, 1995 [#3768]); 1 PT (EJCB): “China N, Sichuan, Jiuzhaigou, 2000m, 16.Jun 1992, M. Bok lgt.” (JENDEK, 1995 [#3768]); 1 (EJCB): “China, Ne Sichuan, 22.vi., road Wanyuan Chengkou, pass 20 km NE Wanyuan, 32.2N, 108.28E 1500 m, J. Turna leg., 2000”. Yunnan: 1 PT (MNHN): “Yunnan Sen, coll. Guerry” (JENDEK, 1995 [#3768]); 4 PT (EJCB): “China, Yunnan, 3500 m, 30 km N of Lijiang, Ganhaizi pass, 5.VII., M. Bocák leg. 1990” (JENDEK, 1995 [#3768]); 4 PT (EJCB): “China: N. Yunnan, 30 km N of Lijiang, 3000 m, 3.VII.1990, L.& M. Bocák lgt.” (JENDEK, 1995 [#3768]); 2 PT (EJCB): “China: N. Yunnan, Lijiang, 2600 m, 30.6. 2.7.1990, L.& M. Bocák lgt.” (JENDEK, 1995 [#3768]); 1 PT (EJCB): “China, Yunnan, 6. 10.VII., Ninglang env., 27.19N, 100.55E, E. Jendek leg. 1992” (JENDEK, 1995 [#3768]); 7 PT (EJCB): “China, Yunnan, 1800 m, Lijiang, 23.6. 21.7., 26.53N, 100.18E, E. Jendek leg. 1992” (JENDEK, 1995 [#3768]); 2 PT (EJCB): “Yunnan, 1500 2800 m, 26.07N, 103.14E, Dongchuan, 28.6. 3.7., Vít Kubáň leg., 1994” (JENDEK, 1995 [#3768]); 1 PT (EKCS): “China Yunnan, 10 15.6.[19]94, Litiang [ Lijiang], lgt. E. Kučera” (JENDEK, 1995 [#3768]); 3 PT (EJCB): “China, Yunnan, Daju, 17.VI.1995, Šiška & Spevár & Pekarovič” (JENDEK, 1995 [#3768]); 59 PT (EJCB): “China, Yunnan, Lijiang, 19.VI.1995, Šiška & Spevár & Pekarovič” (JENDEK, 1995 [#3768]); 2 (ZIN): “China, Yunnan, Loc. 4, Er Hai Lake, 15 km NE of Xiaguan, 25°45’32.6’’N, 100°13’43.0’’E, 2261m, 20.V.2002, Konstantinov & Volkovitsh leg.”; 1 (ZIN): “China, Yunnan, Loc. 26, Road #214, Dali Lijiang, 72 km NNW of Xiaguan, 26°09’41.0’’N, 100°10’14.1’’E, 2500m, 5.VI.2002, Konstantinov & Volkovitsh leg.”.

Adult occurence: 5-6-7 (n=15). Altitude range: 1100-2500 m (n=6). Host plants: Rosa: MAKHNOVSKIJ, 1958 [#6090]; MAKHNOVSKIJ, 1959 [#5829]; GURYEVA, 1974 [#5428]; KADYROV, 1976 [#3461]. Erroneous: Quercus rotundifolia: GHAHARI et al., 2008 [#8160]. Distribution: IRAN. KAZAKHSTAN. KYRGYZSTAN. TAJIKISTAN. TURKMENISTAN. UZBEKISTAN. Remarks: This species has not been recorded from East Asia yet but it is possible in Mongolia or Tyva.

Agrilus pekinensis accolus Jendek Fig. 22E (habitus); Fig. 37N (aedeagus); Fig. 42A (type) accolus Jendek, 1995 (Agrilus; subspecies of pekinensis) JENDEK, 1995: 142 [#3768] (description). JENDEK, 2006: 393 [#7441] (subgenus Robertius; subspecies of pekinensis; Palaearctic catalog). BELLAMY, 2008: 2230 [#8576] (subspecies of pekinensis; world catalog).

Adult occurence: 5-6-7 (n=17). Altitude range: 700-3500 m (n=13). Host plants: See A. pekinensis pekinensis. Distribution: CHINA: Anhui, Sichuan, Yunnan.

Agrilus pekinensis pekinensis Obenberger Fig. 22D (habitus); Fig. 37M (aedeagus); Figs. 43T, 44F, 44H, 47X, 50P (types) pekinensis Obenberger, 1924 (Agrilus)

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA OBENBERGER, 1924: 55 [#2040] (description). OBENBERGER, 1926: 658 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1021 [#2191] (world catalog). MIWA & CHÛJÔ, 1940: 74 [#1810] (faunal records; south Korea). PENG ZHONGLIANG, 1987: 357 [#2429] (checklist; China). ALEXEEV, 1989: 479 [#64] (key to species; Khabarovsk; Amursk; Primorsk; North Korea; northeastern China; northeastern Mongolia). ALEXEEV & VOLKOVITSH, 1989: 318, 345, 356 [#3241] (characters in key; faunal records; Mongolia). JENDEK, 1995: 141 142 [#3768] (lectotype designation; synonymy; notes; Russia: Chita; Amurskiy, Khabarovskiy, Primorskiy regions; Mongolia; Korea; northeastern and eastern China). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). NONNAIZAB et al., 1999: 113 [#8247] (checklist; China: Inner Mongolia). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Beijing). JENDEK, 2006: 393 [#7441] (subgenus Robertius; Palaearctic catalog). VOLKOVITSH, BELLAMY, 2008: 2229 [#8576] (world catalog). 2009: 136 [#8850] (subgenus Robertius; faunal records; Russia: Primorskiy kray). boreooccidentalis Obenberger, 1936 (Agrilus; subspecies of pekinensis) OBENBERGER, OBENBERGER, 1936: 107 [#2205] (description). 1936: 1021 [#2191] (subspecies of pekinensis; world catalog). JENDEK, 1995: 141, 142 [#3768] (synonym of pekinensis; lectotype designation). JENDEK, 2006: 393 [#7441] (synonym of pekinensis pekinensis; Palaearctic catalog). BELLAMY, 2008: 2229 [#8576] (synonym of pekinensis; world catalog). cerskii Obenberger, 1936 (Agrilus) OBENBERGER, OBENBERGER, 1936: 104 [#2205] (description). 1936: 986 [#2191] (world catalog). ALEXEEV, 1989: 479 [#64] (synonym of pekinensis). ALEXEEV & VOLKOVITSH, 1989: 318 [#3241] (synonym of pekinensis; types examination). JENDEK, 1995: 141, 142 [#3768] (synonym of pekinensis; lectotype designation). NONNAIZAB et al., 1999: 113 [#8247] (synonym of pekinensis). JENDEK, 2006: 393 [#7441] (synonym of pekinensis pekinensis; Palaearctic catalog). BELLAMY, 2008: 2229 [#8576] (synonym of pekinensis; world catalog). klapperichi Obenberger, 1940 (Agrilus) PENG OBENBERGER, 1940: 183 184 [#2273] (description). ZHONGLIANG, 1987: 356 [#2429] (checklist; China). ALEXEEV & VOLKOVITSH, 1989: 318 [#3241] (synonym of pekinensis; types examinated). JENDEK, 1995: 141, 142 [#3768] (synonym of pekinensis; lectotype designation). NONNAIZAB et al., 1999: 113 [#8247] (synonym of pekinensis). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China). PENG ZHONGLIANG, 2002: 269 [#6628] (notes). JENDEK, 2006: 393 [#7441] (synonym of pekinensis pekinensis; Palaearctic catalog). BELLAMY, 2008: 2229 [#8576] (synonym of pekinensis; world catalog). charbinensis Théry, 1942 (Agrilus) OBENBERGER, THÉRY, 1942: 256 258 [#2875] (description). 1943: 9 [#2333] (synonym of pekinensis). JENDEK, 1995: 141 [#3768] (synonym of pekinensis). BELLAMY, 2008: 2230 [#8576] (synonym of pekinensis; world catalog).

Type material: Agrilus pekinensis Obenberger, 1924 Type locality: Pekin. Type specimens: Primary: Lectotype (Fig. 50P), ♂, designated by JENDEK (1995: 141-142) [#3768], (NMPC): “PECHINO [ = Beijing] S. Folchini 190 [p] 5 [h] \ Typus [p] [red label] \ Agrilus pekinensis m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24937 [h] [orange label]”. Secondary: 1 PLT (MCSN). Described from unknown number of syntypes. Agrilus pekinensis boreooccidentalis Obenberger, 1936

149 Type locality: Transbaicalia. Type specimens: Primary: Lectotype (Fig. 43T), sex not examined, designated by JENDEK (1995: 141, 142) [#3768], (NMPC): “Transbaikalia [h] \ Typus [p] [red label] \ Agrilus pekinensis var. [sic!] boreooccidentalis m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24278 [h] [orange label]”. Described from unknown number of syntypes. Agrilus cerskii Obenberger, 1936 Type locality: Primorje: Lacus Chanka; Ussuri: Nižne Michajlovskaya; Kamen-Rybolov. Type specimens: Primary: Lectotype (Fig. 44F), ♂, designated by JENDEK (1995: 141, 142) [#3768], (NMPC): “Ussuri Nizhne Mikhailovskaya [= Nizhnemikhailovka, 50 km N of Dal’nerechensk] 1899 G. Suvorov [p] [in Russian] \ Typus [p] [red label] \ Mus. Nat. Pragae Inv. [p] 24330 [h] [orange label] \ Agrilus Čerskii m. Type [h] Det. Dr Obenberger [p]”. Secondary: 11 PLT (ZIN); 3 PLT (NMPC). Described from unknown number of syntypes. Agrilus klapperichi Obenberger, 1940 Type locality: China.: Fukien; Shaowu, Kwangtsch. Type specimens: Primary: Lectotype (Fig. 47X), ♂, designated by JENDEK (1995: 141, 142) [#3768], (NMPC): “Kwangtseh - Fukien [ = Fujian province] J. Klapperich 17.9. 1937 [p] [violet label] \ Typus [p] [red label] \ Agrilus Klapperichi m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24303 [h] [orange label]”. Secondary: 2 PLT (NMPC). Described from unknown number of syntypes. Agrilus charbinensis Théry, 1942 Type locality: Charbin; Mandchourie. Type specimens: 1 ST (Fig. 44H) (MNHN), labeled as paratype. Described from 2 syntypes. Diagnosis: 5.4-7.8 mm; prehumerus filamentary, in basal part sometimes carinal, reaching to anterior pronotal angle, sometimes obsolete in middle of pronotum. Species group: Pekinensis. Relevant species: A. alashanensis Obenberger, 1936. Specimens examined: CHINA: 1 (NMPC): “Liao che China b.” (JENDEK, 1995 [#3768]); 1 (MNHN): “Chine A. David” (JENDEK, 1995 [#3768]). Beijing: 1 (MNHN): “Env. Pekin Duchaine” (JENDEK, 1995 [#3768]); 1 (AGCM): “China, Beijing, Fragrant Hill, Bot. Garden, 2.VII.1992, L. Borowiec leg.”; 1 (AGCM): “China, Beijing, Bodaling, The Great Wall Area, 5.VIII.1992, L. Borowiec leg.”; 1 ♂ (EJCB): “China, Beijing distr., Yan Shan, Dongling Mts., 39º58’N, E 115º26’E, 1400 m, Xiaolongmen, 15 16.VI.2001”. Shaanxi: 1 (EJCB): “China, Shaanxi province, Mt. Hua Shan, 17 21.VI.1991, leg. R. Sauer” (JENDEK, 1995 [#3768]). Zhejiang: 1 (MNHN): “Chekiang [ Zhejiang province] Hangchow [ Hangzhou Shi] Chine” (JENDEK, 1995 [#3768]); 1 (MNHN): “30. VII.[19]36 O. Piel coll. \ T’jenmu Shan, Museé Heude” (JENDEK, 1995 [#3768]). KAZAKHSTAN: East Kazakhstan: 2 (HHCR): “E. Kazakhstan, S Altaj env. mt. Kamenogorsk, 20.06.1995”. KOREA NORTH: 1 (MNHN): “Coreé” (JENDEK, 1995 [#3768]); 1 (VSCS): “Korea, 15 km südwest Pjôngiang, 8.8.1974, Josifov leg.”; 1 (VSCS): “Korea, Tesôngsan bei Pjôngiang, 23.7.1974, Josifov leg.”. MONGOLIA: 6 (EJCB): “Mongolia or., Chentejský ajmak, dol. Mörön gol, B. Malec lgt. / 5 31.7.1985” (JENDEK, 1995 [#3768]). RUSSIA: Amurskaya oblast’: 2 (ZIN): “Simonovo, Amur. obl., 75 km W Svobodnovo Zinovjev 18.VII.[1]959” (JENDEK, 1995 [#3768]); 4 (AGCM): “Russia, Amurskaja reg., near Kundur vill. 25.7.2001, leg. Pavlov. M.”. Khabarovskiy kray:

150 1 (EJCB): “Russia: Khabarovskiy region: Khekhtsir Mt., 30 km SW Khabarovsk VII.1977, leg. Gotwald” (JENDEK, 1995 [#3768]); 4 (AGCM): “Russia, Khabarovsk reg., near Tarmanchukan st. 30.6.1997, leg. Pavlov. M.”. Primorskiy kray: 1 (EJCB): “Primorsk. krai, Klimoutsy, 40 km W Svobodnyi, 4.VII.1949, leg. Zinovjev” (JENDEK, 1995 [#3768]); 1 (EJCB): “Primorsk. krai, Klimoutsy, 40 km W Svobodnyi, 9.VII.1957, leg. Zinovjev” (JENDEK, 1995 [#3768]); 5 (EJCB): “SU, Ussuri region, Benevskoe, 20 22. VII.1989 leg. S. Jákl, M. Nikodým, R. Červenka” (JENDEK, 1995 [#3768]); 1 ♀ (EJCB): “Russia: Primorskiy region, Dal’nerechensk, 31.VII.1989, 1989 leg. S. Jákl” (JENDEK, 1995 [#3768]); 1 (MNHE): “Russia, 02.07.1989, Primorje Distr., Dukhovskoye, leg. U. Roosileht”; 1 ♀ (EJCB): “Russia, Prrimorsk, Vladivostok, 28.VII.1990, leg. A. Pütz” (JENDEK, 1995 [#3768]); 3 (EJCB): “Russia: Primorskiy region, Novochuguevka, 16 18.VII.1990, leg. M. Snížek” (JENDEK, 1995 [#3768]); 1 (JMCZ): “Ussuri country m., Sergejevka by Chanka, 28 30.7.1990, lgt. D. Boukal”; 8 (EJCB): “Russia, Primorsk, Pogranichnyi, Sergeevka, 29 31.VII.1990, leg. M. Snížek” (JENDEK, 1995 [#3768]); 1 (JMCZ): “SSSR or., 29 31.7., Primorskij Kraj, Pograničnyj, Sergejevka, 1990, lgt. M. Snížek”; 1 ♂ (EJCB): “Sib. or. m., Primorje, Ussuri res., 20.7.1990, Kadlec + Voříšek lg.” (JENDEK, 1995 [#3768]); 1 (EJCB): “Russia, Primorski kray, Khankajski distr. 1995, lake Khanka, 17 19.vii., Kamen’ Rybolov”; 1 (EJCB): “Russia, Primorski kray, Chassanski district, Rjazanovka env., 27.vi. 8.vii.1995”; 6 (FACF): “Russia Far East, S Primorie, Hanka lake, 30.vii.2002”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Preobrazhenie env. 14.07.06 leg.М.& L. Smirnov’s”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Preobrazhenie env., 14.07.06 leg. L. Smirnov”; 1 ♂ (EJCB): “Russia, Far East, Nakhodka, 25.vii.2007, V. Grebennikov”. Zabaykalskiy kray: 4 (EJCB): “Russia: Chita region: Chita, Karinskaya, 29.6.1992, leg. Z. Kletečka” (JENDEK, 1995 [#3768]); 3 (EJCB): “Russia SE Chita rep., 50 km SE of Chita, Makkaveevo, 28.vi.1993, Z. Kletečka leg.”.

Adult occurence: 6-7-8 (n=34). Faunal records published by MIWA & CHÛJÔ, 1940 [#1810] are also included. Altitude range: ~1400 m (n=1). Host plants: Lespedeza: ALEXEEV, 1989 [#64]; ALEXEEV & VOLKOVITSH, 1989 [#3241]. Distribution: CHINA: Beijing, Fujian, Heilongjiang, Nei Mongol, Shaanxi, Xinjiang, Zhejiang. KAZAKHSTAN. KOREA NORTH. MONGOLIA. RUSSIA: Amurskaya oblast’, Buryatiya Respublika, Khabarovskiy kray, Primorskiy kray, Zabaykalskiy kray.

Agrilus peregrinus Kiesenwetter Fig. 15G (habitus); Fig. 34U (aedeagus); Figs. 43V, 50Q (types) peregrinus Kiesenwetter, 1879 (Agrilus) KIESENWETTER, 1879: 145 [#3347] (description). HEYDEN, 1881: 119 [#5548] (catalog). KERREMANS, 1885: 153 [#1381] (catalog). MARSEUL, 1887: 252 [#4707] (catalog). KERREMANS, 1892: 267 [#1397] (catalog). KERREMANS, 1903: 289 [#1441] (catalog). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 657 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1021 [#2191] (world catalog). ALEXEEV, 1979: 129 [#57] (subgenus Arquagrilus; characters; notes; USSR: Primorskiy kray; ?Amur). ALEXEEV, 1989: 485 [#64] (key to species; Primorsk; North Korea). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). JENDEK, 2003: 184 [#5823] (lectotype designation). JENDEK, 2006: 393 [#7441] (subgenus Robertius; Palaearctic catalog). BELLAMY, 2008: 2231 [#8576] (subgenus Robertius; world catalog). VOLKOVITSH,

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA 2009: 136 [#8850] (subgenus Robertius; faunal records; Russia: Primorskiy kray). brodjaga Obenberger, 1914 (Agrilus) OBENBERGER, 1914: 46 47 (in Czech), 52 (in German) [#1965] (description). OBENBERGER, 1926: 657 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1021 [#2191] (variety of peregrinus; world catalog). ALEXEEV, 1979: 129 [#57] (synonym of peregrinus). ALEXEEV, 1989: 485 [#64] (synonym of peregrinus). JENDEK, 2006: 393 [#7441] (synonym of peregrinus; Palaearctic catalog). BELLAMY, 2008: 2231 [#8576] (synonym of peregrinus; world catalog).

Type material: Agrilus peregrinus Kiesenwetter, 1879 Type locality: Amur. Type specimens: Primary: Lectotype (Fig. 50Q), ♀, designated by JENDEK (2003: 184) [#5823], (ZSMC): “Amur [h] \ Kiesenwetter [p] \ Sammlung Cl. Müller [p] \ Type von [p] Agrilus peregrinus Kiesw. [h] [red label] \ Holotypus [p] Agrilus peregrinus Kiesenw. [h] Zool. Staatsammlg. München [red label]”. Described from unknown number of syntypes. Agrilus brodjaga Obenberger, 1914 Type locality: Sibirien: Amurgebiet. Type specimens: Primary: Lectotype (Fig. 43V) by present designation, ♀, (NMPC): “Sibirien Amur Christoph 77 [p] \ Typus [p] [red label] \ Agrilus brodjaga m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24781 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 5.9-7.7 mm; elytra golden-green with dark sutural part and pair of pubescent spot in apical half; aedeagus slightly expanded apically. Species group: Smaragdinus. Relevant species: A. smaragdinus Solsky, 1876. Specimens examined: CHINA: Hebei: 1 ♂, 2 ♀ (EJCB): “China, Hebei/Nei Mongol, pass Chengde Chifeng, 41.6N, 118.2E, 14 16.VI., Jaroslav Turna leg. 2001”; 8 (EJCB): “China, Hebei/Nei Mongol, pass Chengde Chifeng, 41.6N, 118.2E, 14 16.VI.2001, O. Šafránek leg.”. KOREA SOUTH: 2 (JLCS): “Gyeongsan si, Gyeongsanbuk do, Korea, 1.v.1992, coll. Gyeong hwa Kang”; 1 (JLCS): “Sagimakgol, Jungwon gu, Seongnam si, Gyeonggi do, Korea, 1.vi.2001, coll. Hee ah Lee”; 1 (JLCS): “Mt. Cheonggyeasan, Seoul si, Korea, 23.v.2004, coll. Jeong eun Ban”. RUSSIA: Khabarovskiy kray: 1 (ZIN): “SZ Khabarov. okr. Kukan, 25.9.1978 Kabakov [in Russian]”. Primorskiy kray: 1 ♂ (ZIN): “Vladimiro Aleksandrovo, Suchan. Sukhanov 3.vii.1912”; 1 (ZIN): “Yuzhn. Primor’e, Suputinskii zapovednik 4.6.1969 Kabakov [in Russian]”; 1 ♂ (MNHE): “Russia, 09 11.07.1989, Primorye Distr., Kamenushka, leg. U. Roosileht”; 1 ♀ (EJCB): “USSR, Primorsk. kraj, Arsenev env., VI.1991, 44.11N, 133.19E, O. Šausa leg.”; 2 ♀ (EJCB): “Sib. or., Primorje, Komarovka flum., Kamenushka env., 300 m, Vořišek lgt., V.1992”; 1 (HMMG): “Russia, Prim., Partizansk distr. Alexeyevski khreb., 20km E Sergeyevka, forest near Andreyevka river 400 m, 26 29.vii.1993, Pütz & Wrase”; 3 ♂ (EJCB): “Russia E, Primorje reg, Anisimovka, 7 13.vii.1996, Mt. Litovka, 200 m, Plutenko leg.”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., 12 кm S Staraja Kamenka, 10.07.04 leg. М.& L. Smirnov’s”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Lazo env., 19.07.06 leg. М.& L. Smirnov’s”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Tachingousa (Prosiolotchnaja) bay 1 3.07.06 leg. М.& L. Smirnov’s”; 1 ♂ (EJCB): “Russia, Ussuri Primor’e region, Pedan Mts., 28 29.7.2006, leg. K. Hadulla”.

Adult occurence: 5-6-7-9 (n=17). Altitude range: 200-400 m (n=3).

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Host plants: Alnus japonica: ALEXEEV, 1979 [#57]. − Ulmus: ALEXEEV, 1989 [#64]. Distribution: CHINA: Hebei, Nei Mongol. KOREA NORTH. KOREA SOUTH. RUSSIA: Amurskaya oblast’, Primorskiy kray.

Agrilus perroti Descarpentries & Villiers Fig. 20B (habitus of HT); Fig. 36O (aedeagus) perroti Descarpentries & Villiers, 1963 (Agrilus) DESCARPENTRIES & VILLIERS, 1963: 108, 118 [#700] (description). PENG ZHONGLIANG, 2002: 269, 281 [#6628] (characters; new record for China; Fujian). JENDEK, 2006: 400 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2233 [#8576] (world catalog).

Type material: Agrilus perroti Descarpentries & Villiers, 1963 Type locality: Tonkin: Thanh-Moï. Type specimens: Primary: Holotype (Fig. 20B), ♀, (MNHN): “Thanh-Moi Tonkin H. Perrot [p] \ Agrilus perroti n. sp. Holotype ♀ nob. A. Descarpentries et [h] A. Villiers det. 19 [p] 62 [h] \ Muséum Paris Coll. générale [p] [yellow label]”. Described from 1 specimen (HT). Diagnosis: 10-14 mm; frons and vertex medially less impressed; vertex obscurely pubescent; preapical transverse pubescent stria on elytra wide, not undulated; margins of elytral apices distinctly denticulated. Species group: Occipitalis. Relevant species: A. zanthoxylumi Li Meng Lou, 1989. Specimens examined: CHINA: Guangxi: 1 ♀ (EJCB): “Gangxi”. Yunnan: 1 ♂ (NMPC): “Yun’nan’, 30 km SW Czin’pina, 400m, 24.iv.1956, Khuan Kè zhen’ i dr. [in Russian]”.

Adult occurence: 4 (n=1). Altitude range: ~400 m (n=1). Host plants: Unknown. Distribution: VIETNAM.

CHINA:

Fujian,

Guangxi,

Yunnan.

Agrilus pilosovittatus Saunders Fig. 31B (habitus); Fig. 41P (aedeagus); Figs. 50S, 51N, 53D, 54W (types) pilosovittatus Saunders, 1873 (Agrilus) SAUNDERS, 1873: 515 [#2554] (description). LEWIS, 1879: 15 [#3345] (catalog; Japan). KERREMANS, 1885: 153 [#1381] (catalog). SCHÖNFELDT, 1887: 113 [#8901] (Anambus; catalog; Japan). KERREMANS, 1892: 267 [#1397] (catalog). KERREMANS, 1903: 276 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 19 [#1809] (catalog; Japan). OBENBERGER, 1936: 1022 [#2191] (world catalog). CHÛJÔ & MATUDA, 1940: 65 [#500] (checklist; faunal records; Japan (Kyushu)). IGA, 1947: 17 [#3273] (notes). KUROSAWA, 1949: 15 [#5028] (cited as pilosovittus; characters). CHÛJÔ & KUROSAWA, 1950: 11 [#501] (catalog; faunal records; Japan (Shikoku)). KUROSAWA, 1950: 1114 [#3274] (note). IGA, 1955: 80 [#3828] (iconography; Japan). KUROSAWA, 1956: 1114 [#8107] (characters; Japan). IGA, 1962: 80 [#3351] (iconography; Japan). KUROSAWA, 1963: 154 [#1618] (characters; Japan). HESPENHEIDE, 1968: 77 80 [#1233] (new

151 immigrant to USA; characters; USA: New Jersey, New York, Pennsylvania). KUROSAWA, 1974: 2 [#4575] (characters; notes). KUROSAWA, 1974: 4 [#4577] (note). TÔYAMA, 1985: 22 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). PENG ZHONGLIANG, 1992: 400 [#2431] (Pallas is cited as the author; characters; notes; Hunan). JENDEK, 1995: 138 139 [#3768] (lectotype designation; synonymy; charaters; Japan; China: Jiangxi). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 185 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 37 [#4091] (checklist; Japan: Honshu, Shikoku, Kyushu, Tsushima; China; USA (culturized)). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). NONNAIZAB et al., 1999: 113 [#8247] (checklist; China: Inner Mongolia). AKIYAMA & OHMOMO, 2000: 88 [#6240] (faunal records; Japan (Hiroshima)). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Hunan). GATES, 2004: 3 [#7136] (new parasitoid Henryana magnifica (Hymenoptera, Chalcidoidea); USA (Maryland)). JENDEK, 2006: 393 [#7441] (subgenus Robertius; Palaearctic catalog). ANONYMOUS, 2007: 32 [#8250] (checklist; Japan: Honshu). BELLAMY, 2008: 2236 [#8576] (world catalog). JENDEK & GREBENNIKOV, 2009: 243 [#8724] (notes). zemani Obenberger, 1925 (Agrilus) OBENBERGER, OBENBERGER, 1925: 34 [#2048] (description). 1926: 655 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 21 [#1809] (catalog; Japan). OBENBERGER, 1936: 1022 [#2191] (synonym of pilosovittatus; world catalog). HESPENHEIDE, 1968: 77 [#1233] (synonym of pilosovittatus). PENG ZHONGLIANG, 1992: 400 [#2431] (synonym of pilosovittatus). JENDEK, 1995: 138, 139 [#3768] (synonym of pilosovittatus; lectotype designation). AKIYAMA & OHMOMO, 1997: 37 [#4091] (synonym of pilosovittatus). HUA LI ZHONG, 2002: 90 [#6696] (synonym of pilosovittatus). JENDEK, 2006: 393 [#7441] (synonym of pilosovittatus; Palaearctic catalog). BELLAMY, 2008: 2236 [#8576] (synonym of pilosovittatus; world catalog). spesivcevi Obenberger, 1927 (Agrilus) syn. nov. OBENBERGER, OBENBERGER, 1927: 16 [#2060] (description). 1936: 1049 [#2191] (cited as spessivcevi; world catalog). PENG ZHONGLIANG, 1987: 358 [#2429] (cited as spessivcevi; checklist; China). ALEXEEV & VOLKOVITSH, 1989: 319, 347, 356 [#3241] (cited as spessivcevi; characters in key; faunal records; Mongolia). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). HUA LI ZHONG, 2002: 90 [#6696] (cited as spessivcevi; checklist; N China). JENDEK, 2005: 19 20 [#7217] (correct original spelling is spesivcevi; [Note: ICZN, Article 32.5.1]; lectotype designation). JENDEK, 2006: 394 [#7441] (subgenus Robertius; Palaearctic catalog). BELLAMY, 2008: 2302 [#8576] (subgenus Robertius; world catalog). pusillipubis Gebhardt, 1929 (Agrilus) OBENBERGER, GEBHARDT, 1929: 94 95 [#3349] (description). 1936: 1026 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 357 [#2429] (checklist; China). JENDEK, 1995: 138, 139 [#3768] (synonym of pilosovittatus; lectotype designation). AKIYAMA & OHMOMO, 1997: 37 [#4091] (synonym of pilosovittatus). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Jiangsu, Zhejiang). JENDEK, 2006: 393 [#7441] (synonym of pilosovittatus; Palaearctic catalog). BELLAMY, 2008: 2236 [#8576] (synonym of pilosovittatus; world catalog).

Type material: Agrilus pilosovittatus Saunders, 1873 Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 50S), ♂, designated by JENDEK (1995: 138-139) [#3768], (BMNH): “Type H.T. [p] [round label with red border] \ Japan G. Lewis 1910-320 [p] \ Agrilus tempestivus Lewis Type [h]”.

152 Secondary: 1 PLT (BMNH). Described from unknown number of syntypes. Agrilus zemani Obenberger, 1925 Type locality: Japan: Kobe. Type specimens: Primary: Lectotype (Fig. 54W), ♂, designated by JENDEK (1995: 138, 139) [#3768], (NMPC): “Kobe -1918 Japonsko Dr. Jureček [p] \ Typus [p] [red label] \ Agrilus Zemani m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24884 [h] [orange label]”. Secondary: 1 PLT (NMPC). Described from unknown number of syntypes. Agrilus spesivcevi Obenberger, 1927 Type locality: China. bor. or.: Lac Tai-hu. Type specimens: Primary: Lectotype (Fig. 53D), ♀, designated by JENDEK (2005: 19-20) [#7217], (NMPC): “China: Lac Tai-hu [h] \ Typus [p] [red label] \ Agrilus spessivcevi [sic!] m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24938 [h] [orange label]”. Described from unknown number of syntypes. Agrilus pusillipubis Gebhardt, 1929 Type locality: China: Soochow, Hangchow. Type specimens: Primary: Lectotype (Fig. 51N), ♂, designated by JENDEK (1995: 138, 139) [#3768], (NMPC): “China Hangchow [ = Hanghzou, Zhejiang province] [p] \ Typus [p] [red label] \ Agrilus pusillipubis m. Type [h] Det. Dr. A. v. Gebhardt [p]”. Secondary: PLT (MNHN); PLT (HNHM). Described from unknown number of syntypes. Diagnosis: 4.3-5 mm; prehumerus arcuate carinal, reaching to half of pronotum, anterior end distant from pronotal marginal carina. Species group: Unassigned. Relevant species: A. improcerus sp. nov. Specimens examined: CHINA: Jiangxi: 1 (EJCB): “Kiang Si [ Jiangxi province]” (JENDEK, 1995 [#3768]). JAPAN: Honshu: 1 (EJCB): “Hattoji, Okayama Prefecture, 28.V. 1979, leg. H. Jamazaki” (JENDEK, 1995 [#3768]); 3 (JMCZ): “Japan, 10.5.1979, Shishizuka, Tsuchiura city, K. Akiyama lgt.”; 1 (EJCB): “Koshiwa City, Chiba Prefecture, Japan, 25.V. 1982, leg. N. Kobayashi” (JENDEK, 1995 [#3768]); 6 (EJCB): “Japan, Tsukubasan Mt., Ibaraki Prefecture, 17.VI.1984, leg. S. Ohmomo” (JENDEK, 1995 [#3768]). KOREA NORTH: 1 (EJCB): “Korea, Sujang san bei Hedzu, 4.6.1975, Josifov”.

Adult occurence: 5-6 (n=7). Faunal records published by CHÛJÔ & MATUDA, 1940 [#500] are also included. Host plants: Wisteria floribunda: AKIYAMA & OHMOMO, 1997 [#4091]. − W. sinensis: HESPENHEIDE, 1968 [#1233]; GATES, 2004 [#7136]. Erroneous: Caragana spinosa: ALEXEEV & VOLKOVITSH, 1989 (as spesivcevi) [#3241]; Fagus: HUA LI ZHONG, 2002 [#6696]. Distribution: CHINA: Jiangsu, Jiangxi, Zhejiang. JAPAN: Honshu, Kyushu, Shikoku, Tsushima. KOREA NORTH. NORTH AMERICA (introduced): UNITED STATES. Remarks: Distributional records without supporting data from Hunan and Nei Mongol cited by Chinese authors are omitted. Records under the name of A. spesivcevi cited by ALEXEEV & VOLKOVITSH, 1989 [#3241] and ALEXEEV, 1998 [#4322] are misidentifications of A. ulaangomiensis Cobos, 1972.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Agrilus plagiatus Ganglbauer Fig. 13E (habitus); Fig. 33S (aedeagus); Figs. 49H, 50V (types) plagiatus Ganglbauer, 1889 (Agrilus) GANGLBAUER, 1889: 22, 31 33 [#3427] (description). HEYDEN, 1893: 90 [#3450] (catalog). KERREMANS, 1903: 289 [#1441] (catalog). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 657 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1022 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 357 [#2429] (checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Gansu). JENDEK, 2006: 400 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2238 [#8576] (world catalog). mentor Kerremans, 1912 ( Agrilus ) syn. nov. KERREMANS, 1912: 207 [#1377] (description). MIWA, 1931: 124 [#6101] (catalog; Formosa). MIWA & CHŰJÔ, 1936: 18 [#1809] (catalog; Japan). OBENBERGER, 1936: 1091 [#2191] (world catalog). CHŰJÔ & KUROSAWA, 1950: 9 [#501] (synonym of discalis ; catalog). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Taiwan). MÜHLE, 2003: 47 [#6155] (checklist; Taiwan). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2185 [#8576] (world catalog).

Type material: Agrilus plagiatus Ganglbauer, 1889 Type locality: Kan-ssu [Fluss Hei-Ho] [Note: Locality was complemented from the list of localities on page 21-22]. Type specimens: Holotype by monotypy (Fig. 50V), ♀, (ZIN): “Kan-ssu 1885 G. Patanin [sic!] [p] 21. VI. 85 [h] [underside] \ Agrilus plagiatus Gglb. typ [h] \ Coll. SemenovTian-Shansky [p]”. Described from 1 specimen. Agrilus mentor Kerremans, 1912 Type locality: [Formosa] Taihorinsho. Type specimens: Primary: Lectotype (Fig. 49H) by present designation, ♂, (MNHN): “Taihorinsho Formosa H. Sauter [p] \ Mentor Kerr Type [h] \ Museum Paris Coll. Ch. Kerremans 1923 [p][yellow label]”. Described from unknown number of syntypes. Diagnosis: 5.5-7 mm; prehumerus carinal at least at posterior pronotal angle; elytral apices separately arcuate; male with impressed prosternal process with elevated sides. Species group: Plagiatus. Relevant species: A. pumi sp. nov.; A. stigma sp. nov. Specimens examined: CHINA: Shaanxi: 1 ♀ (EJCB): “China S, Shaanxi, 14 16.6.1995, Qinling Mts. N. slope, 33.56 59’/108.50’ 800 1200 m, Changan Co., L.+R. Businský lgt.”; 1 ♀ (EJCB): “China, Shaanxi prov., 21 23.June 1998 \ Qing Ling Shan mts, 1000 m road Baoji Taibai vill., pass 40 km S Baoji, Zd. Jindra lgt.”; 7 (EJCB): “China, Shaanxi prov., 21 23.vi.1998, Quin Ling Shan mts, road Baoji Taibai vill., pass 35 km S Baoji, O. Šafránek et M. Trýzna lgt.”. Sichuan: 2 ♂, 2 ♀ (EJCB): “China, Sichuan, 8 13.VI.[19]91, Abazhou, 2000 m, Nanping, Jiuzhaigou, 33.15N / 104.15”; 1 (EJCB): “China, Sechuan, Nanping, 6 9.6.2001, leg. E. Kučera”.

Adult occurence: 6 (n=5). Altitude range: 800-2000 m (n=3). Host plants: Unknown. Distribution: CHINA: Shaanxi, Sichuan, Taiwan. Remarks: The distributional record without supporting data

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA cited from Gansu (China) by HUA LI ZHONG, 2002 [#6696] was omitted.

Agrilus planipennis Fairmaire Fig. 8D (habitus); Fig. 32B (aedeagus); Figs. 46D, 49A, 50X, 54G (types) planipennis Fairmaire, 1888 (Agrilus) FAIRMAIRE, 1888: 121 [#887] (description). KERREMANS, 1892: 268 [#1397] (catalog). KERREMANS, 1903: 289 [#1441] (catalog). JAKOBSON, 1913: 796 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1022 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 357 [#2429] (checklist; China). JENDEK, 1994: 21 [#1355] (lectotype designation; synonymy; Russian Far East; Mongolia; Japan; China; Taiwan). AKIYAMA & OHMOMO, 1997: 37 [#4091] (checklist; Mongolia; North China; Korea). ALEXEEV, 1998: 372 [#4322] (subgenus Uragrilus; subgeneric classification). AKIYAMA & OHMOMO, 2000: 275 [#5370] (characters; distributional summary; host plants). HAACK et al., 2002: 1 5 [#5632] (new pest in America; USA: Michigan). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Beijing). MCCULLOUGH & ROBERTS, 2002: 1 2 [#5726] (introduction to Michigan; characters; biology; United States). SCARR et al., 2002: 1 4 [not paginated] [#5981] (life cycle). HERMS et al., 2003: 2 10 [#6801] (economic and ecological impact; taxonomy; biology; diagnosing; plan to eradicate; USA: Michigan, Ohio, Maryland; Canada: Ontario). LIU HOUPING et al., 2003: 191 203 [#7112] (exploratory survey; natural enemies; China (Hebei, Harbin, Heilongjiang, Jilin, Liaoning, Tianjin)). MÜHLE, 2003: 44, 47 [#6155] (checklist; Taiwan). BARTELS, 2004: 19 [#6781] (remote sensing for detection of infestations). BAUER et al., 2004: 36 37 [#6794] (laboratory rearing). BAUER et al., 2004: 8 [#6772] (life cycle). BAUER et al., 2004: 31 32 [#6791] (microbial control). BAUER et al., 2004: 33 34 [#6792] (natural enemies: fungi, hymenopteran parasitoids). BAUER et al., 2004: 9 [#6773] (flight potential). BEAN, 2004: 4 [#6769] (detection and quarantine; Maryland). BROWN RYTLEWSKI & WILSON, 2004: 11 [#6775] (emergence of adult). FRANCESE et al., 2004: 22 [#6784] (survey tools). FUKUTOMI & HORI, 2004: 14 [#6851] (checklist; faunal records; Japan (Hokkaido)). GATES & GIBSON, 2004: 12 [#7132] (new parasitoid of Mymaromatidae (Hymenoptera, Chalcidoidea); USA (Michigan)). GOULD et al., 2004: 35 [#6793] (biocontrol; parasitoids and predators attacking related borers). HAACK & PETRICE, 2004: 26 [#6787] (tests of insecticides). HAACK & PETRICE, 2004: 23 [#6785] (survival in firewood). HAACK & PETRICE, 2004: 14 [#6778] (attraction to trap logs). HAACK & PETRICE, 2004: 10 [#6774] (adult dispersal; Michigan). HAACK et al., 2004: 38 [#6795] (host range evaluation by foliage of several trees and shrubs as food for adults). HERMS et al., 2004: 41 [#6797] (resistance of Asian and North American ashes). LEWIS, 2004: 29 [#6789] (control by trunk injection). LIU HOUPING et al., 2004: 20 [#6782] (natural enemy complex; 50 parasitation of larvae by Spathius sp. (Hymenoptera, Braconidae); China: Heilongjiang; Jilin, Liaoning; Hebei, Tianjin; unconfirmed in Shandong). LYONS et al., 2004: 5 [#6770] (biology and phenology). MACFARLANE & FRIEDMAN, 2004: 21 [#6783] (spatial distribution and abundance model). MARCHANT, 2004: 3 [#6768] (delimitation and detection survey; Ontario). MCCULLOUGH et al., 2004: 39 40 [#6796] (host range and host preference; alternate hosts). MCCULLOUGH et al., 2004: 27 28 [#6788] (control of adults and larvae with insecticides). MCCULLOUGH et al., 2004: 6 7 [#6771] (dispersal of one generation; Michigan). MCCULLOUGH et al., 2004: 24 25 [#6786] (survival in wood chips). OLIVER et al., 2004: 17 18 [#6780] (traps). POLAND et al., 2004: 15 16 [#6779] (attractants and trapping techniques). RAUSCHER & MASTRO, 2004: 1 2 [#6767] (eradication plan;

153 Michigan, Ohio, Virginia, Maryland). REARDON, 2004: 30 [#6790] (natural enemies). SCHAEFER, 2004: 13 [#6777] (faunal record; Japan: Honshu). SCHRÖDER, 2004: 25 26 [#6857] (placed on list of regulated pests by EPPO; Germany). SCHRÖDER, 2004: 222 226 [#7072] (overview on biology and risk analysis for Europe). SMITH et al., 2004: 12 [#6776] (genetic analysis to determinate point of origin in North American infestation). TIMMS et al., 2004: 42 [#6798] (susceptibility and within tree distribution). WEI et al., 2004: 679 685 [#7133] (characters; biology; parasitoids; distribution; damages; China (Harbin, Heilongjiang, Jilin, Liaoning, Shandong, Hebei, Tianjin)). ANONYMOUS, 2005: 436 438. [#8844] (quarantine pest datasheet). CAPPAERT et al., 2005: 152 165 [#7891] (research history in North America; regulatory challenge). CHORNESKY et al., 2005: 336 337 [#8289] (significance to sustianable forestry; USA). FRANCESE et al., 2005: 93 95 [#7110] (color traps evaluation). GIBSON, 2005: 31 36 [#7138] (parasitoid Balcha indica (Hymenoptera: Chalcidoidea: Eupelmidae) accidental introduction from the Oriental region). LIU HAI JUN et al., 2005: 348 352 [#7282] (biological characteristics). LIU HOUPING & BAUER, 2005: 594 598 [#8298] (host range). MACFARLANE & MEYER, 2005: 15 23 [#7149] (potential ash tree hosts). WANG XIAOYI et al., 2005: 98 102 [#7275] (larval instars). YANG ZHONGQI et al., 2005: 636 641 [#7263] (new species of parasitoid Spathius agrili). ZHANG YAN ZHOU et al. 2005: 253 259 [#7126] (new species of egg parasitoids). GROOT et al., 2006: 1 16 [#8068] (life stages; signs and symptoms of infested trees; measures; Canada). HAACK, 2006: 270, 271 [#8288] (summary data about introduction to North America). JENDEK, 2006: 395 [#7441] (subgenus Uragrilus; Palaearctic catalog). LIU HOUPING & BAUER, 2006: 1096 1103 [#8283] (susceptibility to entomopathogenic fungi Beauveria and Metarhizium). OHMOMO, 2006: 6 [#8002] (checklist; Japan). RODRIGUEZ SAONA et al., 2006: 75 84 [#8239] (host plants findings; induced volatiles). BARTELT et al., 2007: 1299 1301 [#8241] (pheromonal function of lactone). LELITO et al., 2007: 537 551 [#8171] (trapping; mating behavior). LIU HOUPING et al., 2007: 61 71 [#8183] (seasonal abundance; population dynamics; biological control; hymenopteran parasitoids: Oobius agrili (Encyrtidae); Tetrastichus planipennisi (Eulophidae); northeastern China (Liaoning, Jilin)). LYONS et al., 2007: 1 58 [#8066] (biology; life cycle; host plants; monitoring; Canada). MCCULLOUGH & SIEGERT, 2007: 1577 1585 [#8280] (estimated surface area of infested plants; eradication). MCCULLOUGH et al., 2007: 1304 1315 [#8285] (phytosanitary treatment). MOZOLEVSKAYA, 2007: 137 138 [#8376] (new pest discovered in 2004; Russia: Moscow region). YURCHENKO, TUROVA & KUZMIN, 2007: 94 98 [#8892] (faunal records; parasitoids; Russia: Primorsk, Khabarovsk). WEI XIA et al., 2007: 367 372 [#8182] (damage traits; life history; northeastern China (Heilongjiang, Tianjin, Jilin)). ZHAO TONG HAI et al., 2007: 826 831 [#8297] (induced outbreak by an introduction of exotic tree species; China). ANULEWICZ et al., 2008: 230 240 [#8180] (host range; oviposition preference; host selection). BELLAMY, 2008: 2238 2239 [#8576] (subgenus Uragrilus; world catalog). CROOK et al., 2008: 1103 1111 [#8284] (antennal sensilla structure). LELITO et al., 2008: 668 673 [#8728] (effectivity of the visual cue based sticky traps). LINDELL et al., 2008: 434 442 [#8172] (woodpecker predation). REBEK et al., 2008: 242 245 [#8181] (host plant resistance). SMITLEY et al., 2008: 1643 1650 [#8281] (progression ash canopy thinning). VASANTHAKUMAR et al., 2008: 1344 1351 [#8292] (gut microbial communities). WESSELS BERK & SCHOLTE, 2008: 165 168 [#8544] (phytosanitary risk; Europe). CAPPAERT & MCCULLOUGH, 2009: 16 28 [#8896] (occurence, seasonal abundance of parasitoid Atanycolus planipennis; USA: Michigan, Ohio). DUAN et al., 2009: 588 592 [#8842] (parasitoids; biocontrol) JENDEK & GREBENNIKOV, 2009: 243 [#8724] (notes). MARSH, STRAZANAC & LAURUSONIS, 2009: 8 14 [#8895] (new species of parasitoid Atanycolus cappaerti

154 (Hymenoptera, Braconidae)). VOLKOVITSH, 2009: 136 [#8850] (subgenus Uragrilus; faunal records; notes; Russia: Primorskiy kray). POLAND & MCCULLOUGH, 2010: 4 8 [#8898] (biological control). BARANCHIKOV, 2010: 870 871 [#8916] (biology; habitat; introduced range; pathways; impact; management). marcopoli Obenberger, 1930 (Agrilus) OBENBERGER, 1930: 108 109 [#2088] (description). MIWA & CHÛJÔ, 1940: 74 [#1810] (cited as malcopoli; faunal record; Japan: Sapporo). IGA, 1955: 78 [#3828] (iconography; Japan). KUROSAWA, 1956: 40 [#1614] (faunal records; Korea). IGA, 1962: 78 [#3351] (iconography; Japan). KUROSAWA, 1963: 153 [#1618] (characters; Japan). KUROSAWA, 1974: 3 [#4577] (notes). ALEXEEV, 1979: 127 128 [#57] (subgenus Anambus; cited as markopoli; faunal records; characters; Russia; northeastern China). TÔYAMA, 1985: 23 [#3678] (iconography; Japan). ALEXEEV, 1989: 482 [#64] (key to species; Primorsk; Japan; northeastern China). LI JINGKE, 1992: 92 [#6755] (checklist; northeastern China). YU CHENGMIN, 1992: 400 401 [#5982] (characters; biology; China: Heilongjiang; Jilin; Liaoning; Shandong; Inner Mongolia; Mongolia; Korea; Japan). JENDEK, 1994: 21 [#1355] (synonym of planipennis; lectotype designation). AKIYAMA & OHMOMO, 1997: 37 [#4091] (synonym of planipennis). NONNAIZAB et al., 1999: 113 [#8247] (checklist; China: Inner Mongolia). HUA LI ZHONG, 2002: 90 [#6696] (cited as macropoli; checklist; China: Inner Mongolia, Hebei, Shandong). XU GONGTIAN, 2003: 321 [#6836] (biology; instars). JENDEK, 2006: 395 [#7441] (synonym of planipennis; Palaearctic catalog). BELLAMY, 2008: 2238 [#8576] (synonym of planipennis; world catalog). feretrius Obenberger, 1936 (Agrilus) MIWA & OBENBERGER, 1936: 37 38 [#2200] (description). CHÛJÔ, 1936: 16 [#1809] (catalog; Japan). OBENBERGER, 1936: 1105 [#2191] (cited as teretrius; world catalog). KUROSAWA, 1974: 3 [#4577] (cited as teretrius; related to A. marcopoli). PENG ZHONGLIANG, 1987: 355 [#2429] (checklist; China). JENDEK, 1994: 21 [#1355] (synonym of planipennis; lectotype designation). AKIYAMA & OHMOMO, 1997: 37 [#4091] (synonym of planipennis). HUA LI ZHONG, 2002: 89 [#6696] (cited twice, as feretrius on page 89 and as teretrius on page 90; checklist; China: Taiwan). MÜHLE, 2003: 47 [#6155] (cited as teretrius; checklist; Taiwan). JENDEK, 2006: 395 [#7441] (synonym of planipennis; Palaearctic catalog). BELLAMY, 2008: 2238 [#8576] (synonym of planipennis; world catalog). ulmi Kurosawa, 1956 (Agrilus; subspecies of marcopoli) KUROSAWA, KUROSAWA, 1956: 40 41 [#1614] (description). 1963: 153 [#1618] (subspecies of marcopoli; notes; Japan). KUROSAWA, 1974: 3 [#4577] (subspecies of marcopoli; notes). KUROSAWA, 1974: 1 [#4575] (characters; notes). ALEXEEV, 1979: 127 [#57] (subspecies of marcopoli cited as markopoli; notes). TÔYAMA, 1985: 23 [#3678] (subspecies of marcopoli; iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (subspecies of marcopoli; checklist; Japan). JENDEK, 1994: 21 [#1355] (synonym of planipennis). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (subspecies of marcopoli; checklist; Japan). AKIYAMA & AKIYAMA, 1996: 185 [#3962] (subspecies of planipennis; faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 5, 37 [#4091] (subspecies of planipennis; checklist). AKIYAMA & OHMOMO, 2000: 275 [#5370] (subspecies of planipennis; faunal records; Japan). JENDEK, 2006: 395 [#7441] (synonym of planipennis; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (synonym of planipennis). BELLAMY, 2008: 2238 [#8576] (synonym of planipennis; world catalog).

Type material: Agrilus planipennis Fairmaire, 1888 Type locality: Pekin.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Type specimens: Primary: Lectotype (Fig. 50X), ♀, designated by JENDEK (1994: 21) [#1355], (MNHN): “Chine A. David [p] \ Agrilus planipennis Fairm. [by Fairmaire’s hand]”. Described from unknown number of syntypes. Agrilus marcopoli Obenberger, 1930 Type locality: Mongolia or.: Chan-Heu. Type specimens: Primary: Lectotype (Fig. 49A), ♂, designated by JENDEK (1994: 21) [#1355], (NMPC): “Mongol. or. Chan-heou [p] \ Typus [p] [red label] \ Agrilus MarcoPoli m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Agrilus feretrius Obenberger, 1936 Type locality: Formosa. Type specimens: Primary: Lectotype (Fig. 46D), ♀, designated by JENDEK (1994: 21) [#1355], (NMPC): “Formosa H. Sauter, 1911 [p] \ Typus [p] [red label] \ Agrilus Teretrius [sic!] m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Agrilus marcopoli ulmi Kurosawa, 1956 Type locality: Sapporo. Hokkaidô. Type specimens: Primary: Holotype, ♂, (NSMT). Secondary: PT (NSMT), 1 PT (Fig. 54G) (MNMS). Described from 13 specimens (HT, AT, PT). Diagnosis: 12-15 mm; body robust, golden-green; eyes large; pronotum widest at base; prehumerus tubercular. Species group: Cyaneoniger. Relevant species: A. cyaneoniger Saunders, 1873. Specimens examined: CHINA: Heilongjiang: 1 (EJCB): “Primorsk 13.8.1935 Mai He [river]” (JENDEK, 1994 [#1355]). Nei Mongol: 2 ♀ (EJCB): “Chahar Yangkiaping 8 10.VII.1937” (JENDEK, 1994 [#1355]). Sichuan: 1 (NMPC): “Szechuan” (JENDEK, 1994 [#1355]). Tianjin: 1 (MNHN): “Tianjin” (JENDEK, 1994 [#1355]). JAPAN: Honshu: 1 (EJCB): “Ogawadani Vall., Okutuma, Tokyo Met., 2.VI.1964, J. Komiya” (JENDEK, 1994 [#1355]); 1 (MNCA): “Japan, Todai Nagano, 16.VIII.1972, S. Okuno” (JENDEK, 1994 [#1355]); 1 (EJCB): “1981.VIII.1, Koganezawa, Yamanashi Pref. Ikeda leg.” (JENDEK, 1994 [#1355]); 1 (MNCA): “Koganezawa, Ohtsuki City, Yamanashi Pref. 4.VIII.81 leg. M. Horikawa” (JENDEK, 1994 [#1355]). LAOS: Xiangkhoang: 2 ♀ (EJCB): “Laos NE, Xieng Khouang prov., 45 km (by road) E of Phonsavan: ~1000 m, Ban Namseung, April 2008, Ch. Keomaravong leg.”. RUSSIA: Primorskiy kray: 1 ♀ (EJCB): “SU Ussuri reg., Benevskoe, 20.7.1989, M. Nikodým leg.” (JENDEK, 1994 [#1355]); 1 ♂ (JMCZ): “Ross. or., Prim. kr., Novočugujevka, 26 31.vii.1992, Snížek leg.”; 1 (ZIN): “Yu Primor’e Chernye gory, prav. pritok r. Poima, 30.vii. 5. viii.1999 [in Russian]”. UNITED STATES: Michigan: 5 (EJCB): “USA, Michigan, Detroit env., vi.2002, on Fraxinus”.

Adult occurence: 4-6-7-8 (n=21). Faunal records published by AKIYAMA & OHMOMO, 2000 [#5370] (as planipennis ulmi); ALEXEEV, 1979 [#57] (as marcopoli); KUROSAWA, 1956 [#1614] (as marcopoli and marcopoli ulmi); MIWA & CHÛJÔ, 1940 [#1810] (as marcopoli) and SCHAEFER, 2004 [#6777] are also included. Altitude range: 800-1000 m (n=2). Host plants: Fraxinus: MÜHLE, 2003 [#6155]; GATES & GIBSON, 2004 [#7132]; WEI XIA et al., 2007 [#8182]. – F. americana: MCCULLOUGH & ROBERTS, 2002 [#5726]; HERMS et al., 2003 [#6801]; LIU HOUPING & BAUER, 2005 [#8298]. – F. chinensis: YU CHENGMIN, 1992 (as marcopoli) [#5982]; WEI et al., 2004 [#7133]; LIU HOUPING et al., 2007 [#8183].

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA – F. mandshurica: YU CHENGMIN, 1992 (as marcopoli) [#5982]; WEI et al., 2004 [#7133]; LIU HOUPING & BAUER, 2005 [#8298]; RODRIGUEZ SAONA et al., 2006 [#8239]; LIU HOUPING et al., 2007 [#8183]. – F. mandshurica var. japonica: AKIYAMA & OHMOMO, 1997 (as ulmi) [#4091]. – F. nigra: MCCULLOUGH & ROBERTS, 2002 [#5726]; HERMS et al., 2003 [#6801]; LIU HOUPING & BAUER, 2005 [#8298]. – F. pennsylvanica: MCCULLOUGH & ROBERTS, 2002 [#5726]; HERMS et al., 2003 [#6801]; LIU HOUPING et al., 2007 [#8183]. – F. pennsylvanica var. subintegerrima: LIU HOUPING & BAUER, 2005 [#8298]. – F. quadrangulata: HERMS et al., 2003 [#6801]. – F. rhynhophylla: YU CHENGMIN, 1992 (as marcopoli) [#5982]; WEI et al., 2004 [#7133]; LIU HOUPING & BAUER, 2005 [#8298]; LIU HOUPING et al., 2007 [#8183]. – F. velutina: LIU HOUPING & BAUER, 2005 [#8298]. – Juglans: MÜHLE, 2003 [#6155]. – J. mandshurica var. sieboldiana: AKIYAMA & OHMOMO, 1997 (as ulmi) [#4091]. – Pterocarya: MÜHLE, 2003 [#6155]. – P. rhoifolia: AKIYAMA & OHMOMO, 1997 (as ulmi) [#4091]. – Ulmus: ALEXEEV, 1989 (as marcopoli) [#64]; MÜHLE, 2003 [#6155]. – U. davidiana var. japonica: AKIYAMA & OHMOMO, 1997 (as ulmi) [#4091]. – U. propinqua: KUROSAWA, 1956 (as marcopoli ulmi) [#1614]. Distribution: CHINA: Beijing, Hebei, Heilongjiang, Jilin, Liaoning, Nei Mongol, Shandong, Sichuan, Tianjin. JAPAN: Hokkaido, Honshu, Kyushu, Shikoku. KOREA SOUTH. LAOS: Xiangkhoang. MONGOLIA. RUSSIA: Moscow region (introduced), Primorskiy kray. NORTH AMERICA (introduced): CANADA. UNITED STATES. Remarks: Among the east-Asian species, A. planipennis Fairmaire, 1888 is most similar and likely related to the members of Cyaneoniger group, but outside this region it seems to have closer relation to other robust and mainly undescribed species from the Oriental region. Specimens from Laos (see Specimens examined) differ from specimens collected from other countries by its sharper prehumerus, white perisutural elytral pubescence and by two whitish, adsutural spots at apical fourth of elytra.

Agrilus plasoni plasoni Obenberger Fig. 12D (habitus); Fig. 33N (aedeagus); Figs. 47P, 51A (types) plasoni Obenberger, 1917 (Agrilus) OBENBERGER, 1917: 212 213 [#1997] (description). OBENBERGER, 1926: 660 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1022 [#2191] (world catalog). OBENBERGER, 1940: 173 174 [#2273] (type locality emended to “China, Fokien. (China bor. occ.)”; faunal records). OBENBERGER, 1958: 237 [#2389] (faunistic redords; China). HOU, 1987: 589 590 [#3490] (notes; Yunnan). PENG ZHONGLIANG, 1987: 357 [#2429] (checklist; China). HOU, 1988: 279 [#4900] (checklist; China: Xizang). PENG ZHONGLIANG, 1992: 400 [#2431] (characters; notes; Hunan). NIEHUIS & TEZCAN, 1993: 43 [#1910] (not in Turkey). JENDEK, 2001: 6 8, 10 [#5211] (synonymy; faunal records; key; China: Shaanxi, Zheijang, Guizhou, Hunan, Jiangxi, Fujian, Yunnan; Vietnam; Laos). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Hubei, Fujian, Sichuan, Yunnan, Xizang). PENG ZHONGLIANG, 2002: 267 [#6628] (characters; Fujian). MÜHLE, 2003: 47 [#6155] (checklist). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2239 [#8576] (world catalog). LEE, KIM & AHN, 2010: 264 266 [#8939] (characters; faunal records; South Korea).

155 jeanvoinei Descarpentries & Villiers, 1963 (Agrilus) DESCARPENTRIES & VILLIERS, 1963: 52, 54 55 [#699] (description). JENDEK, 2001: 6 [#5211] (synonym of plasoni). JENDEK, 2006: 401 [#7441] (synonym of plasoni; Palaearctic catalog). BELLAMY, 2008: 2239 [#8576] (synonym of plasoni; world catalog).

Type material: Agrilus plasoni Obenberger, 1917 Type locality: Constantinople [Note: Locality emended to “China-Fukien” by OBENBERGER, 1940 [#2273]]. Type specimens: Holotype by monotypy (Fig. 51A), ♂, (NMPC): “Constantinopel [sic!], c. Plason [p] \ Typus [p] [red label with black border] \ Agrilus Plasoni m. [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae [p] 26479 [h] Inv. [p] [orange label]”. Described from 1 specimen. Agrilus jeanvoinei Descarpentries & Villiers, 1963 Type locality: Tonkin: Than-Moï. Type specimens: Primary: Holotype (Fig. 47P), ♀, (MNHN): “TONKIN [p] Than Moo [sic!] II. IV. [h] 191 [p] 8 [h] JEANVOINE [leg.] [p] \ A. jeanvoinei n. sp. Holotype nobis A. Descarpentries et [h] A. Villiers det. 195 [p] [number 5 is overwritten to 6] 2 [h] \ Muséum Paris 1935 Coll. A. Théry [p]”. Secondary: 1 PT (MHNG); 3 PT (MNHN); 1 PT (NMPC). Described from unknown number of specimens (HT, PT). Diagnosis: 6.2-9.5 mm; body dorsally brightly bicolored; pronotum from carmine to golden-orange, elytra goldenbluish-green, with two adsutural spots of whitish tomentum in apical third; spine on outer margin of elytral apices with long, attenuate tip. Species group: Plasoni. Relevant species: A. plasoni dichrosomus Obenberger, 1924. Specimens examined: CHINA: Fujian: 1 (NMPC): “Kuatun (2300 m) 27,40n[ordlich]. Br[eite]. 117,40ö[stlich]. L[änge]. J. Klapperich [leg.] 22.6.1938 (Fukien)” (JENDEK, 2001 [#5211]); 3 (NMPC): “Kuatun [27°40’N, 117°40’E], Fukien, China, 1.VI., 24.VI., 25.VI.1946 (Tschung Sen)” (JENDEK, 2001 [#5211]); 1 (EJCB): “China, Fujian prov., Shaowu env., 13 16.6.1991” (JENDEK, 2001 [#5211]); 1 (EJCB): “China, N Fujian, 1 4.vi., Fengshui Guan, 27.9N, 117.85E, 1700m, Jaroslav Turna leg., 2004”. Guangxi: 1 (EJCB): “S China, NE Guangxi, Jiaojiang: Baogai mt., 1571 m, N25º32,598’, E110º57,502’, VI 2009, lgt. Häckel & Sehnal”, 1 ♂ (EJCB): “S China, NE Guangxi, Quanzhou/Guanyang border, Baogai mt., 500 900 m, VI 2007, lgt. Häckel & Sehnal”. Guizhou: 1 ♂ (EJCB): “CH Guizhou NE, 27.V. 3.VI., 20 km NW of Jiangkou, 1995, Fanjing Shan Kuaichang, E. Jendek & O. Šauša leg.” (JENDEK, 2001 [#5211]); 1 (EJCB): “S China, SE Guizhou, Dushan: Gengding Shan env., N25º52,5’, E107º38’, 1445 m, VI 2009, lgt. Häckel & Sehnal”. Hubei: 1 (EJCB): “P.R. China,, Hubei, Dabieshan, N31˚07’06’’ E115˚48’57”, 11 21.vi.2008, 640 m, V. Grebennikov leg.”. Hunan: 1 ♂, 1 ♀ (EJCB): “China Hunan prov., Zhang Jia Jie [National Park, nearby Dayong, NW Hunan, approximate coordinates 29°07’N, 110°28’E, C. Holzschuh pers. com.], VI.1993, C. Holzschuh leg.” (JENDEK, 2001 [#5211]); 1 ♂ (EJCB): “CHINA, Hunan S, 1994, CHENZHOU env., 19 21.VI., 25.49N 112.59E, E. Jendek & O. Šauša leg.”.” (JENDEK, 2001 [#5211]). Jiangxi: 1 ♂ (EJCB): “China, Jiangxi W, Jinggang Shan, Ciping, 2 14.VI.1994, E. Jendek & O. Šauša leg.” (JENDEK, 2001 [#5211]). Shaanxi: 1 ♂ (EJCB): “CHINA, 1000 1300 m, Shaanxi, Qinling mts., XUN YANGBA (6 km E) [33°33’N, 108°37’E], 23.v. 13.vi.1998, I. H. Marshal leg.” (JENDEK, 2001 [#5211]). LAOS: Borikhamxai: 1 ♂ (EJCB): “Laos centr., Bolikhamsai prov., Ban Nape Kaew Nua Pass, 18.4. 1.5.1998, alt. 600±100 m, N18º22.3, E105º09.1 (GPS),

156 E. Jendek & O. Šauša leg.” (JENDEK, 2001 [#5211]); 2 ♀ (EJCB): “Laos CE., 1 18.v.2001, Boli Kham Xai prov., 18º21’N, 105º08’E, Ban Nape (8 km NE), ~600m, Kubáň leg.”. Louang Namtha: 1 ♂, 1 ♀ (EJCB): “Laos, Louang Namtha pr., 21°09’N, 101°19’E, Namtha Muang Sing, 5 31.v.1997, 900 1200 m, Vit Kubáň leg.” (JENDEK, 2001 [#5211]). Phongsali: 1 ♂ (EJCB): “Lao N, Phongsaly prov., 21°41’ 2’N, 102°06’ 08’E, 28.v. 20.vi.2003 Phongsaly env., ~1500m, Vít Kubáň leg.”; 1 ♂ (EJCB): “Lao, Phongsaly prov., 21°41’N, 102°06’E, Phongsaly env., 6 17.v.2004, 1500 m, Vít Kubáň leg.”. VIETNAM: 1 (MNHN): “Hoa Binh [20°20’N, 105°15’E] Tonkin, de Cooman [leg.]” (JENDEK, 2001 [#5211]); 2 PT (MNHN): “Tonkin, reg. de Hoa Binh, Man Son” (JENDEK, 2001 [#5211] [paratypes of jeanvoinei); 2 (MNHN): “Tonkin région de Hoa Binh, Man Son [? 21°31’N, 105°15’E]” (JENDEK, 2001 [#5211]); 2 (EJCB): “Vietnam N, Hoa binh 4 7/6 1986, Ha Son Binh prov., Strnad Jan lgt.” (JENDEK, 2001 [#5211]). 1 (EJCB): “Vietnam, sept[entrional]., 1990 Sa Pa, 11 19.VI., (Houang Lien Son) Mir. Dvořák legit” (JENDEK, 2001 [#5211]); 11 (EJCB): “N Viet Nam (Tonkin), pr. Houang Lien Son, Sa Pa, 11 15.V.1990, Vít Kubáň leg.” (JENDEK, 2001 [#5211]); 1 (EJCB): “SaPa 11 18.VI., N. Vietnam, A. Olexa [leg.] 1990” (JENDEK, 2001 [#5211]); 1 (EJCB): “N Viet Nam (Tonkin) pr. Vin Phu, 1990, TAM DAO [21°28’N, 105°38’E] 17 21.V., Vít Kubáň leg.” (JENDEK, 2001 [#5211]); 1 ♂ (EJCB): “Vietnam, 1100 1700 m, 22.18N 103.50E, W SaPa, 29.V. 11.VI., lg[t]. K. W. Anton 1996” (JENDEK, 2001 [#5211]).

Adult occurence: 4-5-6-7 (n=26). Faunal records published by OBENBERGER, 1940 [#2273] and 1958 [#2389] are also included. Altitude range: 500-2300 m (n=14). Host plants: Unknown. Distribution: CHINA: Fujian, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Shaanxi, Yunnan, Zhejiang. KOREA SOUTH. LAOS. VIETNAM. Remarks: Species distribution in South Korea was complemented by the recent publication of LEE, KIM & AHN, 2010 [#8939] and is not included in the chapter Analysis.

Agrilus plavilstshikovi Alexeev Fig. 21K (habitus of PT); Fig. 37H (aedeagus)

plavilstshikovi Alexeev, 1989 (Agrilus) ALEXEEV in ALEXEEV & VOLKOVITSH, 1989: 315 317, 344 [#3241] (description). ALEXEEV, 1998: 375 [#4322] (subgenus Xeragrilus; subgeneric classification). JENDEK, 2006: 396 [#7441] (subgenus Xeragrilus; Palaearctic catalog). BELLAMY, 2008: 2239 [#8576] (subgenus Xeragrilus; world catalog).

Type material: Agrilus plavilstshikovi Alexeev, 1989 Type locality: Mongolia, Vostochno-Gobiiskii aimak, 25 km V oz. Shokhoi-Nur [in Russian]. Type specimens: Primary: Holotype, sex not examined, (ZIN): “Mongoliya, V. - Gobiisk. 25 km V oz. Shokhoi-Nyur 13. VIII. [sic!] Emelyanov [h] [in Russian] \ Holotypus [p] [red label] \ male sign Pr. Ag.-84-05 [h] [red label] \ Agrilus plavilstshikovi m. sp. n. [h] Alexeev det. [p] [in Russian]”. Secondary: 5 PT (ZIN). Described from 13 specimens (HT, PT). Paratype (Fig. 21K) from ZIN is illustrated. Diagnosis: 3.6-4.0 mm; body sparsely, longly pubescent, without tomentum; vertex with feeble medial impression;

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA pronotum obviously convex; prehumerus absent; prosternal lobe medially distinctly emarginated. Species group: Albogularis. Relevant species: A. aerosus Ganglbauer, 1889. Specimens examined: Known only from the type specimens. Host plants: Unknown.

Distribution: MONGOLIA.

Agrilus pleuralis Jendek, sp. nov. Fig. 21E (habitus of HT); Fig. 37B (aedeagus)

Description: 6.2-7.0 mm (Holotype 6.2 mm). BODY: Shape: cuneiform, Build: robust, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Medial impression: present, Impression (extent): vertex and frons, Impression (depth): deep, Epistoma: with raised upper margin. Frons: Shape: flat or moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae. Eyes: Size: small, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: long, Sexual modification in male: antennae longer. PRONOTUM: Shape: visually square or transverse, Sides: arcuate, Maximal width: at anterior margin, rarely at middle, Anterior margin: subequal to posterior. Anterior lobe: absent or vague, Shape: arcuate or subangulate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: flat, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow, Lateral impressions (width): wide. Prehumerus: absent or obsolete. Marginal and submarginal carinae: Interspace: wide, Convergence: subparallel, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate or large, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Width: narrow, Shape: arcuate. Elytral pubescence: entire, Color: unicolored, Character: homogenous. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: narrow or moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): moderate. Prosternal process: moderate, Shape: narrowed, Sides: straight, Angles: absent, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): shorter or subequal to 2-3 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): markedly dorsoventrally bent, Ovipositor: elongate. Diagnosis: vertex with deep medial carina; pronotum widest in anterior half; entire abdominal pleura obviously pubescent. Species group: Albogularis. Relevant species: A. ecarinatus Marseul, 1866.

157

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Type locality: Juzhaigou.

China,

Sichuan

province,

Nanping-

Type specimens: Primary: Holotype (Fig. 21E), ♂, (EJCB): “China-Sechuan, Nanping-Juzhaigou, 7-12.6.2009, leg. E. Kučera”. Secondary: CHINA: Sichuan: 6 PT (EJCB): “China-Sechuan, Nanping-Juzhaigou, 7-12.6.2009, leg. E. Kučera”. Adult occurence: 6 (n=1). Host plants: Unknown. Distribution: CHINA: Sichuan. Etymology: The specific name is derived from the Greek noun pleura (side). It refers to the dense tomentum on abdominal pleura of the species.

than 2-4 combined. GENITALIA: Ovipositor: elongate. Diagnosis: elytra glabrous except for 3 pairs of tomentose spots and small patch of pubescence on apices. Species group: Maculifer. Relevant species: A. nelsonodaimio Ohmomo, 2006. Type locality: China, Shaanxi province, Qinling mts., 6 km east Xunyangba, altitude 1000-1300 m. Type specimens: Primary: Holotype (Fig. 11J), ♀, (EJCB): “China, 1000-1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.-13.vi.1998, I. H. Marshal leg.”. Adult occurence: 5-6 (n=2). Altitude range: 1000-1300 m (n=2). Host plants: Unknown. Distribution: CHINA: Shaanxi.

Agrilus pleurostigmus Jendek, sp. nov. Fig. 11J (habitus of HT)

Description: 7.1-7.2 mm (Holotype 7.1 mm). BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): black. HEAD: Medial impression: present, Impression (extent): vertex, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture (aspect): subparallel, Sculpture (density): dense, Sculpture (intensity): superficial. Eyes: Size: small, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: flat, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (width): wide. Prehumerus: filamentary, Shape: bisinuate, Extent: to anterior pronotal angle, Posterior end: joining posterior pronotal angle, Anterior end: joining anterior pronotal angle, Arc: weak. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Shape: arcuate. Elytral pubescence: distal only, Distal: apical, Color: unicolored. Tomentum: spots, Spots (locations): humeral, premedial or postmedial. STERNUM: Metasternal projection: impressed. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): deep, Emargination (width): moderate. Prosternal process: moderate, Shape: narrowed, Sides: straight, Angles: absent, Disk: convex. ABDOMEN: Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer

Etymology: The specific name is a combination of the Greek noun pleura (side) and the Latin noun stigma (small mark, spot). It refers to the obvious pubescent spot at pleural abdominal portions of the species.

Agrilus pliculipennis Obenberger Fig. 28K (habitus); Fig. 40O (aedeagus); Fig. 51B (type) pliculipennis Obenberger, 1940 (Agrilus) OBENBERGER, 1940: 182 183 [#2273] (description). PENG ZHONGLIANG, 1987: 357 [#2429] (checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China). PENG ZHONGLIANG, 2002: 269 [#6628] (notes). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2239 [#8576] (world catalog).

Type material: Agrilus pliculipennis Obenberger, 1940 Type locality: China: Fokien: Taining-li. Type specimens: Primary: Lectotype (Fig. 51B) by present designation, ♂, (NMPC): “Fokien, Tainingli [h] \ Typus [p] [red label] \ Agrilus pliculipennis m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24988 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 3.6-4.9 mm; eyes smaller; elytra with humeral carina; sides of aedeagus without extending membraneous lobes. Species group: Viridiobscurus. Relevant species: A. lobatus sp. nov. Specimens examined: CHINA: Fujian: 1 (NMPC): “Kuatun (2300m) 27,40n. Br 117,40ö. L. J. Klapperich, 31.5.1938 Fukien”; 1 (NMPC): “Kuatun (2300m) 27,40n. Br 117,40ö. L. J. Klapperich, 4.6.1938 Fukien”; 2 ♂ (EJCB): “China, Fujian prov. bor. occ., Shaowu env. 25 km road, Shaowu Taining 6 10.7., R. Červenka lgt. 1991”; 1 ♀ (EJCB): “China, prov. Fujian bor. occ., Shaowu env., 25km road Shaowu Taining, 13 16.6., M. Nikodým lgt., 1991”; 1 ♂ (EJCB): “China, Fujian prov. bor. occ., Shaowu env. 25 km road, Shaowu Taining, 6 10.7., R. Červenka lgt. 1991”; 3 ♂ (EJCB): “China, Fujian prov., Shaowu env., 23 27.vi.1991, leg. Nikodým”; 4 ♂ (EJCB): “China, Fujian prov. Shaowu env., 5 10. VII.1991”. Guizhou: 1 ♂ (EJCB): “China Guizhou 21 26.V., 60 km N of Kaili, 1995, Shibing Yuntai Shan, E. Jendek & O. Šauša leg.”; 1 ♂ (EJCB): “CH Guizhou NE, 27.V. 3.VI., 20 km NW of Jiangkou, 1995, Fanjing Shan Kuaichang, E. Jendek & O. Šauša leg.”; 2 ♂ (EJCB): “China, Guizhou prov., Leigongshan, Xijiang,

158 29 May 2 June 1997, 1200 1900 m, Bolm lgt.”. Henan: 1 ♂ (EJCB): “China, W Henan, 1.vii., 20 km SE Luanchan “tunel”, 33.7ºN 111.8ºE, Jaroslav Turna leg., 2001”. Hunan: 13 (EJCB): “China, Hunan SE, 1994, Guidong env. 26 31.V., 26.04N 113.56E, E. Jendek & O. Šauša leg.”. Jiangxi: 8 (EJCB): “China, Jiangxi W, Jinggang Shan, Ciping, 2 14.VI.1994, E. Jendek & O. Šauša leg.”. Shaanxi: 8 (EJCB): “China S, Shaanxi, 14 16.6.1995, Qinling Mts. N. slope, 33.56 59’/108.50’ 800 1200 m, Changan Co., L.+R. Businský lgt.”; 1 ♀ (EJCB): “China, 1000 1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v. 13.vi.1998, I. H. Marshal leg.”; 1 ♀ (EKCS): “China Shaanxi, Lüeang 33°07’N 106°05’E, 18.6. 24.6.[19]97 lgt. E. Kučera”; 1 ♂ (EJCB): “China Shaanxi, Lueang env., 15 km NW, 26.5. 31.5.2001, lgt. E. Kučera”.

Adult occurence: 5-6-7 (n=17). Altitude range: 800-2300 m (n=5). Host plants: Unknown. Distribution: CHINA: Fujian, Guizhou, Henan, Hunan, Jiangxi, Shaanxi.

Agrilus pratensis (Ratzeburg) Figs. 17I, 17J (habitus); Fig. 35N (aedeagus); Figs. 44G, 45N, 51I, 52A (types) pratensis Ratzeburg, 1837 (Buprestis) RATZEBURG, 1837: 56 57 [#2498] (description). GORY & LAPORTE, 1839: 55 56 [#3738] ([Note: Misidentification: KIESENWETTER, 1857: 146, 147 [#3015] cited linearis Gory & Laporte as a synonym of pratensis; Gory & Laporte did not describe the species as new, he attributed it to Fabricius]). KIESENWETTER, 1857: 125, 145 147 [#3015] (characters in key, redescription; notes; Germany). MARSEUL, 1866: 435, 456 [#676] (characters in key; redescription; distribution overview). GEMMINGER & HAROLD, 1869: 1445 [#1057] (synonym of robertii cited as roberti; catalog). SAUNDERS, 1871: 119 [#2552] (catalog). HOCHHUTH, 1873: 158 [#3453] (faunal records; Gouvernements Kiew und Volhynien). MARSEUL, 1887: 251 [#4707] (catalog). KERREMANS, 1892: 270 [#1397] (synonym of robertii cited as roberti; catalog). HEYDEN, 1897: 194 [#5286] (distributional summary). CSIKI, 1901: 89 [#505] (faunal records; Rossia: Kazan). KERREMANS, 1903: 275 [#1441] (synonym of robertii; catalog). JAKOBSON, 1913: 797 [#4386] (synonym of robertii; catalog). BAU, 1914: 256 257 [#8668] (characters in key). BEDEL, 1921: 210 [#170] (synonym of robertii). PLANET, 1923: 256 [#8562] (characters). OBENBERGER, 1924: 43 [#2040] (variety of robertii). OBENBERGER, 1926: 655 [#2057] (synonym of robertii). OBENBERGER, 1936: 1028 1029 [#2191] (aberration of robertii cited as roberti; world catalog). SCHAEFER, 1937: 83 84 [#3570] (synonym of robertii). RIKHTER, 1945: 166 [#3507] (aberration of robertii). RIKHTER, 1948: 394 [#3879] (aberration of robertii). SCHAEFER, 1949: 355, 391 392, 490 [#2565] (characters; references; faunal records; biology; notes; ditributional overview; notes; France). HORION, 1955: 87 88 [#1285] (distribution overview; faunal records; biology; Deutschland). ALEXEEV, 1959: 8 [#3229] (synonym of robertii; characters in key). RIKHTER & ALEXEEV, 1965: 301 [#2522] (synonym of robertii; cited as roberti). ALEXEEV, 1975: 152 [#52] (faunal records; distributional overview; Mongolia). LOMPE, 1979: 237 [#3628] (characters in key; Central Europe). BÍLÝ, 1982: 64,79 [#295] (characters in key; distribution overview; biology). GOBBI, 1983: 39 [#1098] (note). ALEXEEV, 1989: 487 [#64] (key to species; south Siberia.; central and south parts of European USSR; central China, central Mongolia; west Asia; western Europe). ALEXEEV & VOLKOVITSH, 1989: 318, 346, 356 [#3241] (characters in key; faunal records; Mongolia). SPARACIO, 1990: 75 [#2652] (notes; Italy (including Corsica,

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Sardinia, Sicilia)). GUTOWSKI, 1993: 302 [#3638] (characters). NIEHUIS & TEZCAN, 1993: 23, 46, 47 [#1910] (subgenus Anambus; faunal records from Turkey; distribution and host plants summary; Turkey). CURLETTI, 1994: 23, 190 191 [#658] (subgenus Robertius; catalog; faunal records; Italia; Corsica; Sicilia; Sardegna). JENDEK, 1994: 10 11 [#1355] (synonymy). ALEXEEV, 1998: 369, 370 [#4322] (subgenus Arquagrilus; genotype of the subgenus Arquagrilus). TLEPPAEVA, 1999: 184 [#6551] (subspecies of zigzag; notes; Kazakhstan). BÍLÝ, 2002: 23 24 [#5877] (bionomy; host plants; distribution). BRECHTEL & KOSTENBADER, 2002: 288 290 [#5766] (biology; ecology; distribution; Germany). JENDEK, 2002: 5 8 [#5709] (nomenclatural and taxonomic notes). GEORGIEV et al., 2004: 238 [#6736] (notes; Bulgaria). BERNHARD et al., 2005: 601, 602 [#7261] (mitochondrial DNA; molecular systematics). DEDYUKHIN, NIKITSKY & SEMENOV, 2005: 303 [#8860] (checklist; Russia: Udmurtia). JENDEK, 2006: 393 [#7441] (subgenus Robertius; Palaearctic catalog). BELLAMY, 2008: 2243 [#8576] (subgenus Robertius; world catalog). robertii Chevrolat, 1840 (Agrilus) CHEVROLAT, 1840: 89 90 [#489] (description). KIESENWETTER, 1857: 146 [#3015] (synonym of pratensis). MARSEUL, 1857: 92 [#4371] (cited as roberti; checklist; Europe). JACQUELIN DU VAL, 1860: 143 [#8086] (synonym of pratensis). MARSEUL, 1866: 456 [#676] (cited as roberti; synonym of pratensis). GEMMINGER & HAROLD, 1869: 1445 [#1057] (cited as roberti; catalog). SAUNDERS, 1871: 119 [#2552] (cited as roberti; synonym of pratensis). MARSEUL, 1887: 251 [#4707] (cited as roberti; synonym of pratensis). REITTER, 1891: 200 [#5278] (cited as roberti; synonym of pratensis). KERREMANS, 1892: 270 [#1397] (cited as roberti; catalog). KERREMANS, 1903: 275 [#1441] (cited as roberti; catalog). REITTER, 1911: 194 [#2518] (cited as roberti; characters in key; Germany). CAILLOL, 1913: 512 [#4053] (cited as roberti; biology; faunal records; France). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). BEDEL, 1921: 202, 210 211 [#170] (cited as roberti; characters in key; references; faunal records; distribution; France). OBENBERGER, 1924: 43 [#2040] (note). OBENBERGER, 1926: 655 [#2057] (cited as roberti; Palaearctic catalog). JAKOBSON, 1927: 179 [#8655] (cited as roberti; characters in key). JAKOBSON & OGLOBLIN, 1931: 162 [#8314] (cited as roberti; characters in key). OBENBERGER, 1935: 58 59 [#2178] (cited as roberti; catalog; summary of distribution; Bulgaria). OBENBERGER, 1936: 1027 1029 [#2191] (cited as roberti; world catalog). SCHAEFER, 1937: 84 [#3570] (cited as roberti). THÉRY, 1942: 133, 155 156 [#2873] (subgenus Agrilus; cited as roberti). MÉQUIGNON, 1943: 160 161 [#3661] (synonym of pratensis). RIKHTER, 1945: 166 [#3507] (cited as roberti; characters in key; European part of USSR except northernmost region, eastward to Primorsk). THÉRY, 1945: 65 66 [#2884] (synonym of pratensis). PALMÉN, 1946: 120 [#7730] (cited as roberti; faunal records; Russia (Karelia)). THÉRY, 1947: 670 671 [#2891] (cited as roberti; type species of the subgenus Robertius). RIKHTER, 1948: 394 [#3879] (cited as roberti). STEPANOV, 1948: 243 [#3535] (cited as roberti; checklist; faunal record; Chelyabinskaya oblast’). RIKHTER, 1950: 158, 164 [#3881] (cited as roberti; adult and larval characters in key). MEDVEDEV et al., 1951: 311 [#8639] (cited as roberti; notes; Ukraine). STARK, 1955: 775 [#3533] (cited as roberti; faunal record; USSR: Saratov). TER MINASYAN, 1955: 453 [#7893] (host plants; distribution; references). ALEXEEV, 1957: 153 [#3226] (cited as roberti; characters in key). ALEXEEV, 1958: 185 [#3982] (cited as roberti; checklist; Moskovsk). ALEXEEV, 1959: 8 [#3229] (cited as roberti; characters in key). PORTA, 1959: 197 [#2488] (cited as roberti; synonym of pratensis). KHARAZISHVILI, 1960: 146 [#8630] (pest; Georgia). ZAGAIKEVICH, 1960: 257 [#5615] (cited as roberti; checklist; Ukraine). ZAGAIKEVICH, 1962: 60 [#5616] (cited as roberti;

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA checklist; Ukraine). ZAGAIKEVICH, 1962: 103 [#3249] (cited as roberti; faunal records; biology; Ukraine; Krym). RIKHTER & ALEXEEV, 1965: 301 [#2522] (cited as roberti; characters in key; European part of USSR). KOSTIN, 1973: 110 [#3693] (cited as roberti; characters in key; Kazakhstan). ALEXEEV, 1975: 152 [#52] (cited as roberti; synonym of pratensis). VOLKOVITSH, 1977: 26 [#2985] (cited as roberti; faunal record; notes; USSR: Belgorod; Voronezh; Lugansk). LOMPE, 1979: 237 [#3628] (cited as roberti; synonym of pratensis). COBOS, 1986: 316 [#609] (cited as roberti; synonym of pratensis). CURLETTI, 1994: 190 [#658] (cited as roberti; synonym of pratensis). LAGUNOV & NOVOZHENOV, 1996: 47 [#7461] (cited as roberti; checklist; Russia (Chelyabinsk)). CURLETTI, 1998: 133 [#4357] (subgenus Robertius; cited as roberti). MÜHLE et al., 2000: 160 [#4970] (cited as roberti; type of subgenus Robertius Théry). BÍLÝ, 2002: 23 [#5877] (cited as roberti; synonym of pratensis). BRECHTEL & KOSTENBADER, 2002: 288 [#5766] (cited as roberti; synonym of pratensis). JENDEK, 2002: 6 [#5709] (synonym of pratensis; taxonomic notes). JENDEK, 2006: 35 [#7359] (synonym of pratensis; type species of Agrilus subgenus Robertius; robertii is correct original spelling). JENDEK, 2006: 393 [#7441] (synonym of pratensis pratensis; Palaearctic catalog). BELLAMY, 2008: 2244 [#8576] (synonym of pratensis; world catalog). NIKITSKY et al., 2008: 148 [#8610] (synonym of pratensis). praeclarus Krogerus, 1925 (Agrilus; var. of robertii) syn. nov. KROGERUS, 1925: 73, 75 76 [#3472] (description). OBENBERGER, 1926: 655 [#2057] (variety of robertii; Palaearctic catalog). OBENBERGER, 1936: 1029 [#2191] (synonym of robertii aberration fennicus; world catalog). OBENBERGER, 1937: 58 [#2226] (synonym of robertii aberration fennicus). COBOS, 1986: 316 [#609] (synonym of pratensis fennicus). CURLETTI, 1994: 190 [#658] (synonym of pratensis). BÍLÝ, 2002: 23 [#5877] (synonym of pratensis). FARRUGIA, 2002: 107 [#6511] (synonym of pratensis). JENDEK, 2002: 6 [#5709] (subspecies of pratensis; nomenclatural and taxonomic notes). JENDEK, 2006: 393 [#7441] (subgenus Robertius; subspecies of pratensis; Palaearctic catalog). BELLAMY, 2008: 2245 [#8576] (subspecies of pratensis; world catalog). pseudocoeruleus Obenberger, 1930 (Agrilus) OBENBERGER, 1930: 112 [#2088] (description). ALEXEEV & VOLKOVITSH, 1989: 318 [#3241] (synonym of pratensis; type examination). JENDEK, 1994: 10 11 [#1355] (synonym of pratensis; lectotype designation). JENDEK, 2002: 6 [#5709] (synonym of pratensis; taxonomic notes). JENDEK, 2006: 393 [#7441] (synonym of pratensis pratensis; Palaearctic catalog). BELLAMY, 2008: 2244 [#8576] (synonym of pratensis; world catalog). djukini Obenberger, 1935 (Agrilus; ssp. of robertii) syn. nov. OBENBERGER, OBENBERGER, 1935: 169 [#2158] (description). 1936: 1029 [#2191] (subspecies of robertii cited as roberti; world catalog). SCHAEFER, 1937: 84 [#3570] (subspecies of robertii cited as roberti). SCHAEFER, 1949: 392 [#2565] (subspecies of pratensis; notes). COBOS, 1986: 242, 316 [#609] (subgenus Anambus; subspecies of pratensis; notes). ALEXEEV, 1989: 487 [#64] (subspecies of pratensis; key to species; Chita; Buryatia). JENDEK, 1994: 11 [#1355] (subspecies of pratensis; lectotype designation; synonymy; characters; Mongolia; Russian Far East). BÍLÝ, 2002: 24 [#5877] (subspecies of pratensis). JENDEK, 2002: 6, 7 [#5709] (subspecies of pratensis; taxonomic notes). JENDEK, 2006: 393 [#7441] (subgenus Robertius; subspecies of pratensis; Palaearctic catalog). BELLAMY, 2008: 2244 2245 [#8576] (subspecies of pratensis; world catalog). VOLKOVITSH, 2009: 136 [#8850] (subgenus Robertius; subspecies of pratensis; faunal records; Russia: Primorskiy kray).

159 chankae Obenberger, 1935 (Agrilus; subspecies of betuleti) OBENBERGER, OBENBERGER, 1935: 169 [#2158] (description). 1936: 961 [#2191] (subspecies of betuleti; world catalog). THÉRY, 1942: 156 [#2873] (subgenus Agrilus; subspecies of betuleti). SCHAEFER, 1949: 388 [#2565] (subspecies of betuleti; notes). COBOS, 1972: 56 [#591] (uncertain synonym of betuleti). JENDEK, 1994: 11 [#1355] (synonym of pratensis djukini; lectotype designation). BÍLÝ, 2002: 13 [#5877] (subspecies of betuleti; distribution; China (north territory); RUSSIA (East Siberia)). JENDEK, 2002: 6, 7 [#5709] (synonym of pratensis djukini; taxonomic notes). JENDEK, 2006: 393 [#7441] (synonym of pratensis djukini; Palaearctic catalog). BELLAMY, 2008: 2245 [#8576] (synonym of pratensis djukini; world catalog). fennicus Obenberger, 1936 (Agrilus; unnecessary replacement name for praeclarus Krogerus) OBENBERGER, 1936: 1029 [#2191] (replacement name proposal). OBENBERGER, 1937: 58 [#2226] ( subspecies of robertii; cited as nomen novum). SCHAEFER, 1937: 84 [#3570] (variety of robertii cited as roberti). KASZAB, 1940: 102 [#1374] (aberration of robertii cited as roberti; Kingdom of Hungary). THÉRY, 1942: 155 [#2873] (subgenus aberration of pratensis). SCHAEFER, 1949: 392 [#2565] (subspecies of pratensis; notes). COBOS, 1986: 242, 316 [#609] (subgenus Anambus; subspecies of pratensis). JENDEK, 2002: 6, 7 [#5709] (synonym of pratensis praeclarus; nomenclatural and taxonomic notes). JENDEK, 2006: 393 [#7441] (synonym of pratensis praeclarus; Palaearctic catalog). BELLAMY, 2008: 2245 [#8576] (synonym of pratensis praeclarus; world catalog).

Type material: Buprestis pratensis Ratzeburg, 1837 Type locality: not given [Wäldern Preussens are cited in the title of the publication]. Type specimens: Not located. Described from 1 specimen. Agrilus robertii Chevrolat, 1840 Type locality: Belgia. ... des environs de Liège. Type specimens: Primary: Lectotype (Fig. 52A) by present designation, sex not examined, (BMNH): “SYN-Type [p] [round label with blue border] \ Type. [p] \ Belgia Robert [h] \ Roberti [sic!] Chevr. [h] \ Saunder 74.18 [p]”. Described from unknown number of syntypes. Agrilus robertii variety praeclarus Krogerus, 1925 Type locality: ... südlichen Finnland, St. Ylöjärvi. Type specimens: Not located. Described from unknown number of syntypes. Agrilus pseudocoeruleus Obenberger, 1930 Type locality: Asia centralis: Sefid-rud. Type specimens: Primary: Lectotype (Fig. 51I), ♀, designated by JENDEK (1994: 10-11) [#1355], (NMPC): “Asia centr. Safid-rud [h] \ Typus [p] [red label] \ Agrilus pseudocoeruleus m. Type [h] Det. Dr Obenberger [p] \ LECTOType [p] Agrilus pseudocoeruleus Obnb. [h]”. Described from unknown number of syntypes. Agrilus robertii djukini Obenberger, 1935 Type locality: Primorje. Ussuri. Type specimens: Primary: Lectotype (Fig. 45N), ♂, designated by JENDEK (1994: 11) [#1355], (NMPC): “s. Trojckoe, oz. Chanka, Primors. Tscherskij [p] 4. VI. [h] [19]09 [p] [in Russian] \ Typus [p] [red label] \ Agrilus Roberti ssp. Djukini m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24791 [h] [orange label]”. Secondary: 2 PLT

160 (ZIN); 1 PLT (NMPC). Described from unknown number of syntypes. Agrilus betuleti chankae Obenberger, 1935 Type locality: Ussuri. Lac Chanka. Type specimens: Primary: Lectotype (Fig. 44G), ♂, designated by JENDEK (1994: 11) [#1355], (NMPC): “Chanka Ussuri [h] \ Kamen-Rybolov oz. Chanka, Yuzhnouss Djukin [p] 24 [h] V [19]08 [p] [in Russian] \ Typus [p] [red label] \ Mus. Nat. Pragae Inv. [p] 24270 [h] [orange label] \ Agrilus betuleti ssp. chankae m. Type [h] Det. Dr. Obenberger [p] \”. Secondary: 1 PLT (ZIN); 1 PLT (NMPC). Described from unknown number of syntypes. Agrilus fennicus Obenberger, 1936 Type locality and specimens: See: Agrilus robertii variety praeclarus Krogerus, 1925. Diagnosis: 4.1-6.9 mm; body bicolored (Fig. 17J) or unicolored (Fig. 17I); elytral apices subtruncate. Species group: Betuleti. Relevant species: A. betuleti (Ratzeburg, 1837). Specimens examined: CHINA: Hebei: 4 (EJCB): “China, Hebei/ Nei Mongol, pass Chengde Chifeng, 41.6N, 118.2E, 14 16. VI., Jaroslav Turna leg. 2001”. Nei Mongol: 2 (EJCB): “China, Nei Mongol, Wuhai, 6.5. 13.5.1996, E. Kučera leg.”. Shaanxi: 1 ♂ (EJCB): “China SE, Shaanxi, Danfeng NE env., 900 1500m, 28 29.5.1995, 33°45’ 52’/110°22’ 37’, L. + R. Businský lgt.”; 2 (EJCB): “China, Shaanxi prov., 21 23.June 1998 \ Qing Ling Shan mts, 1000 m road Baoji Taibai vill., pass 40 km S Baoji, Zd. Jindra lgt.”; 5 (EJCB): “China, Shaanxi prov., 21 23.vi.1998, Quin Ling Shan mts, road Baoji Taibai vill., pass 35 km S Baoji, O. Šafránek et M. Trýzna lgt.”. Xinjiang: 1 ex (EJCB): “China, Xinjiang, 1993, road Narat Kuytun, Erenhaberga Shan, 40 km S Maytag, 1500m, J. Turna leg., 22/VII.”. IRAN: 1 ♂ (EJCB): “N Iran, Golestan Prov., Golestan N.P., Tangegol, 37°22N, 55°56E, 700m, 18 22 V.2001, E. Heiss”. KAZAKHSTAN: 2 ♂, 1 ♀ (EJCB): “W Kazakhstan, Aktyubinsk, Chalkov env., 9.6.1995, leg. Dolin”. MONGOLIA: Dornod: 1 ♂, 1 ♀ (EJCB): “Mongolia, East ajmak Numregin Gol river 32 km SE Salchit town 8.VII.1976” (JENDEK, 1994 [#1355] as pratensis djukini). RUSSIA: Adygea Respublika: 1 ♀ (EJCB): “W Caucasus, Adygeia, Nikel vill., 28.vi.2000”. Buryatiya Respublika: 1 ♀ (ZIN): “Bass. r. Vitim, sr. tech. r. B. Amalat 17.8.1968 Kabakov [in Russian]”. Chelyabinskaya oblast’: 1 (ZIN): “pos. Sokolovskii, Chelyab., u. Orenb. g. Infant’ev 19.v.1912 [in Russian]”; 1 ♂ (EJCB): “Russia, South Ural Mts., Cheliabinsk, 3.vii.1990”. Ivanovskaya oblast’: 1 (MSCI): “C. Russia, Ivanovskaja ar., Ivanovo env., Semenovskoje vill. 14.07.1994 leg. O. Kashin”. Kaluzhskaya oblast’: 1 (MSCI): “C. Russia, Kaluzhskaja ar., Malojaroslavetsky dist., Afanasovo vill. 11 19.06.2005 leg. М. Smirnov”. Lipetskaya oblast’: 1 (MCCD): “37км СВ г. Елец, с. Лески, 10.6.2007 Мазуров”. Primorskiy kray: 1 (MSCI): “SE Maritime Prov., Lazovsky dist., 12 кm S Staraja Kamenka , 10.07.04 leg. М.& L. Smirnov’s”. Vladimirskaya oblast’: 1 (MSCI): “C. Russia, Vladimir ar., Koltschugino dist., Litvinovo vill. V.99 leg. S. Svetlov”. Zabaykalskiy kray: 2 (ZIN): “YuV. Chitinsk. obl. Sherlovaya gora, 20.vi.1989 Kadyrov [in Russian]”; 1 ♂, 1 ♀ (ZIN): “YuV Chitinsk. obl., Sherlovaya gora, 15 VII 1989 Kabakov [in Russian]”.

Adult occurence: 5-6-7-8 (n=22). Altitude range: 7001500 m (n=4). Host plants: Populus: DAEHNE, 1912 (as robertii) [#3609]; SAINTE CLAIRE, 1935 (as robertii) [#8551]; TER MINASYAN, 1955 (as robertii) [#7893]; ALEXEEV, 1959 (as robertii)

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA [#3229]; ALEXEEV, 1959 [#3229]; KHARAZISHVILI, 1960 (as robertii) [#8630]; RIKHTER & ALEXEEV, 1965 (as robertii) [#2522]; ALEXEEV, 1975 [#52]; VOLKOVITSH, 1977 (as robertii) [#2985]; ALEXEEV, 1989 [#64]; ALEXEEV & VOLKOVITSH, 1989 [#3241]; WEIDLICH, 1989 [#6165]; CURLETTI, 1994 [#658]; KLETECKA, 1998 [#4416]; HASTIR & GASPAR, 2002 [#6211]. – P. alba: CAILLOL, 1913 (as robertii) [#4053]; SCHAEFER, 1949 [#2565]; HORION, 1955 [#1285]; BÍLÝ, 2002 [#5877]; BRECHTEL & KOSTENBADER, 2002 [#5766]. – P. nigra: CAILLOL, 1913 (as robertii) [#4053]; BEDEL, 1921 (as robertii) [#170]; SCHAEFER, 1949 [#2565]; HORION, 1955 [#1285]; NIEHUIS, 1988 [#1912]; BÍLÝ, 2002 [#5877]; BRECHTEL & KOSTENBADER, 2002 [#5766]. – P. nigra italica: SPARACIO, 1990 [#2652]. – P. tremula: CHEVROLAT, 1840 (as robertii) [#489]; MARSEUL, 1866 [#676]; HEYDEN, 1897 [#5286]; CAILLOL, 1913 (as robertii) [#4053]; BEDEL, 1921 (as robertii) [#170]; SCHAEFER, 1949 [#2565]; HORION, 1955 [#1285]; NIEHUIS, 1988 [#1912]; BÍLÝ, 2002 [#5877]; BRECHTEL & KOSTENBADER, 2002 [#5766]; GUTOWSKI & MILKOWSKI, 2008 [#8244]. Dubious: Salix: TER MINASYAN, 1955 (as robertii) [#7893]; ALEXEEV, 1959 (as robertii) [#3229]; ALEXEEV, 1959 [#3229]; RIKHTER & ALEXEEV, 1965 (as robertii) [#2522]; ALEXEEV, 1975 [#52]; ALEXEEV, 1989 [#64]; ALEXEEV & VOLKOVITSH, 1989 [#3241]. Erroneous: Cornus sanguinea: DAEHNE, 1912 (as robertii) [#3609]. – Quercus: MARSEUL, 1866 [#676]; DAEHNE, 1912 (as robertii) [#3609]; TER MINASYAN, 1955 (as robertii) [#7893]. Distribution: Local: CHINA: Hebei, Nei Mongol, Shaanxi, Xinjiang. MONGOLIA. RUSSIA: Amurskaya oblast’, Buryatiya Respublika, Irkutskaya oblast’, Primorskiy kray, Zabaykalskiy kray. Global: ASIA: CHINA. GEORGIA. IRAN. KAZAKHSTAN. RUSSIA. TURKEY. EUROPE: ALBANIA. AUSTRIA. BELARUS. BELGIUM. BOSNIA & HERZEGOVINA. BULGARIA. CROATIA. CZECH REPUBLIC. ESTONIA. FINLAND. FRANCE. GERMANY. GREECE. HUNGARY. ITALY. LATVIA. LIECHTENSTEIN. LITHUANIA. LUXEMBOURG. MACEDONIA. MOLDOVA. NETHERLANDS. NORWAY. POLAND. ROMANIA. RUSSIA. SERBIA & MONTENEGRO. SLOVAKIA. SPAIN. SWEDEN. SWITZERLAND. TURKEY. UKRAINE. UNITED KINGDOM. Remarks: This species is very variable in color, body size and in the shape of head and pronotum (Figs. 17I, 17J). Unicolored specimens occur mainly in Mongolia.

Agrilus presbys Jendek, sp. nov. Fig. 14H (habitus of HT)

Description: 7.2-7.5 mm (Holotype 7.3 mm). BODY: Shape: cuneiform, Build: robust, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture

161

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA on vertex: rugae, Sculpture (aspect): subparallel. Eyes: Size: large, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short, Width: stout. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: absent, Position to anterior angles: not reaching level of anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: strongly convex, Disk impressions: absent or lateral only, Lateral impressions (deepness): shallow. Prehumerus: absent. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior third of pronotal length, Modifications: submarginal carina divergent at posterior angles. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Width: wide, Shape: arcuate. Elytral pubescence: absent. STERNUM: Metasternal projection: impressed. Prosternal lobe: large, Anterior margin: arcuate. Prosternal process: moderate, Shape: narrowed, Sides: straight, Angles: absent or obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Ovipositor: uritiform. Diagnosis: body robust, less elongate; elytra glabrous; elytral apices broad, sharply denticulate. Species group: Subrobustus. Relevant species: A. yunnanicola Obenberger, 1936. Type locality: China, Kouy-Tschéou [= Guizhou province]. Type specimens: Primary: Holotype (Fig. 14H), ♀, (EJCB): “Kouy-Tschéou, R.P.J.R. Chaffanjon, 1903”. Secondary: CHINA: Guizhou: 1 ♀ PT (EJCB): “Kouy-Tschéou, R.P.J.R. Chaffanjon, 1903”; 1 PT (MNHN): “Kouy-Tschéou, P. Cavalerie 1910”. Adult occurence: Unknown. Host plants: Unknown. Distribution: CHINA: Guizhou. Etymology: The specific name is the Greek noun presbys (old man, elder) in apposition. It refers to the age of the type specimens, which were collected more than one hundred years ago.

Agrilus priamus Kerremans Fig. 15H (habitus); Fig. 34V (aedeagus); Fig. 51G (type) priamus Kerremans, 1912 (Agrilus) KERREMANS, 1912: 207 208 [#1377] (description). MIWA, 1931: 124 [#6101] (catalog; Formosa). MIWA & CHÛJÔ, 1936: 19 [#1809] (catalog; Japan). OBENBERGER, 1936: 1099 [#2191] (world catalog). BELLAMY, 1994: 373 [#237] (two presumable paralectotypes in USNM). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Taiwan). MÜHLE, 2003: 47 [#6155] (checklist; Taiwan). JENDEK,

2006: 401 [#7441] (Palaearctic catalog). [#8576] (world catalog).

BELLAMY, 2008: 2246

Type material: Agrilus priamus Kerremans, 1912 Type locality: Kosempo et Alikang. Type specimens: Primary: Lectotype (Fig. 51G) by present designation, ♂, (MNHN): “Kosempo Formosa H. Sauter, 1911 [p] \ Priamus Kerr Type [h] \ Museum Paris coll. Ch. Kerremans 1923 [p] [yellow label]”. Secondary: 2 PLT (USNM); 5 PLT (ISNB); 2 PLT (NMPC); 6 PLT (MNHN); 14 PLT (DEI). Described from unknown number of syntypes. Diagnosis: 11-13 mm; prehumerus absent; paramerae of aedeagus with inconspicuous sensoric zone. Species group: Smaragdinus. Relevant species: A. loosdregti Baudon, 1965. Specimens examined: CHINA: Taiwan: 1 ♂ (EJCB): “Sengpei, Kaohsiung Hsien, Taiwan, 2.x.1980, W. Chen”.

Adult occurence: 10 (n=1). Host plants: Unknown. Distribution: CHINA: Taiwan. Remarks: Not recorded from East Asia but possible in South China. Record from Yunnan (China) cited by JENDEK, 2006 [#7441] pertains to A. loosdregti.

Agrilus proditor Obenberger Fig. 9D (habitus of LT) proditor Obenberger, 1917 (Agrilus) OBENBERGER, 1917: 53 54 [#2000] (description). OBENBERGER, 1924: 38 [#2040] (note). OBENBERGER, 1926: 653 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1023 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 357 [#2429] (checklist; China). JENDEK, 1995: 144 [#3768] (lectotype designation; characters). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2247 [#8576] (world catalog).

Type material: Agrilus proditor Obenberger, 1917 Type locality: China. Type specimens: Primary: Lectotype (Fig. 9D), ♀, designated by JENDEK (1995: 144) [#3768], (NMPC): “China [h] \ Typus [p] [red label] \ Agrilus proditor m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 26216 [h] [orange label] \”. Described from unknown number of syntypes. Diagnosis: 9.6 mm; prehumerus absent, pronototum with obvious anterior medial lobe; humeri deeply impressed; prosternal lobe widely arcuate; prosternal process wide, without lateral angles. Species group: Biguttatus. Relevant species: A. magnus sp. nov. Specimens examined: Known only from the lectotype. Host plants: Unknown. Distribution: CHINA.

162

Agrilus prolatus Jendek, sp. nov. Fig. 29J (habitus of HT); Fig. 41A (aedeagus)

Description: 3.7-4.8 mm (Holotype 4.1 mm). BODY: Shape: cuneiform, Build: slender, Color (dorsally): unicolored, Color (dominant): green. HEAD: Medial impression: absent, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: punctures, Sculpture (density): sparse, Sculpture (intensity): superficial. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles or projecting beyond anterior pronotal angles. Posterior angles: Shape: obtuse, rarely rectangular, Apex: sharp. Disk: Convexity: strongly convex, Disk impressions: medial and lateral, Medial impression (shape): entire or anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow, Lateral impressions (width): narrow. Prehumerus: obsolete or carinal, Shape: arcuate, Extent: to half of pronotal length, Posterior end: distant from pronotal angle or margin, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: moderate, Convergence: strongly convergent, Junction: present, Narrowest point: at posterior third of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: obsolete or present. Apices: Arrangement: separate, Width: narrow, Shape: arcuate. Elytral pubescence: entire except epipleura, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate or large, Anterior margin: arcuately emarginate, Emargination (deepness): shallow, Emargination (width): narrow. Prosternal process: moderate, Shape: dilated, Sides: straight, Angles: obtuse, Angles (tips): sharp, Disk: impressed. ABDOMEN: Sexual modifications in male: pair of medial tubercles on ventrite 1, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate or subtruncate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: elongate. Diagnosis: body lustrous; pronotal sides feebly arcuate; humeral carina long; elytral apices finely separately arcuate; ventrite 1 in male with tubercles; aedeagus expanded apically. Species group: Prolatus. Relevant species: A. qiang sp. nov. Type locality: China, Yunnan province, 100 km west of Kunming, Diaolin Natural Reservation.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Type specimens: Primary: Holotype (Fig. 29J), ♂, (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”. Secondary: CHINA: Yunnan: 2 PT (EJCB): “Yunnan, 8-10.Jun, Yipinglang, 1800 m, 25.05 N 101.53 E, Bolm lgt., 1993”; 2 PT (EJCB): “Yunnan 2000m, 25.03N 101.55E, Yipinglang 8-10/6.,Vít Kubáň leg. 1993”; 6 PT (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”. Adult occurence: 5-6 (n=3). Altitude range: 1800-2000 m (n=2). Host plants: Unknown. Distribution: CHINA: Yunnan. Etymology: The specific name is the Latin adjective prolatus (extended, prolonged), referring to the markedly elongate body of the species.

Agrilus prosternalis Jendek, sp. nov. Fig. 11D (habitus of HT); Fig. 33D (aedeagus)

Description: Holotype 6.6 mm. BODY: Shape: subparallel, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Medial impression: absent, Epistoma: with raised upper margin. Frons: Shape: flat or moderately convex, Outline: not protruding from head outline. Vertex: Sculpture on vertex: rugae, Sculpture (aspect): irregular, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: flat, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial. Prehumerus: obsolete or filamentary, Shape: arcuate, Extent: to third of pronotal length, Modifications: obsolete in anterior part, Posterior end: joining posterior pronotal angle, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate, Width: wide, Shape: subtruncate, Truncation: transverse. Elytral pubescence: proximal and distal, Proximal: perisutural stripes, Distal: perisutural stripe, Color: unicolored, Character: with spots of denser pubescence. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: narrow, Anterior margin: arcuately emarginate, Emargination (deepness): deep, Emargination (width): wide. Prosternal process:

163

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA wide, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: impressed with sides protruding, Modification: sides crenulate. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): distinctly longer than mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed.

“Mongolia, Vostochn. aimak, Tamsag Bulak, 1976, Kerzhner, 21.vi. [in Russian]”; 1 ♂ (ZIN): “Mongolia, Vostochnyi aimak, Tamsag Bulak, 1976, M. Kozlov, 21.vi. [in Russian]”. 2 ♂ (ZIN): “Mongoliya, Sukhè Bator. aim., peski Ongon Èls, 15 km YuYuV Khongora, Kerzhner, 5 6.vii.[1]971 [in Russian]”; 1 ♂ (ZIN): “Mongoliya, Sukhè Batorskii aimak, 25 km SZ Bayan Dèlgèra, Emel’yanov, 19.viii.[1]975 [in Russian]”; 1 ♂ (ZIN): “Mongoliya, Sukhè Batorskii aimak, 80 km SSV Barun Urta, Gur’eva, 19.viii. [1]975 [in Russian]”.

Adult occurence: 5-6-7-8 (n=6).

Diagnosis: prosternal process in male deeply impressed with exserted sides. Species group: Quadrisignatus. Relevant species: A. sospes Lewis, 1893.

Host plants: Ajania fruticulosa: ALEXEEV & VOLKOVITSH, 1989 [#3241] (as A. aerosus). – Nitraria: ALEXEEV, 1975 [#52] (as A. aerosus); ALEXEEV & VOLKOVITSH, 1989 [#3241] (under A. aerosus).

Type locality: China, northwest Yunnan province, Dequen, altitude 3900 m.

Distribution: CHINA: Nei Mongol. MONGOLIA.

Type specimens: Holotype (Fig. 11D), ♂, (EJCB): “China, pr. Yunnan b. occ., Dequen, 3900 m, 7.VI.1993, R. Červenka lgt.”.

Remarks: All specimens of A. przewalskii, examined in ZIN, have been misdetermined by Alexeev as A. aerosus.

Adult occurence: 6 (n=1). Altitude range: 3900 m (n=1).

Agrilus pseudobscuricollis Jendek

Host plants: Unknown.

Fig. 24D (habitus of HT); Fig. 38K (aedeagus)

Distribution: CHINA: Yunnan.

pseudobscuricollis Jendek, 2007 (Agrilus) JENDEK, 2007: 38 39 [#8006] (description). [#8576] (world catalog).

Etymology: The specific name is derived from the Latin noun prosternum (the ventral sclerite of the prothorax of insects). It refers to the deeply impressed prosternal process of the species.

Agrilus przewalskii Obenberger Fig. 21I (habitus of LT); Fig. 37F (aedeagus) przewalskii Obenberger, 1936 (Agrilus) OBENBERGER, 1936: 112 113 [#2205] (description). OBENBERGER, 1936: 1023 [#2191] (Agrilus; world catalog). ALEXEEV, 1975: 156 157 [#52] ([Note: Misidentified as aerosus]; notes; faunal records; distributional overview; Mongolia). ALEXEEV & VOLKOVITSH, 1989: 315, 344, 356 [#3241] ([Note: Misidentified as aerosus]; characters in key; faunal records; Mongolia). ALEXEEV, 1998: 375 [#4322] (subgenus Xeragrilus; [Note: Misidentified as aerosus ; subgeneric classification). JENDEK, 2006: 401 [#7441] (Agrilus; Palaearctic catalog). BELLAMY, 2008: 2248 [#8576] (Agrilus; world catalog).

Type material: Agrilus przewalskii Obenberger, 1936 Type locality: Asia centralis: Ala-Shan. Type specimens: Primary: Lectotype (Fig. 21I) by present designation, ♂, (NMPC): “Asia centr. Ala-Shan [h] \ Typus [p] [red label] \ Agrilus Przewalskii m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24320 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 4.4-5.4 mm; pronotum and elytra obviously pubescent; elytral humeral part with longer pubescence; ventral side with tomentum on thoracal parts; prehumerus absent. Species group: Albogularis. Relevant species: A. aerosus Ganglbauer, 1889. Specimens examined: MONGOLIA: 1 ♂ (EJCB): “Mongolia, Bayankhongor, 6.7.1975, 20 km SW lake Bon Cagan”. 6 (ZIN):

BELLAMY, 2008: 2249

Type material: Agrilus pseudobscuricollis Jendek, 2007 Type locality: China, Yunnan, 100 km W of Kunming, Diaolin Nat. Reservation. Type specimens: Primary: Holotype (Fig. 24D), ♂, (EJCB): “China,Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation”. Secondary: 1 ♂, 1 ♀ PT (CBCS); 2 ♂ PT (SOCI); 27 PT (EJCB). Described from 32 specimens (HT, PT). Diagnosis: 3.7-4.8 mm; body slender; elytral apices narrower; aedeagus feably asymmetrical. Species group: Sulcicollis. Relevant species: A. fissus Obenberger, 1917. Specimens examined: CHINA: Sichuan: 1 ♀ (NMW): “China: Sichuan, 15.6.1996, S Chengdu, Emei Shan, 700m 1000m above Fuhu Monastery, leg. Ji & Wang (CWBS 238)”. Yunnan: 31 PT (EJCB): “China,Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation” (JENDEK, 2007 [#8006]). Adult occurence: 5 - 6 (n=2). Altitude range: 700-1000 m (n=1). Host plants: Unknown. Distribution: CHINA: Yunnan, Sichuan.

Agrilus pseudoconstantini Alexeev pseudoconstantini Alexeev, 1981 (Agrilus; subgenus Xeragrilus) ALEXEEV, 1981: 7 9 [#62] (description). ALEXEEV, 1989: 485 [#64] (key to species; Yu Primorsk). JENDEK, 2006: 396 [#7441] (subgenus Xeragrilus; Palaearctic catalog). BELLAMY, 2008: 2249 [#8576] (subgenus Xeragrilus; world catalog).

164 Type material: Agrilus pseudoconstantini Alexeev, 1981 Type locality: Ussuriisk [in Russian]. Type specimens: Not located. Holotype not in ZIN as declared in the description. Described from 5 specimens (HT, PT). Diagnosis: 6.1-7 mm; see ALEXEEV, 1981[#62] and Remarks. Species group: Unassigned. Relevant species: Unknown. Specimens examined: none.

Host plants: Caragana: ALEXEEV, 1989 [#64]. – C. arbo rescens: ALEXEEV, 1981 [#62]. Distribution: RUSSIA: Primorskiy kray. Remarks: Based on the characters given by ALEXEEV, 1981[#62], this species might be related to those in the Albogularis or Hyperici species-groups.

Agrilus pseudolimoniastri Cobos Fig. 21L (habitus); Fig. 37I (aedeagus); Fig. 51J (type) pseudolimoniastri Cobos, 1968 (Agrilus; subgenus Agrilus) COBOS, 1968: 401 404 [#580] (description). COBOS, 1972: 51 52, 56 [#591] (subgenus Agrilus; faunal record; Mongolia). ALEXEEV, 1975: 159 [#52] (notes; faunal records; Mongolia). ALEXEEV & VOLKOVITSH, 1989: 322, 351, 357 [#3241] (characters in key; faunal records; Mongolia). ALEXEEV, 1998: 382 [#4322] (subgenus Spiragrilus; subgeneric classification). NONNAIZAB et al., 1999: 113 [#8247] (checklist; China: Inner Mongolia). JENDEK, 2006: 395 [#7441] (subgenus Spiragrilus; Palaearctic catalog). BELLAMY, 2008: 2250 [#8576] (subgenus Spiragrilus; world catalog).

Type material: Agrilus pseudolimoniastri Cobos, 1968 Type locality: Ostgobi aimak, 54 km. NNW. v. Sainschand, 970 m. alt. Type specimens: Primary: Holotype (Fig. 51J), ♂, (HNHM): “MONGOLIA, Ostgobi aimak, 54 km. NNW. v. Sainschand, 970 m. Exp. Dr. Z. KASZAB, 1963 [p] \ Nr. 50, .29. VI. 63 [p] \ HOLOTYPUS A. Cobos [p] [red label] \ Agrilus pseudolimoniastri nov. sp. Holotypus [h] A. Cobos det. 196 [p] 8 [p]”. Secondary: PT (MNMS); PT (NHMB); 2 PT (MNHN); 29 PT (HNHM). Described from 47 specimens (HT, PT). Diagnosis: 4.7-7.6 mm; posterior pronotal angles obtuse or subrectangular; extent of pubescence and tomentum varies considerably from discrete to obvious. Species group: Hyperici. Relevant species: A. unguiculosus Obenberger, 1936. Specimens examined: CHINA: Gansu: 1 ♀ (EJCB): “China, Gansu, 18.vi.1988, Lanzhou #226, 1700 1900m, Eckweiler leg.”; 1 ♂ (EJCB): “China, vi.1991, Linxia, Gansu, 2000 m”; 1 ♂ (EJCB): “China, S. Gansu, Dengkaguin, 3 9.6.1996, Beneš & Štepař lgt”; 3 ♂, 1 ♀ (EJCB): “China Gansu, Dingxi, 23.5. 30.5.1996, E. Kučera leg.”. KAZAKHSTAN: 1 ♀ (EJCB): “Kazachstan (USSR), Zailijskij Ala Tau, Zsalanasch, 1 3.6.[19]91, Beneš lgt.”. MONGOLIA: 1 ♂, 1 ♀ (EJCB): “MGL Bayankhongor, 75 km S Bayankhongor, N45º20’, E100º48.5’, 1330m, 8.VII.2004, J. Halada leg.”. 7 (NMPC): “Mongolia, Bulgan ajmak, 19.vi.1975, river Tuul, 15 km NE “Telengijn Bajtin [from Russian]”“; 2 (NMPC):

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA “Mongolia, Bulgan, 16.6.1975, river Tuul, “Telengijn Bajtin [from Russian]”“. 1 ♀ (EJCB): “Mongolia: Südgobi aimak, Zöölön ul., 58 km WSW von Somon Bajandalaj, 1500 m, Exp. Dr. Z. Kaszab, 1967, Nr. 808, 16.vi.1967”; 1 ♀ (ZIN): “MNR Centr., Undisul, 29.7.1975 [in Russian]”; 1 ♂, 1 ♀ (ZIN): “Mongolia, 25.vi.1976, Vostochnyi aymak, oz. Khukh Nur, M. Kozlov leg. [in Russian]”; 1 ♂ (EJCB): “Mongoliya, Vostochnyi aimak, oz. Khukh Nur, 25.vi.1976 [in Russian], on Caragana microphyla”; 1 ♂ (ZIN): “MNR Centr., Undisul, 16.7.[19]76 [in Russian]”; 1 ♀ (ZIN): “Mongolia, 25.vi.1976, Vostochnyi aymak, oz. Khukh Nur, Kerzhner leg. [in Russian]”; 1 ♀ (ZIN): “Mongolia, r. Tola, 23.6.1978, Guryeva [in Russian]”; 1 ♂ (EJCB): “Mongolia bor., Džargalant, 10.7.[19]87, O. Majzlan lgt.”; 1 ♂, 2 ♀ (EJCB): “Mongolia mer., Šara chulany Bul., 7.7.[19]88, O. Majzlan leg.”; 1 ♀ (ZIN): “Daurskie stepi, Vost. bereg o. Barun Gorei, Teli, 20.vi.[19]89, I. Kostyuk [in Russian]”. RUSSIA: Altay Kray: 4 ♂, 2 ♀ (EJCB): “SE Altay, Kosh Agach, 49º59’N, 88º38’E, 15 16.vi.1989”. Tyva Respublika: 1 ♀ (EJCB): “Tuva, Kyzyl, right bank, 18.vi.1979, on Caragana splendens”; 1 ♂ (EJCB): “Tuva, 12.vi.1979, 25 km NW of Kyzyl, Seserlig ( Germanovka), Caragana sp., dried steppe”; 1 ♀ (EJCB): “Tuva, 18 20.vi.1994, W of Tanu Ola, 30 km N Samagaltaj, Z. Kletečka leg.”; 1 ♀ (EJCB): “Russia, Tuva, right bank of Biy Khem riv. 800 m, Ujukskiy Mts. 22.V. 6.VI.1999, S. N. Vashchenko leg.”.

Adult occurence: 5-6-7 (n=23). Altitude range: 800-2000 m (n=6). Host plants: Caragana leucophloea ALEXEEV & VOLKOVITSH, 1989 [#3241]. – C. microphylla: ALEXEEV & VOLKOVITSH, 1989 [#3241]. Distribution: CHINA: Gansu. KAZAKHSTAN. MONGOLIA. RUSSIA: Altay Kray, Tyva Respublika. Remarks: The distributional record without supporting data cited from Inner Mongolia (China) by NONNAIZAB et al., 1999 [#8247] was omitted. Specimens from China (Gansu) are assigned to this species rather tentatively as they somewhat differ by the shape of prosternal process and prosternal lobe.

Agrilus pseudosericans Obenberger Fig. 21J (habitus of LT); Fig. 37G (aedeagus); Figs. 21J, 42S (types) pseudosericans Obenberger, 1936 (Agrilus) OBENBERGER, 1936: 107 108 [#2205] (description). OBENBERGER, 1936: 1025 [#2191] (world catalog). ALEXEEV & VOLKOVITSH, 1989: 319, 348, 356 [#3241] (lectotype designation; characters in key; faunal records; Mongolia; Tuva; Kazakhstan; Kyrgyzstan; east Gruzia). OVTCHINNIKOV, 1996: 132 [#8621] (checklist; Kyrgyzstan). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). JENDEK, 2006: 394 [#7441] (subgenus Robertius; Palaearctic catalog). BELLAMY, 2008: 2251 [#8576] (subgenus Robertius; world catalog). angulicollis Cobos, 1968 (Agrilus; subgenus Agrilus) COBOS, 1968: 393 396 [#580] (description). COBOS, 1972: 56 [#591] (subgenus Agrilus; checklist; Mongolia). ALEXEEV, 1975: 152 [#52] (faunal records; Mongolia). ALEXEEV & VOLKOVITSH, 1989: 319 [#3241] (synonym of pseudosericans; type examination). OVTCHINNIKOV, 1996: 132 [#8621] (synonym of pseudosericans). JENDEK, 2006: 394 [#7441] (synonym of pseudosericans; Palaearctic catalog). BELLAMY, 2008: 2251 [#8576] (synonym of pseudosericans; world catalog).

165

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Type material: Agrilus pseudosericans Obenberger, 1936 Type locality: Rossia asiatica: Věrnenskij Újezd: Bajzerka. Type specimens: Primary: Lectotype (Fig. 21J), ♂, designated by ALEXEEV & VOLKOVITSH (1989: 319, 348, 356) [#3241], (NMPC): “Върнепскій үъздь Урочище Байсерке 1907 В. Недзељцкій [h] \ Typus [p] [red label] \ Agrilus pseudosericans m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24952 [h] [orange label]”. Described from unknown number of syntypes. Agrilus angulicollis Cobos, 1968 Type locality: Bajanchongor aimak, 8 km S von somon Zinst, 1.400 m. alt. Type specimens: Primary: Holotype (Fig. 42S), ♂, (HNHM): “MONGOLIA, Bajanchongor aimak, 8 km S von somon Žinst, 1.400 m. Exp. Dr. Z. KASZAB, 1964 [p] \ Nr. 197, 25.VI. 1964 [p] \ HOLOTYPUS A. Cobos [p] [red label] \ Agrilus angulicollis nov. sp. Holotypus [h] A. Cobos det. 196 [p] 8 [p]”. Secondary: PT (MNMS); 1 PT (HNHM). Described from 4 specimens (HT, AT, PT).

357 [#2429] (checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Zhejiang). JENDEK, 2005: 16 [#7217] (lectotype designation). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2251 [#8576] (world catalog).

Diagnosis: 4.6-6.2 mm; head markedly convex; vertex without or with very fine medial impression; eyes very small; pronotum widest in anterior third; posterior pronotal angles sharp; prehumerus long, reaching to anterior pronotal angle; median lobe of aedeagus shorter than parameres, sharp on apex. Species group: Albogularis. Relevant species: A. albogularis Gory, 1841; A. sericans Kiesenwetter, 1857.

Distribution: CHINA: Zhejiang.

Specimens examined: KAZAKHSTAN: 1 (ZIN): “r. Kendyrlichka, 10 km VYUV Przhevalskogo, khr. Saur, V. Kazakh., Volkovich, 26 27.VII. [19]83 [in Russian]”; 5 (ZIN): “khr. Tarbagatai, 45 km VSV p. Tarbagatai, Kazakhstan, Volkovich 21 22.vii.[1]983 [in Russian]”; 6 (ZIN): “khr. Tarbagatai, 30km ZSZ p. Tarbagatai, Kazakhstan, Volkovich 19 20.vii.[1]983 [in Russian]”; 4 (ZIN): “khr. Tarbagatai, 40 km V p. Tarbagatai, V. Kazakhstan, Volkovich, 14 15.07.[1]988 [in Russian]”; 2 ♂ (EJCB): “E Kazakhstan, 400 m, Ust Kamenogorsk env., Siinka riv., Bazombai, M. Danilevsky 19.6.2002”. KYRGYZSTAN: 1 ♀ (NMPC): “USSR, Kirghizia, 30 km S of Frunze, 2 4.7.1981, 600 m, Vít Kubáň leg.”; 2 (ZIN): “40 km V pos. Tyup, yuzh. sklon khr. Kungei Alatoo, Kirgiziya, Volkovich 14.vii.[1]986 [in Russian]”; 1 ♂ (ZIN): “okr. pos. Kyzyl Oi, dolina r. Susamyr, Kirgizia, 1986, Volkovich, 5 6.VII. [in Russian]”; 6 (MKCY): “Kirgizstan, env. Przhevalsk, Dzhety Oguz, 29.07.1991, Napolov leg.”. MONGOLIA: Bayanhongor: 2 (EJCB): “Mongolia, Bayankhongor, 6.7.1975, 20 km SW lake Bon Cagan”.

Adult occurence: 6-7 (n=10). Altitude range: 400-1400 m (n=3). Host plants: Caragana: ALEXEEV & VOLKOVITSH, 1989 [#3241]. Distribution: KAZAKHSTAN. KYRGYZSTAN. MONGOLIA. RUSSIA: Tyva Respublika.

Agrilus pterostigma Obenberger Fig. 26F (habitus of LT); Fig. 39H (aedeagus) pterostigma Obenberger, 1927 (Agrilus) OBENBERGER, 1927: 17 18 [#2060] (description). OBENBERGER, 1936: 1025 [#2191] (world catalog). PENG ZHONGLIANG, 1987:

Type material: Agrilus pterostigma Obenberger, 1927 Type locality: China: Ning-Po. Type specimens: Primary: Lectotype (Fig. 26F), ♂, designated by JENDEK (2005: 16) [#7217], (NMPC): “China: Ning Po [h] \ Typus [p] [red label] \ A. pterostigma m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24886 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 6.2 mm; body more elongate; pronotum visually transverse; distal part of elytral pubescence limited; medial lobe of aedeagus wide. Species group: Cyaneovirens. Relevant species: A. terracotta sp. nov. Specimens examined: Known only from the lectotype.

Host plants: Unknown.

Agrilus pumi Jendek, sp. nov. Fig. 13G (habitus of HT); Fig. 34B (aedeagus)

Description: 6.0-6.4 mm (Holotype 6.2 mm). BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Medial impression: absent, rarely present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Sculpture on vertex: punctures or rugae, Sculpture (aspect): irregular or subparallel. Eyes: Size: moderate, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square or transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles or at level with anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, rarely entire, Medial impression (deepness): shallow, Anteromedial and posteromedial impressions: subequal in size, Lateral impressions (deepness): shallow, Lateral impressions (width): narrow. Prehumerus: obsolete, filamentary or carinal, Shape: arcuate or bisinuate, Extent: to third of pronotal length or to anterior pronotal angle, Modifications: rarely with rudiment at anterior pronotal angle or obsolete in anterior part, Posterior end: joining posterior pronotal angle, Anterior end: distant from pronotal marginal carina, joining pronotal marginal carina or joining anterior pronotal angle, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior

166 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, rarely unicolored with darker hue in apical half, Humeral carina: absent. Apices: Arrangement: separate, Width: narrow, Shape: arcuate, subangulate, rarely cuspidate, Position of dominant cusp (spine): medial. Elytral pubescence: entire except glabrous part at apical half, Color: unicolored, Character: with patches of denser pubescence. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate or large, Anterior margin: arcuately emarginate, Emargination (deepness): moderate or deep, Emargination (width): moderate or wide. Prosternal process: moderate, Shape: narrowed, Sides: straight, Angles: absent, Disk: flat, Modification: disk in male with two smooth glabrous protuberances. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): shorter or subequal to 2-3 combined or subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed, Ovipositor: elongate. Diagnosis: prosternal process narrowed apically, without angles, disk flat, with two smooth glabrous protuberances in male. Species group: Plagiatus. Relevant species: A. miao sp. nov. Type locality: China, Yunnan province, Jizushan, 25°58’N, 101°21’E, altitude 2500-2700 m. Type specimens: Primary: Holotype (Fig. 13G), ♂, (EJCB): “Yunnan, 2500-2700m, 25.58N, 100.21E, Jizushan, 6-10. vii.1994, Vít Kubáň leg.”. Secondary: CHINA: Yunnan: 1 ♂ PT (EJCB): “Yunnan, 2500-2700m, 25.58N, 100.21E, Jizushan, 6-10.vii.1994, Vít Kubáň leg.”. Adult occurence: 7 (n=1). Altitude range: 2500-2700 m (n=1). Host plants: Unknown. Distribution: CHINA: Yunnan. Etymology: Noun in apposition; the specific name refers to the Pumi nationality living in Yunnan province of China.

Agrilus pusillesculptus Obenberger Fig. 28J (habitus); Fig. 40N (aedeagus); Fig. 51M (type) pusillesculptus Obenberger, 1940 (Agrilus) PENG OBENBERGER, 1940: 177 178 [#2273] (description). ZHONGLIANG, 1987: 357 [#2429] (checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2256 [#8576] (world catalog).

Type material: Agrilus pusillesculptus Obenberger, 1940 Type locality: China bor. occ. Tien-mu-shan.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Type specimens: Primary: Lectotype (Fig. 51M) by present designation, ♂, (NMPC): “Tienmushang China b. occ. [h] \ Typus [p] [red label] \ Agrilus pusillesculptus m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24987 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 4-5.6 mm; anterior margin of prosternal lobe subtruncate; prosternal process enlarged, with sharp lateral angles. Species group: Viridiobscurus. Relevant species: A. tai sp. nov. Specimens examined: CHINA: Shaanxi: 17 (EJCB): “China S, Shaanxi, 14 16.6.1995, Qinling Mts. N. slope, 33.56 59’/108.50’ 800 1200 m, Changan Co., L.+R. Businský lgt.”; 1 ♂ (EJCB): “China, Shaanxi prov., 21 23.vi.1998, Quin Ling Shan mts, road Baoji Taibai vill., pass 35 km S Baoji, O. Šafránek et M. Trýzna lgt.”; 2 ♀ (EJCB): “China, 1150 1300 m, Shaanxi, Qinling mts., Foping (6 km N), 20 21.VI.1998, I. H. Marshal leg.”; 7 (EJCB): “C China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000 1300m, 23.V. 13.VI. leg. C. Holzschuh 2000”; 1 ♀ (EJCB): “China, Shaanxi prov., 21 23.June 1998 \ Qing Ling Shan mts, 1000 m road Baoji Taibai vill., pass 40 km S Baoji, Zd. Jindra lgt.”.

Adult occurence: 6 (n=5). Altitude range: 800-1300 m (n=4). Host plants: Unknown. Distribution: CHINA: Shaanxi, Zhejiang.

Agrilus qiang Jendek, sp. nov. Fig. 29K (habitus of HT); Fig. 41B (aedeagus)

Description: 3.4-4.6 mm (Holotype 4.0 mm). BODY: Shape: subparallel, Build: slender, Color (dorsally): unicolored, Color (dominant): green. HEAD: Medial impression: absent or present, Impression (extent): vertex, Impression (depth): shallow. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: protruding from head outline or not protruding from head outline, Sculpture on vertex: punctures, Sculpture (density): sparse. Eyes: Size: moderate, Shape: not protruding from head outline, rarely protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: lateral only, Lateral impressions (deepness): shallow. Prehumerus: obsolete or carinal, Shape: straight or arcuate, Extent: to half of pronotal length, Posterior end: distant from pronotal angle or margin, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior angles or at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Shape:

167

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA arcuate, subangulate or subtruncate, Truncation: oblique with projecting pleural angle. Elytral pubescence: entire, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: narrow, Anterior margin: arcuately emarginate, Emargination (deepness): moderate. Prosternal process: narrow, Shape: narrowed, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: subtruncate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: uritiform. Diagnosis: body less lustrous; eyes more convex; pronotal sides markedly arcuate; pronotal disk less convex; humeral carina absent; elytral apices subangulately arcuate or obliquely subtruncate; ventrite 1 in male without tubercles; aedeagus expanded apically. Species group: Prolatus. Relevant species: A. prolatus sp. nov. Type locality: China, Sichuan province, Luding env., about 30 km southestern-estern of Kangding, 30°00’N, 102°20’E. Type specimens: Primary: Holotype (Fig. 29K), ♂, (EJCB): “China, Sichuan, 10.vii.1995, Luding env., cca 30 km SEE of Kangding, 30°00’N, 102°20’E, M. Trýzna et O. Šafránek”. Secondary: CHINA: Sichuan: 3 PT (JMCZ): “China, Sichuan, 10.vii.1995, Luding env., cca 30 km SEE of Kangding, 30°00’N, 102°20’E, M. Trýzna et O. Šafránek”; 49 PT (EJCB): “China, Sichuan, 10.vii.1995, Luding env., cca 30 km SEE of Kangding, 30°00’N, 102°20’E, M. Trýzna et O. Šafránek”; Yunnan: 2 ♂ PT (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”. Adult occurence: 5-6-7 (n=3). Host plants: Unknown. Distribution: CHINA: Sichuan, Yunnan. Etymology: Noun in apposition; the specific name refers to the ethnic group Qiang living in the northwestern part of Sichuan province of China.

Agrilus qinling Jendek Fig. 8H (habitus of HT); Fig. 32E (aedeagus) qinling Jendek, 2000 (Agrilus) JENDEK, 2000: 190, 192 [#4852] (description). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2257 [#8576] (world catalog).

Type material: Agrilus qinling Jendek, 2000 Type locality: China, 1000-1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E). Type specimens: Primary: Holotype (Fig. 8H), ♂, (EJCB): “China, 1000 - 1300 m, Shaanxi [prov.], Qinling mts.,

Xunyangba (6 km E), 23. v.-13.vi.1998, I. H. Marhshal leg.”. Secondary: 6 PT (EJCB); 1 PT (TSCB). Described from 8 specimens (HT, PT). Diagnosis: 10.3-13.6 mm; vertex convex with distinct medial carina; eyes protruding beyond head outline; elytral apices widely subarcuate; pronotal sides with large golden tomentose patches. Species group: Cyaneoniger. Relevant species: A. agnatus Kerremans, 1892; A. lafertei Kerremans, 1892. Specimens examined: Known only from the type specimens.

Adult occurence: 5-6 (n=1). Altitude range: 1000-1300 m (n=1). Host plants: Unknown. Distribution: CHINA: Shaanxi.

Agrilus qinlingcola Jendek, sp. nov. Fig. 27L (habitus of HT); Fig. 40E (aedeagus)

Description: Holotype 4.5 mm. BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): green or brown. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (intensity): superficial. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: in line with antennal socket or below antennal socket. Antennae: Length: short or moderate. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: vague, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles or at level with anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Lateral impressions (deepness): shallow, Lateral impressions (width): narrow. Prehumerus: carinal, Shape: arcuate, Extent: to half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: close to pronotal marginal carina or joining pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: conjoint, Shape: subtruncate, Truncation: transverse. Elytral pubescence: absent. Tomentum: absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: rectangular, Angles (tips): sharp, Disk: impressed. ABDOMEN: Apical

168 margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed. Diagnosis: antennae shorter; elytra glabrous; elytral apices truncate with spinuliform sutural rim; sternal groove on last ventrite sinuate. Species group: Cyaneovirens. Relevant species: A. gussakovskiji Alexeev, 1981. Type locality: China, Shaanxi province, Qinling mts., 6 km east of Xunyangba, altitude 1000-1300 m. Type specimens: Primary: Holotype (Fig. 27L), ♂, (EJCB): “China, 1000-1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.-13.vi.1998, I. H. Marshal leg.”. Adult occurence: 5-6 (n=1). Altitude range: 1000-1300 m (n=1). Host plants: Unknown. Distribution: CHINA: Shaanxi. Etymology: The specific name is a combination of Qinling (the mountain range of the type locality) and the Latin verb colo (to dwell, inhabit).

Agrilus quadrisignatus Marseul Fig. 11E (habitus); Fig. 33E (aedeagus); Figs. 48K, 49N, 51Q (types) quadrisignatus Marseul, 1866 (Agrilus) MARSEUL, 1866: 434, 444 445 [#676] (description). POPOFF, 1853: 102 [#7853] (Mannerheim is cited as the author; faunal record; notes; no characters cited; Buryatia). GEMMINGER & HAROLD, 1869: 1444 [#1057] (catalog). SAUNDERS, 1871: 119 [#2552] (catalog). HEYDEN, 1881: 118 [#5548] (catalog). MARSEUL, 1887: 251 [#4707] (Mannerheim is cited as the author; catalog). KERREMANS, 1892: 270 [#1397] (catalog). ABEILLE DE PERRIN, 1897: 3 [#19] (characters in key). KERREMANS, 1903: 290 [#1441] (catalog). JAKOBSON, 1913: 796 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1934: 14 [#2146] (note). MIWA & CHÛJÔ, 1936: ADDENDA P. 4 [#1809] (catalog; Japan). OBENBERGER, 1936: 1026 [#2191] (world catalog). ALEXEEV, 1975: 151 [#52] (faunal records; distributional overview; Mongolia). KUROSAWA, 1975: 2 [#4578] (note). PENG ZHONGLIANG, 1987: 358 [#2429] (Mannerheim is cited as the author; checklist; China). ALEXEEV, 1989: 487 [#64] (key to species; Primorsk; Chita; Buryatia; North Korea; northeastern China, Mongolia). ALEXEEV & VOLKOVITSH, 1989: 319, 348, 356 [#3241] (characters in key; faunal records; Mongolia). JENDEK, 1998: 329 [#4481] (lectotype designation). NONNAIZAB et al., 1999: 113 [#8247] (checklist; China: Inner Mongolia). HUA LI ZHONG, 2002: 90 [#6696] (Mannerheim is cited as the author; checklist; China: Inner Mongolia, Gansu, Jiangsu). JENDEK, 2005: 17 [#7217] (synonymy; notes; faunal records; China (Hebei, Shanxi); Russia (Buryatiya, Primorsk); not in Japan). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (errata for Palaearctic catalog). BELLAMY, 2008: 2259 [#8576] (world catalog). VOLKOVITSH, 2009: 135 [#8850] (faunal records; Russia: Primorskiy kray).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA mongoliae Obenberger, 1922 (Agrilus) OBENBERGER, 1922: 26 27 [#2021] (description). OBENBERGER, 1924: 53 [#2040] (emendation of type locality). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1934: 14 [#2146] (synonym of quadrisignatus). OBENBERGER, 1936: 1026 [#2191] (synonym of quadrisignatus). NONNAIZAB et al., 1999: 113 [#8247] (cited as mongolian; synonym of quadrisignatus). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Inner Mongolia). JENDEK, 2005: 17 [#7217] (synonym of quadrisignatus subspecies quadrisignatus; lectotype designation; notes). JENDEK, 2006: 401 [#7441] (synonym of quadrisignatus quadrisignatus; Palaearctic catalog). BELLAMY, 2008: 2259 [#8576] (synonym of quadrisignatus; world catalog). lama Obenberger, 1935 (Agrilus; subspecies of quadrisignatus) OBENBERGER, 1935: 164 [#2158] (description). OBENBERGER, 1936: 1026 [#2191] (subspecies of quadrisignatus; world catalog). KUROSAWA, 1975: 2 [#4578] (subspecies of quadrisignatus). PENG ZHONGLIANG, 1987: 358 [#2429] (subspecies of quadrisignatus; checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (subspecies of quadrisignatus; checklist; N China). JENDEK, 2005: 17 [#7217] (synonym of quadrisignatus; lectotype designation). JENDEK, 2006: 401 [#7441] (synonym of quadrisignatus quadrisignatus; Palaearctic catalog). BELLAMY, 2008: 2259 [#8576] (synonym of quadrisignatus; world catalog).

Type material: Agrilus quadrisignatus Marseul, 1866 Type locality: Sibérie. Type specimens: Primary: Lectotype (Fig. 51Q), sex not examined, designated by JENDEK (1998: 329) [#4481], (MNHN): “Transcriptio [p] 4-signatus (Manh.) Mars. Type Sibérie [h] \ Muséum Paris 1952 Coll. R. Oberthur [p] [yellow label]”. Described from unknown number of syntypes. Agrilus mongoliae Obenberger, 1922 Type locality: China bor. or.: Chingan [Note: Locality emended to Kiachta by OBENBERGER, 1924 [#2040]]. Type specimens: Primary: Lectotype (Fig. 49N), ♂, designated by JENDEK (2005: 17) [#7217], (NMPC): “Kiachta [h] \ Typus [p] [red label] \ Agrilus Mongoliae m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 26217 [h] [orange label]”. Described from unknown number of syntypes. Agrilus quadrisignatus lama Obenberger, 1935 Type locality: China bor. Type specimens: Primary: Lectotype (Fig. 48K), ♂, designated by JENDEK (2005: 17) [#7217], (NMPC): “China bor. [h] \ Typus [p] [red label] \ A. quadrisignatus subsp. lama m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24283 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 4.9-7.1 mm; body wider; elytra with 1-2 pairs of pubescent spots; apical pair almost joined. Species group: Quadrisignatus. Relevant species: A. sospes Lewis, 1893. Specimens examined: CHINA: Hebei: 3 ♂, 1 ♀ (EJCB): “China, Hebei/Nei Mongol, pass Chengde Chifeng, 41.6N, 118.2E, 14 16. VI., Jaroslav Turna leg. 2001” (JENDEK, 2005 [#7217]). Shaanxi: 1 ♀ (JMCZ): “China c., 17 18.V., Ganguyi 110,3’ 36,8’, 35 km NE of Yanan, leg. J. Halada 1996”. Shanxi: 1 (EJCB): “China, Shanxi, 9.June.2000, Lüliang Shan, 1000m, road Fangshan Jiaocheng, Hengjian env., Zd. Jindra” (JENDEK, 2005 [#7217]); 12 (EJCB): “China, W Shanxi, 9.VI.2000, 37.7N, 111.6E, Lüliang Shan, road Fangshan Jiaocheng, Hengjian env., ~1000m, Jaroslav Turna leg.” (JENDEK, 2005 [#7217]); 1 ♀ (EJCB): “China, centr. Shanxi, Chao

169

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Shan 37.1N, 112.4E, 25 km SE Pingyao., 4 5.VII., Jaroslav Turna leg. 2001” (JENDEK, 2005 [#7217]). MONGOLIA: 4 (EJCB): “Mongolia, Bayankhongor, 6.7.1975, 20 km SW lake Bon Cagan”. 2 (EJCB): “Mongolia, Bulgan, 16.6.1975, river Tuul, “Telengijn Bajtin [from Russian]”“. 4 (EJCB): “Mongolia, Sukhbaatar, Darichanga, 17.vi.1975, 1200 m”; 1 (EJCB): “Mongolia, Arkhangal, 5 km S of lake Bulgana, 2.7.1975”; 6 (EJCB): “Mongolia, Calgant, 10.vii.1975”; 1 (NMPC): “Mongolia, Chilgant, 29.7.1975”. RUSSIA: Buryatiya Respublika: 1 (EJCB): “Transbaicalia mr. occ. fl. Tshikoj: Sharagol 1.VII.1928 Th. Lukjanovitsh” (JENDEK, 2005 [#7217]); 1 (MSCI): “S Burjatia, Kjachtinsky dist., 5 km NE Kiran vill., 50º21’N,106º46’E, 25 30.06.99 leg. Igor Melnik”. Primorskiy kray: 1 (ZIN): “Vinogradovka Ussur. kr. Kirichenko 5.vii.1929 [in Russian]”; 1 ♂ (JMCZ): “SU Far East, Kamenushka, 19.7.1983”; 1 (EJCB): “USSR Sibiria or., Ussuri region, 450 m, Yasnoe, 11 14.7.1989, R. Dunda lgt.” (JENDEK, 2005 [#7217]); 2 (MNHE): “Russia, 27 29.06.1989, Primorye Distr., Buyanki, leg. U. Roosileht”; 8 (EJCB): “Sib. or. m., Primorje, Sichote Alin Mts., Sokolči, 1 15.7.1990, Kadlec + Voříšek lg.” (JENDEK, 2005 [#7217]); 1 ♂ (EJCB): “Russia, Primorsk reg., 30.7. 8.8.1992, Kajmanovka, E. Kučera leg. ex Ulmus” (JENDEK, 2005 [#7217]); 2 ♂, 1 ♀ (EJCB): “Sib. or. Primorje, Komarovka flum., Kamenushka env. 300 m, Voříšek, J. leg. VI.1992” (JENDEK, 2005 [#7217]); 2 (EJCB): “SSSR, Primorskij k., Kajmanovka 10 22.vii.1993, Z. Kletečka leg.” (JENDEK, 2005 [#7217]); 3 (EJCB): “Russia, Primorski kray, Khankajski distr. 1995, lake Khanka, 17 19.vii., Kamen’ Rybolov” (JENDEK, 2005 [#7217]); 6 (EJCB): “Russia, Primorski kray, Ussurijski district, Baranovski volcano env. 4.ix.1995” (JENDEK, 2005 [#7217]); 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Lazo env., 3.07.04 leg. М.& L. Smirnov’s”; 1 ♀ (ZIN): “Primorskii kr. s. Lazo 8 12.vi.2007 V. Shokhrin leg [in Russian]”.

Adult occurence: 5-6-7-8-9 (n=26). Altitude range: 3001200 m (n=5). Host plants: Ulmus: ALEXEEV, 1989 [#64]. – U. macrocarpa: ALEXEEV & VOLKOVITSH, 1989 [#3241]. – U. pumila: ALEXEEV & VOLKOVITSH, 1989 [#3241]. Distribution: CHINA: Gansu, Hebei, Jiangsu, Nei Mongol, Shanxi, Shanxi. KOREA NORTH. MONGOLIA. RUSSIA: Buryatiya Respublika, Primorskiy kray, Zabaykalskiy kray.

Agrilus quentini Descarpentries & Villiers Fig. 15D (habitus); Fig. 34S (aedeagus); Figs. 51S, 54R (types) quentini Descarpentries & Villiers, 1963 (Agrilus) DESCARPENTRIES & VILLIERS, 1963: 4, 16 [#701] (description). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2260 [#8576] (subgenus Sinagrilus; world catalog). wilsoni Baudon, 1968 (Agrilus) syn. nov. BAUDON, 1968: 109, 128 [#162] (description). BAUDON, 1963: 53 [#148] ([Note: Unavailable name, cited without characters]). BELLAMY, 2008: 2363 [#8576] (subgenus Sinagrilus; world catalog).

Type material: Agrilus quentini Descarpentries & Villiers, 1963 Type locality: Tonkin: Hoa Binh. Type specimens: Primary: Holotype (Fig. 51S), ♂, (MNHN): “Hoa Binh Tonkin de Cooman [h] \ Agrilus quentini n. sp. Holotype ♂ nob. A. Descarpentries et [h] A. Villiers det. 19 [p] 63 [h]”. Secondary: 1 PT (MHNG); 1 PT (ISNB); 7 PT (MNHN). Described from unknown number of specimens (HT, AT, PT).

Agrilus wilsoni Baudon, 1968 Type locality: Laos: Vientiane, Muong Panh. Type specimens: Primary: Holotype (Fig. 54R), ♀, (MHNB): “Vientiane 7.iv.61 [h] Laos (Baudon) [p] \ Type [p] [red label] \ Agrilus wilsoni mihi Type [h] A. Baudon det. [p] [blue label]”. Secondary: 1 PT (HHCC); 9 PT (MHNB). Described from unknown number of specimens (HT, AT, PT). Diagnosis: 3.4-5 mm; body less elongate, black; aedeagus flat. Species group: Subrobustus. Relevant species: A. ineptus Kerremans, 1892. Specimens examined: CHINA: Yunnan: 1 ♂ (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”; 6 (MNCA): “China: S Yunnan (Xishuangbanna) 23 km NW Jinghong, vic. Na Ban Village \ N22 10.04E, 100 39.52 (NNR) 680 m, ex Liane, leg. A. Weigel VI/VII.2008”. LAOS: Borikhamxai: 1 ♂ (EJCB): “Laos centr., Bolikhamsai prov., Ban Nape Kaew Nua Pass, 18.4. 1.5.1998, alt. 600±100 m, N 18º22.3, E 105º09.1 (GPS), E. Jendek & O. Šauša leg.”; 1 (EJCB): “Laos C., Bolikhamsai pr., Ban Nape env., 7 16.V.2004, alt., 400±100 m, 18º20’N, 105º08’E, E. Jendek & O. Šauša leg.”. Louang Namtha: 15 (EJCB): “N Laos, 14 22.vi.1996, Louang Namtha pr. 10 30 km NW Louang Namtha, C. Holzschuh leg. 800 m”; 2 ♀ (EJCB): “Laos north, 24 30.V.1997, 20 km NW Louang Namtha, N21°09.2, E101°18.7, alt. 900±100 m, E. Jendek & O. Šauša leg.”; 1 ♂, 2 ♀ (EJCB): “Laos north, 5 11.V.1997, 20 km NW Louang Namtha, N 21°09.2, E 101°18.7, alt. 900±100 m, E. Jendek & O. Šauša leg.”. Louangphrabang: 1 ♂ (EJCB): “Laos N, 23.iv.1999, Louangphrabang prov. 20°42’N 102°54’E, 25 km E Muang Ngoy, 1000 m, Vít Kubáň leg.”. Oudomxai: 1 ♀ (EJCB): “Laos N (Oudomxai), 1 9.v.2002, ~1100m, 20º45’N, 102º09’E, Oudom Xai (17 km NEE), Vít Kubáň leg.”. Phongsali: 1 ♂ (EJCB): “Lao N, Phongsaly prov., 21°41’ 2’N, 102°06’ 08’E, 28.v. 20.vi.2003 Phongsaly env., ~1500m, Vít Kubáň leg.”. NEPAL: Kosi: 1 ♂, 1 ♀ (EJCB): “E Nepal, Koshi, Tamur River, Phulvari Waku, 1200 1600 m, 9.VI., leg. Holzschuh, 1985”; 1 ♂ (EJCB): “Nepal, East Kosi Zone, Dhankuta distr., Arun valley, Hille Shidua, Bhedetar, 2000 2700m, 24 28.v.1996, P. Čechovský lgt.”. THAILAND: 3 ♂ (EJCB): “Thai, 18 24.IV.1991, Lansang n. p. 500 m, 16°48’N, 98°57’E, Vít Kubáň leg.”; 1 (EJCB): “Thailand occ., 08 12.04.1991, Sangklaburi, Jan Farkač leg.”. Kanchanaburi: 5 ♂ (EJCB): “Thai, 9 13.IV.1991, Thimonghta, 350 m, 15°02’N, 98°35’E, Vít Kubáň leg.”. Mae Hong Son: 1 ♀ (EJCB): “NW Thailand, 1 7.V.1992, Mae Hong Son, Ban Si Lang, 1000 m, S. Bílý leg.”; 1 ♂ (EJCB): “Thai, Mae Hong Son pr., 19°27’N, 98°20’E, 1500 m, Soppong 7 12.V., Vít Kubáň leg., 1996”. VIETNAM: 2 ♀ (EJCB): “N Vietnam, 21º27N, 105º39E, 70 km NW of Hanoi, Tam Dao, 1 8.vi.1996, 900 1200 m, Dembický & Pacholátko leg.”.

Adult occurence: 4-5-6-7 (n=18). Altitude range: 3002700 m (n=16). Host plants: Unknown. Distribution: CHINA: Yunnan. LAOS. NEPAL. THAILAND. VIETNAM.

Agrilus recticollis Jendek, sp. nov. Fig. 29L (habitus of HT); Fig. 41C (aedeagus)

Description: 4.2-4.3 mm (Holotype 4.2 mm). BODY: Shape: cuneiform, Build: slender, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Medial impression:

170 absent, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): subparallel, Sculpture (density): dense. Eyes: Size: small, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate or long, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square or transverse, Sides: straight, Maximal width: at posterior margin, Anterior margin: narrower than posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: rectangular, Apex: sharp. Disk: Convexity: strongly convex, Disk impressions: lateral only, Lateral impressions (deepness): shallow, Lateral impressions (width): narrow. Prehumerus: obsolete, Shape: arcuate, Extent: to half of pronotal length, Posterior end: distant from pronotal angle or margin, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior third of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Width: wide, Shape: arcuate. Elytral pubescence: entire, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: large, Anterior margin: arcuately emarginate, Emargination (deepness): shallow. Prosternal process: moderate, Shape: narrowed, Sides: arcuate, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate, Sinuosity (deepness): very shallow, Sinuosity (width): very narrow. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): shorter or subequal to 2-3 combined. GENITALIA: Aedeagus (symmetry): symmetrical. Diagnosis: pronotal sides straight; aedeagus subparallel. Species group: Prolatus. Relevant species: A. prolatus sp. nov.; A. qiang sp. nov. Type locality: China, Yunnan province, 100 km west of Kunming, Diaolin Natural Reservation. Type specimens: Primary: Holotype (Fig. 29L), ♂, (EJCB). Secondary: CHINA: Yunnan: 1 ♀ PT (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”. Adult occurence: 5-6 (n=1). Host plants: Unknown. Distribution: CHINA: Yunnan. Etymology: The specific name was derived from the Latin adjective rectus (right, stright) and the noun collum (neck); it refers to the stright pronotal sides of the species.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Agrilus reichardti Obenberger Fig. 23C (habitus); Fig. 37W (aedeagus); Fig. 51T (type) reichardti Obenberger, 1935 (Agrilus) OBENBERGER, 1935: 165 [#2158] (description). OBENBERGER, 1936: 1026 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 358 [#2429] (checklist; China). ALEXEEV, 1998: 374 [#4322] (subgenus Quercuagrilus; subgeneric classification). JENDEK, 2000: 505 [#4851] (lectotype designation; remarks on type locality). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Sichuan). JENDEK, 2006: 392 [#7441] (subgenus Quercuagrilus; Palaearctic catalog). BELLAMY, 2008: 2263 [#8576] (subgenus Quercuagrilus; world catalog).

Type material: Agrilus reichardti Obenberger, 1935 Type locality: Tatsienlu, Kiulung, China. Type specimens: Primary: Lectotype (Fig. 51T), ♂, designated by JENDEK (2000: 505) [#4851], (NMPC): “China [p] \ Tatsienlu [p] \ Kiulung [p] \ Reitter [p] \ Typus [p] [red label] \ Agrilus Reichardti m.Type [h] [Obenberger’s MS] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24889 [h] [ora Described from unknown number of syntypes. Diagnosis: 4.7-7.4 mm; body slender, subparallel, aedeagus asymmetrical. Species group: Sulcicollis. Relevant species: A. adelphinus Kerremans, 1895. Specimens examined: CHINA: Sichuan: 1 (GNCW): “China Pr. Sichuan nord, Jiuzhaigon [sic!], 4 7.Juli 1994, 80 km N Songpan, 3000 m, leg. Beneš”; 1 ♂ (EJCB): “China (Sichuan), n. Kangding: See Mu ge cuo, 3500 3900m, 22.vii.1994, Heinz leg.”. Xizang: 1 ♀ (EJCB): “China, Tibet, Zayu Co, upper basins of the W & E branch of Taron river (Irrawaddy), 3800 4600 m, 28º35’ 43’N, 97º40’ 58’E, 22 29.6.1999, L. & R. Businský leg.”. Yunnan: 1 ♀ (EJCB): “China, Yunnan, 2600m, Lijiang, 2 3.7.1990, 26.53N, 100.18E, M. Bocák lgt.”; 1 ♀ (EJCB): “China, Yunnan, 1 19.vii., Heishui, 27.13N, 100.19E, 35 km N of Lijiang, E. Jendek leg., 1992”; 1 ♂ (EJCB): “China, Yunnan, 6. 10.VII., Ninglang env., 27.19N, 100.55E, E. Jendek leg., 1992”; 1 ♀ (EJCB): “Yunnan, 3300 2500m, 27.14N, 100.15E, Yulongshan mts., N slope, 5/7.92, Vít Kubáň leg.”; 4 (EJCB): “Yunnan, Xishan Mts., 24.57N, 102.38E, 19 May 1993, Bolm lgt., 2300 m”; 7 (EJCB): “China, Yunnan, 24 29.VI., 50 km N of Lijiang, 1993, Yulongsjhan Nat. Res., E. Jendek & O. Šauša leg.”; 13 (EJCB): “Yunnan, 2300 m, 24.57N, 102.38E, Xishan Mts., 19/5, Vít Kubáň 1993”; 1 ♂ 1 ♂ (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”; 1 ♂, 2 ♀ (EJCB): “China, Yunnan 1994 Zhongdian 16 20.vi. leg. Šiška T. Spevár”; 1 ♀ (EJCB): “China, Yunnan prov., Lijiang, Bai Shui, 2900 3300m, 7 18.vii.1994, leg. Holzschuh”; 1 ♂ (EJCB): “China, Yunnan, 2.5 3.8km, 27.20N, 100.11E, Habashan mts., SE slope, 3 6.6., lgt. S. Bečvář, 1995”; 1 ♀ (EJCB): “China, Yunnan, Lijiang, 19.VI.1995, Šiška & Spevár & Pekarovič”; 1 ♀ (EJCB): “China NW, Yunnan, cca 3600 m, road Deqen Yanjing, 10 km W of Deqen, 1997, 28°28’N 98°53’E, 21 22.vi. M. Trýzna et O. Šafránek lgt.”; 7 (EKCS): “China Yunnan, Haba Shan, 1.7. 6.7.2005, lgt. E. Kučera”. LAOS: Houaphan: 1 ♂ (EJCB): “Laos NE, Hua Phan prov., Ban Saluei, Phu Phan Mt., 20°15’N, 104°02’E, 1500 2000m, D. Hauck leg., 26.iv. 11.v.2001”. THAILAND: Chiang Mai: 1 ♂ (EJCB): “N Thailand, Chiang Mai, Doi Suthep, 1 XI [19]87 E. Heiss”. Mae Hong Son: 1 ♀ (EJCB): “Thai, Mae Hong Son pr., 19°27’N, 98°20’E, 1500 m, Soppong 7 12.V., Vít Kubáň leg., 1996”.

Adult occurence: 4-5-6-7-11 (n=20). Altitude range: 1500-4600 m (n=11).

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Host plants: This species was collected by various collectors in China on Quercus foliage. Distribution: CHINA: Sichuan, Xizang, Yunnan. LAOS. THAILAND.

Agrilus ribbei Kiesenwetter Figs. 22K 22L (habitus); Fig. 37T (aedeagus); Figs. 43H, 44T, 46M, 46T, 47E, 47O, 51H, 51V, 53O, 53U (types) ribbei Kiesenwetter, 1879 (Agrilus) Kiesenwetter in KRAATZ & KIESENWETTER, 1879: 255 [#1595] (description). HEYDEN, 1881: 119 [#5548] (catalog). HEYDEN, 1884: 290 [#8482] ([Note: Misidentified as graminis]; faunal record; Amur region). KERREMANS, 1885: 154 [#1381] (catalog). MARSEUL, 1887: 252 [#4707] (catalog). KERREMANS, 1892: 270 [#1397] (catalog). KERREMANS, 1903: 290 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1026 [#2191] (world catalog). ALEXEEV, 1989: 480 [#64] (presumed synonym of friebi; key to species). JENDEK, 2003: 185 [#5823] (lectotype designation; synonymy). JENDEK, 2006: 392 [#7441] (subgenus Quercuagrilus; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (checklist; Japan). BELLAMY, 2008: 2265 2266 [#8576] (subgenus Quercuagrilus; world catalog). VOLKOVITSH, 2009: 135 [#8850] (subgenus Quercuagrilus; faunal records; Russia: Primorskiy kray). gracilipes Lewis, 1893 (Agrilus; [preoccupied]) LEWIS, 1893: 335 [#1722] (description). KERREMANS, 1903: 287 [#1441] (synonym of lewisiellus; catalog). JAKOBSON, 1913: 798 [#4386] (synonym of lewisiellus; catalog). MIWA & CHÛJÔ, 1936: 17 [#1809] (synonym of lewisiellus). ALEXEEV, 1979: 131 [#57] (synonym of tibialis). JENDEK, 1994: 16, 17 [#1355] (synonym of tibialis; lectotype designation). AKIYAMA & OHMOMO, 1997: 41 [#4091] (synonym of tibialis). JENDEK, 2003: 185 [#5823] (synonym of ribbei). JENDEK, 2006: 392 [#7441] (synonym of ribbei; Palaearctic catalog). BELLAMY, 2008: 2265 [#8576] (synonym of ribbei; world catalog). tibialis Lewis, 1893 (Agrilus) KERREMANS, 1903: LEWIS, 1893: 335 [#1722] (description). 291 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1924: 44 45 [#2040] ([Note: Misidentified as asiaticus according to JENDEK, 1994: 15 [#1355]]; characters; E Siberia; Amur). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 21 [#1809] (catalog; Japan). OBENBERGER, 1936: 1054 [#2191] (world catalog). KUROSAWA, 1963: 153 [#1618] (characters; Japan). KUROSAWA, 1974: 1, 2, 3 [#4575] (characters; notes). KUROSAWA, 1975: 2 [#4591] (note). KUROSAWA, 1976: 133 [#1628] (notes; color variability; type in BMNH examined). ALEXEEV, 1979: 131 133 [#57] (subgenus Quercuagrilus; characters; notes; USSR: Chita, Amursk, Khabarovsk, Primorsk). TÔYAMA, 1985: 29 [#3678] (iconography; Japan). ALEXEEV, 1989: 480 [#64] (key to species). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). ANONYMOUS, 1991: 106 [#8552] (faunal records; Japan: Aomori). LI JINGKE, 1992: 92 [#6755] (checklist; China: Liaoning). JENDEK, 1994: 16 17 [#1355] (synonymy; characters; notes; Japan, Kuril Islands). JENDEK, 1995: 147 [#3768] (lectotype designation). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 186 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 41 [#4091] (checklist; Japan: Hokkaido, Honshu, Shikoku, Kyushu, Kunashiri Island, Sado Island, Tsushima; Korea; northeastern China; Rusia: East Siberia). ALEXEEV, 1998: 374 [#4322] (subgenus Quercuagrilus;

171 subgeneric classification). NONNAIZAB et al., 1999: 113 [#8247] (cited as tibia; checklist; China: Inner Mongolia). AKIYAMA & OHMOMO, 2000: 88 [#6240] (faunal records; Japan (Hiroshima)). HUA LI ZHONG, 2002: 90 [#6696] (checklist; northeastern China). JENDEK, 2003: 185 [#5823] (synonym of ribbei). FUKUTOMI & HORI, 2004: 13 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 392 [#7441] (synonym of ribbei; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (synonym of ribbei). ANONYMOUS, 2007: 32 [#8250] (checklist; Japan: Honshu). BELLAMY, 2008: 2265 [#8576] (synonym of ribbei; world catalog). japonicus Kerremans, 1898 (Agrilus) KERREMANS, 1898: 178 179 [#1427] (description). KERREMANS, 1903: 276 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1003 [#2191] (world catalog). KUROSAWA, 1963: 152 [#1618] (characters; Japan). KUROSAWA, 1975: 4 [#4591] (notes). KUROSAWA, 1976: 133 [#1628] (synonym of tibialis; type in MNHN and syntypes in BMNH examined). ALEXEEV, 1979: 131 [#57] (synonym of tibialis). JENDEK, 1994: 16, 17 [#1355] (synonym of tibialis; lectotype designation). AKIYAMA & OHMOMO, 1997: 41 [#4091] (synonym of tibialis). JENDEK, 2003: 185 [#5823] (synonym of ribbei). JENDEK, 2006: 392 [#7441] (synonym of ribbei; Palaearctic catalog). BELLAMY, 2008: 2265 [#8576] (synonym of ribbei; world catalog). lewisiellus Kerremans, 1903 (Agrilus; replacement name for gracilipes Lewis not Waterhouse) KERREMANS, 1903: 287 [#1441] (replacement name proposal). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 17 [#1809] (catalog; Japan). OBENBERGER, 1936: 1008 [#2191] (world catalog). KUROSAWA, 1949: 14 [#5028] (characters). KUROSAWA, 1950: 1115 [#3274] (note). IGA, 1955: 79 [#3828] (iconography; Japan). KUROSAWA, 1956: 1115 [#8107] (characters; Japan). IGA, 1962: 79 [#3351] (iconography; Japan). KUROSAWA, 1963: 153 [#1618] (form of tibialis; characters; Japan). KUROSAWA, 1976: 133 [#1628] (synonym of tibialis; type in BMNH examined). ALEXEEV, 1979: 131 [#57] (synonym of tibialis). ANONYMOUS, 1986: 177 [#8848] (insect pest; Korea). AKIYAMA & OHMOMO, 1997: 41 [#4091] (synonym of tibialis). NONNAIZAB et al., 1999: 112 [#8247] (checklist; China: Inner Mongolia). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Inner Mongolia, Shandong, Henan, Gansu, Shaanxi). LIN GUI RUI, 2002: 233 234 [#7926] (checklist; pest). JENDEK, 2003: 185 [#5823] (synonym of ribbei). JENDEK, 2006: 392 [#7441] (synonym of ribbei; Palaearctic catalog). BELLAMY, 2008: 2265 [#8576] (synonym of ribbei; world catalog). corax Obenberger, 1917 (Agrilus) OBENBERGER, 1917: 39 40 [#1995] (description). OBENBERGER, 1924: 44 45 [#2040] (note). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 978 [#2191] (world catalog). KUROSAWA, 1976: 133 134 [#1628] (subspecies of tibialis; type in NMPC examined). ALEXEEV, 1979: 132 [#57] (synonym of tibialis asiaticus). ALEXEEV, 1989: 480 [#64] (synonym of tibialis). JENDEK, 1994: 17 18 [#1355] (subspecies of tibialis; lectotype designation; synonymy; characters; Russian Far East; Korea; China). AKIYAMA & OHMOMO, 1997: 41 [#4091] (synonym of tibialis). JENDEK, 2000: 506 [#4851] (subspecies of tibialis; synonymy). JENDEK, 2003: 185 [#5823] (synonym of ribbei). JENDEK, 2006: 392 [#7441] (synonym of ribbei; Palaearctic catalog). BELLAMY, 2008: 2265 [#8576] (synonym of ribbei; world catalog). ignoratus Obenberger, 1924 (Agrilus; cited as ingnoratus on page 44 and ignoratus on pages 45 and 46)

172 OBENBERGER, 1924: 44, 45, 46 [#2040] (description; [Note: Multiple original spelling: Precedence of the name ignoratus has been fixed by the original author (OBENBERGER, 1926: 655 [#2057]) as the first reviser (Article 24.2.4). The name ingnoratus is incorect original spelling and unavailable name (Article 32.4)]). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1000 [#2191] (world catalog). MIWA & CHÛJÔ, 1940: 74 [#1810] (faunal records; South Korea). THÉRY, 1942: 252 [#2875] (note). TER MINASYAN, 1955: 451 [#7893] (synonym of foveicollis). KRIVOLUTSKAYA, 1973: 177, 191, 193 [#8102] (notes; Kuril Islands). KUROSAWA, 1976: 133 134 [#1628] (synonym of tibialis corax; type in NMPC examined). ALEXEEV, 1979: 132 [#57] (synonym of tibialis asiaticus). ALEXEEV, 1989: 480 [#64] (synonym of tibialis). JENDEK, 1994: 17, 18 [#1355] (synonym of tibialis corax; [Note: The name ignoratus deemed to be unjustified emendation]; lectotype designation). AKIYAMA & OHMOMO, 1997: 41 [#4091] (synonym of tibialis). NONNAIZAB et al., 1999: 112 [#8247] (checklist; China: Inner Mongolia). JENDEK, 2003: 185 [#5823] (synonym of ribbei). JENDEK, 2006: 392 [#7441] (synonym of ribbei; Palaearctic catalog). taigicola Obenberger, 1924 (Agrilus) OBENBERGER, 1924: 45 46 [#2040] (description). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1053 [#2191] (world catalog). JENDEK, 1994: 17, 18 [#1355] (synonym of tibialis corax; lectotype designation). AKIYAMA & OHMOMO, 1997: 41 [#4091] (synonym of tibialis). JENDEK, 2003: 185 [#5823] (synonym of ribbei). JENDEK, 2006: 392 [#7441] (synonym of ribbei; Palaearctic catalog). BELLAMY, 2008: 2266 [#8576] (synonym of ribbei; world catalog). prinadai Fisher, 1925 (Agrilus) FISHER, 1925: 6 7 [#975] (description). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1023 [#2191] (world catalog). KUROSAWA, 1976: 133 134 [#1628] (synonym of tibialis corax; notes). ALEXEEV, 1979: 132 [#57] (synonym of tibialis; cited as prunadai). ALEXEEV, 1989: 480 [#64] (synonym of tibialis). BELLAMY, 1994: 362 [#237] (holotype (alotype) in USNM). JENDEK, 1994: 17 [#1355] (synonym of tibialis corax). AKIYAMA & OHMOMO, 1997: 41 [#4091] (synonym of tibialis). JENDEK, 2000: 506 [#4851] (synonym of tibialis corax). JENDEK, 2003: 185 [#5823] (synonym of ribbei). JENDEK, 2006: 392 [#7441] (synonym of ribbei; Palaearctic catalog). BELLAMY, 2008: 2266 [#8576] (synonym of ribbei; world catalog). bakinensis Obenberger, 1935 (Agrilus) OBENBERGER, OBENBERGER, 1935: 168 [#2158] (description). 1936: 958 [#2191] (world catalog). JENDEK, 1994: 17, 18 [#1355] (synonym of tibialis corax; lectotype designation; type locality confused). JENDEK, 2003: 185 [#5823] (synonym of ribbei). JENDEK, 2006: 392 [#7441] (synonym of ribbei; Palaearctic catalog). BELLAMY, 2008: 2266 [#8576] (synonym of ribbei; world catalog). freyi Théry, 1939 (Agrilus) OBENBERGER, THÉRY, 1939: 153 154 [#2861] (description). 1943: 10 [#2333] (synonym of ignoratus). ALEXEEV, 1979: 132 [#57] (synonym of tibialis asiaticus). ALEXEEV, 1989: 480 [#64] (synonym of tibialis). JENDEK, 1994: 17 [#1355] (synonym of tibialis corax). AKIYAMA & OHMOMO, 1997: 41 [#4091] (synonym of tibialis). JENDEK, 2003: 185 [#5823] (synonym of ribbei). JENDEK, 2006: 392 [#7441] (synonym of ribbei; Palaearctic catalog). BELLAMY, 2008: 2266 [#8576] (synonym of ribbei; world catalog). iturupicus Alexeev, 1979 (Agrilus; subgenus Quercuagrilus) ALEXEEV, 1979: 133 134 [#57] (description). ALEXEEV, 1989: 480 [#64] (key to species; Kuril Islands (Iturup)). JENDEK, 1994: 17 [#1355] (synonym of tibialis; notes). AKIYAMA & OHMOMO, 1997: 41 [#4091] (synonym of tibialis). ALEXEEV, 1998: 374

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA [#4322] (subgenus Quercuagrilus; subgeneric classification). JENDEK, 2003: 185 [#5823] (synonym of ribbei). JENDEK, 2006: 392 [#7441] (synonym of ribbei; Palaearctic catalog). BELLAMY, 2008: 2265 [#8576] (synonym of ribbei; world catalog).

Type material: Agrilus ribbei Kiesenwetter, 1879 Type locality: Amur. Type specimens: Primary: Lectotype (Fig. 51V), ♀, designated by JENDEK (2003: 185) [#5823], (ZSMC): “Sibirien Amur Christoph [18]77 [p] \ Kiesenwetter [p] \ Sammlung Cl. Müller [p] \ Type von [p] Agrilus Ribbei Kiesw. [h] [red label] \ Holotypus [p] Agrilus ribbei Kiesenw. [h] Zool. Staatssammlg Described from unknown number of syntypes. Agrilus gracilipes Lewis, 1893 Type locality: Main island. Nikko, Nara, and Miyanoshita. Type specimens: Primary: Lectotype (Fig. 46T), ♂, designated by JENDEK (1994: 16, 17) [#1355], (BMNH): “Type H.T. [p] [round label with red border] \ Japan. G. Lewis 1910-320 [p] \ Nikko. 3.VI.-21.VI. [18]80 [p] \ Agrilus gracilipes Lewis Type [male symbol] [by Lewis hand]”. Described from unknown number of syntypes. Agrilus tibialis Lewis, 1893 Type locality: Yezo ... at Junsai and Sapporo. Type specimens: Primary: Lectotype (Fig. 53U), ♂, designated by JENDEK (1995: 147) [#3768], (BMNH): “Type H.T. [p] [round label with red border] \ Junsai Lake [p] \ G. Lewis 1910-320. [p] \ Agrilus tibialis Lewis, Type [+male symbol] [by Lewis hand] \ LECTOType Agrilus tibialis LEWIS, 1893 Ed. Jendek design. 1994 [p] [red label]”. Secondary: 4 PLT (BMNH). Described from unknown number of syntypes. Agrilus japonicus Kerremans, 1898 Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 47O), ♂, designated by JENDEK (1994: 16, 17) [#1355], (BMNH): “SYN-Type [p] [round label with blue border] \ Japan Staud [h] \ japonicus Kerr. Type [h] \ Kerremans 1903-59 [p]”. Secondary: 1 PLT (MNHN); 1 PLT (BMNH). Described from unknown number of syntypes. Agrilus lewisiellus Kerremans, 1903 Type locality and specimens: See: Agrilus gracilipes Lewis, 1893. Agrilus corax Obenberger, 1917 Type locality: Ostsibirien: Kitajskaja Sterana [= Chinese side of Amur river near Radde, Jewish Autonomous Oblast, Russia. See BODEMEYER, 1915: 438, 439 [#8261] and BODEMEYER, 1928:72-73, 79 [#7705]]. Type specimens: Primary: Lectotype (Fig. 44T), ♀, designated by JENDEK (1994: 17-18) [#1355], (NMPC): “Kitajskaja Sterana Sib. or. [h] \ Typus [p] [red label] \ Agrilus corax m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24878 [h] [orange label]”. Described from unknown number of syntypes. Agrilus ignoratus Obenberger, 1924 Type locality: Sibiria or. (Šilka montes); Ussuri (Vladivostok). Type specimens: Primary: Lectotype (Fig. 47E), ♂,

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA designated by JENDEK (1994: 17, 18) [#1355], (NMPC): “Ussuri [h] \ Typus [p] [red label] \ Agrilus ingnoratus m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24264 [h] [orange label]”. Secondary: 2 PLT (NMPC); 1 PLT (NHMB). Described from unknown number of syntypes. Agrilus taigicola Obenberger, 1924 Type locality: Vladivostok. Type specimens: Primary: Lectotype (Fig. 53O), ♀, designated by JENDEK (1994: 17, 18) [#1355], (NMPC): “COLLECTIO DR. OBENBERGER MUS. PRAGENSE [p] \ Vladivostok [h] \ Typus [p] [red label] \ Agrilus taigicola m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24262 [h] [orange label]”. Described from unknown number of syntypes. Agrilus prinadai Fisher, 1925 Type locality: Vladivostok, Siberia. Type specimens: Primary: Holotype (Fig. 51H), ♀, (USNM): “Vladivostok Siber [sic!] 1923 V. Prinada [p] \ TypeNo. [p] 27563 [h] U. S. N. M. [p] [red label] \ Agrilus prinadai Fisher [h]”. Secondary: 1 PT (USNM). Described from 2 specimens (HT, PT). Agrilus bakinensis Obenberger, 1935 Type locality: Caucasus: Bakinskaja Gubernia: Apšeron [Note: Locality confused, see JENDEK, 1994 [#1355]]. Type specimens: Primary: Lectotype (Fig. 43H), ♂, designated by JENDEK (1994: 17, 18) [#1355], (NMPC): “Apšeron Bakinská g. \ Typus [p] [red label] \ Agrilus bakinensis m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 26833 [h] [orange label]”. Described from unknown number of syntypes. Agrilus freyi Théry, 1939 Type locality: Maoerschan, Mandchourie. Type specimens: 2 ST (MNHN); 1 ST (NHMB); 1 ST (NMPC). Described from 8 syntypes. Syntype (Fig. 46M) from MNHN is illustrated. Agrilus iturupicus Alexeev, 1979 Type locality: Kuril’skie ostrova, o-v Iturup, okr. pos. Rybaki [in Russian]. Type specimens: Primary: Holotype (AAOR). Secondary: 1 PT (AAOR). Described from 2 specimens (HT, PT). Diagnosis: 4.4-7.6 mm; pronotum visually transverse; medial pronotal impression weak; aedeagus without angular sides. Species group: Sulcicollis. Relevant species: A. adelphinus Kerremans, 1895. Specimens examined: CHINA: Hebei: 7 (EJCB): “China, 1. 2. VI.2000, border Hebei Nei Mongol, road Chengde Chifeng, pass 1600 m, 41.6N, 118.2E, Jaroslav Turna leg.”; 4 (EJCB): “China, 1. 2. VI.2000, border Hebei Inner Mongol, road Chengde Chifeng, Zd. Jindra lgt., pass 1600 m”; 3 (EJCB): “China, N Hebei, 13.vi., 20 km NNE Qijia, 41.6N, 118.15E, Jaroslav Turna, 2005”. Heilongjiang: 1 (NMPC): “E.v. Bodemeyer Sibiria orient. Chitaizki Sterani” (JENDEK, 1994 [#1355] as tibialis corax). Henan: 4 (EJCB): “China, W Henan, 15 17.v., Funiu Shan, 33.5N, 111.9E, Baotianman, Jaroslav Turna, 2005”. Hubei: 1 ♂, 1 ♀ (EJCB): “China, W Hubei prov., Shennongjia Co, Yannzi Pass, 31º43’/110º28’, 2200m, 23 26.6.1995, L.+R. Businský lgt.”. Jilin: 1 (EJCB): “China, Jilin province., ca 2 km S Bai He, c a 750 m, N42º24.093’, E128º06.431’, 1 6.vi.2004, leg. J. Cooter”. Shaanxi: 1 ♂ (EJCB): “China, S Shaanxi, Qinling Mts. S slope, Xunyangba

173 S+W env., 33º28 37’ / 108º23 33’, 1400 2100m, 5 9.6.[19]95, L.+R. Businský lgt.”; 4 (EJCB): “China, Shaanxi prov., 21 23. vi.1998, Quin Ling Shan mts, road Baoji Taibai vill., pass 35 km S Baoji, O. Šafránek et M. Trýzna lgt.”; 1 ♂ (EJCB): “China, 1150 1300 m, Shaanxi, Qinling mts., Foping (6 km N), 20 21.VI.1998, I. H. Marshal leg.”; 1 ♀ (EJCB): “40 km SE of Taibai Shan Mts., Hou Zen Zi vill. env., Z. Jindra lgt. cca 1200m, China, Shaanxi prov., 11 July 1998, Qing Ling Shan mts.”; 2 ♀ (EJCB): “China, Shaanxi, 26.VI.1998, Qing Ling Shan mts., 1500 m, Hou Zen Zi vill. env., 30 km SE of Taibai Shan Mts., O. Šafránek et M. Trýzna lgt.”; 9 (EJCB): “C China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000 1300m, 23.V. 13.VI. leg. C. Holzschuh 2000”; 1 (EKCS): “China Shaanxi, Lueyang, 29.5. 2.6.2005, lgt. E. Kučera”; 1 (EKCS): “China Shaanxi, Lueyang, 23 30.5.2009, lgt. E. Kučera”. Shanxi: 10 (EJCB): “China, W Shanxi, 9.VI.2000, 37.7N, 111.6E, Lüliang Shan, road Fangshan Jiaocheng, Hengjian env., ~1000m, Jaroslav Turna leg.”. Sichuan: 45 (EJCB): “China, Sichuan, 8 13.6.1991, Abazhou (102) 2000m, Nanping Jiuzhaigou, 33.15N/104.15E” (JENDEK, 1994 [#1355] as tibialis corax); 1 (EJCB): “China Sechuan, Jiuzhaigou [sic!], 11.6. 16.6.2001, leg. E. Kučera”; 7 (EKCS): “China Sechuan, Juizhaigou, 13 19.6.2009 leg. E. Kučera”. JAPAN: Hokkaido: 1 (EJCB): “Asahi 1.VIII. [19]54 Hokkaido Jap” (JENDEK, 1994 [#1355] as tibialis). Honshu: 1 ♂ (EJCB): “Japon: Yumoto pres Nikko 3 Aout 1911 Edme Gallois” (JENDEK, 1994 [#1355] as tibialis); 1 (EJCB): “Kiyomi, Gifu (VI 4 1949) K. Ohbayashi” (JENDEK, 1994 [#1355] as tibialis); 1 (EJCB): “Japan 25.5.[19]75 Takahashi City T. Aomo lgt.” (JENDEK, 1994 [#1355] as tibialis); 3 (JMCZ): “Japan, 10.5.1979, Shishizuka, Tsuchiura city, K. Akiyama lgt.”; 1 ♂ (EJCB): “Japan, Gunma pref. 24.7.1982 A. Torikata leg.” (JENDEK, 1994 [#1355] as tibialis); 1 ♂ (EJCB): “Jap. Shizuoka pref. Sumata 2.VI.1984 A. Torikata leg.” (JENDEK, 1994 [#1355] as tibialis); 1 ♂ (EJCB): “Japan, Iwate, Gosho dam, Moroika shi, 16.vi.1993”; 40 (EJCB): “Japan, Fukushima, Namie town, 26.vi.1995, N. Sugiura leg.”. Kyushu: 2 ♂ (JMCZ): “Japan, Kyushu, Oita pref., Mt. Kurodake, 1.vi.1986”. KOREA NORTH: 1 (NMPC): “Hakugan, N.Korea (Chosen) NIPPON june 1936 (F. Yano)” (JENDEK, 1994 [#1355] as tibialis corax); 1 (VSCS): “Korea, Onpha ri, Kreis Kjôngsông, 12.6.1975, Josifov leg.”; 1 ♀ (EJCB): “N Korea, 19.7.1990, Toxong env., Omdouri”. RUSSIA: Amurskaya oblast’: 1 (NMPC): “Amur Stjernkreutz” (JENDEK, 1994 [#1355] as tibialis corax); 1 (ZIN): “Klimoutsi Amur. obl. 40 km W Svobodnovo Zinovjev 2 20.VI. [1]957 [1]959 [in Russian]” (JENDEK, 1994 [#1355] as tibialis corax); 1 (ZIN): “Simonovo, Amur. obl., 75 km W Svobodnovo Zinovjev 18.VII.[1]959” (JENDEK, 1994 [#1355] as tibialis corax); 2 (EJCB): “Amurskaja obl., Kundur, 4.V.[19]75, plk duba, Danilevskij leg.” (JENDEK, 1994 [#1355] as tibialis corax); 1 (ZIN): “Amurskaya obl., Arkharinskii r n, s. Antonovka, S. Vinter 5 9.vi.1976 [in Russian]”; 1 (EJCB): “USSR rez. Chingan, VII.1990, Ing. Lorenc” (JENDEK, 1994 [#1355] as tibialis corax). Khabarovskiy kray: 1 (NMPC): “Sibiria orient., Sotka Gora, B. v. Bedemeyer”; 1 (EJCB): “Chabarovskij rajon GUR 15.9.[19]75 plk duba Danilevskij leg.” (JENDEK, 1994 [#1355] as tibialis corax); 1 (MSCI): “Khabarovsky reg., Komsomolsky reserve, Zolotoj cordon 26.VII.1991 leg. М. Smirnov”. Primorskiy kray: 1 (NMPC): “Romanovka, 23 VI [19]27, distr. Vladivostok, Kistiakovsky leg.” (JENDEK, 1994 [#1355] as tibialis corax); 1 ♂ (EJCB): “Primorskij kraj, 1951, Ussurij. okr., 23.VI., Voroshilov, D. Kononov leg.” (JENDEK, 1994 [#1355] as tibialis corax); 1 (EJCB): “USSR Siberia, Primorje, Ussurijskij rajon, Kajmanovka, 7. 13.VII.1988 S. Chvilja lgt.” (JENDEK, 1994 [#1355] as tibialis corax); 1 (MNHE): “Russia, 01.07.1989, Primorye Distr., Limonniki, leg. U. Roosileht”; 1 (MNHE): “Russia, 27 29.06.1989, Primorye Distr., Buyanki, leg. U. Roosileht”; 4 (EJCB): “USSR, Primorsk. kraj Vladivostok env., 22.VII.1990 leg. Snížek” (JENDEK, 1994 [#1355] as tibialis corax); 1 (EJCB): “USSR, Primorsk. kraj, Novocugujevka env., 16 18.

174 VII.1990, leg. Snížek” (JENDEK, 1994 [#1355] as tibialis corax); 1 (JMCZ): “USSR / Sibiria Primorskij Kraj. Novocugujevka 15 20.7.1990 leg. Boukal”; 1 (ZIN): “Vladivostok Morskoje kladbyschce les 17.06.1990 Belokobylskij” (JENDEK, 1994 [#1355] as tibialis corax); 15 (EJCB): “Sib.or. m., Primorje Sichote Alin Mts. Sokolči 1. 15.7.1990 Kadlec + Voříšek lg” (JENDEK, 1994 [#1355] as tibialis corax); 15 (EJCB): “USSR, Ussuri prov. Arsenev env., VII.1990, leg. B. Šiška & R. Čermák” (JENDEK, 1994 [#1355] as tibialis corax); 1 (EJCB): “Primorje, Ussurijsk, Kamenushka, 2.VII.[19]90” (JENDEK, 1994 [#1355] as tibialis corax); 1 (ZIN): “Prim., Lazov., Kijevka, 18.6.[19]91 Filimonov” (JENDEK, 1994 [#1355] as tibialis corax); 25 (EJCB): “USSR, Primorskij kraj Arseneev env. VI.1991 M. Štrba leg.” (JENDEK, 1994 [#1355] as tibialis corax); 1 (EJCB): “Prim., Khasansk, Riazanovka, 5.6.91 Filimonov” (JENDEK, 1994 [#1355] as tibialis corax); 50 (EJCB): “USSR, Primorsk. kraj, Arsenev env., VI.1991, 44.11N, 133.19E, O. Šausa leg.” (JENDEK, 1994 [#1355] as tibialis corax); 1 (MKCY): “Russia, Primorski reg. Khasan distr. Ryazanovka 05.06.1991 Filimonov leg.”; 1 (ZIN): “Prim., Khasan, Zamadvorovka, 24.8.[19]91, Namolov” (JENDEK, 1994 [#1355] as tibialis corax); 1 (EJCB): “Russia or., Primorsk. Reg., Artyom env., 5.VI. 21.VI.1992 Plutenko leg” (JENDEK, 1994 [#1355] as tibialis corax); 1 ♂ (JMCZ): “Ross. or, Prim. kr., Kraskino, Azajsan, 13 16.VII.1992, Snížek leg.”; 25 (EJCB): “Sib. or., Primorje, Komarovka flum., Kamenushka env., 300 m, Vořišek lgt., VI.1992” (JENDEK, 1994 [#1355] as tibialis corax); 1 ♀ (EJCB): “Rossia, Primorsk, Kraskino env. mer., 13 16. VII.1992, P. Smrž leg.” (JENDEK, 1994 [#1355] as tibialis corax); 1 (MNCA): “Russia: Primorskiy Kray, Sergeyevka, 30 km N Partizansk, 43.22N, 133.23E, 04.VI.1993, 300 m, leg. L. Zerche et al.”; 7 (EJCB): “Russia, Primorski reg., Ussuri distr., Kaymanovka, 05.07.1993, Khvylya leg.”; 1 (MNCA): “Russia: Primorskiy Kray, Ryazanovka 14 km SW Slavyanka, 42.48N, 131.12E, 16.VI.1993, 50 m, leg. L. Zerche et al.”; 1 (MNCA): “Russia: Primorskiy Kray, 4 km S Poshiga, 37 km SE Dalnerechensk, 45.45N, 130.07E, 28.V.1993, 150 m, leg. L. Zerche et al.”; 1 (MNCA): “Russia: Primorskiy Kray, Sikhote Alin, Biol. Stat. 35 km SE Chuguevka, 44.05N, 134.12E, 31.V.1993, 650 m, leg. L. Zerche et al.”; 5 (EJCB): “Russia, Primorski kray, Ternejski distr., 12 19.viii., Sikhote Alin res., riv. Serebrianka env., 1995”; 2 (EJCB): “Russia E, Primorje reg, Anisimovka, 7 13.vii.1996, Mt. Litovka, 200 m, Plutenko leg.”; 4 ♂ (EJCB): “Russia Primorskij kraj, Vladivostok, Botanical garden, 26.vi.1997, leg. L. Roller, L. Vidlička”; 1 (MKCY): “Russia, Far East, Khanka lake NW Novokachalinsk vill. env. 5.7.2003, Lg. A. Napolov”; 2 (MKCY): “Russia Far East, 20 km SE Ussuriysk, Gornotaezhnoje vill. env., 28.06.2003, Lg. A. Napolov”; 1 (MKCY): “Russia, Primorski reg. Khankayski distr., Novokachalinsk, 07.07.2003, Filimonov leg.”; 2 (MKCY): “Russia, Far East, 93 km E Partizansk, Valentin vill. env., 22. VII.2003, Lg. A. Napolov”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Tachingousa (Prosiolotchnaja) bay 30.06.04 leg. М.& L. Smirnov’s”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Tachingousa (Prosiolotchnaja) bay, 26.07.06, leg. L. Smirnov”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Preobrazhenie env. 14.07.06 leg. М.& L. Smirnov’s”; 2 (IMCM): “Yu Primor’e, Lazovskii raion, p. Preobrazhenie na usokhshikh Quercus mongolica, 9 14.vii.2006, leg. M. L. Smirnovy [in Russian]”; 1 ♀ (ZIN): “Primorskii kr., Lazovskii z k, b. Proselochnaya, 2 5.VII.2007 Y. Sundukov, V. Khokhrin [in Russian]”; 19 (ZIN): “Primorskii kr. s. Lazo 8 12. vi.2007 V. Shokhrin leg [in Russian]”. Sakhalinskaya oblast’: 1 ♂ (ZIN): “Kuril. ostr. [ Kuril Islands], 10.VII.[19]63, Kol. Krivolutskaja, o. Iturup [in Russian]” (JENDEK, 1994 [#1355] as tibialis); 1 (ZIN): “Dubovoje bliz Golovnino, Kunashir, Kerzhner 22.VII.[1]973 [in Russian]” (JENDEK, 1994 [#1355] as tibialis); 1 ♀ (EJCB): “Kuril isl., Iturugo Kurilsk env., 19.7.1987, leg. Dolin”; 1 (MNHE): “Kunašir, Tretjakovo, 05 19.08.1991, leg. U. Roosileht”;

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA 1 ♂ (EJCB): “SU Far East (Kunashir), Kunashir isl. C., cape Stolbchatiy env., 1991, Lg. A. Napolov”; 1 ♂ (EJCB): “Russia or., Sakhalin, Novoaleksandrovska, 17.vii.1992, Babarukov leg.”; 2 (IMCM): “Kunashir, kordon Alëkhinskii, ur. Danilovo, solki, 43°57’16’’N, 145°35’38’’E, h~45, 29.vii. 6.viii.2008 leg. I. Mel’nik, na Quercus crispula et Q. dentata [in Russian]”. Yevreyskaya avtonomnaya oblast’: 1 (NMPC): “v. Bodemayer Sibiria orient. Sotka Gora” (JENDEK, 1994 [#1355] as tibialis corax).

Adult occurence: 5-6-7-8-9 (n=85). Altitude range: 502200 m (n=17). Host plants: Carpinus: AKIYAMA & OHMOMO, 1997 (as tibialis) [#4091]. – Quercus: ALEXEEV, 1979 (as tibialis and iturupicus) [#57]; ALEXEEV, 1989 (as iturupicus) [#64]; ALEXEEV, 1989 [#64]; AKIYAMA & OHMOMO, 1997 (as tibialis) [#4091]. Distribution: CHINA: Hebei, Heilongjiang, Henan, Hubei, Jilin, Liaoning, Nei Mongol, Shaanxi, Shanxi, Sichuan. JAPAN: Hokkaido, Honshu, Kyushu, Shikoku, Tsushima. KOREA NORTH. KOREA SOUTH. RUSSIA: Amurskaya oblast’, Khabarovskiy kray, Primorskiy kray, Sakhalinskaya oblast’, Yevreyskaya avtonomnaya oblast’, Zabaykalskyi kray.

Agrilus ribesi Schaefer Fig. 17A (habitus); Fig. 35E (aedeagus); Fig. 51W (type) ribesi Schaefer, 1946 (Agrilus; variety of viridis) SCHAEFER, 1946: 107 108 [#3585] (description). DEKHTIAREV, 1928: 20 [#8663] ([Note: Misidentified as viridis]; pest; Ukraine). SCHAEFER, 1949: 397, 399, 400, 491 [#2565] (variety of viridis; cited as ribesii; characters). RIKHTER, 1950: 158, 165 [#3881] ([Note: Misidentified as chrysoderes]; adult and larval characters in key). HORION, 1955: 94 [#1285] (variety of viridis; cited as ribesii; notes). TER MINASYAN, 1955: 454 [#7893] ([Note: Misidentified as viridis variety fagi]; host plant; distribution; references). SCHAEFER, 1968: 77 79 [#2577] (cited as ribesii and valid species; redescription; bionomy; France; Germany, Italy). BÍLÝ, 1977: 40 [#283] (key to species; Slovakia). VOLKOVITSH, 1977: 25 [#2985] ([Note: Misidentified as viridis aubei]; faunal record; notes; USSR: Belgorod). LOMPE, 1979: 240 [#3628] (cited as ribesii; characters in key; Central Europe). SAVKOVSKII & RYBALOV, 1981: 82 86 [#6092] ([Note: Misidentified as viridis]; bionomy; Ukraine). COBOS, 1986: 254, 317 [#609] (synonym of viridis). ALEXEEV, 1989: 488 [#64] (key to species; Khabarovsk; Primorsk; Chita; Buryatia; Irkutsk; Tuva; Altay; W Siberia, north Kazakhstan; European USSR; North Korea; west Europe). CURLETTI, 1994: 22, 176 [#658] (subgenus Agrilus; cited as ribesii; catalog; single record from Italia). ALEXEEV, 1998: 381 [#4322] (subgenus Agrilus; cited as ribesii; subgeneric classification). JANICKI, 1998: 31, 32 [#4786] (Ukraine). LISKENNE, 1998: 79 [#6018] (cited as ribesii; faunal records; France). BURAKOWSKI et al., 2000: 76 77 [#5188] (note). BÍLÝ, 2002: 24 [#5877] (bionomy; host plants; distribution). BRECHTEL & KOSTENBADER, 2002: 290 294 [#5766] (biology; ecology; distribution; Germany). GOBBI, 2002: 27 28 [#6898] (faunal records; Italy). JENDEK, 2003: 15, 19 [#5878] (presumable ecological form of viridis; lectotype designation; notes). BERNHARD et al., 2005: 601, 602 [#7261] (mitochondrial DNA; molecular systematics). JENDEK, 2006: 389 [#7441] (subgenus Agrilus; Palaearctic catalog). BELLAMY, 2008: 2266 [#8576] (subgenus Agrilus; world catalog). VOLKOVITSH, 2009: 134 [#8850] (subgenus Agrilus; checklist; Russia: Primorskiy kray).

175

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Type material: Agrilus viridis variety ribesi Schaefer, 1946 Type locality: Thénissey et Volnay (Cote-d’Or). Type specimens: Primary: Lectotype (Fig. 51W), sex not examined, designated by JENDEK (2003: 15, 19) [#5878], (MNHN): “Thénissey, 15. 6. [19]38”. Described from unknown number of syntypes.

Agrilus rivalieri Descarpentries & Villiers, 1963

Diagnosis: 4.9-7.5 mm; pronotum widest at anterior half, with sides almost straight in most of pronotal length and distinctly arcuate just before anterior angles; disk with very rough sculpture and distinct medial impression. Species group: Viridis. Relevant species: A. cuprescens (Ménétriés, 1832); A. viridis (Linné, 1758).

Specimens examined: CHINA: Guizhou: 1 ♂ (EJCB): “CH Guizhou NE, 27.V. 3.VI., 20 km NW of Jiangkou, 1995, Fanjing Shan Kuaichang, E. Jendek & O. Šauša leg.”. Hunan: 2 ♀ (EJCB): “China, N Hunan, 4 7.vii., Wuling Shan, 29.4N, 110.4E, Zhangjiajie, 700 m, Jaroslav Turna leg. 2003”; Shaanxi: 1 ♀ (EKCS): “China Shaanxi, Lueyang, 29.5. 2.6.2005, lgt. E. Kučera”. LAOS: Borikhamxai: 15 (EJCB): “Laos centr., Bolikhamsai prov., Ban Nape Kaew Nua Pass, 18.4. 1.5.1998, alt. 600±100 m, N 18°22.3, E 105°09.1 (GPS), E. Jendek & O. Šauša leg.”; 4 (EJCB): “Laos CE., 1 18.v.2001, Boli Kham Xai prov., 18°21’N, 105°08’E, Ban Nape (8 km NE), ~600m, Pacholátko leg.”. Champasak: 2 ♂ (EJCB): “Laos south., Champasak prov., Bolaven Plateau, 10 18.IV.1999, Route (No. 23) Pakse Paksong, Ban Itou env. (km.35), alt. 800m, N 15°10.4’, E 106°05.8’ (GPS), E. Jendek & O. Šauša leg.”. Houaphan: 1 ♂ (EJCB): “Lao NE, Hua Phan pr., ~20°12’N,104°01’E, Phu Phan Mt., 1500 1900m, 17.v. 3.vi.2007, Vít Kubáň leg.”. Phongsali: 1 ♀ (EJCB): “Lao N, Phongsaly prov., 21°41’ 2’N, 102°06’ 08’E, Phongsaly env., 28.v. 20.vi.2003, 1500m, P. Pacholátko leg.”. THAILAND: Nan: 1 ♀ (EJCB): “Thailand N, Nan prov., Doi Phu Kha N.P. Headq., 19°13’N 101°07’E, 1500m, 22 26. IV.1999, D. Hauck & M. Říha leg.”. VIETNAM: 1 ♀ (EJCB): “Vietnam N, Tonkin, Cuc Phuong nat. park, 2 12.V.1991, E. Jendek leg.”; 1 ♀ (EJCB): “S Vietnam, Gia Lai Kon Tum pr., 40 km N Ankhé, 800 m, Buon Loi, 15.VI.1983”; 3 (EJCB): “N. Vietnam (Tonkin), Tamdao (pr. (Vinphu), 2 11.6.1985, A. Olexa leg.”; 2 (EJCB): “N. Vietnam, 5 10.vi.1989, Tam Dao, Brantlová leg.”; 1 ♂ (EJCB): “N Viet Nam (Tonkin) pr. Vin Phu, 1990, Tam Dao, 17 21.V., Vít Kubáň leg.”; 1 ♀ (EJCB): “N Viet Nam (Tonkin), pr. Houang Lien Son, Sa Pa, 11 15.V.1990, Vít Kubáň leg.”; 1 (EJCB): “11 16.5.1990, Sa Pa, Hoang Lien Son Dist., N. Vietnam, 1600 m, J. Horák leg.”; 3 (EJCB): “N Viet Nam (Tonkin) pr. Vin Phu, 1990, Tam Dao, 6 9.V., Vít Kubáň leg.”; 1 ♂ (EJCB): “6 10.5.1990, Tam Dao, Vinh Phu Distr., 900 m, Jan Horák leg.”; 4 (EJCB): “17 21.5.1990, Tam Dao, Vinh Phu Distr., 900 m, Jan Horák leg.”; 1 ♀ (EJCB): “N Viet Nam (Tonkin) pr. Vin Phu, Tam Dao, 2 11.6.1985, Vít Kubáň leg.”; 1 ♀ (EJCB): “Vietnam N, 15.V. 16.VI., 75 km NW Hanoi, Tam Dao, 1991, E.Jendek leg.”; cca 80 (EJCB): “N Vietnam, 21°27N, 105°39E, 70 km NW of Hanoi, Tam Dao, 1 8.vi.1996, 900 1200 m, Dembický & Pacholátko leg.”.

Specimens examined: RUSSIA: Altay Kray: 1 (EJCB): “Altai, g. Biisk, 19.vi.1952 [in Russian]”. Ivanovskaya oblast’: 1 (MSCI): “C. Russia, Ivanovo dist., Ivanovo city, on a Ribes nigrum 8.07.2008 leg. Smirnov”. Lipetskaya oblast’: 1 (MCCD): “1ex 72км ЮВ г. Липецка, 2км ЮВ пос. Добринка 16.6.2000 Цуриков”. Primorskiy kray: 3 (EJCB): “Russia, Primorski reg., Shkotov distr. env., Anisimovka 01.07.1997 (on Ribes sp.) Kharin leg.”. Voronezhskaya oblast’: 1 (MCCD): “25км NNO г. Воронежа, Усманский Бор, 9.6.1988 Цуриков”; 2 (MCCD): “Воронежская обл., 30км Ю г. Острогожска 10.6.1996 Цуриков”.

Adult occurence: 6-7 (n=6). Host plants: Grossularia: RIKHTER, 1950 [#3881]; ALEXEEV, 1989 [#64]. – Ribes: DEKHTIAREV, 1928 [#8663]; RIKHTER, 1950 [#3881]; HORION, 1955 [#1285]; TER MINASYAN, 1955 [#7893]; SCHAEFER, 1968 [#2577]; VOLKOVITSH, 1977 [#2985]; SAVKOVSKII & RYBALOV, 1981 [#6092]; ALEXEEV, 1989 [#64]; LISKENNE, 1998 [#6018]. – R. alipnum: BÍLÝ, 2002 [#5877]. – R. aureum: BÍLÝ, 2002 [#5877]; BRECHTEL & KOSTENBADER, 2002 [#5766]. – R. grossularia: BÍLÝ, 2002 [#5877]; BRECHTEL & KOSTENBADER, 2002 [#5766]. – R. nigrum: SCHAEFER, 1946 [#3585]; CURLETTI, 1994 [#658]; KRÓLIK & MAJEWSKI, 1994 [#3471]; BÍLÝ, 2002 [#5877]; GUTOWSKI & MILKOWSKI, 2008 [#8244]. – R. rubrum: SCHAEFER, 1946 [#3585]; KRÓLIK & MAJEWSKI, 1994 [#3471]; BÍLÝ, 2002 [#5877]; BRECHTEL & KOSTENBADER, 2002 [#5766]. – R. uvacrispa: SCHAEFER, 1946 [#3585]; BÍLÝ, 2002 [#5877]; BRECHTEL & KOSTENBADER, 2002 [#5766]. Erroneous: (misdetermination of A. cuprescens (Ménétriés, 1832)): Rosa: RIKHTER, 1950 [#3881]. – Rubus idaeus: LISKENNE, 1998 [#6018]. Distribution: Local: KOREA NORTH. RUSSIA: Buryatiya Respublika, Irkutskaya oblast’, Khabarovskiy kray, Primorskiy kray, Tyva Respublika, Zabaykalskiy kray. Global: ASIA: GEORGIA. KAZAKHSTAN. KOREA NORTH. RUSSIA. EUROPE: AUSTRIA. BELARUS. BULGARIA. CZECH REPUBLIC. FRANCE. GERMANY. ITALY. POLAND. RUSSIA. SLOVAKIA. SWITZERLAND. UKRAINE.

Type locality: Tonkin. - Tam-Dao. Type specimens: Primary: Holotype (Fig. 51X), (MNHN). Described from 1 specimen (HT). Diagnosis: 5.2-8.2 mm; elytra with narrow perisutural stripes not reaching pleural sides on apices. Species group: Imitans. Relevant species: A. sikkimensis Obenberger, 1928.

Adult occurence: 4-5-6-7 (n=22). Altitude range: 5001900 m (n=12). Host plants: Unknown. Distribution: CHINA: Guizhou, Hunan, Shaanxi. LAOS. THAILAND. VIETNAM.

Agrilus rivalieri Descarpentries & Villiers

Agrilus rougeoti Descarpentries & Villiers

Fig. 13D (habitus); Fig. 33Y (aedeagus); Fig. 51X (type)

Fig. 26H (habitus); Fig. 39I (aedeagus); Fig. 52E (type)

rivalieri Descarpentries & Villiers, 1963 (Agrilus) DESCARPENTRIES & VILLIERS, 1963: 4, 15 [#701] (description). BELLAMY, 2008: 2267 [#8576] (subgenus Sinagrilus; world catalog).

rougeoti Descarpentries & Villiers, 1963 (Agrilus) DESCARPENTRIES & VILLIERS, 1963: 107, 115 116 [#700] (description). BELLAMY, 2008: 2271 [#8576] (world catalog).

176 Type material: Agrilus rougeoti Descarpentries & Villiers, 1963 Type locality: Tonkin: Hoa Binh. Type specimens: Primary: Holotype (Fig. 52E), ♂, (MNHN): “Tonkin Région de Hoa-Binh \ Type [p] ♂ [h] [red label] \ Agrilus rougeoti n. sp. Holotype ♂ nob. A. Descarpentries et [h] A. Villiers det. 19 [p] 62 [h] \ Muséum Paris 1935 coll. A. Théry [p]”. Secondary: 1 ♀ PT (MNHN). Described from 2 specimens (HT, AT). Diagnosis: 4.2-5.8 mm; elytral pubescence bicolored; elytra slightly attenuate before apices; apices separately arcuate; aedeagus slender. Species group: Cyaneovirens. Relevant species: A. tsushimanus Kurosawa, 1963. Specimens examined: CAMBODIA: 7 (EJCB): “SW Cambodia, 20km SE Koh Kong, Tatai river 11°34’N,103°07’E, 3 19.v.2005, 50 300m, E.Jendek & O.Šauša leg.”. CHINA: Yunnan: 1 ♂ (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”. INDIA: Andhra Pradesh: 1 ♀ (MNHN): “Khammameth Div., Hyderabad (Deccan), D.F.O. Coll., 30.iii.1934 \ Ex Chloroxylon swietenia”. Tamil Nadu: 10 (EJCB): “South India, 12 22.V.1994, Nilgiri Hills Kunchapanai, 11°20’N, 77°00’E, R. Sauer lgt.”; 3 (EJCB): “S India, Tamil Nadu, 17 22.V.1997, 15 km SE Kotagiri, 11°22’N, 76°56’E, Kunchappanai, Dembický & Pacholátko leg.”; 24 (EJCB): “India S, Tamil Nadu, Nilgiris, 15 km SE of Kotagiri, Kunchappanai, 900 m, 11.22 N 76.56 E, 7 22.v.2000, leg. P. Pacholátko”; 1 (EJCB): “India S, Tamil Nadu, Nilgiris, 15 km SE Kotagiri, Kunchappanai, 900 m, 11°22’N, 76°56’E, 7 22.V.2000, leg. D. Hauck”; 1 (EJCB): “India S, Tamil Nadu, 7 28.IX, Nilgiri Hills, 15 km SE Kotagiri, Kunchappanai, 900 m, D. Boukal & Z. Kejval leg., 1993”; 13 (EJCB): “S India, Tamil Nadu, Nilgiri Hills, 11 km SE Kotagiri, 1100±100m, 11°24’N, 76°56’E, Kunchappanai, L. Dembický leg., 3 15.v.2002”. Uttarakhand: 2 ♂ (EJCB): “Nakronda, iii.1934, Dehra Dun, dir. Bahadur”. LAOS: Khammouan: 2 ♀ (EJCB): “Laos centr., Khammouan prov., 4 16.XI., 25 30.XI. 2000, Ban Khoun Ngeun env., N 18°07’, E 104°29’, alt. 250m, E. Jendek & P. Pacholátko leg.”. NEPAL: Janakpur: 1 ♀ (EJCB): “C Nepal, Janakpur, 1987 Tamba Koshi Khola, SE Charikot, 900 1200 m, leg. C. Holzschuh, 5. 10.VI.”. THAILAND: Chumphon: 1 (EJCB): “Thailand, 1 11.v.1998, Chumphon prov., Pha To env., P. Prudek & R. Sigut leg.”. Mae Hong Son: 2 ♂ (EJCB): “Thailand, 1 8.5.1993, Soppong Pai, 1800 m, Pacholátko & Dembický leg.”; 1 ♀ (EJCB): “Thai, Mae Hong Son pr., 19°27’N, 98°20’E, 1500 m, Soppong 7 12.V., Vít Kubáň leg., 1996”. VIETNAM: Gia Lai: 1 ♀ (EJCB): “S Vietnam, 14.10N 108.30E, 40 km NW Ankhe, Buon Luoi, 620 750 m, 28.3. 12.4.1995, Pacholátko & Dembický leg.”.

Adult occurence: 3-4-5-6-11 (n=16). Altitude range: 501800 m (n=11). Host plants: Specimen from Hyderabad (India) (see Specimens examined) was reared from Chloroxylon swietenia. Distribution: CAMBODIA. CHINA: Yunnan. INDIA. LAOS. NEPAL. THAILAND. VIETNAM.

Agrilus rudicollis Alexeev Fig. 31C (habitus); Fig. 41Q (aedeagus); Fig. 52G (type) rudicollis Alexeev, 1979 (Agrilus; subgenus Dentagrilus) ALEXEEV, 1979: 136 [#57] (description). ALEXEEV, 1981: 56 58

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA [#59] (subgenus Dentagrilus; redescription). ALEXEEV, 1989: 484 [#64] (key to species; Yu Primorsk; northeastern China). ALEXEEV, 1998: 378 [#4322] (subgenus Dentagrilus; subgeneric classification). JENDEK, 2006: 390 [#7441] (subgenus Dentagrilus; Palaearctic catalog). BELLAMY, 2008: 2272 [#8576] (subgenus Dentagrilus; world catalog). VOLKOVITSH, 2009: 135 [#8850] (subgenus Dentagrilus; faunal records; Russia: Primorskiy kray).

Type material: Agrilus rudicollis Alexeev, 1979 Type locality: S.V. Kitai, Vost. liniya KVZhD, st. Gaolinczy [in Russian]. Type specimens: Primary: Holotype (Fig. 52G), ♂, (ZIN): “St. Gaolinczy, Vost. liniya KVZhD Kitai 2. VII. [1]940 [h] [in Russian] \ Agrilus rudicollis Alex. [h] Alexeev det. [p] Holotypus [h] Alexeev det. [p]”. Secondary: 1 PT (ZIN). Described from 5 specimens (HT, PT). Diagnosis: 4.1-6 mm; head small; pronotum with obvious, deep lateral impressions; anterior pronotal lobe subangular, extending beyond anterior pronotal angles; prehumerus carinal and short. Species group: Unassigned. Relevant species: Unknown. Specimens examined: RUSSIA: Primorskiy kray: 1 ♀ (EJCB): “USSR, Ussuri prov. Arsenev env., VII.1990, leg. B. Šiška & R. Čermák”; 1 (EJCB): “Russia: Primorskiy Kray, 13 km S Ussuriysk, 43.42 N, 132.00E, 15.vi.1993, 100 m, leg. L. Zerche et al.”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., ~10 кm SW Sokoltchi 28.06.06 leg. М.& L. Smirnov’s”.

Adult occurence: 5-6-7 (n=7). Faunal records published by ALEXEEV, 1981 [#59] are also included. Altitude range: ~100 m (n=1). Host plants: Ulmus: ALEXEEV, 1979 [#57]; ALEXEEV, 1981 [#59]; ALEXEEV, 1989 [#64]. Distribution: CHINA: Heilongjiang. RUSSIA: Primorskiy kray.

Agrilus sachalinicola Obenberger Fig. 9E (habitus); Fig. 32K (aedeagus) sachalinicola Obenberger, 1940 (Agrilus) OBENBERGER, 1940: 115 [#2297] (description). KUROSAWA, 1954: 92 [#1612] (faunal records; Japan). AKIYAMA, 1974: 8 [#3176] (faunal record; Japan: Shizouka). KUROSAWA, 1974: 4 [#4577] (note). TÔYAMA, 1985: 23 [#3678] (iconography; Japan). ALEXEEV, 1989: 482 [#64] (key to species; Sakhalin). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). LI JINGKE, 1992: 92 [#6755] (checklist; China: Heilongjiang). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 37 38 [#4091] (checklist; Japan: Hokkaido, Honshu; Russia: Sakhalin). ALEXEEV, 1998: 372 [#4322] (subgenus Uragrilus; subgeneric classification). FUKUTOMI & HORI, 2004: 11 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 395 [#7441] (subgenus Uragrilus; Palaearctic catalog). BELLAMY, 2008: 2275 [#8576] (subgenus Uragrilus; world catalog). VOLKOVITSH, 2009: 137 [#8850] (subgenus Uragrilus; faunal records; Russia: Primorskiy kray). tamanukii Stepanov, 1954 (Agrilus; subgenus Epinagrilus) STEPANOV, 1954: 116 117, 119 [#2662] (description). ALEXEEV, 1989: 482 [#64] (synonym of sachalinicola). JENDEK, 2006: 395 [#7441] (synonym of sachalinicola; Palaearctic catalog).

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Type material: Agrilus sachalinicola Obenberger, 1940 Type locality: Sachalin: Uchirda Okada. Jawamoto. Type specimens: Not located. Type is missing in NMPC. Described from unknown number of syntypes. Note: Akiyama (pers. com.) has examined the type from NMP labeled: “Saghalien, Uchida, Okada, Sawamoto / [reverse side] Konuma [Japanese grapheme] (Konuma), 27.vii.1933”. Agrilus tamanukii Stepanov, 1954 Type locality: na Yuzhnom Sakhaline, v raione Kanume [in Russian]. Type specimens: Not located. Described from 1 specimen. Type not found in ZIN. Diagnosis: 7.8-9.9 mm; body robust; eyes moderate; anterior pronotal lobe moderate; posterior pronotal angles subangular or feebly acute; prehumerus tubercular or short carinal. Species group: Spinipennis. Relevant species: A. spinipennis Lewis, 1893 Specimens examined: JAPAN: Honshu: 1 ♂ (EJCB): “1992.7.30. Masuzawa, Tateiwa V., S Aizu, Fukushima, NE Japan, K. Shindo leg.”. RUSSIA: Primorskiy kray: 1 ♂ (MSCI): “SE Primorskii kr., Lazovskii zap., 12 кm S pos., Staraja Kamenka, okr. kor. Amerika, 43°15’N, 134°1’E, 6.07.04 leg. М.& L. Smirnov’s [in Russian]”. Sakha Respublika: 1 ♀ (EJCB): “Jakutia, Agajakan, 5.7.1990”.

Adult occurence: 6-7-8 (n=8). Faunal records published by KUROSAWA, 1954 [#1612] are also included. Host plants: Alnus hirsuta AKIYAMA & OHMOMO, 1997 [#4091]. – Salix bakko: AKIYAMA & OHMOMO, 1997 [#4091]. Erroneous: Populus: STEPANOV, 1954 (as tamanukii) [#2662]. Distribution: JAPAN: Hokkaido, Honshu. RUSSIA: Primorskiy kray, Sakhalinskaya oblast’, Sakha Respublika. Remarks: The concept of this taxon was judged from the Japanese specimen (see Specimens examined) determined by S. Ohmomo. The distributional record for this species without supporting data from Heilongjiang (China) cited by LI JINGKE, 1992 [#6755] was omitted. The specimen from Sakha differs from the specimen from Honshu (see Specimens examined) by the tubercular prehumerus, cuspidate elytral apices and by the greenish tinge of the elytra.

Agrilus sagittifer Jendek, sp. nov. Fig. 27B (habitus of HT); Fig. 39O (aedeagus)

Description: 4.3-5.2 mm (Holotype 4.8 mm). BODY: Shape: subparallel, Build: robust, Color (dorsally): unicolored, Color (dominant): black, green or brown. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (density): dense. Eyes: Size: large, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: transverse, Sides:

177 arcuate, rarely straight, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: obtuse, rarely acute or rectangular, Apex: sharp, rarely blunt. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial or posteromedial only, Medial impression (deepness): shallow, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial. Prehumerus: carinal, Shape: bisinuate, Extent: beyond half of pronotal length or to anterior pronotal angle, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina or joining anterior pronotal angle, Arc: moderate or weak. Marginal and submarginal carinae: Interspace: moderate or wide, Convergence: moderately convergent or strongly convergent, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: conjoint, Width: wide, Shape: subtruncate, Truncation: oblique with projecting sutural angle, Modifications: sutural rim spinuliform. Elytral pubescence: proximal and distal or proximal only, Proximal: like V or upturned T, Distal: preapical, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: large, Anterior margin: arcuately emarginate, Emargination (deepness): shallow, Emargination (width): narrow. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: acute, rarely rectangular, Angles (tips): sharp, rarely blunt, Disk: impressed. ABDOMEN: Sexual modifications in male: last ventrite impressed on apex, rarely medial sulcus on ventrite 1, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate or angulately sinuate, Sinuosity (deepness): markedly deep, Sinuosity (width): very narrow. LEGS: Male sexual modifications: metafemur markedly incrassate. Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed, Ovipositor: elongate. Diagnosis: elytral pubescence often vanished posteriorly; apices subtruncate with lateral angle subangulate; medial lobe of aedeagus with sharp apex. Species group: Cyaneovirens. Relevant species: A. tsushimanus Kurosawa, 1963. Type locality: China, Shaanxi province, Qinling mts., 6 km east of Xunyangba, altitude 1000-1300 m. Type specimens: Primary: Holotype (Fig. 27B), ♂, (EJCB): “China, 1000-1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.-13.vi.1998, I. H. Marshal leg.”. Secondary: CHINA: Henan: 1 ♀ PT (EJCB): “China, W Henan, 1.vii.2001, 20 km SE Luanchan “tunel”, 33.7°-111.8°E, O. Šafránek leg.”; Shaanxi: 6 PT (EJCB): “China, 1000-1300

178 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.-13. vi.1998, I. H. Marshal leg.”; 1 ♀ PT (EJCB): “China, Shaanxi prov., 21-23.vi.1998, Quin Ling Shan mts, road Baoji-Taibai vill., pass 35 km S Baoji, O. Šafránek et M. Trýzna lgt.”; 1 ♂ PT (EJCB): “C-China, Shanxi [sic! should be Shaanxi], Qinling Shan, 6km E of Xunyangba, 1000-1300 m, 23.v.-13. vi., leg. C. Holzschuh, 2000”. Adult occurence: 5-6-7 (n=4). Altitude range: 1000-1300 m (n=2). Host plants: Unknown. Distribution: CHINA: Henan, Shaanxi. Etymology: The specific name was derived from the Latin noun sagitta (arrow) and the the verb fero (bear or carry); it refers to the sharp shape of of the apical portion of the aedeagus.

Agrilus salicis Frivaldszky Fig. 17D (habitus); Fig. 35I (aedeagus); Figs. 42O, 46F, 48Q, 52M (types) salicis Frivaldszky, 1877 (Agrilus; variety of viridis) FRIVALDSZKY, 1877: 319 [#3933] (description). JAKOBSON, 1913: 797 [#4386] (variety of viridis; catalog; Russia and Europe). OBENBERGER, 1916: 272 [#1983] (variety of viridis; notes). OBENBERGER, 1926: 656 [#2057] (variety of chrysoderes; Palaearctic catalog). OBENBERGER, 1936: 1068 [#2191] (aberration of viridis; world catalog). RIKHTER, 1945: 166 [#3507] ([Note: Misidentified as aurichalceus]; charactes). SCHAEFER, 1949: 398 [#2565] (variety of viridis; characters). ALEXEEV, 1957: 152 [#3226] ([Note: Misidentified as aurichalceus]; characters in key). ALEXEEV, 1958: 185 [#3982] ([Note: Misidentified as aurichalceus]; checklist; Moskovsk). STEPANOV, 1958: 172 [#3664] ([Note: Misidentified as aurichalceus]; Moldavia; Ukraine; Stalingradsk; Saratovsk; Kuibyshevsk). ALEXEEV, 1959: 15 [#3229] ([Note: Misidentified as aurichalceus]; characters in key; Tatarstan; Moskovsk; Kievsk; Khar’kovsk; Belgorodsk; Kyrsk; Voronezhsk; Saratovsk; Kamensk; Lugansk; Voroshilovgradsk; Kuibyshevsk; Stalingradsk; west Kazakhstan; ?Transcaucasia). ALEXEEV, 1960: 1508 [#45] ([Note: Misidentified as aurichalceus]; larval characters in key). ALEXEEV, 1961: 9 [#3243] ([Note: Misidentified as aurichalceus]; larval characters in key). ZAGAIKEVICH, 1962: 107 [#3249] ([Note: Misidentified as aurichalceus]; faunal records; bionomy; Ukraine). RIKHTER & ALEXEEV, 1965: 300 [#2522] ([Note: Misidentified as aurichalceus]; characters in key; European part of USSR). KOSTIN, 1973: 107 [#3693] ([Note: Misidentified as aurichalceus]; characters in key; Kazakhstan). GURYEVA, 1974: 98 [#5428] ([Note: Misidentified as aurichalceus according to host plant cited]; pest of agriculture cultures; references; Soviet Union). ALEXEEV, 1975: 158 [#52] ([Note: Misidentified as aurichalceus according to JENDEK, 2003 [#5878]]; distribution overview). VOLKOVITSH, 1977: 26 [#2985] ([Note: Misidentified as aurichalceus]; faunal record; notes; USSR: Belgorod, Voronezh; Kursk; Kharkov). NIEHUIS & TEZCAN, 1993: 34 35, 48 [#1910] (subgenus Agrilus; faunal records from Turkey; distribution and host plants summary; Turkey). CURLETTI, 1994: 22, 172 [#658] (subgenus Agrilus; catalog; faunal records; Italia; Sardegna). ALEXEEV, 1998: 381 [#4322] (subgenus Agrilus; subgeneric classification). BÍLÝ, 2002: 25 [#5877] (bionomy; host plants; distribution). BRECHTEL & KOSTENBADER, 2002: 296 297 [#5766] (biology; ecology; distribution; Germany). JENDEK, 2002: 2, 3 4 [#5348] (lectotype designation; taxonomic and nomenclatural notes). JENDEK, 2003: 6, 19 [#5878] (notes;

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA host plants). GOTTWALD & HORNBURG, 2004: 49 50 [#7007] (faunal records; Germany). JENDEK, 2006: 389 [#7441] (subgenus Agrilus; Palaearctic catalog). LÖBL & SMETANA, 2007: 28 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2276 [#8576] (subgenus Agrilus; world catalog). NIKITSKY et al., 2008: 149 150 [#8610] (faunal record; Krasnodar; Karachayevo Cherkessia). alutaceus Obenberger, 1916 (Agrilus; variety of aurichalceus) OBENBERGER, OBENBERGER, 1916: 273 [#1983] (description). 1926: 656 [#2057] (aberration of aurichalceus; Palaearctic catalog). OBENBERGER, 1927: 212, 222, 244 [#2064] (variety of aurichalceus). OBENBERGER, 1935: 69 [#2178] (variety of aurichalceus; catalog). OBENBERGER, 1936: 956 [#2191] (variety of aurichalceus; world catalog). SCHMIDT, 1937: 262 [#5323] (variety of aurichalceus; notes; faunal records; Germany). SCHAEFER, 1949: 393 [#2565] (variety of acutangulus). HORION, 1955: 96 [#1285] (variety of acutangulus; note). COBOS, 1986: 251 [#609] (subgenus Agrilus; form of acutangulus). JENDEK, 2002: 2, 3 4 [#5348] (synonym of salicis; lectotype designation; taxonomic and nomenclatural notes). JENDEK, 2006: 389 [#7441] (synonym of salicis; Palaearctic catalog). BELLAMY, 2008: 2276 [#8576] (synonym of salicis; world catalog). NIKITSKY et al., 2008: 148 [#8610] (synonym of salicis). filigranus Obenberger, 1916 (Agrilus) syn. nov. OBENBERGER, 1916: 274 275 [#1983] (description). OBENBERGER, 1926: 656 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 990 [#2191] (world catalog). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2093 2094 [#8576] (world catalog). acutangulus Schaefer, 1949 (Agrilus) SCHAEFER, 1949: 355, 393 [#2565] (new name proposal [Note: Established as a new name for an unavailable name acutangulus Théry (Article 45.5.1)]). HORION, 1955: 96 [#1285] (Théry is cited as the author; distribution overview; biology; notes; Deutschland). ZAGAIKEVICH, 1962: 107 [#3249] (Théry is cited as the author; synonym of aurichalceus). BALACHOWSKY, DAVATCHI & DESCARPENTRIES, 1962: 286 [#96] (Théry is cited as the author; economic importance). ZAGAIKEVICH, 1962: 60 [#5616] (Théry is cited as the author; checklist; Ukraine). HELLRIGL, 1978: 267 [#3442] (Théry is cited as the author; critical overview of host plants). LOMPE, 1979: 237 [#3628] (Théry is cited as the author; characters in key; Central Europe). ALEXEEV, 1981: 76 [#61] (Théry is cited as the author; larval characters in key). COBOS, 1986: 251, 317 [#609] (Théry is cited as the author; subgenus Théry is cited as the author; characters in key). NIEHUIS & Tezcan, 1993: 35 [#1910] (Théry is cited as the author; synonym of salicis). RUICANESCU, 1993: 211 [#3257] (Théry is cited as the author; synonym of salicis). CURLETTI, 1994: 172 [#658] (Théry is cited as the author; synonym of salicis). FARRUGIA, 2002: 107 [#6511] (Théry is cited as the author; synonym of salicis). BÍLÝ, 2002: 25 [#5877] (synonym of salicis). JENDEK, 2002: 2, 3 4 [#5348] (synonym of salicis; nomenclatural notes). JENDEK, 2006: 389 [#7441] (synonym of salicis; Palaearctic catalog). BELLAMY, 2008: 2277 [#8576] (synonym of salicis; world catalog). lugovoi Stepanov, 1958 (Agrilus) syn. nov. STEPANOV, 1958: 172 173, 174 [#3664] (description). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2171 [#8576] (world catalog).

Type material: Agrilus viridis variety salicis Frivaldszky, 1877 Type locality: Comitata Temesiensi ad pagum Grebenácz. Type specimens: Primary: Lectotype (Fig. 52M), ♂, designated by JENDEK (2002: 2, 3-4) [#5348], (HNHM):

179

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA “KRASSÓ- M. [p] \ Typus [p] [red ink] Agrilus salicis J. Frivaldszky [h]”. Secondary: 5 PLT (HNHM). Described from unknown number of syntypes. Agrilus aurichalceus variety alutaceus Obenberger, 1916 Type locality: Südrußland: Sarepta, Kroatien: Vinkovci. Type specimens: Primary: Lectotype (Fig. 42O), sex not examined, designated by JENDEK (2002: 2, 3-4) [#5348], (NMPC): “Sarepta [h] \ Typus [p] [red label] \ Agrilus aurichalceus ssp. alutaceus m. Det. Dr. Obenberger [h] \ Mus. Nat. Pragae Inv. [p] 24819 [h] [pink label]”. Secondary: 1 PLT (NMPC). Described from unknown number of syntypes. Agrilus filigranus Obenberger, 1916 Type locality: Rußland. Type specimens: Holotype by monotypy (Fig. 46F), ♂, (NMPC): “Rossia [h] \ Typus [p] [red label] \ Agrilus filigranus mihi Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae [p] 24822 [h] Inv. [p] [orange label]”. Described from 1 specimen. Agrilus acutangulus Schaefer, 1949 Type locality: Le Plantay (Ain). Type specimens: The specific name acutangulus Schaefer is based on the type of Agrilus aurichalceus aberration acutangulus Théry, 1942. Described from unknown number of syntypes. The type was not found in MNHN. Agrilus lugovoi Stepanov, 1958 Type locality: v raione B. Barsukov, st. Cholkar, Aktyubinskoi oblasti Kazakskoi SSR [in Russian]. Type specimens: Primary: Lectotype (Fig. 48Q) by present designation, ♂, (ZIN): “Kazakh. SSR Aktyubinsk. ob. B. Barsuki 22-V-1950 A. V. Lugovoi [h] \ Typus [h] [red label] \ Agrilus lugowoi [sic!] sp. n. 52 [h] \ V. Stepanov det. [p] \ Syntypus [p] [red label]”. Secondary: 2 PLT (ZIN). Described from 9 syntypes. Diagnosis: 4.7-5.8 mm; body with silky tinge, head and pronotum with disctinct microsculpure; eyes small, not reaching to antennal sockets (lateral view); elytral apices broad. Species group: Viridis. Relevant species: A. cuprescens (Ménétriés, 1832); A. viridis (Linné, 1758). Specimens examined: KAZAKHSTAN: 2 (ZIN): “g. Alutau Karagand. obl. Loginova [in Russian] 26.vi.1958”; 1 (ZIN): “g. Aktau, 60 km S Zhana Arka, Karag. Loginova 19.vii.1959 [in Russian]”. 1 (ZIN): “Alma Ata okr., 19 vi 1949, Skopin N. G. [in Russian]”; 1 ♂ (EJCB): “Yanvarcevo, prav. b. Urala, Kazakhstan, Romadina, 28 vi [1]950 [in Russian]”; 30 (ZIN): “Yanvarcevo, prav. b. Urala, Kazakhst. Romadina 27.v 28.vi.1950 [in Rusian]”; 1 (ZIN): “r. Ural u Khar’kina, Kazakhst., Romadina, 17.v.1951 [in Russian]”; 1 ♀ (ZIN): “khr. Tarbagatai, 40 km V p. Tarbagatai, V. Kazakhstan, Volkovich, 14 15.07.[1]988 [in Russian]”. RUSSIA: Altay Respublika: 2 (MNCA): “Altaj, Gorno Altajsk, 15.7.1984, ex Salix, Beloborodov”. Belgorodskaya oblast’: 1 (ZIN): “uchkhoz les na Vorskle, Belgorodsjaya obl., Geraseb, 7.vii.1971”. Voronezhskaya oblast’: 100 (ZIN): “Saval’skoe l. Voronezhskaya obl. 29.vi.1950 V. Stark leg. [in Russian]”.

Adult occurence: 5-6-7 (n=12). Host plants: Salix: THÉRY, 1942 [#2873] (as acutangulus); ALEXEEV, 1958 [#3982]; ALEXEEV, 1961 [#3243]; ZAGAIKEVICH, 1962 [#3249]; GURYEVA, 1974 [#5428]; ALEXEEV, 1975 [#52];

VOLKOVITSH, 1977 [#2985]; ALEXEEV, 1981 (as acutangulus) [#61]; CURLETTI, 1994 [#658]; NIKITSKY et al., 2008 [#8610]. – S. acutifolia: GUTOWSKI & MILKOWSKI, 2008 [#8244]. – S. americana: BALACHOWSKI et al., 1962 (as acutangulus) [#96]; HELLRIGL, 1978 (as acutangulus) [#3442]. – S. amygdalina: BALACHOWSKI et al., 1962 (as acutangulus) [#96]; HELLRIGL, 1978 (as acutangulus) [#3442]. – S. aurita: SCHAEFER, 1949 (as acutangulus) [#2565]; FRANZ, 1974 (as acutangulus) [#4334]; HELLRIGL, 1978 (as acutangulus) [#3442]; NIEHUIS, 1988 [#1912]; BÍLÝ, 2002 [#5877]; BRECHTEL & KOSTENBADER, 2002 [#5766]. – S. caprea: SCHAEFER, 1949 (as acutangulus) [#2565]; FRANZ, 1974 (as acutangulus) [#4334]; HELLRIGL, 1978 (as acutangulus) [#3442]; BÍLÝ, 2002 [#5877]; BRECHTEL & KOSTENBADER, 2002 [#5766]. – S. cinerea: NIEHUIS & TEZCAN, 1993 [#1910]; BÍLÝ, 2002 [#5877]; BRECHTEL & KOSTENBADER, 2002 [#5766]. – S. incana: HELLRIGL, 1978 (as acutangulus) [#3442]. – S. purpurea: BALACHOWSKI et al., 1962 (as acutangulus) [#96]; HELLRIGL, 1978 (as acutangulus) [#3442]. – S. repens: − S. repens: FRIVALDSZKY, 1877 [#3933]; STEPANOV, 1958 (as lugovoi) [#3664]; JENDEK, 2003 [#5878]. – S. rosmarinifolia: ALEXEEV, 1959 [#3229]; RIKHTER & ALEXEEV, 1965 [#2522]; RUICANESCU, 1993 [#3257]. – S. rossica: RIKHTER & ALEXEEV, 1965 [#2522]. – S. triandra: ALEXEEV, 1959 [#3229]; RIKHTER & ALEXEEV, 1965 [#2522]. – S. viminalis: HORION, 1955 (as acutangulus) [#1285]; BALACHOWSKI et al., 1962 (as acutangulus) [#96]; HELLRIGL, 1978 (as acutangulus) [#3442]; GOTTWALD & HORNBURG, 2004 [#7007]; GUTOWSKI & MILKOWSKI, 2008 [#8244]. Erroneous: Populus: STEPANOV, 1958 (as lugovoi) [#3664]. Distribution: ASIA: KAZAKHSTAN. RUSSIA. TURKEY. UZBEKISTAN. EUROPE: ALBANIA. AUSTRIA. BELARUS. BOSNIA & HERZEGOVINA. BULGARIA. CROATIA. CZECH REPUBLIC. FRANCE. GERMANY. GREECE. HUNGARY. ITALY. MACEDONIA. MOLDOVA. POLAND. ROMANIA. RUSSIA. SERBIA & MONTENEGRO. SLOVAKIA. SPAIN. SWITZERLAND. UKRAINE. Remarks: This species has not yet been recorded from East Asia but it is possible in Mongolia and Respublika Tyva (Russia).

Agrilus satoi Kurosawa Fig. 10L (habitus); Fig. 32AA (aedeagus); Fig. 52K (type) satoi Kurosawa, 1954 (Agrilus) KUROSAWA, 1954: 85 87 [#1613] (description). PENG ZHONGLIANG, 1987: 358 [#2429] (checklist; China). TÔYAMA, 1988: 756 [#2932] (note). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). OHMOMO & AKIYAMA, 1994: 37 [#3822] (faunal records; Japan: Kyushu). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 38 [#4091] (checklist; Japan: Kyushu; Taiwan). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Taiwan). MÜHLE, 2003: 47 [#6155] (subspecies of rokuyai; checklist; Taiwan). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). ANONYMOUS, 2007: 32 [#8250] (checklist; Japan: Honshu). BELLAMY, 2008: 2279 2280 [#8576] (world catalog).

180 Type material: Agrilus satoi Kurosawa, 1954 Type locality: Mt. Rarasan, N. Formosa. Type specimens: Primary: Holotype (Fig. 52K), ♂, (NSMT): “Rarasan Mt. N. Formosa 12. VI. 1939 T. Kaneko [h] \ Holotype [p] Agrilus satoi Y. Kurosawa, 1954 [h] [red label]”. Described from 2 specimens (HT, AT). Diagnosis: 6.9 9.4 mm; apical third of elytra feebly attenuate; elytral apices sharply cuspidate. Species group: Niveoguttatus. Relevant species: A. niveoguttatus Kerremans, 1892. Specimens examined: CHINA: Fujian: 1 ♂, 1 ♀ (EJCB): “Chian, Fujian. 7.v., Baishi Feng, 500 700 m, 26°47’N, 116°57’E, Jaroslav Turna leg., 2008”. Guizhou: 16 (NMW): “China: Guizhou, Leishan Co., SE Kaili, 15 km E Leishan, Leigong Shan, E slope, 26º23.39’N, 108º13.33’E / ca 2.5 km E of pass between Leishan and Fangxiang vill., ca 1600 m, 23/24.6.2001, leg. Schillhammer (17)”. Jiangxi: 5 (EJCB): “China, Jiangxi W, Jinggang Shan, Ciping, 2 14.VI.1994, E. Jendek & O. Šauša leg.”. Shaanxi: 30 (EJCB): “China, 1000 1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v. 13.vi.1998, I. H. Marshal leg.”; 1 ♀ (EJCB): “C China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000 1300m, 23.V. 13.VI. leg. C. Holzschuh 2000”. Taiwan: 1 (EJCB): “Nashanchi (Formosa), 29 31.v.1971, T. Mizunuma”; 1 ♀ (EJCB): “(Tsuifeng) Nantou Hsien, Taiwan, July 11th, 1978, Coll. Tshimomura”; 1 (PMCL): “Rep. of China Formosa (Taiwan) A Li Shan 17 26.6.1995, P. Moravec”. JAPAN: Honshu: 1 ♂ (EJCB): “Japan, 29.7.1992, Miyagawa, Mie pref., Otobe leg.”. Adult occurence: 5 6 7 (n 11). Faunal records published by KUROSAWA, 1954 [#1613] and by TÔYAMA, 1988 [#2932] are also included. Altitude range: 1000 1600 m (n 4). Host plants: Unknown. Distribution: CHINA: Fujian, Guizhou, Jiangxi, Shaanxi, Taiwan. JAPAN: Honshu, Kyushu.

Agrilus schillhammeri Jendek, sp. nov. Fig. 26I (habitus of HT); Fig. 39J (aedeagus)

Description: 5.4-6.3 mm (Holotype 6.3 mm). BODY: Shape: subparallel, Build: robust, Color (dorsally): unicolored, Color (dominant): black or brown. HEAD: Medial impression: present, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: flat, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): semispherical, Sculpture (intensity): superficial. Eyes: Size: large, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate or long, Sexual modification in male: antennae longer. PRONOTUM: Shape: visually square or transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate or obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Lateral impressions (deepness): shallow, Lateral impressions (width): narrow. Prehumerus: carinal, Shape: arcuate, Extent: to half of pronotal length, Modifications: obsolete

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA in anterior part, Posterior end: joining posterior pronotal angle, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: narrow or moderate, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior third of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: conjoint, rarely separate, Width: wide, Shape: arcuate. Elytral pubescence: proximal and distal, Proximal: like V or upturned T, Distal: preapical, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate or large, Anterior margin: arcuately emarginate, Emargination (deepness): shallow or moderate. Prosternal process: moderate or wide, Shape: dilated, Sides: straight, Angles: rectangular, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate, Sinuosity (deepness): very shallow, Sinuosity (width): markedly wide. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe markedly obtuse, Ovipositor: elongate. Diagnosis: body brown-black; antennae longer; elytral apices shallowly arcuate; sternal groowe on apex of last ventrite shallowly emarginate. Species group: Cyaneovirens. Relevant species: A. cyaneovirens Bourgoin, 1922. Type locality: China, Guizhou province, Jiangkou Co., about 50 km southwest Jiangkou, surrounding of Shidu village, 27°32.83’N, 108°36.45’E, altitude about 650 m. Type specimens: Primary: Holotype (Fig. 26I), ♂, (NMW): “China: Guizhou, Jiangkou Co., ca 50 km SW Jiangkou, surr. of Shidu vill. \ 27°32.83’N, 108°36.45’E, ca 650 m, 29.66.7.2001, leg. Wang & Shillhammer”. Secondary: CHINA: Guizhou: 3 PT (NMW): “China: Guizhou, Jiangkou Co., ca 50 km SW Jiangkou, surr. of Shidu vill. \ 27°32.83’N, 108°36.45’E, ca 650 m, 29.6-6.7.2001, leg. Wang & Shillhammer”. Adult occurence: 6-7 (n=1). Altitude range: 650 m (n=1). Host plants: Unknown. Distribution: CHINA: Guizhou. Etymology: Patronymic; the species was named in honour of Harald Schillhammer, an eminent expert on Staphylinidae, who collected the type series.

Agrilus sentis Jendek, sp. nov. Fig. 25A (habitus of HT); Fig. 38T (aedeagus)

Description: 6.8-10. mm (Holotype 9.2 mm). BODY: Shape: subparallel, Build: slender, Color (dorsally): unicolored

181

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA or bicolored, Color (dominant): brown. HEAD: Medial impression: present, Impression (extent): vertex or vertex and frons, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (density): dense, Sculpture (intensity): superficial. Eyes: Size: large, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: visually elongate or visually square, Sides: arcuate, Maximal width: at anterior margin or at middle, Anterior margin: wider than posterior or subequal to posterior. Anterior lobe: obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: acute or rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial. Prehumerus: obsolete or carinal, Shape: straight, Extent: to third of pronotal length, Posterior end: distant from pronotal angle or margin, Anterior end: distant from pronotal marginal carina. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior angles, Modifications: rarely submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate or conjoint, Shape: arcuate, subangulate or subtruncate, Truncation: transverse or oblique with projecting pleural angle, Modifications: sutural rim spinuliform. Elytral pubescence: distal only, Distal: apical, Color: unicolored. Tomentum: absent. Prosternal lobe: narrow, Anterior margin: angulately emarginate. Prosternal process: wide, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Apical margin of pygidium: extended into long spine, Apical margin of last ventrite: laterally emarginate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Ovipositor: elongate. Diagnosis: prehumerus obsolete; elytra glabrous except of apicalmost part; apices arcuate, subacuminate or subtruncate with spinuliform sutural rim. Species group: Tempestivus. Relevant species: A. asiaticus Kerremans, 1898. Type locality: Central China, Shaanxi province, Qinling Shan, 6 km east of Xunyangba, altitude 1000-1300 m. Type specimens: Primary: Holotype (Fig. 25A), ♂, (EJCB): “C-China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000-1300m, 23.V.-13.VI. leg. C. Holzschuh 2000”. Secondary: CHINA: Shaanxi: 2 ♀ PT (EJCB): “China, Shaanxi pr., Hua Shan, 17-21.VI.1991, R. Sauer lgt.”; 1 ♀ PT (EKCS): “China - Shaanxi, Lüeang 33°07’N 106°05’E, 18.6.-24.6.[19]97 lgt. E. Kučera”; 1 ♂ PT (EKCS): “China - Shaanxi, Lüeang 33°07’N 106°05’E, 19.7.-27.7.[19]97 lgt. E. Kučera”; 1 ♀ PT (EJCB): “China, 1000-1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.-13.vi.1998, I. H. Marshal leg.”; 8 PT (EJCB): “C-China, Shaanxi, Qinling

Shan, 6 km E of Xunyangba, 1000-1300m, 23.V.-13.VI. leg. C. Holzschuh 2000”. Adult occurence: 5-6-7 (n=5). Altitude range: 1000-1300 m (n=2). Host plants: Unknown. Distribution: CHINA: Shaanxi. Etymology: The specific name is the Latin noun sentis (thorn) in apposition. It refers to the spinuliform sutural rim of elytral apices of this species.

Agrilus shan Jendek, sp. nov. Fig. 14F (habitus of HT); Fig. 34K (aedeagus)

Description: 5.3-7.3 mm (Holotype 5.4 mm). BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): subparallel, Sculpture (intensity): superficial. Eyes: Size: large, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short, Shape: stout, Sexual modification in male: not apparent. PRONOTUM: Sides: arcuate, Maximal width: at anterior margin, rarely at middle, Anterior margin: wider than posterior. Anterior lobe: absent, rarely vague, Position to anterior angles: not reaching level of anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: flat, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow. Prehumerus: absent, obsolete or tubercular. Marginal and submarginal carinae: Interspace: wide, Convergence: strongly convergent, Junction: absent, Narrowest point: at posterior third of pronotal length, Modifications: submarginal carina posteriorly obliterate or submarginal carina divergent at posterior angles. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Shape: arcuate. Elytral pubescence: perisutural stripes, distal only, rarely absent, Color: unicolored. Tomentum: rarely absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: large, Anterior margin: arcuate. Prosternal process: moderate, Shape: narrowed, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe markedly obtuse, Ovipositor: uritiform.

182 Diagnosis: prehumerus obsolete or tubercular; elytral apices less elongate; medial lobe of aedeagus wider, broadly arcuate on apex. Species group: Viduus. Relevant species: A. viduus Kerremans, 1914. Type locality: Central China, Shaanxi province, Qinling Shan, 6 km east of Xunyangba, altitude 1000-1300 m. Type specimens: Primary: Holotype (Fig. 14F), ♂, (EJCB): “C-China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000-1300m, 23.V.-13.VI. leg. C. Holzschuh 2000”. Secondary: CHINA: Shaanxi: 1 ♂ PT (EJCB): “China, Shaanxi pr., Hua Shan, 17-21.VI.1991, R. Dunda lgt.”; 9 PT (EJCB): “C-China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000-1300m, 23.V.-13.VI. leg. C. Holzschuh 2000”; 1 ♂ PT (EJCB): “China-Shaanxi, Zhongchuan, 3.65.6.2005, lgt. E. Kučera”. Adult occurence: 5-6 (n=3). Altitude range: 1000-1300 m (n=1). Host plants: Unknown.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA KUROSAWA, KUROSAWA, 1963: 104 106 [#1617] (description). 1963: 153 [#1618] (characters; Japan). KUROSAWA, 1974: 4 [#4577] (note). TÔYAMA, 1985: 21 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 38 [#4091] (checklist; Japan: Honshu, Shikoku, Kyushu). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). ANONYMOUS, 2007: 32 [#8250] (checklist; Japan: Honshu). BELLAMY, 2008: 2291 [#8576] (world catalog).

Type material: Agrilus shashamboe Kurosawa, 1963 Type locality: Kôchi Pref.: Mt. Kajigamori. Type specimens: Primary: Holotype not in NSMT (S. Nomura pers. com. 2001). Secondary: 3 PT (NSMT). Described from unknown number of specimens (HT, AT, PT). Paratype (Fig. 52T) from NSMT is illustrated. Diagnosis: 4.4-6.3 mm; pronotum widest in anterior third; aedeagus subparallel. Species group: Maculifer. Relevant species: A. nelsonodaimio Ohmomo, 2006.

Distribution: CHINA: Shaanxi.

Specimens examined: JAPAN: Honshu: 2 ♂ (EJCB): “Mt. Sawaguchi, Shizuoka pref., 15.vii.1978, S. Morita leg.”.

Etymology: The specific name is the Chinese noun shan (mountain, hill) in apposition. It refers to the occurence of this species in the mountains of Qinling Shan.

Adult occurence: 7-8 (n=3). Faunal records published by KUROSAWA, 1963 [#1617] are also included.

Agrilus shaowensis Jendek

Host plants: Rhaphiolepis umbellata: AKIYAMA & OHMOMO, 1997 [#4091]; Vaccinium bracteatum: KUROSAWA, 1963 [#1617]; AKIYAMA & OHMOMO, 1997 [#4091].

Fig. 31D (habitus of HT); Fig. 41R (aedeagus)

Distribution: JAPAN: Honshu, Kyushu, Shikoku.

shaowensis Jendek, 1994 (Agrilus) JENDEK, 1994: 33 34 [#1356] (description). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2290 [#8576] (world catalog).

Remarks: The concept of this taxon was judged from three paratypes sent for examination by S. Nomura (NSMT) in 2001.

Type material: Agrilus shaowensis Jendek, 1994 Type locality: China, prov. Fujian, Shaowu env. Type specimens: Primary: Holotype (Fig. 31D), ♂, (EJCB): “China, Fujian prov., Shaowu env., 11-13.6.1991”. Secondary: 2 PT (EJCB). Described from 3 specimens (HT, PT). Diagnosis: 6.3-8.8 mm; eyes markedly small and flat; elytra white pubescent except for glabrous spots at proximal and distal third. Species group: Unassigned. Relevant species: Unknown, but it might be close to the species around A. pekinensis Obenberger, 1924. Specimens examined: CHINA: Fujian: 2 ♀ PT (EJCB): “China, Fujian prov., Shaowu env., 11 13.6.1991” (JENDEK, 1994 [#1356]); 3 (EJCB): “China, Fujian prov., Wuyi Shan Nat. Par., Sangan env. (900 m), 30.v. 12.vi.2001, Hlaváč, P. & Cooter, J. lgt.”.

Adult occurence: 6 (n=2). Altitude range: ~900 m (n=1). Host plants: Unknown. Distribution: CHINA: Fujian.

Agrilus shashamboe Kurosawa Fig. 12B (habitus); Fig. 33L (aedeagus); Fig. 52T (type) shashamboe Kurosawa, 1963 (Agrilus)

Agrilus she Jendek, sp. nov. Fig. 26A (habitus of HT)

Description: 6.3-7.2 mm (Holotype 6.3 mm). BODY: Shape: cuneiform, Build: robust, Color (dorsally): unicolored, Color (dominant): black or green. HEAD: Medial impression: present, Impression (extent): vertex, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (density): dense, Sculpture (intensity): superficial. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short, Sexual modification in male: not apparent. PRONOTUM: Shape: visually elongate, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate or obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: acute or rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): deep. Prehumerus: carinal, Shape: arcuate, Extent: beyond half of pronotal length, Anterior end: joining pronotal marginal carina, Arc: obvious. Marginal

183

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Shape: arcuate. Elytral pubescence: sutural wedge or proximal and distal, Proximal: quadrangular, Distal: perisutural stripe, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: arcuate or arcuately emarginate, Emargination (deepness): shallow. Prosternal process: moderate, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: laterally emarginate. Sternal groove: Shape on apex of last ventrite: angulately sinuate, Sinuosity (deepness): very shallow, Sinuosity (width): markedly wide. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus lost. Diagnosis: antennae short; postero-medial impression on pronotum deep; elytra anteriorly with white pubescence, in posterior half reduced into perisutural pubescent strip; male without sexual modifications on ventrite 1. Species group: Tempestivus. Relevant species: A. tempestivus Lewis, 1893. Type locality: China, west Jiangxi province, Jinggang Shan, Ciping. Type specimens: Primary: Holotype (Fig. 26A), ♂, (EJCB): “China, Jiangxi W, Jinggang Shan, Ciping, 2-14. VI.1994, E. Jendek & O. Šauša leg.” [Remark: Aedeagus is lost]. Secondary: CHINA: Fujian: 1 ♀ PT (EJCB): “China, Fujian prov., Shaowu env., 23-27.vi.1991, leg. Nikodým”. Adult occurence: 6 (n=2).

141 [#6850] (not conspecific with hokkaidoensis). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2291 [#8576] (world catalog). hokkaidoensis Tôyama, 1985 (Agrilus) TÔYAMA, 1985: 45 47 [#2924] (description). TÔYAMA, 1985: 32 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 34 [#4091] (checklist; Japan: Hokkaido). JENDEK, 2003: 186 [#5823] (synonym of shibatai). FUKUTOMI & HORI, 2004: 12 [#6851] (checklist; Japan (Hokkaido)). OHMOMO, 2004: 141 [#6850] (cited as valid species; not conspecific with shibatai; faunal records; Hokkaido). JENDEK, 2006: 60, 401 [#7441] (synonym of shibatai; Palaearctic catalog). BELLAMY, 2008: 2291 [#8576] (synonym of shibatai; world catalog).

Type material: Agrilus hornianus shibatai Kurosawa, 1964 Type locality: Hatsuno, Amami-Oshima. Type specimens: Primary: Holotype (Fig. 52U), ♂, (NSMT): “Hatsuno, AMAMI IS. [p] 25. V. [h] 19 [p] 60 [h] T. Shibata [p] \ Holotype [p] Agrilus hornianus shibatai Y. Kurosawa, 1964 [h] [red label with black border]”. Described from 8 specimens (HT, AT, PT). Agrilus hokkaidoensis Tôyama, 1985 Type locality: Horoka, Hokkaido. Type specimens: Primary: Holotype (Fig. 47C), ♂, (NSMT): “Horoka [h] Hokkaido [p] 25. VII. [h] 19 [p] 69 [h] H. Kawai [p] [scratched] \ Holotype [p] Agrilus hokkaidoensis Tôyama, 1985 [h] [red label]”. Described from 1 specimen (HT). Diagnosis: 6.8-8.2 mm; prehumerus carinal; anterior pronotal lobe obvious, extending beyond anterior pronotal angles; aedeagus subparallel. Species group: Tempestivus. Relevant species: A. esakii Kurosawa, 1964. Specimens examined: JAPAN: Ryukyu isl. (Okinawa incl.): 1 (EJCB): “Hatsuno, Amami Oshima, leg. J. Nagao”.

Adult occurence: 5-6-7-8 (n=5). Faunal records published by KUROSAWA, 1964 [#1619] are also included.

Host plants: Unknown.

Host plants: Castanopsis lutchuensis: AKIYAMA & OHMOMO, 1997 [#4091].

Distribution: CHINA: Fujian, Jiangxi.

Distribution: JAPAN: Hokkaido, Ryukyu Islands.

Etymology: Noun in apposition; the specific name refers to the Chinese ethnic group She living in provinces Fujian, Zhejiang, Anhui, Jiangxi and Guangdong.

Agrilus sibiricus fukushimenis Jendek

Agrilus shibatai Kurosawa Fig. 25I (habitus); Fig. 39B (aedeagus); Figs. 47C, 52U (types) shibatai Kurosawa, 1964 (Agrilus; subspecies of hornianus) KUROSAWA, 1964: 323 [#1619] (description). KUROSAWA, 1974: 2 [#4576] (subspecies of hornianus). AKIYAMA, 1975: 4 [#3178] (subspecies of hornianus; faunal records; Ryûkyû Archipelago). TÔYAMA, 1985: 26 [#3678] (subspecies of hornianus; iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (subspecies of hornianus; checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (subspecies of hornianus; checklist; Japan). AKIYAMA & OHMOMO, 1997: 4, 38 [#4091] (checklist; Japan: Ryukyus (Amami oshima I., Okinawa I., Ishigaki jima I., Iriomote jima I., Yonaguni jima I.)). JENDEK, 2003: 186 [#5823] (synonymy). OHMOMO, 2004:

Fig. 18G (habitus of HT); Fig. 35W (aedeagus) fukushimenis Jendek, 1994 (Agrilus; subspecies of sibiricus) JENDEK, 1994: 12 13 [#1355] (description). AKIYAMA & AKIYAMA, 1996: 185 [#3962] (subspecies of sibiricus; faunal record; Japan: Hiroshima Prefecture). AKIYAMA & OHMOMO, 1997: 39 [#4091] (subspecies of sibiricus; checklist; Japan: Hokkaido, Honshu, Shikoku, Kyushu, Tsushima). FUKUTOMI & HORI, 2004: 9 [#6851] (subspecies of sibiricus; checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 394 [#7441] (subgenus Robertius; subspecies of sibiricus; Palaearctic catalog). BELLAMY, 2008: 2292 [#8576] (subspecies of sibiricus; world catalog).

Type material: Agrilus sibiricus fukushimenis Jendek, 1994 Type locality: Eda, Fukushima Prefecture, Honshu, Japan.

184 Type specimens: Primary: Holotype (Fig. 18G), ♂, (EJCB): “Eda Fukushima Pref. 19.V.1981 S. Ohmomo leg.”. Secondary: 3 PT (MNCA); 6 PT (NMPC); 5 PT (EJCB). Described from 15 specimens (HT, PT). Diagnosis: 4.6-7 mm; elytra regularly attenuate to apices; apices widely and separately arcuate or subtruncate. Species group: Betuleti. Relevant species: A. sibiricus Obenberger. Specimens examined: JAPAN: Hokkaido: 1 PT (NMPC): “collec tor H. Nishijima \ Mt. Moswa Joussankei Hokkaido 20.aug.1937” (JENDEK, 1994 [#1355]). Honshu: 1 PT (NMPC): “Daisen, Japan 27.vi.1944 coll. Y. Kuroda” (JENDEK, 1994 [#1355]); 3 PT (NMPC): “Yunohana S. Aizu Japan 20.VII.[19]47” (JENDEK, 1994 [#1355]); 1 PT (EJCB): “Yunohana S. Aizu Japan 8.VIII.[19]49” (JENDEK, 1994 [#1355]); 1 PT (MNCA): “(Hinoemata) Fukushima Pref. JAPAN 25.VII.1977 A. Nishiyama leg.” (JENDEK, 1994 [#1355]); 2 PT (MNCA): “Akanerindou, Haronomachi City, Fukushima Pref. 18.VI.1979 leg. Nishiyama” (JENDEK, 1994 [#1355]); 1 PT (EJCB): “Eda Fukushima Pref. 21.VI.1979 S. Ohmomo leg.” (JENDEK, 1994 [#1355]); 3 PT (EJCB): “Eda Fukushima Pref. 19.V.1981 S. Ohmomo leg” (JENDEK, 1994 [#1355]); 1 PT (EJCB): “Mts. Daibusatsu, Yamanashi Pref. 21.VI.1985 S. Ohmomo” (JENDEK, 1994 [#1355]); 1 ♀ (EJCB): “Japan, Fukushima, Namie town, 26.vi.1995, N. Sugiura leg.”. Kyushu: 1 ♀ (JMCZ): “Japan, Kyushu, Oita pref., Mt. Kurodake, 1.vi.1986”. RUSSIA: Sakhalinskaya oblast’: 1 ♀ (MNHE): “Russia, 27.07.1989, Kunashiri Island, 13 km, leg. U. Roosileht”; 1 ♀ (IMCM): “Kunashir, Tret’yakovo, 43°59’17’’N, 143°39’05’’E, 19 25.viii.2008 leg. I. Mel’nik, na Acer mayrii Schwer. ( A. pictum auct) [in Russian]”.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA [#4091] (synonym of sibiricus). NONNAIZAB et al., 1999: 112 [#8247] (checklist; China: Inner Mongolia). JENDEK, 2006: 394 [#7441] (synonym of sibiricus sibiricus; Palaearctic catalog). BELLAMY, 2008: 2291 2292 [#8576] (synonym of sibiricus; world catalog). turdus Théry, 1942 (Agrilus) OBENBERGER, THÉRY, 1942: 255 256 [#2875] (description). 1943: 9 [#2333] (synonym of insuspectus). JENDEK, 1994: 11, 12 [#1355] (synonym of sibiricus). AKIYAMA & OHMOMO, 1997: 38 [#4091] (synonym of sibiricus). JENDEK, 2006: 394 [#7441] (synonym of sibiricus sibiricus; Palaearctic catalog). BELLAMY, 2008: 2292 [#8576] (synonym of sibiricus; world catalog).

Host plants: Acer: AKIYAMA & OHMOMO, 1997 [#4091].

Type material: Agrilus sibiricus Obenberger, 1912 Type locality: Sibiria. Type specimens: Holotype by monotypy (Fig. 52W), ♂, (NMPC): “Sibiria [h] \ Typus [p] [red label] \ Agrilus sibiricus m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24795 [h] [orange label]”. Described from 1 specimen. Agrilus insuspectus Obenberger, 1924 Type locality: Vladivostok (Ussuri mer.). Type specimens: Primary: Lectotype (Fig. 47L), ♂, designated by JENDEK (1994: 11-12) [#1355], (NMPC): “Vladivostok Ussuri mer. Dr. Jureček 1919 [p] \ Typus [p] [red label] \ Mus. Nat. Pragae Inv. [p] 24256 [h] [orange label] \ Agrilus insuspectus m.Type [h] Det. Dr. Obenberger [p] \”. Described from unknown number of syntypes. Agrilus turdus Théry, 1942 Type locality: Mandchourie, Weischadre. Type specimens: Not located. Described from 6 syntypes.

Distribution: JAPAN: Hokkaido, Honshu, Kyushu, Shikoku, Tsushima. RUSSIA: Sakhalinskaya oblast’ (Kunashir Island).

Diagnosis: 5.5-8.2 mm; elytra distinctly expanded just before apex; apices subtruncate. Species group: Betuleti. Relevant species: A. sibiricus fukushimenis Jendek, 1994.

Adult occurence: 5-6-7-8 (n=13).

Agrilus sibiricus sibiricus Obenberger Fig. 18F (habitus); Fig. 35V (aedeagus); Figs. 47L, 52W (types) sibiricus Obenberger, 1912 (Agrilus) OBENBERGER, OBENBERGER, 1912: 70 [#1957] (description). 1924: 47 [#2040] (notes). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1041 [#2191] (world catalog). ALEXEEV, 1989: 485 [#64] (key to species; Primorsk; North Korea; northeastern China). JENDEK, 1994: 11 12 [#1355] (lectotype designation; synonymy; characters; Russian Far East). AKIYAMA & OHMOMO, 1997: 38 [#4091] (checklist; Korea; Russia). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). JENDEK, 2006: 394 [#7441] (subgenus Robertius; Palaearctic catalog). ANONYMOUS, 2007: 32 [#8250] (checklist; Japan: Honshu). BELLAMY, 2008: 2291 [#8576] (subgenus Robertius; world catalog). VOLKOVITSH, 2009: 136 [#8850] (subgenus Robertius; faunal records; Russia: Primorskiy kray). insuspectus Obenberger, 1924 (Agrilus) OBENBERGER, OBENBERGER, 1924: 43 [#2040] (description). 1926: 656 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1001 [#2191] (world catalog). KUROSAWA, 1963: 153 [#1618] (characters; Japan). KUROSAWA, 1974: 2 [#4576] (note). TÔYAMA, 1985: 30 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). LI JINGKE, 1992: 93 [#6755] (checklist; northeastern China). JENDEK, 1994: 11 12 [#1355] (synonym of sibiricus; lectotype designation). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 38

Specimens examined: CHINA: Heilongjiang: 1 (NMPC): “Mandjourie” (JENDEK, 1994 [#1355]). RUSSIA: Primorskiy kray: 1 (NMPC): “Primorje” (JENDEK, 1994 [#1355]); 1 (ZIN): “Vladivostok; Prim. kraj 11.VI.1968” (JENDEK, 1994 [#1355]); 2 (EJCB): “Russia or., 18.6.1978, Primor., Artyom, Plutenko leg.”; 1 (MNHE): “Russia, 01.07.1989, Primorye Distr., Limonniki, leg. U. Roosileht”; 1 ♀ (EJCB): “USSR, Ussuri prov. Arsenev env., VII.1990, leg. B. Šiška & R. Čermák” (JENDEK, 1994 [#1355]); 1 ♂ (EJCB): “USSR, Primorsk. kraj, Arsenev env., VI.1991, 44.11N, 133.19E, O. Šausa leg.” (JENDEK, 1994 [#1355]); 1 (MNCA): “Russia (Primorye) env. Vladivostok 14.VII.1991 A. Schröder [leg.]”; 1 ♀ (EJCB): “Russia, Primorsk. prov., Arsenev env., VII.1992, B. Šiska leg.” (JENDEK, 1994 [#1355]); 1 (EJCB): “Sib. or., Primorje, Komarovka flum., Kamenushka env., 300 m, Vořišek lgt., V.1992” (JENDEK, 1994 [#1355]); 1 ♀ (EJCB): “RUSSIA or. Primorsk.Reg. Artyom env. 5.VI. 21.VII.1992 Plutenko leg.;” (JENDEK, 1994 [#1355]); 1 (MNCA): “Russia: Primorskij Kray, Vladivostok Sedanka 100 m, 20.VI.1993, 43.09N, 131.53E, Leg. L. Zerche et al.”; 1 (EJCB): “Russia, Primorski kray, Ternejski distr., 12 19.viii., Sikhote Alin res., riv. Serebrianka env., 1995”; 2 (EJCB): “Russia, Primorski kray, Khankajski distr. 1995, lake Khanka, 17 19.vii., Kamen’ Rybolov”; 1 (EJCB): “Russia, Primorski kray, Chassanski district, Res. Kedrovja Padj, 22 27. vi.1995”; 4 (EJCB): “Russia, Primorski kray, Partizanski distr., 1995, Partizanski ridge, riv. Frolovka, 19 25.vii”; 1 ♀ (EJCB): “Russia E, Primorje reg, Anisimovka, 7 13.vii.1996, Mt. Litovka, 200 m, Plutenko leg.”; 13 (EJCB): “Russia, Primorskyi Kray, Artem, VI.1998, Plutenko leg.”; 1 (EJCB): “Russia Far East, 20

185

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA km SE Ussuriysk, Gornotaezhnoje vill. env., 29.06.2003, Lg. A. Napolov”.

Adult occurence: 5-6-7-8 (n=17). Altitude range: 100-300 m (n=3).

with narrow medial row of erect setae. Species group: Imitans. Relevant species: A. blatteicollis Bourgoin, 1922; A. nepalensis Tôyama, 1988.

Host plants: Unknown.

Type locality: China, Su-Tchuen [= Sichuan province] Chasseurs Indigènes.

Distribution: CHINA: Heilongjiang, Nei Mongol. KOREA NORTH. KOREA SOUTH. RUSSIA: Primorskiy kray.

Type specimens: Primary: Holotype (Fig. 12J), ♂, (EJCB): “Su-Tchuen Chasseurs Indigènes 1903 [p]”. Adult occurence: Unknown.

Agrilus sichuanus Jendek, sp. nov.

Host plants: Unknown.

Fig. 12J (habitus of HT); Fig. 33S (aedeagus)

Distribution: CHINA: Sichuan.

Description: 10-10.1mm (Holotype 10.1 mm). BODY: Shape: cuneiform, Build: slender, Color (dorsally): bicolored, Color (dominant): green or brown. HEAD: Medial impression: absent, Epistoma: with raised upper margin. Frons: Shape: flat, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): irregular, Sculpture (density): sparse, Sculpture (intensity): superficial. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square, Sides: arcuate, Maximal width: at anterior margin or at middle, Anterior margin: wider than posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles or at level with anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp or blunt. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): entire. Prehumerus: filamentary, rarely carinal, Shape: bisinuate, Extent: to anterior pronotal angle, Posterior end: joining posterior pronotal angle, Anterior end: joining anterior pronotal angle, Arc: weak. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior angles or at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Shape: arcuate. Elytral pubescence: perisutural stripes, Color: unicolored. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with medial row of setae. Prosternal lobe: large, Anterior margin: arcuately emarginate, Emargination (deepness): shallow, Emargination (width): narrow. Prosternal process: moderate, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Male sexual modifications: metacoxa medially with cluster of setae and posterior margin of metacoxa with dent. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: elongate.

Etymology: The specific name refers to the Sichuan province of China, the type locality of this species.

Diagnosis: elytra greenish black, with wide perisutural pubescent stripe reaching to sutural margin; prosternum

Agrilus sikkimensis Obenberger Fig. 13C (habitus); Fig. 33X (aedeagus); Fig. 52X (type) sikkimensis Obenberger, 1928 (Agrilus) OBENBERGER, 1928: 69 70 [#2075] (description). OBENBERGER, 1936: 1102 [#2191] (world catalog). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2292 [#8576] (world catalog).

Type material: Agrilus sikkimensis Obenberger, 1928 Type locality: Sikkim. Type specimens: Primary: Lectotype (Fig. 52X) by present designation, ♂, (NMPC): “Sikkim [h] \ Typus [p] [red label] \ Agrilus sikkimensis m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24912 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 4.5-7 mm; antennae short; frons distinctly convex; elytra black with silky tinge; elytra with wide perisutural stripes or entirely pubescent, elytral apices entirely pubescent; aedeagus subparallel. Species group: Imitans. Relevant species: A. rivalieri Descarpentries & Villiers, 1963; A. smaragdifrons Ganglbauer, 1889. Specimens examined: CHINA: Yunnan: 1 ♀ (EJCB): “China, Yunnan, 22.V. 2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”. INDIA: Meghalaya: 1 ♀ (EJCB): “NE India, Meghalaya state, West Garo Hills, Nokrek Nat.Park, 9 17.V.1996 alt.1100±150m, GPS N25°29.6’, E90°19.5’ (WGS 84), E. Jendek & O. Šauša leg.”; 1 ♀ (EJCB): “NE India, Meghalaya state, West Garo Hills reg., Tura, 5 7.V.1996, alt. 700±100 m, GPS N25°30.7’ E90°13.9’ (WGS 84), E.Jendek & O.Šauša leg.”; 1 ♀ (EJCB): “NE India, Meghalaya, 1999, 3 km E of Tura, 1150 m, 25°30’N, 90°14’E, 6 12.v., Tryzna & Benda leg., 2002”. LAOS: Champasak: 1 ♂ (EJCB): “Laos south., Champasak prov., Bolaven Plateau, 10 18.IV.1999, Route (No. 23) Pakse Paksong, Ban Itou env. (km.35), alt. 800m, N 15º10.4’, E 106º05.8’ (GPS), E. Jendek & O. Šauša leg.”. Louang Namtha: 1 ♀ (EJCB): “Laos north, 5 11.V.1997, 20 km NW Louang Namtha, N 21°09.2, E 101°18.7, alt. 900±100 m, E. Jendek & O. Šauša leg.”. Louangphrabang: 3 ♀ (EJCB): “Laos N, 23.iv.1999, Louangphrabang prov. 20°42’N 102°54’E, 25 km E Muang Ngoy, 1000 m, Vít Kubáň leg.”. NEPAL: 2 ♂, 2 ♀ (EJCB): “E Nepal, Dhankuta, Arun Valley, Lamobagar Gola, 27.5. 3.6.1980, 1000 1400 m, leg. C. Holzschuh”; 1 ♂ (EJCB): “E Nepal, Arun valley, Num Arun river Ruhruma, 1600 900 1200m, 13.6.1992, lgt. J. & J.

186 Probst”; 1 ♂, 1 ♀ (EJCB): “W Nepal, NW Pokhara, Khali Gandaki Khola, 1100 1300 m, Tatopani, 12 14.V.1994, leg. C. Holzschuh”. THAILAND: Chiang Mai: 1 ♂ (EJCB): “Thai, Chiang Mai prov. 18°49’N, 98°54’E, 1600 m, Doi Pui mt., 2 6.V.1996, Vít Kubáň leg.”. Mae Hong Son: 1 ♂ (EJCB): “NW Thailand, 25.4. 5.5.1992, Soppong Pai pass, P. Pacholátko leg.”; 7 (EJCB): “Thailand, 1 8.5.1993, Soppong Pai, 1800 m, Pacholátko & Dembický leg.”; 1 ♂ (EJCB): “NW Thailand, Mae Hong Son pr., 1500m, Soppong, 9 12.V., Sv. Bílý leg., 1996”.

Adult occurence: 4-5-6 (n=14). Altitude range: 600-1800 m (n=12). Host plants: Unknown. Distribution: CHINA: Yunnan. INDIA. LAOS. NEPAL. THAILAND.

Agrilus sinensis sinensis Thomson Fig. 8I (habitus); Fig. 52Y (type) sinensis Thomson, 1879 (Agrilus) THOMSON, 1879: 73 [#2909] (description). KERREMANS, 1892: 272 [#1397] (catalog). KERREMANS, 1903: 278 [#1441] (cited as chinensis; catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). BODEMEYER, 1915: 447 [#8261] (cited as chinensis; faunal record; China: Heilongjiang). OBENBERGER, 1924: 36, 37 [#2040] (note). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1934: 14 [#2146] (note). OBENBERGER, 1935: 163 [#2158] (note). OBENBERGER, 1936: 1041 [#2191] (world catalog). OBENBERGER, 1958: 237 [#2389] (faunistic redords; China). DESCARPENTRIES & VILLIERS, 1963: 8 [#701] (notes). COBOS, 1972: 56 [#591] (subgenus Agrilus; checklist; Mongolia). KUROSAWA, 1975: 1 [#4578] (note). HOU, 1987: 590 [#3490] (notes; Yunnan). PENG ZHONGLIANG, 1987: 358 [#2429] (checklist; China). PENG ZHONGLIANG, 1992: 398 399 [#2431] (characters; notes; Hunan). JENDEK, 1995: 144, 145 [#3768] (lectotype designation; characters; central and southeastern China). AKIYAMA & OHMOMO, 1997: 39 [#4091] (checklist; Japan: Honshu; China; Indochina). ALEXEEV, 1998: 379 [#4322] (subgenus Sinagrilus; genotype of the subgenus Sinagrilus). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Inner Mongolia, Gansu, Hubei, Jiangsu, Jiangxi, Fujian, Hainan, Hunan, Guangxi, Guizhou, Sichuan, Yunnan, Xizang). LIN GUI RUI, 2002: 234 [#7926] (checklist; pest). PENG ZHONGLIANG, 2002: 267 [#6628] (characters; Fujian). JENDEK, 2006: 394 [#7441] (subgenus Sinagrilus; Palaearctic catalog). BELLAMY, 2008: 2294 [#8576] (subgenus Sinagrilus; world catalog). obscuripennis Fairmaire, 1888 (Agrilus; [preoccupied]) FAIRMAIRE, 1888: 120 [#887] (description). KERREMANS, 1892: 256 [#1397] (synonym of fairmairei; catalog). KERREMANS, 1903: 285 [#1441] (synonym of fairmairei). JAKOBSON, 1913: 798 [#4386] (synonym of fairmairei; catalog). OBENBERGER, 1936: 990 [#2191] (synonym of fairmairei). JENDEK, 2006: 398 [#7441] (synonym of fairmairei; Palaearctic catalog). BELLAMY, 2008: 2089 [#8576] (synonym of fairmairei; world catalog). fairmairei Kerremans, 1892 (Agrilus; replacement name for obscuripennis Fairmaire not Gory) syn. nov. KERREMANS, 1892: 256 [#1397] (replacement name proposal). KERREMANS, 1903: 285 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 654, 659 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 990 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 355 [#2429] (checklist; China). HUA LI ZHONG, 2002: 89 [#6696] (checklist;

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA China: Beijing). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2089 [#8576] (world catalog). pseudimitans Miwa & Chûjô, 1940 (Agrilus) MIWA & CHÛJÔ, 1940: 54 [in Japanese], 60 61, 73 [#1810] (description). KUROSAWA, 1947: 2 [#3339] (erroneously cited as the variety of cyaneoniger; lapsus; see Akiyama & Ohmomo, 1997: 31, 39 [#4091]). KUROSAWA, 1975: 1 [#4578] (synonym of sinensis). AKIYAMA & OHMOMO, 1997: 39 [#4091] (synonym of sinensis Saunders). JENDEK, 2006: 394 [#7441] (synonym of sinensis sinensis; Palaearctic catalog). BELLAMY, 2008: 2294 [#8576] (synonym of sinensis; world catalog).

Type material: Agrilus sinensis Thomson, 1879 Type locality: Shang-Haï. Type specimens: Primary: Lectotype (Fig. 52Y), ♀, designated by JENDEK (1995: 144, 145) [#3768], (MNHN): “Schang-Hai [h][yellow label] \ Th. Type [p] [white label with black border] \ Sinensis Laf. M. SS., Th. Type Ap. I, 73 [h] \ Ex Musaeo JAMES THOMSON [p] \”. Described from unknown number of syntypes. Agrilus obscuripennis Fairmaire, 1888 Type locality: Pekin. Type specimens: Not located. Not in MNHN. Described from unknown number of syntypes. Agrilus fairmairei Kerremans, 1892 Type locality and specimens: See: Agrilus obscuripennis Fairmaire, 1888. Agrilus pseudimitans Miwa & Chûjô, 1940 Type locality: Nikkô, Japan. Type specimens: Not located. Described from 1 specimen. Diagnosis: 8-13 mm; head with obvious, deep medial impession; anterior pronotal lobe moderate; prehumerus obsoletely filamentary. Species group: Sinensis. Relevant species: A. coreanus Obenberger, 1935. Specimens examined: CHINA: Fujian: 1 (EJCB): “China, Fujian prov., Shaowu env., 23 27.vi.1991, leg. Nikodým” (JENDEK, 1995 [#3768]). Guizhou: 1 (MNHN): “Kouy Tscheou [ Guizhou province] Abbé Largeteau” (JENDEK, 1995 [#3768]). Hubei: 7 (EJCB): “China, SE Hubei, 7.+17.vi, 20 km NW Tongshan, 29.7N 114.3E, Jaroslav Turna leg. 2004”. Hunan: 1 ♀ (EJCB): “China, N Hunan, 4 7.vii., Wuling Shan, 29.4N, 110.4E, Zhangjiajie, 700 m, Jaroslav Turna leg. 2003”. Jiangxi: 1 (EJCB): “Kiang Si” (JENDEK, 1995 [#3768]); 1 (EJCB): “CHINA: JIANGXI: Suichuan env., 26.27N, 114.31E, 1.VI.1994, leg. E. Jendek & O. Šauša” (JENDEK, 1995 [#3768]). Sichuan: 1 (EJCB): “China, Sichuan, Emei Shan, Emei, 120 km SW of Chengdu, 12 16.VI.1993, leg. Beneš” (JENDEK, 1995 [#3768]); 1 (EJCB): “CHINA: SICHUAN: Emei Shan, Emei Baoguosi, 130 km SW of Chengdu, 29.VII. 3.VIII.1993, leg. Beneš” (JENDEK, 1995 [#3768]); 1 (EJCB): “CHINA: SICHUAN: Jiulonggou env., near Dayi, 70 km W of Chengdu, 23 27.VI.1993, leg. M. Tryzna” (JENDEK, 1995 [#3768]).

Adult occurence: 6-7-8 (n=7). Altitude range: 700-1000 m (n=2). Host plants: Unknown. Distribution: CHINA: Beijing, Fujian, Guizhou, Heilongjiang, Hunan, Jiangxi, Jiangsu, Shanghai, Sichuan, Xizang, Yunnan. JAPAN: Honshu. LAOS.

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Remarks: The conspecifity of A. sinensis sinensis and A. obscuripennis is judged from the description of A. obscuripennis (size, color, character of pronotum and elytra, type locality).

Agrilus sinensis splendidicollis Fairmaire Fig. 8J (habitus); Fig. 32F (aedeagus); Figs. 43L, 48L, 53F (types) splendidicollis Fairmaire, 1889 (Agrilus) FAIRMAIRE, 1889: 348 [#890] (description). KERREMANS, 1892: 273 [#1397] (catalog). KERREMANS, 1900: 341 [#1434] (synonym of sinensis). KERREMANS, 1903: 278 [#1441] (synonym of sinensis cited as chinensis). JAKOBSON, 1913: 798 [#4386] (synonym of sinensis; catalog). OBENBERGER, 1924: 36, 37 [#2040] (subspecies of sinensis). OBENBERGER, 1926: 654 [#2057] (subspecies of sinensis; Palaearctic catalog). OBENBERGER, 1936: 1041 [#2191] (cited as splendicollis; subspecies of sinensis; world catalog). BAUDON, 1963: 54 [#148] (subspecies of sinensis; misspelled as “chinensis s. sp. auricollis”; faunal record; Laos). DESCARPENTRIES & VILLIERS, 1963: 2, 8 [#701] (subspecies of sinensis; characters; faunal records; Tonkin; Laos; Bhoutan). BAUDON, 1968: 106, 110 [#162] (subspecies of sinensis). KUROSAWA, 1975: 1 [#4578] (subspecies of sinensis). HOU, 1987: 591 [#3490] (notes; Yunnan). PENG ZHONGLIANG, 1987: 358 [#2429] (subspecies of sinensis; cited as splendicollis; checklist; China). HOU, 1988: 279 [#4900] (checklist; China: Xizang). JENDEK, 1995: 145 [#3768] (subspecies of sinensis; lectotype designation; synonymy; characters; China: Yunnan; Vietnam; Laos; Thailand; West Bengal). HUA LI ZHONG, 2002: 90 [#6696] (cited twice, as a subspecies of sinensis and as a species; checklist; China: Jiangsu, Sichuan, Yunnan [as a subspecies of sinensis]; China: Shandong, Jiangsu, Sichuan, Yunnan, Xizang [as species]). JENDEK, 2006: 394 [#7441] (subgenus Sinagrilus; subspecies of sinensis; Palaearctic catalog). BELLAMY, 2008: 2294 [#8576] (subspecies of sinensis; world catalog). lameyi Obenberger, 1924 (Agrilus; subspecies of sinensis) OBENBERGER, 1924: 36 37 [#2040] (description). OBENBERGER, 1926: 654 [#2057] (subspecies of sinensis; Palaearctic catalog). OBENBERGER, 1936: 1041 [#2191] (subspecies of sinensis; world catalog). KUROSAWA, 1975: 1 [#4578] (subspecies of sinensis). PENG ZHONGLIANG, 1987: 358 [#2429] (subspecies of sinensis; checklist; China). JENDEK, 1995: 145 [#3768] (synonym of sinensis splendidicollis; lectotype designation). HUA LI ZHONG, 2002: 90 [#6696] (subspecies of sinensis; checklist; C China). JENDEK, 2006: 394 [#7441] (synonym of sinensis splendidicollis; Palaearctic catalog). BELLAMY, 2008: 2294 [#8576] (synonym of sinensis splendidicollis; world catalog). bayeri Obenberger, 1935 (Agrilus) OBENBERGER, 1935: 161 162 [#2158] (description). OBENBERGER, 1936: 959 [#2191] (world catalog). OBENBERGER, 1958: 236 [#2389] (cited as beyeri; faunistic redords; China). HOU, 1987: 591 [#3490] (notes; Yunnan). PENG ZHONGLIANG, 1987: 354 [#2429] (cited as beyeri; checklist; China). JENDEK, 1995: 145 [#3768] (synonym of sinensis splendidicollis; lectotype designation). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Yunnan). JENDEK, 2006: 394 [#7441] (synonym of sinensis splendidicollis; Palaearctic catalog). BELLAMY, 2008: 2294 [#8576] (cited as beyeri; synonym of sinensis splendidicollis; world catalog).

Type material: Agrilus sinensis splendidicollis Fairmaire, 1889 Type locality: Tonkin. Type specimens: Primary: Lectotype (Fig. 53F), ♂, designated by JENDEK (1995: 145) [#3768], (MNHN):

187 “Agrilus splendidicollis Fairm Tonkin [by Fairmaire hand] \ Museum Paris Collection Léon Fairmaire 1906 [p]”. Secondary: 2 PLT (BMNH). Described from unknown number of syntypes. Agrilus sinensis lameyi Obenberger, 1924 Type locality: Na-lang. (Tonkin nord.). Type specimens: Primary: Lectotype (Fig. 48L), ♂, designated by JENDEK (1995: 145) [#3768], (NMPC): “Na - lang. Tonkin Nord. Lamey [h] \ Typus [p] [red label] \ A. sinensis lamei m. Type [h] Det. Dr Obenberger [p] \”. Described from unknown number of syntypes. Agrilus bayeri Obenberger, 1935 Type locality: China, prov. Yunnan, vallis flum. Soling-ho. Type specimens: Primary: Lectotype (Fig. 43L), ♂, designated by JENDEK (1995: 145) [#3768], (NMPC): “China, Prov. Yunnan Vallis flumm. Soling - ho [p] \ Typus [p] [red label] \ Mus. Nat. Pragae Inv. [p] 24293 [h] [orange label] \ Agrilus Bayeri m. Type [h] Det. Dr Obenberger [p]”. Secondary: 1 PLT (NMPC). Described from unknown number of syntypes. Diagnosis: 8.5-12 mm; head and pronotum dark reddishcupreous; elytra black. Species group: Sinensis. Relevant species: A. sinensis sinensis Thomson, 1879. Specimens examined: CHINA: Sichuan: 1 (MNCA): “China Sichuan: Emei Boagou, 700 m, 18.6.[19]96 D. Erber leg.”. Yunnan: 1 (MNHN): “Chine, Yunnan, recu de Lou nan, 1931” (JENDEK, 1995 [#3768]); 1 (EJCB): “S. China, Jinghong, 10 14.7.1990, pr. Yunnan, lgt. S. Bečvář” (JENDEK, 1995 [#3768]); 1 (EJCB): “Yunnan, 1500 2800 m, 26.07N, 103.14E, Dongchuan, 28.6. 3.7., Vít Kubáň leg., 1994” (JENDEK, 1995 [#3768]). INDIA: West Bengal: 1 (MNHN): “British Bootang Maria Basti” (JENDEK, 1995 [#3768]). LAOS: Phongsali: 4 (EJCB): “Lao, Phongsaly prov. 21°41’N, 102°06’E Phongsaly env. 6 17.v.2004, 1500 m, P. Pacholátko leg.”. Xiangkhoang: 1 ♀ (EJCB): “Laos NE, Xieng Khouang prov., 45 km (by road) E of Phonsavan: ~1000 m, Ban Namseung, April 2008, Ch. Keomaravong leg.”. THAILAND: 1 (EJCB): “NW THAI, Nan District, Pha Khap, 15 20.V.1992, leg. P. Pacholátko” (JENDEK, 1995 [#3768]). 1 (GNCW): “Thai[land], 11 15.V.1993, Nan Pha Khab, leg. Dembický + Pachol.[átko]”. VIETNAM: 1 (EJCB): “Vietnam: Tonkin: Cucphuong Natural Reservation, pr. Hasonbinh, 15.VI.1985, leg. Vít Kubáň” (JENDEK, 1995 [#3768]); 1 (EJCB): “Vietnam, Cuc phuong, 24 25.V.1986, Han nam ninh prov., Jan Horák lgt.” (JENDEK, 1995 [#3768]). 1 (MNHN): “Tonkin Dong Van Cap Gadel 1896” (JENDEK, 1995 [#3768]); 1 (MNHN): “Tonkin, Ma cham Recu 26 III 1907” (JENDEK, 1995 [#3768]); 1 (EJCB): “VIETNAM: TONKIN: Hanoj, 10.V. 1989, leg. P. Pacholátko” (JENDEK, 1995 [#3768]); 1 (EJCB): “VIETNAM: TONKIN: Hanoi, V. 1990, leg. O. Šauša” (JENDEK, 1995 [#3768]); 1 (EJCB): “VIETNAM: TONKIN: SaPa, 12 17.V. 1990, leg. M. Kalabza & Z. Košťál” (JENDEK, 1995 [#3768]); 1 (EJCB): “SaPa 11 18.VI., N. Vietnam, A. Olexa [leg.] 1990” (JENDEK, 1995 [#3768]); 1 (EJCB): “Vietnam N, TamDao 900 m, 16 23.V.1991, Strnad Jan lgt.” (JENDEK, 1995 [#3768]); 1 (EJCB): “Vietnam, Chua Huong Huong Son, 60 km SW from Hanoi, 26 29.IV.1991, leg. E. Jendek” (JENDEK, 1995 [#3768]).

Adult occurence: 4-5-6-7 (n=12). Faunal records published by OBENBERGER, 1934 [#2146] and 1958 [#2389] (as beyeri) are also included. Altitude range: 700-2800 m (n=6). Faunal record published by OBENBERGER, 1958 [#2389] (as beyeri) is also included.

188 Host plants: Unknown. Distribution: CHINA: Sichuan, Xizang, Yunnan. INDIA: West Bengal. LAOS. THAILAND. VIETNAM.

Agrilus sinuatus sachalinensis Obenberger stat. nov. Fig. 14C (habitus); Fig. 34H (aedeagus); Figs. 52K, 54X (types) sachalinensis Obenberger, 1935 (Agrilus) OBENBERGER, 1935: 165 [#2158] (description). MIWA & CHÛJÔ, 1936: 20 [#1809] (catalog; Japan). OBENBERGER, 1936: 1039 [#2191] (world catalog). KUROSAWA, 1949: 14 [#5028] (characters). CHÛJÔ & KUROSAWA, 1950: 9 [#501] (catalog; faunal records; Japan (Shikoku)). IGA, 1955: 79 [#3828] (iconography; Japan). IGA, 1962: 79 [#3351] (iconography; Japan). KUROSAWA, 1974: 3 [#4577] (subspecies of mendax). KUROSAWA, 1975: 2 [#4578] (note). TÔYAMA, 1985: 21 [#2924] (subspecies of mendax; faunal records; Japan). ALEXEEV, 1989: 486 [#64] (key to species; Yu Sakh). HIRASHIMA, 1989: 323 [#8078] (subspecies of mendax; checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (subspecies of mendax; checklist; Japan). AKIYAMA & OHMOMO, 1997: 35 [#4091] (subspecies of mendax; checklist; Japan: Hokkaido; Russia: Sachalin). ALEXEEV, 1998: 372 [#4322] (subgenus Sinuatiagrilus; subgeneric classification). FUKUTOMI & HORI, 2004: 9 [#6851] (subspecies of mendax; checklist; Japan (Hokkaido)). JENDEK, 2005: 18 [#7217] (lectotype designation; notes). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2275 [#8576] (world catalog). yokoyamai Iga, 1955 (Agrilus; variety of sachalinensis) syn. nov. IGA, 1955: 79 [#3828] (description). MIWA, 1940: 72 [#4586] (cited as viridri Linné var.; [Note: Misidentification of sinuatus yokoyamai according to KUROSAWA, 1975: 1 2 [#4578]]; checklist; Japan). IGA, 1947: 17 [#3273] (cited as “viridis Linnaeus var.”; [Note: Misidentification of sinuatus yokoyamai according to KUROSAWA, 1975: 1 2 [#4578]]; notes; characters; faunal records; Japan (Shikoku)). KUROSAWA, 1963: 153 [#1618] (valid species; characters; Japan). KUROSAWA, 1974: 2 [#4577] (subspecies of sinuatus). KUROSAWA, 1974: 1 [#4575] (subspecies of sinuatus; characters; notes). KUROSAWA, 1975: 1, 2 [#4578] (subspecies of sinuatus). TÔYAMA, 1985: 23 [#3678] (subspecies of sinuatus; [Note: Kurosawa is cited as the author]; iconography; Japan). HIRASHIMA, 1989: 324 [#8078] (subspecies of sinuatus; checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (subspecies of sinuatus; checklist; Japan). AKIYAMA & AKIYAMA, 1996: 186 [#3962] (subspecies of sinuatus; faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 39 [#4091] (subspecies of sinuatus; checklist; Japan: Hokkaido, Honshu, Shikoku, Kyushu). JENDEK, 2003: 186 [#5823] (subspecies of sinuatus; authorship is discussed). FUKUTOMI & HORI, 2004: 9 [#6851] (subspecies of sinuatus; checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 394 [#7441] (subgenus Sinuatiagrilus; subspecies of sinuatus; Palaearctic catalog). BELLAMY, 2008: 2296 [#8576] (subspecies of sinuatus; world catalog). zhelochovtsevi Alexeev, 1979 (Agrilus; cited as Argilus; subgenus Sinuatiagrilus) syn. nov. ALEXEEV, 1979: 128 129 [#57] (description). ALEXEEV, 1981: 53 56 [#59] (subgenus Sinuatiagrilus; redescription). ALEXEEV, 1989: 486 [#64] (key to species; Primorsk; Chita; Buryatia). ALEXEEV & VOLKOVITSH, 1989: 320, 349, 357 [#3241] (characters in key; faunal records; Mongolia). ALEXEEV, 1998: 372 [#4322] (subgenus Sinuatiagrilus; subgeneric classification). JENDEK, 2006: 394 [#7441] (subgenus Sinuatiagrilus; Palaearctic catalog). BELLAMY, 2008: 2367 [#8576] (subgenus Sinuatiagrilus; world

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA catalog).

Type material: Agrilus sachalinensis Obenberger, 1935 Type locality: Sachalin. Type specimens: Primary: Lectotype (Fig. 52K), ♂, designated by JENDEK (2005: 18) [#7217], (NMPC): “SAGHALIEN MIWA et MINOWA 1927 [p] \ Typus [p] [red label] \ A. sachalinensis m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 26215 [h] [orange label]”. Described from unknown number of syntypes. Agrilus sachalinensis variety yokoyamai Iga, 1955 Type locality: Mt. Asamayama, Nagano Pref. Type specimens: Not located. Described from unknown number of syntypes. Agrilus zhelochovtsevi Alexeev, 1979 Type locality: Reka Ingoda, Nizhn. Klyuchi [in Russian]. Type specimens: Primary: Holotype (Fig. 54X), (ZIN). Described from 3 specimens (HT, PT). Diagnosis: 5.7-10.5 mm. It differs from the nominotypical subspecies by sharper and more elevated prehumerus. Species group: Sinuatus. Relevant species: A. mendax Mannerheim, 1837. Specimens examined: CHINA: Gansu: 1 ♀ (EJCB): “China Gansu, Dingxi, 23.5. 30.5.1996, E. Kučera leg.”; 1 ♂ (EJCB): “China Gansu, Jiuzhaigon [sic!], 12.6. 17.6.2000, leg. E. Kučera”. JAPAN: Hokkaido: 3 ♂, 2 ♀ (EJCB): “Japan, Hokkaido, Ebetsu city, 15.vii., Nopporo virgin forest, 2006, leg. T. Lackner \ on Sorbus”. Honshu: 1 ♀ (EJCB): “Chuzenji (Nikko), 30 VI 1963, Col. K. Kinugasa”; 2 ♂ (EJCB): “1996.7.20, Nasu, Minami gassan, Tochigi, C Jpn, H. Takizawa”; 1 ♀ (EJCB): “1996.7.27., Kiso Ontake, Nagano, C Japan, S. Ohmomo leg.”; 1 ♂ (EJCB): “2000.7.1., Aizu Kanayama, Fukushima, NE Jpn, K. Takahashi leg.”. MONGOLIA: 2 ♂ (EJCB): “Mongolia, 20km S of Ulaanbaatar, 22 23.vi.1990, L. & M. Bocák leg.”. RUSSIA: Primorskiy kray: 1 ♀ (EJCB): “SU Ussuri reg., Novovarvarovka, 6 10.7.1989, M. Nikodým lgt.”; 1 ♀ (EJCB): “Sib. or. m., Primorje, Sichote Alin Mts., Sokolči, 1 15.7.1990, Kadlec + Voříšek lg.”. Zabaykalskiy kray: 1 ♂ (ZIN): “Chitinsk. ob., S. Klyuchi na r. Ingode, 30 vi 1974, Zhelokhovtsev [in Russian]”.

Adult occurence: 5-6-7 (n=13). Faunal records published by ALEXEEV, 1979 [#57] (as zhelochovtsevi) and TÔYAMA, 1985 [#2924] (as mendax sachalinensis) are also included. Host plants: Crataegus: ALEXEEV & VOLKOVITSH, 1989 (as zhelochovtsevi) [#3241]; ALEXEEV, 1989 (as zhelochovtsevi) [#64]. – C. maximowiczii: ALEXEEV, 1979 (as zhelochovtsevi) [#57]. Sorbus: ALEXEEV, 1989 [#64].– S. alnifolia: AKIYAMA & OHMOMO, 1997 (as sinuatus yokoyamai) [#4091]. – S. commixta: AKIYAMA & OHMOMO, 1997 (as sinuatus yokoyamai) [#4091]. Distribution: CHINA: Gansu. JAPAN: Hokkaido, Honshu, Kyushu, Shikoku. MONGOLIA. RUSSIA: Buryatiya Respublika, Primorskiy kray, Sakhalinskaya oblast’, Zabaykalskyi kray. Remarks: Agrilus sinuatus is very variable in the size (see Diagnosis) and color. Except for cupreo-carmineous specimens characteristic for Europe, east-Asian specimens are often golden-greenish, resembling in color those of A.

189

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA mendax Mannerheim, 1837. This similarity was probabbly the reason why subspecies A. sinuatus sachalinensis was considered a subspecies of A. mendax. Agrilus sinuatus may be distinguished from A. mendax by its smaller size, sharp posterior angles of the pronotum and by the sharper, longer, and more arcuate prehumerus reaching to the posterior pronotal angles.

Agrilus siskai Jendek, sp. nov. Fig. 31E (habitus of HT)

Description: 4.5-5.8 mm (Holotype 5.8 mm). BODY: Shape: cuneiform, Color (dorsally): unicolored, rarely bicolored, Color (dominant): green. HEAD: Medial impression: absent, rarely present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Sculpture on vertex: punctures, Sculpture (aspect): regular, rarely irregular, Sculpture (density): sparse or dense, Sculpture (intensity): superficial. Eyes: Size: moderate, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short. PRONOTUM: Shape: visually square, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: posterolateral. Prehumerus: carinal, Shape: arcuate, Extent: beyond half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina, Arc: obvious. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior 1/5-1/4 of pronotal length or at posterior third of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: markedly attenuate. Apices: Arrangement: separate, Width: narrow, Shape: arcuate or subtruncate, Truncation: transverse. Elytral pubescence: entire, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: impressed. Prosternal lobe: moderate, Anterior margin: subtruncate or arcuately emarginate, Emargination (deepness): shallow, Emargination (width): narrow. Prosternal process: narrow, Shape: subparallel, Sides: straight or arcuate, Angles: rectangular or obtuse, Angles (tips): blunt, Disk: impressed with sides protruding. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus or somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Ovipositor: elongate. Diagnosis: prosternal process markedly narrow, disk deeply impressed, sides obviously exserted; metasternal projection

deeply impressed. Species group: Unassigned. Relevant species: Unknown. Type locality: China, Yunnan province, Daju. Type specimens: Primary: Holotype (Fig. 31E), ♀, (EJCB): “China, Yunnan, Daju, 17.VI.1995, Šiška & Spevár & Pekarovič”. Secondary: CHINA: Yunnan: 2 ♀ PT (EJCB): “China, Yunnan, Daju, 17.VI.1995, Šiška & Spevár & Pekarovič”. Adult occurence: 6 (n=1). Host plants: Unknown. Distribution: CHINA: Yunnan. Etymology: Patronymic; the species is named in honor of Bernard Šiška, Slovakia, one of the collectors of the type specimens.

Agrilus smaragdifrons Ganglbauer Fig. 13B (habitus); Fig. 33W (aedeagus); Figs. 47T, 52Z (types) smaragdifrons Ganglbauer, 1889 (Agrilus) GANGLBAUER, 1889: 22, 29 31 [#3427] (description). HEYDEN, 1893: 90 [#3450] (catalog). KERREMANS, 1903: 290 [#1441] (catalog). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1047 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 358 [#2429] (checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Gansu, Sichuan). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2297 [#8576] (world catalog). kanssuanus Obenberger, 1930 (Agrilus) syn. nov. OBENBERGER, OBENBERGER, 1930: 109 [#2088] (description). 1935: 163 [#2158] (note). OBENBERGER, 1936: 1004 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 356 [#2429] (cited as kangsuanus; checklist; China). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2149 [#8576] (world catalog).

Type material: Agrilus smaragdifrons Ganglbauer, 1889 Type locality: Kan-ssu [Fluss Hei-Ho]; Sze-tschuan [Dorf Hunnei-hu (Grenze der Provinzen Kan-Ssu und Szetschuan)] [Note: Locality was complemented from the list of localities on page 21-22]. Type specimens: Primary: Lectotype (Fig. 52Z) by present designation, ♂, (ZIN): “Kan-ssu 1885 G. Patanin [sic!] [p] 20-21. VII. 85 [h] [underside] \ Agr. smaragdifrons Gglb. Typ. [h] \ Coll. Semenov-Tian-Shansky [p] \ Holotypus [p] [red label] \ male det. Alexeev [h] kop. app. vzyat dlya prigotovleniya prep. [h] [in Russian] \ SYNTYPUS Agrilus smaragdifrons Ganglbauer 1890 E. Jendek det. 1993, II [p] [red label]”. Secondary: 1 PLT (NMW). Described from 3 syntypes. Agrilus kanssuanus Obenberger, 1930 Type locality: Kanssu. Type specimens: Primary: Lectotype (Fig. 47T) by present designation, ♀, (NMPC): “Kanssu mer. Hoei-Siee. [h] \ Typus [p] [red label] \ Agrilus kanssuanus m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes.

190

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Diagnosis: 6.4-8.2 mm; antennae long; frons markedly flattened; elytra with greenish tinge; aedeagus enlarged apically. Species group: Imitans. Relevant species: A. sikkimensis Obenberger, 1928.

LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Ovipositor: elongate.

Specimens examined: CHINA: Beijing: 2 ♂, 1 ♀ (EJCB): “China centr., Beijing, 15.4.1989, O. Majzlan leg.”.

Diagnosis: body brightly bicolored, pronotum golden-orange, elytra dark-blue. Species group: Smaragdinus. Relevant species: A. smaragdinus smaragdinus Solsky, 1876.

Adult occurence: 4 (n=1). Host plants: Unknown. Distribution: CHINA: Beijing, Gansu. Remarks: The distributional record without supporting data cited from Sichuan (China) by HUA LI ZHONG, 2002 [#6696] was omitted.

Agrilus smaragdinus habros Jendek, ssp. nov. Fig. 15F (habitus of HT)

Description: 8.6-8.8 mm (Holotype 8.8 mm). BODY: Shape: cuneiform, Build: robust, Color (dorsally): bicolored, Color (dominant): blue. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Sculpture on vertex: punctures, rarely rugae, Sculpture (aspect): irregular, Sculpture (density): sparse. Eyes: Size: moderate, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: transverse, Sides: arcuate or subangulate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: vague or moderate, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles. Posterior angles: Shape: rectangular, Apex: sharp. Disk: Convexity: flat or moderately convex, Disk impressions: medial and lateral, Medial impression (shape): entire or anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): deep, Lateral impressions (width): wide. Prehumerus: carinal, Shape: arcuate, Extent: to third of pronotal length, Posterior end: joining posterior pronotal angle or joining posterior pronotal margin, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: narrow or moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior angles. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Width: wide, Shape: arcuate or subtruncate, Truncation: transverse. Elytral pubescence: absent. Tomentum: absent. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate or deep, Emargination (width): moderate or wide. Prosternal process: moderate, Shape: narrowed, Sides: straight or arcuate, Angles: absent or obtuse, Angles (tips): blunt, Disk: flat or convex. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate.

Type locality: China, Shaanxi province, Qinling mts., 6 km east of Xunyangba, altitude 1000-1300 m. Type specimens: Primary: Holotype (Fig. 15F), ♀, (EJCB): “China, 1000-1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.-13.vi.1998, I. H. Marshal leg.”. Secondary: CHINA: Shaanxi: 1 ♀ PT (EJCB): “China, 1000-1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.-13. vi.1998, I. H. Marshal leg.”. Adult occurence: 5-6 (n=1). Altitude range: 1000-1300 m (n=1). Host plants: Unknown. Distribution: CHINA: Shaanxi. Etymology: The subspecific name is the Greek adjective habros (pretty, elegant). It refers to the appearance of this subspecies.

Agrilus smaragdinus smaragdinus Solsky Fig. 15E (habitus); Fig. 34T (aedeagus); Figs. 51F, 52AA (types) smaragdinus Solsky, 1876 (Agrilus) SOLSKY, 1876: 279 280 [#2644] (description). HEYDEN, 1881: 119 [#5548] (catalog). HEYDEN, 1884: 277, 290 [#8482] (faunal record; Amur region). KERREMANS, 1885: 154 [#1381] (catalog). MARSEUL, 1887: 252 [#4707] (catalog). KERREMANS, 1892: 273 [#1397] (catalog). HEYDEN, 1893: 90 [#3450] (catalog). ABEILLE DE PERRIN, 1897: 4 [#19] (characters in key). KERREMANS, 1903: 290 [#1441] (catalog). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). FISHER, 1925: 4 [#975] (note). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1047 1048 [#2191] (world catalog). TER MINASYAN, 1955: 453 [#7893] (host plant; distribution). ALEXEEV, 1979: 129 [#57] (subgenus Arquagrilus; characters; notes; USSR: Primorskiy kray; South Khabarovsk). ALEXEEV, 1989: 485 [#64] (key to species; Khabarovsk; Primorsk; North Korea). HUA LI ZHONG, 2002: 90 [#6696] (Kerremans is cited as the author; checklist; China: Taiwan). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2297 [#8576] (world catalog). VOLKOVITSH, 2009: 135 [#8850] (faunal records; Russia: Primorskiy kray). prasinulus Obenberger, 1924 (Agrilus; subspecies of peregrinus) OBENBERGER, OBENBERGER, 1924: 48 [#2040] (description). 1926: 657 [#2057] (aberration of peregrinus; Palaearctic catalog). OBENBERGER, 1934: 173 [#2143] (synonym of smaragdinus). OBENBERGER, 1936: 1048 [#2191] (synonym of smaragdinus). ALEXEEV, 1979: 129 [#57] (subspecies of peregrinus; notes). JENDEK, 2006: 401 [#7441] (synonym of smaragdinus; Palaearctic catalog). BELLAMY, 2008: 2297 [#8576] (synonym of smaragdinus; world catalog).

Type material: Agrilus smaragdinus Solsky, 1876

191

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Type locality: les environs de la rivière Souyfoun. Type specimens: Holotype by monotypy (Fig. 52AA), (ZIN). Described from 1 specimen. Agrilus peregrinus prasinulus Obenberger, 1924 Type locality: Vladivostok. Type specimens: Primary: Lectotype (Fig. 51F) by present designation, sex not examined, (NMPC): “Vladivostok [h] \ Typus [p] [red label] \ Agrilus peregrinus ssp. prasinulus m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24782 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 6.5-9.6 mm; dorsall golden-green, glabrous; elytra without pubescent spots and dark sutural part in apical half. Species group: Smaragdinus. Relevant species: A. peregrinus Kiesenwetter, 1879. Specimens examined: KOREA SOUTH: 1 (JLCS): “Gachilbong, Yanggu gun, Gangwon do, Korea, 31.v.1992, coll. J. W. Lee”; 1 (JLCS): “Korea, KW, Bukdaesa, Mt. Odae, 28.v.1998, By Sang Wook, Park”; 1 (JLCS): “Mt. Jirisan, Kyeongnam Prov., Korea, 13.vii.2001, S. L. An leg.”; 1 (JLCS): “Guryongryeong, Gangwon do, Korea, 24.vi.2001, coll. Hae yong Oh”. RUSSIA: Primorskiy kray: 1 (EJCB): “Primorskii krai, 25.vi., Ussuriski zapovednik, 1976, Kononov leg.”; 1 (JMCZ): “Yu. Sikhotè Alin’, okr. Sokolchi 12.vii.1980, S. Murzin [in Russian]”; 5 (EJCB): “USSR Siberia, Primorje, Ussurijskij rajon, Kajmanovka, 7. 13.VII.1988 S. Chvilja lgt.”; 1 (MNHE): “Russia, 27 29.06.1989, Primorye Distr., Buyanki, leg. U. Roosileht”; 3 (MNHE): “Russia, 01.07.1989, Primorye Distr., Limonniki, leg. U. Roosileht”; 1 ♀ (EJCB): “SU, Ussuri reg., 800 m, Oblachnaya Mt., 13 16.7.1989, M. Nikodým lgt.”; 3 (EJCB): “USSR, Ussuri prov. Arsenev env., VII.1990, leg. B. Šiška & R. Čermák”; 2 (EJCB): “Sib. or. m., Primorje, Sichote Alin Mts., Sokolči, 1 15.7.1990, Kadlec + Voříšek lg.”; 1 (JMCZ): “SU or, 18.7.1990, Primorskij reg., Novočugujevka, Ing. Kantner lgt.”; 500 (EJCB): “Russia or, Primorskiy Kray, 44°10’00’’N, 133°15’00’’E, ARSEN’YEV env., June 1991, O. Šauša & M. Štrba leg.”; 7 (EJCB): “Sib. or., Primorje, Komarovka flum., Kamenushka env., 300 m, Vořišek lgt., VI.1992”; 1 (JMCZ): “Russia Far East, 19 20.VII.1993, Primorsk region, Lazo, lgt. M. Trýzna”; 1 (EJCB): “Russia, Primorskij k., Kajmanovka, 10 22.vii.1993, Z. Kletečka leg.”; 2 (EJCB): “Russia, Primorski kray, Khankajski distr. 1995, lake Khanka, 17 19.vii., Kamen’ Rybolov”; 14 (EJCB): “Russia E., Primorskyi reg. Artyom town. env., 3 15.VII.1996, Plutenko leg.”; 1 (EJCB): “Russia, Primorskyi Kray, Artem, VI.1998, Plutenko leg.”; 2 (FACF): “Russia Far East, S Primorie, Ussurijsk, 27.vi.2003”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Tachingousa (Prosiolotchnaja) bay 27.06.04 leg. М.& L. Smirnov’s”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Preobrazhenie env. 14.07.06 leg.М.& L. Smirnov’s”.

Adult occurence: 5-6-7 (n=23). Altitude range: 300-800 m (n=2). Host plants: Quercus mongolica: TER MINASYAN, 1955 [#7893]; ALEXEEV, 1979 [#57]; ALEXEEV, 1989 [#64]. Erroneous: Betula costata: TER MINASYAN, 1955 [#7893]. Distribution: CHINA: Heilongjiang, Jilin, Liaoning. KOREA NORTH. KOREA SOUTH. RUSSIA: Khabarovskiy kray, Primorskiy kray.

Agrilus smetanai Jendek, sp. nov. Fig. 18H (habitus of HT); Fig. 35X (aedeagus)

Description: 5.3-5.9 mm (Holotype 5.9 mm). BODY: Shape: cuneiform, Build: slender, Color (dorsally): unicolored, Color (dominant): blue. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: protruding from head outline or not protruding from head outline, Sculpture on vertex: punctures, Sculpture (aspect): irregular, Sculpture (density): dense. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square or transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: vague, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: flat or moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial or subequal in size, Lateral impressions (deepness): rarely deep. Prehumerus: carinal, Shape: bisinuate, rarely straight or arcuate, Extent: to third of pronotal length, Posterior end: joining posterior pronotal angle or joining posterior pronotal margin, Anterior end: distant from pronotal marginal carina, Arc: weak. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent or subparallel, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate or conjoint, Width: wide, Shape: subtruncate, Truncation: transverse, Modifications: rarely sutural rim spinuliform. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate or large, Anterior margin: arcuately emarginate, Emargination (deepness): moderate, Emargination (width): moderate. Prosternal process: narrow, Shape: narrowed, Sides: straight or arcuate, Angles: absent or obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus or somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: elongate. Diagnosis: body larger; elytral apex markedly elongate; aedeagus longer and faintly attenuate in apical third. Species group: Betuleti. Relevant species: A. nicolanus Obenberger, 1924.

192 Type locality: China, Gansu province, road from Wen Xian to Wudu, northeastern Wen Xian, altitude 2250-2420 m. Type specimens: Primary: Holotype (Fig. 18H), ♂, (EJCB): “China, Prov. Ganshu, Str. v. Wen Xian nach Wudu, NO Wen Xian, 2250-2420m, 30.5-6.6.2006, leg. Puchner”. Secondary: CHINA: Gansu: 2 PT (EJCB): “China, Prov. Ganshu, Str. v. Wen Xian nach Wudu, NO Wen Xian, 22502420m, 30.5-6.6.2006, leg. Puchner”. Adult occurence: 5-6 (n=1). Altitude range: 2250-2420 m (n=1). Host plants: Unknown. Distribution: CHINA: Gansu. Etymology: Patronymic; the species is named in honor of our friend and colleague Aleš Smetana, Ottawa, Canada.

Agrilus sospes Lewis Fig. 11C (habitus); Fig. 33C (aedeagus); Figs. 47V, 51R, 53B (types) sospes Lewis, 1893 (Agrilus) KERREMANS, 1903: LEWIS, 1893: 334 [#1722] (description). 290 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 20 [#1809] (catalog; Japan). OBENBERGER, 1936: 1049 [#2191] (world catalog). KUROSAWA, 1949: 13 14 [#5028] (characters). KUROSAWA, 1950: 1116 [#3274] (note). IGA, 1955: 78 [#3828] (iconography; Japan). KUROSAWA, 1956: 1116 [#8107] (characters; Japan). IGA, 1962: 78 [#3351] (iconography; Japan). KUROSAWA, 1963: 152 [#1618] (characters; Japan). KUROSAWA, 1974: 1, 2 [#4575] (characters; notes). KUROSAWA, 1975: 2 [#4578] (note). TÔYAMA, 1985: 26 [#3678] (iconography; Japan). ANONYMOUS, 1986: 177 [#8848] (insect pest; Korea). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). LI JINGKE, 1992: 92 [#6755] (checklist; China: Liaoning). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 186 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 39 [#4091] (checklist; Japan: Hokkaido, Honshu, Shikoku, Kyushu, Tsushima; Korea). FUKUTOMI & HORI, 2004: 10 11 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2005: 18 19 [#7217] (synonymy; faunal records; China (Shaanxi, Shanxi); Japan (Fukushima, Iwate, Shizuoka, Tokyo, Yamanashi); Korea North). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). ANONYMOUS, 2007: 32 [#8250] (checklist; Japan: Honshu). JENDEK, 2007: 93 [#8008] (synonymy). BELLAMY, 2008: 2300 [#8576] (world catalog). kinoshitae Obenberger, 1936 (Agrilus; subspecies of quadrisignatus) OBENBERGER, 1936: 140 [#2207] (description). MIWA & CHÛJÔ, 1936: ADDENDA P. 4 [#1809] (subspecies of quadrisignatus; catalog; Japan). OBENBERGER, 1936: 1026 [#2191] (subspecies of quadrisignatus; world catalog). KUROSAWA, 1975: 2 [#4578] (subspecies of quadrisignatus). JENDEK, 2005: 18 19 [#7217] (synonym of sospes; lectotype designation). JENDEK, 2006: 401 [#7441] (synonym of sospes; Palaearctic catalog). BELLAMY, 2008: 2300 [#8576] (synonym of sospes; world catalog). quadristictulus Obenberger, 1940 (Agrilus) OBENBERGER, 1940: 179 [#2273] (description). JENDEK, 2005: 18 19 [#7217] (synonym of sospes; lectotype designation). JENDEK, 2006: 401 [#7441] (synonym of quadrisignatus quadrisignatus; [Note: Lapse, should be synonym of sospes]; Palaearctic catalog).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA JENDEK, 2007: 93 [#8008] (synonym of sospes; correction to Palaearctic Catalog). LÖBL & SMETANA, 2007: 29 [#8558] (errata for Palaearctic catalog). BELLAMY, 2008: 2300 [#8576] (synonym of sospes; world catalog).

Type material: Agrilus sospes Lewis, 1893 Type locality: Kiushiu ... from Yuyama in Higo. Type specimens: Primary: Lectotype (Fig. 53B) by present designation, sex not examined, (BMNH): “Type H. T. [p] [round label with red border] \ Japan, G. Lewis, 1910-320 [p] \ Yuyama, 10.V.-14.V.81 [p] \ sospes Lewis Type [h]”. Secondary: 1 PLT (BMNH). Described from four syntypes. Agrilus quadrisignatus kinoshitae Obenberger, 1936 Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 47V), ♀, designated by JENDEK (2005: 18-19) [#7217], (NMPC): “Japan D. Baum [h] \ Typus [p] [red label] \ A. quadrisignatus subsp. Kinoshitae m. Type [h] Det. Dr. Obenberger [p]”. Described from unknown number of syntypes. Agrilus quadristictulus Obenberger, 1940 Type locality: Corea: Soeul. Type specimens: Primary: Lectotype (Fig. 51R), ♂, designated by JENDEK (2005: 18-19) [#7217], (NMPC): “Corée: Séoul [h] \ Typus [p] [red label] \ Agrilus quadristictulus m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24297 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 6.3-8 mm; body more elongate; elytra with 3 pairs of tomentose of spots; spots of apical pair widely separated. Species group: Quadrisignatus. Relevant species: A. quadrisignatus Marseul, 1866 Specimens examined: CHINA: Henan: 3 (EJCB): “China, N Henan, 15.v., 35º42’N, 113º36’E, Wanxianshan, 840 m, Jaroslav Turna leg., 2006”. Hubei: 1 ♂ (EJCB): “China, W Hubei, 1300 2000m, Dashennongjia massif E slope, 31º24’ 30’ / 110º21’ 24’, 28.6. 5.7.1995, L.+R. Businský lgt.”. Shaanxi: 1 (EJCB): “China Shaanxi, Lüeang 33°07’N 106°05’E, 8.6. 14.6.1996, lgt. E. Kučera” (JENDEK, 2005 [#7217]); 1 ♀ (EJCB): “C China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000 1300m, 23.V. 13.VI. leg. C. Holzschuh 2000” (JENDEK, 2005 [#7217]). Shanxi: 1 ♂ (EJCB): “China c. 29 30.v., Jingangling, 111.7’ 50km W Linfen, 36.2’, leg. J. Halada 1996”; 1 ♂ (EJCB): “China, centr. Shanxi, Chao Shan 37.1N, 112.4E, 25 km SE Pingyao., 4 5.VII., Jaroslav Turna leg. 2001” (JENDEK, 2005 [#7217]). Sichuan: 1 (EJCB): “China N Sechuan, Shangliang, 24 29.5.2002, lgt. E. Kučera leg.”. JAPAN: Honshu: 1 ♂ (JMCZ): “15.7.[19]64, Mt. Takao, Japan”; 8 (EJCB): “Mitsukyô, Yokohama city, 5.V.1971, Akiyama leg.” (JENDEK, 2005 [#7217]); 1 (EJCB): “Hikawa rindô, Daibosatsu Mts., Yamanashi pref., 8/9.V.79, leg. Nakamura” (JENDEK, 2005 [#7217]); 1 (EJCB): “Kominawa, Takao machi, Tokyo Met., 18.5.79, leg. Kusakabe” (JENDEK, 2005 [#7217]); 4 ♂ (EJCB): “Japan, Shizuoka pref., Sumata, 2.VI.1984, A. Torikata leg.” (JENDEK, 2005 [#7217]); 2 ♀ (EJCB): “Japan, Iwate, Gosho dam, Morioka shi, 16.VI.1993” (JENDEK, 2005 [#7217]); 7 ♂, 2 ♀ (EJCB): “Japan, Fukushima Pref., Fukushima city, Moniwa, Iizaka town, 2.6.1995, leg. Sugiura” (JENDEK, 2005 [#7217]); 1 ♂, 2 ♀ (EJCB): “Japan, Yamagata, Myozawa, Oguni town, 13.vii.2005, N. Sugiura leg.”. KOREA SOUTH: 1 ♀ (EJCB): “Corée, Kosan, Chass. indigènes” (JENDEK, 2005 [#7217]).

193

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Adult occurence: 5-6-7 (n=15). Altitude range: 840-2000 m (n=3). Host plants: Quercus dentata: AKIYAMA & OHMOMO, 1997 [#4091]. – Zelkova serrata: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: CHINA: Henan, Hubei, Liaoning, Shaanxi, Shanxi, Sichuan. JAPAN: Hokkaido, Honshu, Kyushu, Shikoku, Tsushima. KOREA NORTH. KOREA SOUTH.

Agrilus soudeki Obenberger Fig. 17L (habitus); Fig. 35P (aedeagus); Figs. 52S, 53C (types) soudeki Obenberger, 1926 (Agrilus) OBENBERGER, 1926: 103 [#2054] (description). OBENBERGER, 1926: 656 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1049 [#2191] (world catalog). ALEXEEV, 1979: 129 [#57] (subgenus Arquagrilus; characters; notes; USSR: south part of Primorskiy kray). ALEXEEV, 1989: 484 [#64] (key to species; Primorsk; North Korea). JENDEK, 1994: 20 [#1355] (lectotype designation; synonymy; characters; Russian Far East; Korea). ALEXEEV, 1998: 370 [#4322] (subgenus Arquagrilus; subgeneric classification). JENDEK, 2006: 394 [#7441] (subgenus Robertius; Palaearctic catalog). BELLAMY, 2008: 2300 [#8576] (subgenus Robertius; world catalog). VOLKOVITSH, 2009: 136 [#8850] (subgenus Robertius; faunal records; Russia: Primorskiy kray). sericeus Théry, 1942 (Agrilus) THÉRY, 1942: 266 268 [#2875] (description). OBENBERGER, 1943: 10 [#2333] (synonym of soudeki). ALEXEEV, 1979: 129 [#57] (synonym of soudeki). JENDEK, 1994: 20 [#1355] (synonym of soudeki). JENDEK, 2006: 394 [#7441] (synonym of soudeki; Palaearctic catalog). BELLAMY, 2008: 2300 [#8576] (synonym of soudeki; world catalog).

Type material: Agrilus soudeki Obenberger, 1926 Type locality: Ussuri. Type specimens: Primary: Lectotype (Fig. 53C), ♂, designated by JENDEK (1994: 20) [#1355], (NMPC): “Ussuri [h] \ Typus [p] [red label] \ A. Soudeki m. Typ [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv.[p] 24 809 [h] [orange label]”. Described from unknown number of syntypes. Agrilus sericeus Théry, 1942 Type locality: Maoerschan, Mandchourie. Type specimens: Holotype by monotypy (Fig. 52S), (NHMB): “Mandschurei Maoerschan [p] 7. 7. 40 [h] 19 [p] 39 [p] [scratched] \ Type [p] [red label] \ Sammlung Georg Frey [p] \ Agrilus sericeus Thery [h] Type [p] [red ink] \ Agrilus 1943 soudeki m. comp. Type [h] Det. Dr. Obenberger [p]”. Described from 1 specimen. Diagnosis: 3.9-6.1 mm; pronotum without obvious microsculpture; elytral apices subtruncate without spinuliform sutural margin. Species group: Betuleti. Relevant species: A. alutaceicollis Obenberger, 1930. Specimens examined: CHINA: Shaanxi: 1 ♀ (EJCB): “China Shaanxi, Lüeang, 18.5. 21.5.2000, leg. E. Kučera”. Shanxi: 7 (JMCZ): “China c. 29 30.v., Jingangling, 111.7’ 50km W Linfen, 36.2’, leg. J. Halada 1996”. KOREA NORTH: 1 ♂ (EJCB): “D.P.R. Korea, Ryongaksan Mts., 100 km W of Pyongyang, lgt. M. Slovák \ 14.5.1988” (JENDEK, 1994 [#1355]). RUSSIA: Amurskaya

oblast’: 1 (ZIN): “Amurskaya obl., Arkharinskii r n, s. Ukrainka, S. Vinter 10.vi.1976 [in Russian]”; 1 ♂ (JMCZ): “USSR Amur region, Archara, 11.vii.1989, lgt. Morozinski”. Primorskiy kray: 1 (ZIN): “Russia, Nikolsk Ussurijsk, 1899 G. Suvorov” (JENDEK, 1994 [#1355]); 1 (IMCM): “Primorskii krai, Andreevka, 10 30. vii.1986, Nikireev, C. [in Russian]”; 1 ♂ (EJCB): “Sib.or. m., Primorje Sichote Alin Mts. Sokoltchi 1. 15.7.1990 Kadlec + Voříšek lg.” (JENDEK, 1994 [#1355]); 3 (EJCB): “USSR, Primorskij kraj, Arseneev env., 300 400 m, 11 19.VII.1990, B. Šiška & R. Čermák leg.” (JENDEK, 1994 [#1355]); 12 (EJCB): “USSR, Primorsk. kraj, Arsenev env., VI.1991, 44.11N, 133.19E, O. Šausa leg.” (JENDEK, 1994 [#1355]); 5 (EJCB): “USSR, Primorskij kraj Arseneev env., VI.1991, 44.11N, 133.19E, M. Štrba leg.” (JENDEK, 1994 [#1355]); 1 ♂ (EJCB): “Rossia, Primorsk, Kraskino env. mer., 13 16. VII.1992, P. Smrž leg.”; 1 (JMCZ): “Ross. or, Prim. kr., Kraskino, Azajsan, 13 16.VII.1992, Snížek leg.”; 2 ♂ (EJCB): “Rossia, 2 10. VIII., Primorskij kraj, 1992, Ussurijsk, Kajmanovka, lgt. Smrž”; 2 ♀ (IMCM): “Yu Primor’e, Lazovskii raion, p. Preobrazhenie na usokhshikh Quercus mongolica, 9 14.vii.2006, leg. M. L. Smirnovy [in Russian]”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Preobrazhenie env., on a Quercus mongolica 9 17.07.06 leg. М.& L. Smirnov’s”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Preobrazhenie env. 9 17.07.06 leg. М. & L. Smirnov’s”. Adult occurence: 5 6 7 8 (n 16). Altitude range: 300 400 m (n 1). Host plants: Quercus mongolica: ALEXEEV, 1979 [#57]; ALEXEEV, 1989 [#64]. Distribution: CHINA: Heilongjiang, Shaanxi, Shanxi. KOREA NORTH. RUSSIA: Amurskaya oblast’, Primorskiy kray.

Agrilus souliei Jendek Fig. 22H (habitus of HT); Fig. 37Q (aedeagus) souliei Jendek, 1994 (Agrilus) JENDEK, 1994: 30 31 [#1356] (description). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2301 [#8576] (world catalog).

Type material: Agrilus souliei Jendek, 1994 Type locality: China, Sichuan, Tagu. Type specimens: Primary: Holotype (Fig. 22H), ♂, (EJCB): “China, Szechuan, 20.VII.1992, 130 km SW of Kangding, Tagu, R. Sauer leg.”. Secondary: 2 PT (MNHN); 7 PT (EJCB). Described from 10 specimens (HT, PT). Diagnosis: 6.3-9.6 mm; whole body dorsally roughly granulose; pronotum flat, widest at basal half, markedly transverse. Species group: Pekinensis. Relevant species: A. pekinensis Obenberger, 1924. Specimens examined: CHINA: Sichuan: 1 PT (MNHN): “Se. Tchouen Principaute De Batang Vargong, J. A. Soulie 1907” (JENDEK, 1994 [#1356]); 1 (EJCB): “China, SW Sichuan, Xiangcheng, 2000 m, 15.vii.1994”; 1 ♀ (EJCB): “SE Tibet, 3000 3500 m, ca 20 km N of Yanjing, 23.vii. 8.viii.1998, O. Šafránek et M. Tryzna”. Xizang: 3 PT (EJCB): “China, Thibet, Yaregong, 1900” (JENDEK, 1994 [#1356]). Yunnan: 1 PT (MNHN): “Yun Nan, Tsekou” (JENDEK, 1994 [#1356]); 2 (EJCB): “China, pr. Yunnan b. occ., Dequen, 3900 m, 7.VI.1993, R. Červenka lgt.” (JENDEK, 1994 [#1356]); 1 PT (EJCB): “China Yunnan, 25.VII.1993, Chang Shan Mt., 3000 m, 20 km N of Dali” (JENDEK, 1994 [#1356]); 16 (EJCB): “China,

194

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Yunnan, Lugu Lu (lake), 2900 m, 25 27.vi.1995, Šiška & Spevár & Pekarovič”.

Type locality: China, Yunnan province, 100 km west of Kunming, Diaolin Natural Reservation.

Adult occurence: 6-7-8 (n=5). Altitude range: 2000-3900 m (n=5). Host plants: Unknown.

Type specimens: Holotype (Fig. 14A), ♀, (EJCB): “China, Yunnan, 22.V.-2.VI., 100 km W of Kunming, 1993, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”.

Distribution: CHINA: Sichuan, Xizang, Yunnan.

Adult occurence: 5-6 (n=1). Host plants: Unknown.

Agrilus spathulifer Jendek, sp. nov.

Distribution: CHINA: Yunnan.

Fig. 14A (habitus of HT)

Etymology: The specific name was derived from the Latin diminutive noun spathula (any broad blade) and the verb fero (bear or carry); it refers to the shape of elytral apices that are markedly broaden in this species.

Description: Holotype 5.3 mm. BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: markedly convex, Outline: protruding from head outline. Vertex: Shape: markedly convex, Outline: not protruding from head outline, Sculpture (aspect): irregular. Eyes: Size: moderate, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: visually elongate or visually square, Sides: arcuate, Maximal width: at anterior margin, Anterior margin: subequal to posterior. Anterior lobe: obvious, Shape: arcuate, Position to anterior angles: projecting beyond anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: strongly convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow, Lateral impressions (width): narrow. Prehumerus: obsolete, filamentary or carinal, Shape: arcuate, Extent: to third of pronotal length, Posterior end: joining posterior pronotal angle or distant from pronotal angle or margin, Anterior end: distant from pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: spatulate. Apices: Arrangement: separate, Width: wide, Shape: arcuate. Elytral pubescence: absent. Tomentum: absent or spots, Spots (locations): postmedial. STERNUM: Metasternal projection: flat. Prosternal lobe: moderate or large, Anterior margin: arcuately emarginate, Emargination (deepness): deep, Emargination (width): moderate. Prosternal process: narrow, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus or somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Ovipositor: elongate. Diagnosis: head and pronotum markedly convex, elytra with pair of pubescent adsutural spots at apical third; elytral apices obviously expanded. Species group: Plagiatus. Relevant species: A. majzlani Jendek, 2007.

Agrilus spiculum Jendek, sp. nov. Fig. 29H (habitus of HT); Fig. 40W (aedeagus)

Description: 4.7-5.7 mm (Holotype 5.2 mm). BODY: Shape: cuneiform, Color (dorsally): unicolored, Color (dominant): green or brown. HEAD: Medial impression: absent or present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: in plane with frons. Frons: Shape: moderately convex, rarely flat, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): subparallel, Sculpture (density): dense. Eyes: Size: small or moderate, Shape: not protruding from head outline, Lower margin: in line with antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: transverse, rarely visually square, Sides: arcuate, Maximal width: at anterior margin, Anterior margin: subequal to posterior. Anterior lobe: vague or moderate, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles or at level with anterior pronotal angles. Posterior angles: Shape: acute or rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral or lateral only, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow. Prehumerus: carinal, Shape: bisinuate, Extent: to anterior pronotal angle, Modifications: obsolete in anterior part, Posterior end: joining posterior pronotal angle, Anterior end: joining anterior pronotal angle, rarely joining pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, rarely conjoint, Shape: arcuate. Elytral pubescence: entire, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate or deep, Emargination (width): narrow. Prosternal process: narrow, Shape: subparallel or dilated, Sides: straight,

195

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Angles: rectangular or obtuse, Angles (tips): sharp, Disk: flat. ABDOMEN: Sexual modifications in male: transverse protrusion on ventrite 1, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate with tooth-like projection in male. LEGS: Metatarsus (size to mesotarsus): distinctly longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Ovipositor: elongate. Diagnosis: prosternal lobe deeply emarginate; apex af abdomen in male with sharp teeth. Species group: Roscidus. Relevant species: A. roscidus Kiesenwetter, 1857. Type locality: China, Shanxi province, 50 km west of Linfen, Jingangling. Type specimens: Primary: Holotype (Fig. 29H), ♂, (EJCB): “China c. 29-30.v., Jingangling, 111.7’- 50km W Linfen, 36.2’, leg. J. Halada 1996”. Secondary: CHINA: Shaanxi: 1 PT (EJCB): “China - Yunnan [sic!][should be Shaanxi], Lüyeang, 25.7.-30.7.1996, E. Kučera leg.”; 3 PT (EKCS): “China - Shaanxi, Lüeang 33°07’N 106°05’E, 18.6.24.6.[19]97 lgt. E. Kučera”; 2 PT (EJCB): “China - Shaanxi, Lüeang, 22.5.-25.5.2000, leg. E. Kučera”; 10 PT (EKCS): “China-Shaanxi, Lueyang, 29.5.-2.6.2005, lgt. E. Kučera”; Shanxi: 5 PT (EJCB): “China c. 29-30.v., Jingangling, 111.7’50km W Linfen, 36.2’, leg. J. Halada 1996”; Sichuan: 1 ♀ PT (EJCB): “China N, Sichuan, Jiuzhaigou, 2000m, 16.Jun 1992, M. Bok lgt.”; 1 ♂ PT (EJCB): “China-N, Sechuan, Nanjiang, 21-23.5.2002, lgt. E. Kučera”. Adult occurence: 5-6-7 (n=7). Altitude range: 2000 m (n=1). Host plants: Unknown. Distribution: CHINA: Shaanxi, Shanxi, Sichuan. Etymology: The specific name is the Latin noun spiculum (small needle-like body, part, or process) in apposition. It refers to the small projection of the last ventrite in male of this species.

Agrilus spinipennis Lewis Fig. 9F (habitus); Fig. 32L (aedeagus); Figs. 45I, 53E (types) spinipennis Lewis, 1893 (Agrilus) LEWIS, 1893: 332 [#1722] (description). KERREMANS, 1903: 276 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1924: 36 [#2040] (notes). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). YUASA, 1932: 654 [#3369] (characters; Japan). MIWA & CHÛJÔ, 1936: 20 [#1809] (catalog; Japan). OBENBERGER, 1936: 1049 [#2191] (world catalog). IGA, 1947: 17 [#3273] (notes; characters; faunal records; Japan (Shikoku)). KUROSAWA, 1949: 13 [#5028] (characters). YUASA, 1949: 127 [#8108] (characters; Japan). CHÛJÔ & KUROSAWA, 1950: 8 [#501] (catalog; faunal records; Japan (Shikoku)). KUROSAWA, 1950: 1116 [#3274] (notes). MÜHLMANN, 1954: 83 [#8152] (notes on Biology; Japan). IGA, 1955: 78 [#3828] (iconography; Japan). KUROSAWA, 1956: 1116 [#8107] (characters; Japan). IGA, 1962: 78 [#3351] (iconography; Japan). KUROSAWA, 1963: 152 [#1618] (characters; Japan). KUROSAWA, 1974: 1, 2 [#4575] (characters; notes). KUROSAWA, 1974: 2 [#4576]

(note). KUROSAWA, 1974: 4 [#4577] (note). KUROSAWA, 1975: 2 [#4578] (note). TÔYAMA, 1985: 23 [#3678] (iconography; Japan). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 186 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 39 [#4091] (checklist; Japan: Honshu, Shikoku, Kyushu, Tsushima). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Hubei, Guangdong). OHMOMO, 2004: 142 [#6850] (faunal records; new for Korea and Vietnam). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). ANONYMOUS, 2007: 32 [#8250] (checklist; Japan: Honshu). BELLAMY, 2008: 2302 2303 [#8576] (world catalog). dentiapex Obenberger, 1940 (Agrilus) OBENBERGER, 1940: 179 180 [#2273] (description). KUROSAWA, 1975: 2 [#4578] (synonym of spinipennis). AKIYAMA & OHMOMO, 1997: 39 [#4091] (synonym of spinipennis). JENDEK, 2006: 402 [#7441] (synonym of spinipennis; Palaearctic catalog). BELLAMY, 2008: 2303 [#8576] (synonym of spinipennis; world catalog).

Type material: Agrilus spinipennis Lewis, 1893 Type locality: Main island ... between Yokohama and Oyama. Type specimens: Primary: Lectotype (Fig. 53E) by present designation, ♂, (BMNH): “Type H. T. [p] [round label with red border] \ Japan, G. Lewis, 1910-320 [p] \ Oyama 24.V-26.V.80 [p] \ Agrilus spinipennis Lewis Type [h]”. Secondary: 2 PLT (BMNH); 1 PLT (MHNH). Described from unknown number of syntypes. Note: One paralectotype from BMNH is Agrilus imitans Lewis, 1893. Agrilus dentiapex Obenberger, 1940 Type locality: Japan: Montes Nikko. Type specimens: Primary: Lectotype (Fig. 45I) by present designation, ♂, (NMPC): “Mt. Nikko [h] \ Typus [p] [red label] \ A. dentiapex m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24790 [h] [orange label] \ Agrilus spinipennis Lew. [h] Alexeev det. [p]”. Described from unknown number of syntypes. Diagnosis: 7.6-10.5 mm, body ventrally bicolored; pronotum with obvious anterior lobe; posterior pronotal angles sharply acute; elytra black, glabrous, with pubescent, sometimes inconspicuous, adsutural spot in apical third. Species group: Spinipennis. Relevant species: A. cordicollis sp. now. Specimens examined: JAPAN: Honshu: 1 ♂ (EJCB): “Japan, Numata, Nikko, 22.vii.1953”; 1 (JMCZ): “Japonia, 21.7.[19]64, Inokashira park, M. Suda lgt.”; 1 (EJCB): “Japan, Gunma pref., Kangahara, 6.6.1982, A. Torikata leg.”; 1 (EJCB): “Eda, Fukushima Pref. 21.VI.1989 S. Ohmomo leg.”; 3 (EJCB): “Akane Rindô, Fukushima Pref. 16.VII.1989 S. Ohmomo leg.”; 1 (EJCB): “Akane Rindô, Fukushima Pref. 10.VII.1989 S. Ohmomo leg.”; 1 ♀ (EJCB): “Moniwa, Iizaka Town, Fukushima City, Fukushima Pref., Japan, 25 June 1995, Nobuo Sagiura leg.”; 3 ♂ (EJCB): “Japan, Yamagata, Myozawa, Oguni town, 13.vii.2005, N. Sugiura leg.”.

Adult occurence: 6-7 (n=8). Host plants: Zelkova: MÜHLMANN, 1954 [#8152]. – Z. keaki: LEWIS, 1893 [#1722]. – Z. serrata: AKIYAMA & OHMOMO, 1997 [#4091]. Dubious: Ulmus: MÜHLMANN, 1954 [#8152]. Erroneous: Salix: HUA LI ZHONG, 2002 [#6696]. Distribution: CHINA: Guangdong, Hubei. JAPAN: Honshu,

196

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Kyushu, Shikoku, Tsushima. KOREA SOUTH. VIETNAM.

Type locality: China, Yunnan province, Lijiang, 26.53N 100.18E.

Agrilus stigma Jendek, sp. nov.

Type specimens: Holotype (Fig. 3F), ♂, (EJCB): “China, Yunnan prov., 13-15.VII.1996, Lijiang, 26.53N 100.18E, B. Šiška leg.”.

Fig. 13F (habitus of HT); Fig. 34A (aedeagus)

Description: Holotype 7.4 mm. BODY: Shape: cuneiform, Build: slender, Color (dorsally): unicolored, Color (dominant): green. HEAD: Medial impression: present, Impression (extent): vertex and frons, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: flat, Outline: not protruding from head outline. Vertex: Sculpture on vertex: rugae, Sculpture (aspect): subparallel, Sculpture (density): dense. Eyes: Size: small or moderate, Shape: protruding from head outline, Lower margin: in line with antennal socket. Antennae: Length: long. PRONOTUM: Shape: visually square, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles or at level with anterior pronotal angles. Posterior angles: Shape: rectangular, Apex: sharp. Disk: Convexity: flat or moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Anteromedial and posteromedial impressions: subequal in size, Lateral impressions (deepness): shallow, Lateral impressions (width): wide. Prehumerus: filamentary, Shape: bisinuate, Extent: to anterior pronotal angle, Modifications: obsolete in anterior part, Posterior end: joining posterior pronotal margin, Anterior end: joining anterior pronotal angle, Arc: moderate. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: markedly attenuate. Apices: Arrangement: separate, Shape: subangulate. Elytral pubescence: entire except glabrous part at apical half, Color: unicolored, Character: with patches of denser pubescence. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate or large, Anterior margin: arcuately emarginate, Emargination (deepness): moderate or deep, Emargination (width): moderate. Prosternal process: wide, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: impressed with sides protruding, Modification: sides crenulate. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): distinctly longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed. Diagnosis: prehumerus filamentary; apex of elytra more prolonged, apices distinctly separately subangulate. Species group: Plagiatus. Relevant species: A. plagiatus Ganglbauer, 1889.

Adult occurence: 7 (n=1). Host plants: Unknown. Distribution: CHINA: Yunnan. Etymology: The specific name is the Latin noun stigma (a small mark, spot) in apposition. It refers to the dark, glabrous elytral spot in the species.

Agrilus subauratus amurensis Obenberger Fig. 31F 31G (habitus); Fig. 41S (aedeagus); Figs. 42R, 45G (types) amurensis Obenberger, 1922 (Agrilus) OBENBERGER, 1922: 21 22 [#2021] (description). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 941 [#2191] (world catalog). OGATA & OHMOMO, 1985: 133 140 [#5039] (note). ALEXEEV, 1989: 483 [#64] (subspecies of subauratus; key to species; Primorsk; Chita; Buryatia). ALEXEEV & VOLKOVITSH, 1989: 318, 347, 356 [#3241] (subspecies of subauratus; characters in key; faunal records; notes; Mongolia). AKIYAMA, 1993: 8 [#4875] (faunal record; Japan). JENDEK, 1995: 146 [#3768] (subspecies of subauratus; lectotype designation; synonymy; characters; eastern Russia; Mongolia; Korea; North China). AKIYAMA & OHMOMO, 1997: 39 40 [#4091] (subspecies of subauratus; checklist; Japan: Hokkaido; Russia: East Siberia). HUA LI ZHONG, 2002: 90 [#6696] ([Note: Misidentified as subauratus]; checklist; China: Xinjiang). JENDEK, 2002: 7, 8 [#5744] (subspecies of subauratus; taxonomic notes). FUKUTOMI & HORI, 2004: 9 [#6851] (subspecies of subauratus; checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 394 [#7441] (subgenus Robertius; subspecies of subauratus; Palaearctic catalog). BELLAMY, 2008: 2308 2309 [#8576] (sybspecies of subauratus; world catalog). VOLKOVITSH, 2009: 136 [#8850] (subgenus Robertius; subspecies of subauratus; faunal records; Russia: Primorskiy kray). demuthi Théry, 1942 (Agrilus) THÉRY, 1942: 258 260 [#2875] (description). JENDEK, 1995: 146 [#3768] (synonym of subauratus amurensis). JENDEK, 2002: 7, 8 [#5744] (synonym of subauratus amurensis; taxonomic notes). JENDEK, 2006: 394 [#7441] (synonym of subauratus amurensis; Palaearctic catalog). BELLAMY, 2008: 2309 [#8576] (synonym of subauratus amurensis; world catalog).

Type material: Agrilus subauratus amurensis Obenberger, 1922 Type locality: Siberia orientalis: Amur superior. Type specimens: Primary: Lectotype (Fig. 42R), sex not examined, designated by JENDEK (1995: 146) [#3768], (NMPC): “Oberer Amur [p] \ Typus [p] [red label] \ A. amurensis m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24255 [h] [orange label] \”. Described from unknown number of syntypes. Agrilus demuthi Théry, 1942 Type locality: Kiachta, Sibérie. Type specimens: Holotype by monotypy (Fig. 45G), ♂,

197

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA (MNHN): “Kjachta Sibirien Götzelmann”. Described from 1 specimen. Diagnosis: 6.3-9.2 mm; body unicolored; aedeagus distinctly more expanded than that in the nominotypical subspecies. Species group: Unassigned. Relevant species: A. subauratus (Gebler, 1833). Specimens examined: CHINA: Hebei: 1 ♂ (EJCB): “China, Hebei/Nei Mongol, pass Chengde Chifeng, 41.6N, 118.2E, 14 16.VI., Jaroslav Turna leg. 2001”. Heilongjiang: 1 (ZIN): “st. Gaolinczy, vostochnaya liniya KV zhd., Kitai, 1.vii.1937 [in Rusian]”. Jilin: 1 (EJCB): “China, Jilin province, Changbai Shan Mt., 3.VII.1979” (JENDEK, 1995 [#3768]). MONGOLIA: 1 (ZIN): “r. Tes, Mongoliya, 26.vi.1914 Tomahinskii [in Russian]”; 1 ♀ (EJCB): “Mongolia, 20.June 2003, Ulaanbaatar, near Zaysan, Tuul riv.vall., Z Jindra leg.”. RUSSIA: Amurskaya oblast’: 1 ♀ (ZIN): “Amursk. reg., Klimoutsy, 40 km W Svobodnyi, 9.VII.1958, leg. Zinovjev [in Russian]” (JENDEK, 1995 [#3768]); 1 ♀ (ZIN): “Amursk. reg., rr. M. Pera B. Ergel’ mezhdur., Amur., Zinovjev 22.VI.[19]58 [in Russian]” (JENDEK, 1995 [#3768]); 1 (ZIN): “Samodon na Amure, 100 km W Svobodnogo, Zinov’ev, 7.viii.1959 [in Russian]”; 2 (ZIN): “Amurskaya obl., Skovorodinsk. r n, r. Burgali, 12.7.1959, Kabakov [in Russian]”; 1 (ZIN): “Amur. obl., Skovorodnikov. r n, Ul’ruch’i, 12.6.1961, Kabakov [in Russian]”; 1 (ZIN): “Amur. obl. YuV. Urush. r. Omutnaya, 12.6.1961 Kabakov [in Russian]”; 1 (ZIN): “Amurskaya obl., Arkharinskii r n, s. Ukrainka, S. Vinter 10.vi.1976 [in Russian]”; 1 ♂ (ZIN): “Amursk. obl., Tygdinskii rajon, Tygda, 23.VI.1976, leg. Kuznetsov [in Russian]” (JENDEK, 1995 [#3768]). Buryatiya Respublika: 1 ♀ (EJCB): “V. Ubukunsh [ Verknyiy Ubukun] K. Ehnberg” (JENDEK, 1995 [#3768]); 1 ♂, 1 ♀ (EJCB): “Transbaicalia mr. occ. fl. Tshikoj: Sharagol 1.VII.1928 Th. Lukjanovitsh” (JENDEK, 1995 [#3768]). Primorskiy kray: 1 (ZIN): “Unty, Samanga, Ussur. 19.vii.1925”; 1 (ZIN): “Grigorievka Ussur. 19.vii.1937 Stepanov”; 1 (ZIN): “Amurskaya obl. Skovorodino 10.7.1959 Kabakov [in Russian]”; 1 (EJCB): “SU, Ussuri reg., Jasnoe, 12 19.vii.1989, R. Červenka leg.”; 1 ♂ (EJCB): “USSR Sibiria or., Ussuri region 11 14.7, Jasnoe, 450 m, 1989, leg. St. Jákl” (JENDEK, 1995 [#3768]); 1 ♂ (EJCB): “Sib. or. m., Primorje, Sichote Alin Mts., Sokolči, 1 15.7.1990, Kadlec + Voříšek lg.” (JENDEK, 1995 [#3768]); 1 ♂ (EJCB): “Rezanovka, 19.7.1992, , E. Kučera leg.”; 1 (EJCB): “Sib. or., Primorje, Komarovka flum., Kamenushka env., 300 m, Vořišek lgt., V.1992” (JENDEK, 1995 [#3768]); 1 (MSCI): “SE Maritime Prov., Lazovsky dist., ~10 кm SW Sokoltchi 28.06.06 leg. М. & L. Smirnov’s”; 1 ♀ (EJCB): “Russia, Primorsky Kray, Kaymanovka vil., N43º37’, E132º14’, 7 11.vii.2007, 100m, V. Grebennikov”. Tyva Respublika: 1 (ZIN): “Tuva Èrzin B. Korotyaev 5.vii.1973 [in Russian]”. Zabaykalskiy kray: 1 (NMPC): “Tschita [ Chita], Transbaikalien, Hermann Frieb” (JENDEK, 1995 [#3768]).

Adult occurence: 5-6-7-8 (n=25). Altitude range: 100-450 m (n=3). Host plants: Populus: ALEXEEV, 1989 [#64]; ALEXEEV & VOLKOVITSH, 1989 [#3241]. – Salix: ALEXEEV, 1989 [#64]; ALEXEEV & VOLKOVITSH, 1989 [#3241]. Erroneous: Betula: ALEXEEV, 1989 [#64]; Corylus: ALEXEEV, 1989 [#64]; Quercus: ALEXEEV, 1989 [#64]. Distribution: CHINA: Hebei, Heilongjiang, Jilin, Xinjiang. JAPAN: Hokkaido. KOREA NORTH. MONGOLIA. RUSSIA: Amurskaya oblast’, Buryatiya Respublika, Primorskiy kray, Tyva Respublika, Zabaykalskiy kray.

Remarks: The nominotypical subspecies is bicolored and has less dilated aedeagus. The transitional zone between both subspecies is defined approximately by the river Yenissey.

Agrilus subrobustus Saunders Fig. 15A (habitus); Fig. 34Q (aedeagus); Figs. 43Q, 48D, 48E, 48W, 53A, 53I, (types) subrobustus Saunders, 1873 (Agrilus) SAUNDERS, 1873: 516 517 [#2554] (description). LEWIS, 1879: 15 [#3345] (catalog; Japan). KERREMANS, 1885: 154 [#1381] (catalog). SCHÖNFELDT, 1887: 113 [#8901] (Anambus; catalog; Japan). KERREMANS, 1892: 274 [#1397] (catalog). KERREMANS, 1903: 291 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 21 [#1809] (catalog; Japan). OBENBERGER, 1936: 1052 [#2191] (world catalog). KUROSAWA, 1963: 153 [#1618] ([Note: Other species is illustrated]; characters; Japan). KUROSAWA, 1975: 1 [#4591] (notes). TÔYAMA, 1985: 22 [#3678] (iconography; Japan). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). JENDEK, 1995: 139 140 [#3768] (lectotype designation; characters; Japan; Korea; China: Shaanxi, Sichuan, Hunan, Fujian). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 186 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 40 [#4091] (checklist; Japan: Honshu, Shikoku, Kyushu, Tsushima, Yakushima I., Tokara nakanoshima I., Korea; Taiwan; China). NONNAIZAB et al., 1999: 113 [#8247] (checklist; China: Inner Mongolia). JENDEK, 2000: 506 [#4851] (synonymy). MÜHLE, 2003: 47 [#6155] (checklist; Taiwan). JENDEK, 2005: 20 [#7217] (synonymy). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). JENDEK, 2007: 93 [#8008] (synonymy). WESTCOTT, 2007: 111 [#8118] (new introduction; USA (Georgia)). BELLAMY, 2008: 2313 [#8576] (world catalog). JENDEK & GREBENNIKOV, 2009: 243 [#8724] (notes). HANSEN, MOULTON & KLINGEMAN, 2010: 444 445 [#9057] (flight activity; USA: Tennessee). HATTORI, 2010: 4 [#8899] (faunal records; Tokara Archipelago). sorocinus Kerremans, 1914 (Agrilus) KERREMANS, 1914: 106 107 [#1481] (description). MIWA, 1931: 126 [#6101] (catalog; Formosa). MIWA & CHÛJÔ, 1936: 20 [#1809] (catalog; Japan). OBENBERGER, 1936: 1102 [#2191] (world catalog). KUROSAWA, 1963: 154 [#1618] (characters; Japan). KUROSAWA, 1975: 1 [#4591] (synonym of subrobustus). PENG ZHONGLIANG, 1987: 358 [#2429] (checklist; China). AKIYAMA & OHMOMO, 1997: 40 [#4091] (synonym of subrobustus). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Taiwan; Japan, Korea). JENDEK, 2006: 402 [#7441] (synonym of subrobustus; Palaearctic catalog). BELLAMY, 2008: 2313 [#8576] (synonym of subrobustus; world catalog). kumamotoensis Obenberger, 1935 (Agrilus) OBENBERGER, 1935: 170 171 [#2158] (description). MIWA & CHÛJÔ, 1936: 17 [#1809] (catalog; Japan). OBENBERGER, 1936: 1004 [#2191] (world catalog). AKIYAMA & OHMOMO, 1997: 40 [#4091] (synonym of subrobustus). JENDEK, 2000: 506 [#4851] (synonym of subrobustus; lectotype designation). JENDEK, 2006: 402 [#7441] (synonym of subrobustus; Palaearctic catalog). BELLAMY, 2008: 2313 [#8576] (synonym of subrobustus; world catalog). malyi Obenberger, 1936 (Agrilus) syn. nov. OBENBERGER, OBENBERGER, 1936: 116 [#2205] (description). 1936: 1012 [#2191] (world catalog). COBOS, 1972: 56 [#591] (subgenus Agrilus; checklist; Mongolia). ALEXEEV, 1975: 155 [#52] (faunal record; Mongolia). PENG ZHONGLIANG, 1987: 356 [#2429] (checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Inner Mongolia). JENDEK, 2006: 399 [#7441]

198 (Palaearctic catalog). BELLAMY, 2008: 2177 [#8576] (world catalog). albizziae Kurosawa, 1963 (Agrilus; subspecies of sorocinus; [preoccupied]) KUROSAWA, 1963: 101 102 [#1617] (description). KUROSAWA, 1963: 154 [#1618] (subspecies of sorocinus; characters; Japan). KUROSAWA, 1975: 1 [#4591] (synonym of subrobustus). AKIYAMA & OHMOMO, 1997: 40 [#4091] (synonym of subrobustus). BELLAMY, 1998: 95 [#4505] (synonym of sorocinus kurosawus). JENDEK, 2006: 402 [#7441] (synonym of subrobustus; Palaearctic catalog). BELLAMY, 2008: 2313 [#8576] (synonym of subrobustus; world catalog). boisreymondi Alexeev & Bílý, 1980 (Agrilus) ALEXEEV & BÍLÝ, 1980: 604 605 [#44] (description). JENDEK, 2007: 93 [#8008] (synonym of subrobustus; remark on type locality). BELLAMY, 2008: 2313 [#8576] (synonym of subrobustus; world catalog). kumamotoi Tôyama, 1987 (Agrilus) HUA LI ZHONG, TÔYAMA, 1987: 316 [#2931] (description). 2002: 90 [#6696] (checklist; China: Taiwan). JENDEK, 2005: 20 [#7217] (synonym of subrobustus). JENDEK, 2006: 402 [#7441] (synonym of subrobustus; Palaearctic catalog). BELLAMY, 2008: 2313 [#8576] (synonym of subrobustus; world catalog). kurosawus Bellamy, 1998 (Agrilus; replacement name for albizziae Kurosawa not Fisher) BELLAMY, 1998: 95 [#4505] (replacement name proposal). JENDEK, 2006: 402 [#7441] (synonym of subrobustus; Palaearctic catalog). BELLAMY, 2008: 2313 [#8576] (synonym of subrobustus; world catalog).

Type material: Agrilus subrobustus Saunders, 1873 Type locality: Nagasaki. Type specimens: Primary: Lectotype (Fig. 53I), ♀, designated by JENDEK (1995: 139-140) [#3768], (BMNH): “Type H. T. [p] [round label with red border] \ Oak Nagasaki 1869 [h] \ Japan Described from unknown number of syntypes. Agrilus sorocinus Kerremans, 1914 Type locality: [Formosa] Banshoryo Distr.: Sokutsu. Type specimens: 1 ST (Fig. 53A) (MNHN); 1 ST (DEI). Described from unknown number of syntypes. Agrilus kumamotoensis Obenberger, 1935 Type locality: Japan: Kumamoto. Type specimens: Primary: Lectotype (Fig. 48D), ♀, designated by JENDEK (2000: 506) [#4851], (NMPC): “Japan - Kumamoto [h] \ Typus [p] [red label] \ Agrilus kumamotoensis m. Type [h] [Obenberger’s MS] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24306 [h] [orange label]”. Described from unknown number of syntypes. Agrilus malyi Obenberger, 1936 Type locality: Mongolia or.: Chancheu. Type specimens: Primary: Lectotype (Fig. 48W) by present designation, ♀, (NMPC): “Mongolia or: Chancheu [h] \ Typus [p] [red label] \ Agrilus Malýi m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24335 [h] [orange label]”. Described from unknown number of syntypes. Agrilus sorocinus albizziae Kurosawa, 1963 Type locality: Ôsaka Pref.: Shakudai, Shimamato-chô. Type specimens: Not located. Described from unknown number of specimens (HT, PT).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Agrilus boisreymondi Alexeev & Bílý, 1980 Type locality: Severnaya Birma, Kin-Kiang [Note: Locality emended to China, Kiu Kiang. See JENDEK, 2007 [#8008]]. Type specimens: Primary: Holotype (Fig. 43Q), ♀, (NMPC): “China [p] 1907 Kiu Kiang [h] \ Dr. du BoisReymond Shanghai [p] \ Agrilus bois-reymondi sp. n. [h] Alexeev det. [p] \ Holotype [p] Agrilus boisreymondi [h] Sv. Bílý det. [p] et Alexeev 1977 [h] \ Mus. Nat. Pragae Inv. [p] 65340 [h] [red label]”. Described from 1 specimen (HT). Agrilus kumamotoi Tôyama, 1987 Type locality: Shangping near Liukuei, Kaohsiung Hsien, Taiwan. Type specimens: Primary: Holotype (Fig. 48E), ♀, (NSMT): “(Senpei) Kaohsiung, FORMOSA 5, June, 1976 M. Kubata leg. [p] \ Holotype [p] Agrilus kumamotoi Tôyama, 1986 [h] [red label]”. Described from 1 specimen (HT). Agrilus kurosawus Bellamy, 1998 Type locality and specimens: See: Agrilus sorocinus albizziae Kurosawa, 1963. Diagnosis: 4-6.1 mm; prehumerus short, filamentary; aedeagus with subangulate sides. Species group: Subrobustus. Relevant species: A. ineptus Kerremans, 1892. Specimens examined: CHINA: Anhui: 2 (EJCB): “China, Anhui, 1998, Dabieshan, 65 km SW Huoshan, 1400 m, Bolm lgt. 21 24 Jun”. Fujian: 10 (EJCB): “China, Fujian prov. Shaowu env., 5 10. VII.1991” (JENDEK, 1995 [#3768]); 3 (EJCB): “China, Fujian prov. bor. occ., Shaowu env. 25 km road, Shaowu Taining 6 10.7., R. Červenka lgt. 1991”; 8 (EJCB): “China, Fujian prov., Shaowu env., 23 27.vi.1991, leg. Nikodým” (JENDEK, 1995 [#3768]). Guizhou: 1 ♂ (EJCB): “CH Guizhou NE, 27.V. 3.VI., 20 km NW of Jiangkou, 1995, Fanjing Shan Kuaichang, E. Jendek & O. Šauša leg.”; 4 ♂ (EJCB): “China, Guizhou prov., Fodingshan, Ganshi, 25 km S Shiquaian, 1300 m, 5 9. June 1997, Bolm lgt.”. Hubei: 1 (EJCB): “China, SE Hubei, 6.+19.vi, 20 km NW Tongshan, 29.7N 114.3E, Jaroslav Turna leg. 2002”. Hunan: 2 (EJCB): “China, Hunan, Zhang Jia Jie, VI.1993, leg. C. Holzschuh” (JENDEK, 1995 [#3768]); 1 ♀ (EJCB): “China, N Hunan, 4 7.vii., Wuling Shan, 29.4N, 110.4E, Zhangjiajie, 700 m, Jaroslav Turna leg. 2003”. Shaanxi: 1 ♂ (EJCB): “China, Shaanxi pr., Hua Shan, 17 21.VI.1991, R. Dunda lgt.”; 1 ♂ (EJCB): “China, Shaanxi, pr. Hua Shan, 17. 21.6.1991, R. Dunda lgt.” (JENDEK, 1995 [#3768]); 1 ♀ (EKCS): “China Shaanxi, Lüeang 33°07’N 106°05’E, 18.6. 24.6.[19]97 lgt. E. Kučera”; 1 ♂ (EJCB): “China, Shaanxi, Qinling mts., 500m, Lianghekou, 18 19.vi.1998, I. H. Marshal leg.”; 1 ♀ (EJCB): “China, 1000 1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v. 13.vi.1998, I. H. Marshal leg.”; 1 (EKCS): “China Shaanxi, Lueyang, 29.5. 2.6.2005, lgt. E. Kučera”; 1 (EKCS): “China Shaanxi, Lueyang, 15.7. 22.7.2005, lgt. E. Kučera”; 1 (EKCS): “China Shaanxi, Lüeyang, 22 25.5.2006, lgt. E. Kučera”. Sichuan: 3 (EJCB): “China, NW Sichuan, 103.36/31.00E, Guanxian, 700m, 08 20.07.1990, leg. J. Kolibáč” (JENDEK, 1995 [#3768]). Yunnan: 1 (MNHN): “Chine, Yunnan, recu de Lou nan, 1931”; 1 ♂, 1 ♀ (EJCB): “Yunnan 1800 2000m, 25.04N 101.55E, Yipinglang 17 20.6.1994,Vít Kubáň leg.”; 16 (EJCB): “CH Yunnan, 13 14. VI.1995, 100 km W of Kunming, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”; 2 ♂, 2 ♀ (EJCB): “Yunnan, 1500 2500 m, 25°22’N, 98°49’E, 17 24 / 5 Gaoligong mts, Vít Kubáň leg., 1995”. JAPAN: Honshu: 1 (EJCB): “Japan, 8 10.VI.1979, A. Nishiyama leg., Hananuki keikoku, Takahagi City, Ibaraki Pref.” (JENDEK, 1995 [#3768]); 1 (EJCB): “C Japan, Ibaraki Pref., Ami cho, 17.VI.1987, leg. Ohmomo” (JENDEK, 1995 [#3768]); 1 ♂ (EJCB): “Japan, Tochigi pref., Oashi gawa, Kanuma shi, 29.vii.1992”. Kyushu: 1 (EJCB):

199

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA “Naiju, Fukuoka, 5.VII.1979” (JENDEK, 1995 [#3768]). Tsushima: 3 (EJCB): “Mt. Ohboshiyama, Tsushima Is., 11.VI.1989, leg. K. Adachi” (JENDEK, 1995 [#3768]); 1 (EJCB): “Mt. Ohboshiyama, Tsushima Is., 12.VII.1989, leg. K. Adachi” (JENDEK, 1995 [#3768]). KOREA SOUTH: 1 (EJCB): “Corée Env. de Seoul 1913” (JENDEK, 1995 [#3768]); 1 ♀ (JMCZ): “Korea: Chungnam Prov., Taean gun, Taean eup., Mt. Baehwasan, 6 VII 25 VIII 2001, S J Park, C W Shin e FIT”.

Adult occurence: 5-6-7-8 (n=28). Altitude range: 5002500 m (n=8). Host plants: Albizzia julibrissin: KUROSAWA, 1963 [#1617] (as albizziae); AKIYAMA & OHMOMO, 1997 [#4091]. Erroneous: Quercus: SAUNDERS, 1873 [#2554].

al., 1999: 113 [#8247] (cited as sundai; checklist; China: Inner Mongolia). FUKUTOMI & HORI, 2004: 10 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2315 [#8576] (world catalog).

Type material: Agrilus sudai Kurosawa, 1985 Type locality: Mt. Hayachine, Iwate Pref., Honshu, Japan. Type specimens: Primary: Holotype (Fig. 53J), ♀, (NSMT): “Mt. Hayachine Iwate Japan Jul. 31. 1952 col. M. Suda [p] \ Holotype [p] Agrilus sudai Y. Kurosawa, 1985 [h] [red label with black border]”. Secondary: 2 PT (NSMT). Described from 5 specimens (HT, AT, PT).

Distribution: CHINA: Anhui, Fujian, Guizhou, Hubei, Hunan, Jiangxi, Nei Mongol, Shaanxi, Sichuan, Taiwan, Yunnan. JAPAN: Honshu, Kyushu, Shikoku, Tsushima, Ryukyu Islands. KOREA SOUTH. MONGOLIA. NORTH AMERICA (introduced): UNITED STATES.

Diagnosis: 5.3-7.2 mm; elytral apices subtruncate; proximal part of elytral pubescence entire, distal V-shaped. Species group: Maculifer. Relevant species: A. daimio Obenberger, 1936; A. imasakai Tôyama, 1985.

Agrilus subtilipilis Obenberger

Adult occurence: 7 (n=5). Faunal records published by KUROSAWA, 1985 [#1640] are also included.

Fig. 26G (habitus of LT) subtilipilis Obenberger, 1940 (Agrilus) PENG OBENBERGER, 1940: 180 181 [#2273] (description). ZHONGLIANG, 1987: 358 [#2429] (checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China). PENG ZHONGLIANG, 2002: 268 [#6628] (notes). JENDEK, 2005: 20 [#7217] (lectotype designation). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2314 [#8576] (world catalog).

Type material: Agrilus subtilipilis Obenberger, 1940 Type locality: China: Fukien: Shoa-wu. Kwangtsen. Type specimens: Primary: Lectotype (Fig. 26G), ♀, designated by JENDEK (2005: 20) [#7217], (NMPC): “Kwangtseh-Fukien, J. Klapperich, 0, 22.7.1937 [p] [violet label] \ Typus [p] [red label] \ A. subtilipilis m. Type [h] Det. Dr. Obenberger [p]”. Described from unknown number of syntypes. Diagnosis: 5 mm; prosternal lobe prominent, pronotal disk more convex, prehumerus extending to half of pronotal length; prosternal lobe inconspicuously emarginate, almost subtruncate. Species group: Cyaneovirens. Relevant species: A. yamabusi Miwa & Chûjô, 1940. Specimens examined: Known only from the lectotype.

Host plants: Unknown. Distribution: CHINA: Fujian.

Agrilus sudai Kurosawa

Specimens examined: JAPAN: Honshu: 1 ♂ (EJCB): “2000.7.1., Aizu Kanayama, Fukushima, NE Jpn, K. Takahashi leg.”.

Host plants: Alnus hirsuta: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: JAPAN: Hokkaido, Honshu, Shikoku. Remarks: The distributional record without supporting data cited from Inner Mongolia (China) by NONNAIZAB et al., 1999 [#8247] was omitted.

Agrilus suginoi Tôyama Fig. 23A (habitus); Fig. 37U (aedeagus); Fig. 53L (type) suginoi Tôyama, 1985 (Agrilus) TÔYAMA, 1985: 21 23 [#2924] (description). AKIYAMA & OHMOMO, 1997: 40 [#4091] (checklist; Japan: Ryukyus (Iriomote jima I.)). JENDEK, 2003: 186 187 [#5823] (characters; faunal records; Japan: Fukushima, Tochigi, Ibaraki, Ōita, Okinawa). OHMOMO, 2004: 140 141 [#6850] (notes; known only from the type locality). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 1950 [#8576] (synonym of adelphinus; world catalog).

Type material: Agrilus suginoi Tôyama, 1985 Type locality: Kanpira, Iriomote Is. Type specimens: Primary: Holotype (Fig. 53L), ♀, (NSMT): “(showed) at Nishinomiya 10. V. 1973 K. Sugino leg. [h] \ Kanpira (is. Iriomote) K. Sugino leg. [h] \ Holotype [p] Agrilus suginoi Tôyama, 1985 [h] [red label]”. Described from 2 specimens (HT, PT).

Fig. 12A (habitus); Fig. 33K (aedeagus); Fig. 53J (type)

Diagnosis: 4.8-6.9 mm; body dark-blue; aedeagus with impressed upper part of tegmen. Species group: Sulcicollis. Relevant species: A. adelphinus Kerremans, 1895.

sudai Kurosawa, 1985 (Agrilus) KUROSAWA, 1985: 160 162 [#1640] (description). TÔYAMA, 1985: 25 [#3678] (iconography; Japan). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 40 [#4091] (checklist; Japan: Hokkaido, Honshu, Shikoku). NONNAIZAB et

Specimens examined: JAPAN: Honshu: 2 ♂ (SOCI): “Akané Rindô, Haramomachi City, Fukushima Pref., 23.VI.1980, S. Ohmomo leg.” (JENDEK, 2003 [#5823]); 1 ♂ (SOCI): “1988.6.11, Mt. Hanazono, Kita Ibaraki, Ibaraki, K. Ichige leg.” (JENDEK, 2003 [#5823]); 1 ♂ (SOCI): “1989.5.5., Migimomi, Tsuchiura, Ibaraki

200 Pref. S. Ohmomo leg.” (JENDEK, 2003 [#5823]); 1 ♂ (SOCI): “1994.05.22., Kido, Motegi, Tochigi Pref. S. Ohmomo leg.” (JENDEK, 2003 [#5823]); 1 ♂ (EJCB): “Japan, Fukushima, Namie town, 26.vi.1995, N. Sugiura leg.”; 1 ♂ (SOCI): “1996.06.21., Teraku, Kuroiso, Tochigi Pref. S. Ohmomo leg.” (JENDEK, 2003 [#5823]); 1 ♂ (SOCI): “1998.06.06., Yatakezawa, Tateiwa, Fukushima pref., S. Ohmomo leg.” (JENDEK, 2003 [#5823]); 1 ♂ (SOCI): “2001.04.29., Mt. Tsukuba, Makabe, Ibaraki, S. Ohmomo leg.” (JENDEK, 2003 [#5823]). Kyushu: 2 ♂ (EJCB): “Japan, Kyushu, Oita pref., Kujû, Mts. Kurodake, 25 26.v.1991, Ryô Noda leg.” (JENDEK, 2003 [#5823]); 2 ♂ (SOCI): “Mt. Kurodake, Mts. Kujyusan, Oita Pref., 1.VI.1991, S. Ohmomo leg.” (JENDEK, 2003 [#5823]); 1 ♂ (SOCI): “2001.6 .3., Tsukahara H. S., Yufuin, Oita Pref. Y. Tsutsumiuchi leg.” (JENDEK, 2003 [#5823]).

Adult occurence: 4-5-6 (n=11). Host plants: Unknown. Distribution: JAPAN: Honshu, Kyushu, Ryukyu Islands.

Agrilus sugiurai Jendek Fig. 31H (habitus); Fig. 41T (aedeagus); Fig. 53M (type) sugiurai Jendek, 2007 (Agrilus) JENDEK, 2007: 31, 33 [#8006] (description). BELLAMY, 2008: 2315 [#8576] (world catalog).

Type material: Agrilus sugiurai Jendek, 2007 Type locality: Japan, NE Honshu, Miyagi-ken, Shiroishi-shi, Fukuoka-fukaya, Mizubashô-no-mori, 38.5N, 140.32E, alt. 635m (coordinates and altitude were subsequently updated by the original collector). Type specimens: Primary: Holotype (Fig. 53M), ♂, (NSMT): “Mizubasyouno mori, Fukuokafukaya Shiroishi city, Miyagi Prf. Japan, 28.vii.2005, Nobuo Sugiura leg.”. Secondary: 1 PT (EJCB); 1 PT (SOCI), 1 PT (NSCF). Described from 4 specimens (HT, PT). Diagnosis: 5.6-6.5 mm; dorsally bicolored; anterior pronotal lobe obvious, distinctly reaching beyond anterior pronotal angles. Species group: Unassigned. Relevant species: Agrilus koyoi Ohmomo, 2002. Specimens examined: JAPAN: Honshu: 1 ♀ PT (EJCB): “Mizubasyouno mori, Fukuokafukaya Shiroishi city, Miyagi Prf. Japan, 2.viii.2005, Nobuo Sugiura leg.” (JENDEK, 2007 [#8006]); 1 ♀ PT (EJCB): “Mizubasyouno mori, Fukuokafukaya Shiroishi city, Miyagi Prf. Japan, 1.viii.2005, Nobuo Sugiura leg.” (JENDEK, 2007 [#8006]); 1 ♀ PT (EJCB): “Mizubasyouno mori, Fukuokafukaya Shiroishi city, Miyagi Prf. Japan, 2.viii.2005, Nobuo Sugiura leg.” (JENDEK, 2007 [#8006]).

Adult occurence: 7-8 (n=3). Altitude range: 635 m (n=1). Host plants: Unknown. Distribution: JAPAN: Honshu.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Agrilus sulcinotus Jendek Fig. 23B (habitus of HT); Fig. 37V (aedeagus) sulcinotus Jendek, 2007 (Agrilus) JENDEK, 2007: 38 [#8006] (description). [#8576] (world catalog).

BELLAMY, 2008: 2318

Type material: Agrilus sulcinotus Jendek, 2007 Type locality: Myanmar, SW Shan State, Taunggyi. Type specimens: Primary: Holotype (Fig. 23B), ♂, (EJCB): “Burma (Myanmar), SW Shan State, Taunggyi, J. Rejsek 1-18.6.1997”. Secondary: 2 PT (EJCB), 1 PT (ZIN), 1 PT (USNM). Described from 5 specimens (HT, PT). Diagnosis: 6.6-7.9 mm; posterior V-shaped part of elytral pubescence extending almost to apices; aedeagus feebly asymmetrical. Species group: Sulcicollis. Relevant species: A. reichardti Obenberger, 1935. Specimens examined: CHINA: Yunnan: 1 ♀ PT (EJCB): “China,Yunnan, 22.V. 2.VI., 100km W of Kunming, 1993, Diaolin Nat. Reservation, E.Jendek & O.Šauša leg” (JENDEK, 2007 [#8006]); 1 ♂ PT (ZIN): “China, Yunnan, Loc.31, Yundian, 132th km Hway 320 Kunming Xiaguan, 24°59’06.0’’N, 101°43’43.3’’E, 1618 m, 10.06.2002, Konstantinov & Volkovitsh leg.”. MYANMAR: Shan State: 1 ♀ PT (EJCB): “Burma, Shan State, Taunggyi (backyard), N21°20’ E95°59’, 10VII.1952, G.B. Vogt” (JENDEK, 2007 [#8006]).

Adult occurence: 5-6-7 (n=3). Altitude range: ~1618 m (n=1). Host plants: Unknown. Distribution: CHINA: Yunnan. MYANMAR.

Agrilus suturicuspidatus Jendek Fig. 23I (habitus of HT); Fig. 38D (aedeagus) suturicuspidatus Jendek, 2007 (Agrilus) JENDEK, 2007: 40 [#8006] (description). [#8576] (world catalog).

BELLAMY, 2008: 2320

Type material: Agrilus suturicuspidatus Jendek, 2007 Type locality: CH-Guizhou NE, 20 km NW of Jiangkou, Fanjing Shan-Kuaichang. Type specimens: Primary: Holotype (Fig. 23I), ♂, (EJCB): “CH-Guizhou NE, 27.V.-3.VI., 20 km NW of Jiangkou, 1995, Fanjing Shan-Kuaichang”. Secondary: 1 PT (CBCS); 2 PT (SOCI); 3 PT (NMPC); 3 PT (ZIN); 38 PT (EJCB). Described from 48 specimens (HT, PT). Diagnosis: 4.9-7.0 mm; elytral apices with spinate adsutural projection; basal ventrite in male with tubercles. Species group: Sulcicollis. Relevant species: A. marginicollis Saunders, 1873. Specimens examined: CHINA: Anhui: 9 PT (EJCB): “China, SW Anhui, Yuexi Co., Miaodaoshan Mts., 600 1300 m, 18 20.7.[19]95, 30°48 50’ / 116°05’” (JENDEK, 2007 [#8006]). Beijing: 1 PT (EJCB): “China, Beijing distr., Yan Shan, Dongling Mts., 39°58’N, 115°26’E, 1400 m, Xiaolongmen, 15 16.VI.2001” (JENDEK, 2007 [#8006]). Guizhou: 3 PT (EJCB): “CH Guizhou NE, 27.V. 3.VI.,

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA 20 km NW of Jiangkou, 1995, Fanjing Shan Kuaichang” (JENDEK, 2007 [#8006]). Shaanxi: 26 PT (EJCB): “China, S Shaanxi, Qinling Mts. S slope, Xunyangba S+W env., 33°28 37’ / 108°23 33’, 1400 2100 m, 5 9.6.[19]95” (JENDEK, 2007 [#8006]); 1 PT (EJCB): “China Shaanxi, Lüeang, 8.6. 14.6.1996” (JENDEK, 2007 [#8006]); 2 PT (EJCB): “China Yunnan [sic! should be Shaanxi], Lüyeang, 25.7. 30.7.1996” (JENDEK, 2007 [#8006]); 2 PT (EJCB): “China, 1000 1300 m, Shaanxi, Qinling mts., Xunyangba (6km E), 23.v. 13.vi.1998” (JENDEK, 2007 [#8006]); 2 PT (EJCB): “C CHINA, Shaanxi, Qinling, Shan, 6km E of Xunyangba, 1000 1300m, 23.V. 13.VI. 2000” (JENDEK, 2007 [#8006]); 1 PT (EJCB): “C China, Shaanxi, Qinling, Shan, 6km N of Foping, 1150 1300m, 20 21. VI.2000” (JENDEK, 2007 [#8006]).

Adult occurence: 5-6-7 (n=9). Altitude range: 600-2100 m (n=6). Host plants: Unknown. Distribution: CHINA: Anhui, Beijing, Guizhou, Shaanxi.

Agrilus suvorovi Obenberger Fig. 16J (habitus); Fig. 35D (aedeagus); Figs. 43W, 51E, 53N (types) suvorovi Obenberger, 1935 (Agrilus) OBENBERGER, 1935: 165 [#2158] (description). OBENBERGER, 1936: 1053 [#2191] (world catalog). KUROSAWA, 1949: 15 [#5028] (characters; notes). GRECHKIN, 1951: 51 66 [#8606] ([Note: Misidentified as viridis]; larva; biology; life cycle; parasitoids; USSR: Moscow, Penza, Dnepropetrovsk). OBENBERGER, 1956: 43 47 [#2374] (note). SLIVKINA, 1958: 166 167 [#8660] ([Note: Misidentified as viridis according to host plant cited]; notes; north Kazakhstan). PORTA, 1959: 197 [#2488] (notes). KHARAZISHVILI, 1960: 147 [#8630] ([Note: Misidentified as viridis according to host plant cited]; pest; Georgia). SCHAEFER, 1961: 88 90 [#2574] (note). GRECHKIN & VORONCOV, 1962: 57 60 [#8242] ([Note: Misidentified as viridis according to host plant cited]; biology). KUROSAWA, 1963: 153 [#1618] (characters; Japan). POCHON, 1966: 68 [#3250] (note). POCHON, 1967: 133 [#2473] (faunal record; Mongolia). KUROSAWA, 1974: 2 [#4577] (notes). ALEXEEV, 1975: 157 [#52] (notes; Mongolia). KUROSAWA, 1975: 1 2 [#4591] (note). LOMPE, 1979: 239 [#3628] (characters in key; Central Europe). LUNDBERG, 1980: 96 [#6031] (faunal records; Sweden). LUNDBERG, 1980: 92 [#7208] (faunal records; Sweden). LOHSE, 1984: 290 [#6515] (notes; Germany). ANONYMOUS, 1986: 177 [#8848] (insect pest; Korea). ALEXEEV, 1989: 488 [#64] (synonym of vernadskii). ALEXEEV & VOLKOVITSH, 1989: 321 [#3241] (synonym of vernadskii; type examination). WANG, et al., 1991: 232, 233 [#6398] (cited as surorovi; hymenopteran parasites: Entedon epicharis sp. nov. and E. zanara Walker (Eulophidae); China: Jilin). LI JINGKE, 1992: 92 [#6755] (checklist; northeastern China). NONNAIZAB et al., 1999: 113 [#8247] (checklist; China: Inner Mongolia). JENDEK, 2002: 2 6 [#5392] (valid species; taxonomic and nomenclatural notes; distribution; Russia: Stavropol’sk, Primorsk; China: Shanxi). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2320 2321 [#8576] (subgenus Agrilus; world catalog). VOLKOVITSH, 2009: 134 [#8850] (subgenus Agrilus; checklist; Russia: Primorskiy kray). populneus Schaefer, 1946 (Agrilus; variety of viridis; cited as populnea) SCHAEFER, 1946: 73 [#3288] (description). SCHAEFER, 1949: 397, 399, 400, 491 [#2565] (variety of viridis; cited as populnea; characters). HORION, 1955: 94 [#1285] (variety of viridis;

201 cited as populnea; notes). OBENBERGER, 1956: 44, 46 [#2374] (synonym of suvorovi). PORTA, 1959: 197 [#2488] (synonym of suvorovi; notes). SCHAEFER, 1961: 88 90 [#2574] (subspecies of suvorovi). LOMPE, 1979: 239 [#3628] (synonym of suvorovi). BÍLÝ, 1982: 64, 81 82 [#295] (subspecies of suvorovi; characters in key; distribution overview; biology). GOBBI, 1983: 39 [#1098] (subspecies of suvorovi). CLAYHILLS, 1984: 200 [#4185] (subspecies of suvorovi; faunal record; Finland). COBOS, 1986: 251 253, 317 [#609] (subgenus Agrilus; subspecies of suvorovi; cited as populnea; characters in key). CLAYHILLS, 1988: 151 [#4234] (subspecies of suvorovi; faunal record; Finland). SPARACIO, 1990: 74 75 [#2652] (subspecies of suvorovi; biology; Italy (including Sardinia, Sicilia)). SZAFRANSKI, 1991: 283 285 [#4291] (remarks on taxonomic status; faunal records; Poland). NIEHUIS & TEZCAN, 1993: 36, 48 [#1910] (subgenus Agrilus; faunal records from Turkey; distribution and host plants summary; Turkey). CURLETTI, 1994: 22, 175 176 [#658] (subgenus Agrilus; subspecies of suvorovi; catalog; faunal records; Italia; Sicilia; Corsica; Sardegna). BÍLÝ, 2002: 23 [#5877] (bionomy; host plants; distribution). BRECHTEL & KOSTENBADER, 2002: 280 288 [#5766] (biology; ecology; distribution; parasitoids; predators; forestry; Germany). JENDEK, 2002: 2, 3, 4, 5 [#5392] (synonym of suvorovi; lectotype designation; taxonomic notes). KARAS & KLETEČKA, 2002: 90 [#5869] (faunal records; Czech republic). ALLEGRO, 2004: 1 2 [#6799] (subspecies of suvorovi; life cycle; Italy). KENIS & HILSZCZANSKI, 2004: 489 492 [#8566] (natural enemies). BERNHARD et al., 2005: 601, 602 [#7261] (mitochondrial DNA; molecular systematics). JENDEK, 2006: 402 [#7441] (synonym of suvorovi; Palaearctic catalog). BELLAMY, 2008: 2320 [#8576] (synonym of suvorovi; world catalog). brussae Obenberger, 1956 (Agrilus; subspecies of suvorovi) OBENBERGER, 1956: 45, 47 [#2374] (description). COBOS, 1986: 252, 317 [#609] (subgenus Agrilus; subspecies of suvorovi). JENDEK, 2002: 2, 4, 5 [#5392] (synonym of suvorovi; lectotype designation; taxonomic notes). JENDEK, 2006: 402 [#7441] (synonym of suvorovi; Palaearctic catalog). BELLAMY, 2008: 2320 [#8576] (synonym of suvorovi; world catalog).

Type material: Agrilus suvorovi Obenberger, 1935 Type locality: Ussuri. Type specimens: Primary: Lectotype (Fig. 53N), ♀, designated by SCHAEFER (1961: 88-90) [#2574], (NMPC): “Ussuri Nizhne Mikhailovskaya 1899 G. Suvorov [leg.] [p] [in Russian] \ Typus [p] [red label] \ Mus. Nat. Pragae Inv. [p] 24308 [h] [orange label] \ Agrilus Suvorovi m. Type [h] Det. Dr. Obenberger [p]”. Secondary: 2 PLT (ZIN); 3 PLT (NMPC). Described from unknown number of syntypes. Agrilus viridis variety populneus Schaefer, 1946 Type locality: Bouches-du Rhône, Rhône, Ain, BassésPyrénées. Type specimens: Primary: Lectotype (Fig. 51E), ♂, designated by JENDEK (2002: 2, 3, 4, 5) [#5392], (MNHN): “Pollionnay (Rh) 23. 5. [19]44 [h] L. Schaefer [p] \ d/Populus tremula [h] \ [male sign] [h] \ ssp. populneus [h] \ Type [p] [red label]”. Described from unknown number of syntypes. Agrilus suvorovi brussae Obenberger, 1956 Type locality: Asia Minor: Olympus prope Brussam. Type specimens: Primary: Lectotype (Fig. 43W), ♂, designated by JENDEK (2002: 2, 4, 5) [#5392], (NMPC): “Olymp ad Brussa, Coll Dr. Jureček et H. Jurečková [h] \ Typus [p] [red label] \ Agrilus suvorovi ssp. Brussae m. type [h] Det. Dr. Obenberger [p]”. Described from unknown

202 number of syntypes. Diagnosis: 6.8-10.5 mm; body large; eyes moderate or large; pronotum widest at anterior margin with sides almost straight in posterior half; apical third of elytra markedly protracted with obviously elongate apices. Species group: Viridis. Relevant species: A. viridis (Linné, 1758). Specimens examined: CHINA: Shanxi: 1 (EJCB): “China, W Shanxi, 9.VI.2000, 37.7N, 111.6E, Lüliang Shan, road Fangshan Jiaocheng, Hengjian env., ~1000m, Jaroslav Turna leg.” (JENDEK, 2002 [#5392]). KAZAKHSTAN: 1 ♂ (EJCB): “Kazach., 26.7.1976, Karkalinsk, Vasilenko leg.”; 1 ♀ (EJCB): “Kazach., 4.7.1979, Karaganda, Aktas, Jemosev leg.”; 1 ♀ (MDCM): “Kazakhstan Esil 300 m, 12.6.2001, M. Danilevski”. MONGOLIA: 1 (ZIN): “Mongoliya, Vostochnyi aimak, r. Numregin Gol, 32 km YuV g. Salkhit, M. Kerzhner, 8.vii.1976 [in Russian]”; 2 (EJCB): “Mongolia, 14 20.July.2003, 20km E Terelzh, 60km NE of Ulaanbaatar, Tuul riv.vall., Terelzh Nat. Park, cca 1200 1400m, Zd. Jindra lgt.”; 2 ♀ (EJCB): “Mongolia, 20.vi.2003, 20km E Terelzh, 60km NE of Ulaanbaatar, Tuul riv.valley Terelzh N.P., Z. Jindra lgt., 1200 1400m”. RUSSIA: Amurskaya oblast’: 2 (MNCA): “Asia Amur, reg. Archara, Oblucie env., 28.6 15.7.1991”; 3 (EJCB): “USSR, Siberia, Amurska oblast, Kundur env., 1.7. 10.7.1991, J. Dalihod lgt.”. Kemerovskaya oblast’: 1 (MDCM): “Novokuznetsk Kuzedeevo 350 m, 22.6.1994, M. Danilevski”. Khabarovskiy kray: 3 (ZIN): “Bass. Ussuri, Bikinskii r n, Birskoe, 1.7.1958 Kabakov [in Russian]”; 1 (EJCB): “USSR, 7.7.1981, Chabarovsk, lgt. Mejzlík”. Leningradskaya oblast’: 1 (FACF): “Russia, St. Petersburg, Ivanovskoe, 2.vi.1999”. Orenburgskaya oblast’: 1 (VNCO): “Orenburgskaya obl., 4 km yuzh. r/c Tashpla, s. Trudovoe, Nemkov V.A. leg. [in Russian] 12.6.2008”. Primorskiy kray: 1 ♂ (ZIN): “Priamor’e Pivan’ 20.6.1976 Kabakov [in Russian]”; 1 ♀ (JMCZ): “Yu. Sikhotè Alin’, okr. Sokolchi 12.vi.1980, S. Murzin [in Russian]”; 1 (MNHE): “Russia, 27 29.06.1989, Primorye Distr., Buyanki, leg. U. Roosileht”; 4 (EJCB): “Russia: Primorskiy Kray: Yasnoe, VII.1989” (JENDEK, 2002 [#5392]); 11 (EJCB): “USSR, Ussuri prov. Arsenev env., VII.1990, leg. B. Šiška & R. Čermák”; 1 (EJCB): “Russia: Primorskiy Kray: Sokol’chi, VII.1990” (JENDEK, 2002 [#5392]); 1 ♂ (JMCZ): “SU Far East, Kamenushka, 20.6.1991”; 4 (EJCB): “Russia, Primorsk. prov., Arsenev env., VII.1992, B. Šiska leg.” (JENDEK, 2002 [#5392]); 1 (EJCB): “Sib. or., Primorje, Komarovka flum., Kamenushka env., 300 m, Vořišek lgt., VII.1992” (JENDEK, 2002 [#5392]); 1 (EJCB): “Russia: Primorskij k., Rjazanovka, 4 8.VII.1993, Z. Kletečka leg.” (JENDEK, 2002 [#5392]); 5 (EJCB): “E Russia, S Primorje, Gordeyevka vill., 11.VI.1997” (JENDEK, 2002 [#5392]); 12 (EJCB): “Russia, Primorskij kraj, Černigovka env., Sinnij chrebet Mts., 14 16.vi.1997, leg. L. Roller, L. Vidlička” (JENDEK, 2002 [#5392]); 6 (EJCB): “Russia Primorskij Kraj, Gribnoye env., Sinnij chrebet Mts., 17 19.VI.1997, leg. L. Roller & L. Vidlička” (JENDEK, 2002 [#5392]); 1 (EJCB): “Russia Far East, 20km SE Ussuriysk, Gornotaezhnoje vill. env., 30.vi.2003, lg. A. Napolov”; 1 (EJCB): “Russia Far East, 20km SE Ussuriysk, Gorno Tajozhnoje vill. env., 1.vii.2003, lg. A. Napolov”. Saratovskaya oblast’: 1 (MSCI): “Saratov area, Novoburassky distr., Burasy station env., on a leafs of Ulmus 21.06.08 leg. Smirnov”. Stavropol’skiy kray: 3 (EJCB): “Russia, Stavropol’skiy Kray, Stepnoj, 10 18.VI.1954, reared from Populus” (JENDEK, 2002 [#5392]).

Adult occurence: 6-7 (n=35). Faunal records published by KLETEČKA, 1998 [#4416] (as populneus) are also included. Altitude range: 300-1400 m (n=6). Host plants: Populus: SLIVKINA, 1958 (as viridis) [#8660]; KHARAZISHVILI, 1960 (as viridis) [#8630]; GRECHKIN & VORONCOV, 1962 (as viridis) [#8242]; NIEHUIS, 1988 (as

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA populneus) [#1912]; CURLETTI, 1994 (as populneus) [#658]. – P. alba: GRECHKIN, 1951 (as viridis) [#8606]; HORION, 1955 (as populneus) [#1285]; KLETECKA, 1998 (as populneus) [#4416]; BÍLÝ, 2002 (as populneus) [#5877]; BRECHTEL & KOSTENBADER, 2002 (as populneus) [#5766]; ALLEGRO, 2004 (as populneus) [#6799]. – P. bachofenii : GRECHKIN, 1951 (as viridis) [#8606]. – P. balsamifera: GRECHKIN, 1951 (as viridis) [#8606]. – P. bolleana: GRECHKIN, 1951 (as viridis) [#8606]. – P. canadensis: GRECHKIN, 1951 (as viridis) [#8606]; BÍLÝ, 2002 (as populneus) [#5877]; ALLEGRO, 2004 (as populneus) [#6799]. – P. deltoides: BÍLÝ, 2002 (as populneus) [#5877]. – P. densa: GRECHKIN, 1951 (as viridis) [#8606]. – P. nigra: GRECHKIN, 1951 (as viridis) [#8606]; HORION, 1955 (as populneus) [#1285]; BÍLÝ, 2002 (as populneus) [#5877]. – Populus nigra italica: SPARACIO, 1990 (as populneus) [#2652]. – P. sauveolens: GRECHKIN, 1951 (as viridis) [#8606]. – P. tremula: GRECHKIN, 1951 (as viridis) [#8606]; HORION, 1955 (as populneus) [#1285]; SZAFRANSKI, 1991 (as populneus) [#4291]; KRÓLIK & MAJEWSKI, 1994 (as populneus) [#3471]; BÍLÝ, 2002 (as populneus) [#5877]; BRECHTEL & KOSTENBADER, 2002 (as populneus) [#5766]. Distribution: Local: CHINA: Heilongjiang, Jilin, Nei Mongol, Shaanxi, Shanxi. KOREA NORTH. KO-REA SOUTH. MONGOLIA. RUSSIA: Amurskaya oblast’, Buryatiya Respublika, Khabarovskiy kray, Primorskiy kray, Zabaykalskiy kray. Global: ASIA: CHINA. GEORGIA. KAZAKHSTAN. KOREA NORTH. KOREA SOUTH. MONGOLIA. RUSSIA. TURKEY. EUROPE: AUSTRIA. BELARUS. BOSNIA & HERZEGOVINA. BULGARIA. CROATIA. CZECH REPUBLIC. FINLAND. FRANCE. GERMANY. GREECE. HUNGARY. ITALY. LATVIA. LIECHTENSTEIN. MACEDONIA. NORWAY. POLAND. ROMANIA. RUSSIA. SERBIA & MONTENEGRO. SLOVAKIA. SPAIN. SWEDEN. SWITZERLAND. TURKEY. UKRAINE.

Agrilus tai Jendek, sp. nov. Fig. 28L (habitus of HT); Fig. 40P (aedeagus)

Description: Holotype 5.4 mm. BODY: Shape: cuneiform, Build: robust, Color (dorsally): unicolored, Color (dominant): green. HEAD: Medial impression: absent, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline, Microsculpture: present. Vertex: Outline: not protruding from head outline, Sculpture on vertex: punctures, Sculpture (intensity): superficial. Eyes: Size: large, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: short. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: obvious, Shape: arcuate or subangulate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial. Prehumerus: carinal, Shape: bisinuate,

203

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Extent: beyond half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate, Convergence: strongly convergent, Junction: absent, Narrowest point: at posterior third of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: conjoint, Width: narrow, Shape: arcuate or subtruncate, Truncation: transverse. Elytral pubescence: entire, Color: unicolored. Tomentum: absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: narrow, Anterior margin: arcuately emarginate, Emargination (deepness): shallow, Emargination (width): narrow. Prosternal process: narrow, Shape: subparallel, Sides: straight, Angles: rectangular, Angles (tips): sharp, Disk: impressed. ABDOMEN: Sexual modifications in male: impression at intercoxal projection, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate, Sinuosity (deepness): very shallow, Sinuosity (width): very narrow. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe sharply pointed. Diagnosis: prosternal lobe finely arcuately emarginate on anterior margin; prosternal process very narrow, subparallel and deeply impressed. Species group: Viridiobscurus. Relevant species: A. pusillesculptus Obenberger, 1940.

Type material: Agrilus takahashii Tôyama, 1988 Type locality: Akasai, Hyôgo Pref., Japan,. Type specimens: Primary: Holotype (Fig. 24K), ♂, (NSMT): “Akasai Hyōgo Pref[ecture]. [p] 24.VI. [h] 19 [p] 79 [h] M. Tōyama leg. [p] \ Holotype [p] Agrilus takahashii Tôyama, 1987 [h]”. Described from 7 specimens (HT, AT, PT). Diagnosis: 4.7-6.5 mm; elytra entirely pubescent except for epipleural portions; basal ventrite in male without tubercles. Species group: Sulcicollis. Relevant species: A. foliatus Jendek, 2007. Specimens examined: Known to us only from the holotype.

Host plants: Carpinus: TÔYAMA, 1988 [#2932]; AKIYAMA & OHMOMO, 1997 [#4091]. – Zelkova serrata: AKIYAMA & OHMOMO, 1997 (cited as rerrata) [#4091]. Distribution: JAPAN: Honshu, Kyushu.

Agrilus tazoei Kurosawa Fig. 31I (habitus); Fig. 41U (aedeagus); Fig. 53P (type) tazoei Kurosawa, 1985 (Agrilus) KUROSAWA, 1985: 156 158 [#1640] (description). TÔYAMA, 1985: 23 [#3678] (iconography; Japan). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 41 [#4091] (checklist; Japan: Honshu, Tsushima). FUKUTOMI & HORI, 2004: 11 [#6851] (checklist; Japan (Hokkaido)). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2324 [#8576] (world catalog).

Distribution: CHINA: Yunnan.

Type material: Agrilus tazoei Kurosawa, 1985 Type locality: Tsuchiyu, Fukushima Pref., Honshu, Japan. Type specimens: Primary: Holotype (Fig. 53P), ♂, (NSMT): “Tsuchiyu, Fukushima Pref. 22. VI. 1974, K. Tazoe lgt. [h] \ Holotype [p] Agrilus tazoei Y. Kurosawa, 1985 [h] [red label with black border]”. Secondary: PT (NSMT). Described from 6 specimens (HT, AT, PT).

Etymology: The specific name is a noun in apposition referring to the Tai ethnicity of ethnic groups of southern China and Southeast Asia, stretching from Hainan to eastern India and from southern Sichuan to Laos, Thailand, and parts of Vietnam.

Diagnosis: 6.6-9.2 mm; body dorsally bicolored, glabrous; eyes large; antennae short; prehumerus arcuately carinal, reaching almost to pronotal half, with anterior end not joining marginal carina. Species group: Unassigned. Relevant species: Unknown.

Type locality: China, Yunnan province, Daju. Type specimens: Holotype (Fig. 28L), ♂, (EJCB): “China, Yunnan, 25-27.VI.1994, Daju, B. Šiška & T. Spevár leg.”. Adult occurence: 6 (n=1). Host plants: Unknown.

Agrilus takahashii Tôyama Fig. 24K (habitus of HT); Fig. 38R (aedeagus) takahashii Tôyama, 1988 (Agrilus) TÔYAMA, 1988: 754 756 [#2932] (description). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 40 [#4091] (checklist; Japan: west Honshu, Kyushu). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2322 [#8576] (world catalog).

Specimens examined: CHINA: Shaanxi: 3 ♂ (EJCB): “C China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000 1300m, 23.V. 13.VI. leg. C. Holzschuh 2000”. JAPAN: Honshu: 1 ♂, 1 ♀ (EJCB): “Takatoriyama, Yokosuka city, Kanagawa pref., 24.vi.2001, T. Hattori leg.”.

Adult occurence: 6-7 (n=7). Faunal records published by KUROSAWA, 1985 [#1640] are also included. Altitude range: 1000-1300 m (n=1). Host plants: Pueraria thunbergiana: AKIYAMA & OHMOMO, 1997 [#4091]. – Wisteria floribunda: KUROSAWA, 1985 [#1640]; AKIYAMA & OHMOMO, 1997 [#4091].

204 Distribution: CHINA: Shaanxi. JAPAN: Hokkaido, Honshu, Tsushima.

Agrilus tempestivus Lewis Fig. 25C (habitus); Fig. 38V (aedeagus); Fig. 53Q (type) tempestivus Lewis, 1893 (Agrilus) LEWIS, 1893: 334 335 [#1722] (description). KERREMANS, 1903: 291 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 21 [#1809] (catalog; Japan). OBENBERGER, 1936: 1053 [#2191] (world catalog). KUROSAWA, 1949: 14 [#5028] (characters). CHÛJÔ & KUROSAWA, 1950: 9 [#501] (catalog; faunal records; Japan (Shikoku)). IGA, 1955: 79 [#3828] (iconography; Japan). IGA, 1962: 79 [#3351] (iconography; Japan). KUROSAWA, 1963: 153 [#1618] (characters; Japan). KUROSAWA, 1974: 1, 2 [#4575] (characters; notes). KUROSAWA, 1975: 2 [#4591] (note). TÔYAMA, 1985: 29 [#3678] (iconography; Japan). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). LI JINGKE, 1992: 93 [#6755] (checklist; China: Liaoning). JENDEK, 1995: 139 [#3768] (lectotype designation; characters; Japan; Russian Far East). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 186 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 41 [#4091] (checklist; Japan: Honshu, Shikoku, Kyushu, Tsushima, Yakushima; Far East Siberia). ALEXEEV, 1998: 379, 380 [#4322] (subgenus Orientagrilus; genotype of the subgenus Orientagrilus). AKIYAMA & OHMOMO, 2000: 88 [#6240] (faunal records; Japan (Hiroshima)). FUKUTOMI & HORI, 2004: 12 [#6851] (checklist; Japan (Hokkaido)). JENDEK, 2006: 391 [#7441] (subgenus Orientagrilus; Palaearctic catalog). ANONYMOUS, 2007: 32 [#8250] (checklist; Japan: Honshu). BELLAMY, 2008: 2325 [#8576] (subgenus Orientagrilus; world catalog). VOLKOVITSH, 2009: 135 [#8850] (subgenus Orientagrilus; faunal records; Russia: Primorskiy kray).

Type material: Agrilus tempestivus Lewis, 1893 Type locality: Main island. Usui-Toge, Fukui, and Chiuzenji. Type specimens: Primary: Lectotype (Fig. 53Q), ♂, designated by JENDEK (1995: 139) [#3768], (BMNH): “Type H.T. [p] [round label with red border] \ Japan G. Lewis 1910320 [p] \ Agrilus tempestivus Lewis Type [h]”. Described from unknown number of syntypes. Diagnosis: 4.9-7.7 mm; pronotum with posterolateral impression; margin of elytral apices with fine denticulation; tubercles on basal ventrite in male rounded. Species group: Tempestivus. Relevant species: A. asiaticus Kerremans, 1898. Specimens examined: CHINA: Hebei: 2 ♂ (EJCB): “China, Hebei/Nei Mongol, pass Chengde Chifeng, 41.6N, 118.2E, 14 16. VI.2001, O. Šafránek leg.”. Henan: 2 ♂, 1 ♀ (EJCB): “China, N. Henan, 11 12.VI., Jiuligou, 550 m, 35º12’N, 112º26’E, Jaroslav Turna leg. 2007”. Hunan: 1 ♂ (EJCB): “China, Hunan SE, 1994, Guidong env. 26 31.V., 26.04N 113.56E, E. Jendek & O. Šauša leg.”. Shaanxi: 2 ♂, 1 ♀ (EJCB): “China, 1000 1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v. 13.vi.1998, I. H. Marshal leg.”; 1 ♂ (EJCB): “China, Shaanxi prov., 21 23.vi.1998, Quin Ling Shan mts, road Baoji Taibai vill., pass 35 km S Baoji, O. Šafránek et M. Trýzna lgt.”. JAPAN: Honshu: 1 (MNHN): “Japan, Tokyo env., 1901, leg. J. Harmand” (JENDEK, 1995 [#3768]); 1 (MNHN):

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA “Sendaigawa, 1906, leg. E. Gallois” (JENDEK, 1995 [#3768]); 1 (MNHN): “Chuzenji 21.VIII.1910, leg. E. Gallois” (JENDEK, 1995 [#3768]); 1 ♀ (JMCZ): “Mt. Gagyu, Okayama pref., Japan, May 14, ‘67, T. Aono leg.”; 4 (EJCB): “Eda, Fukushima Prefecture, 21.VI.1979, leg. S. Ohmomo” (JENDEK, 1995 [#3768]); 2 (JMCZ): “Japan, 28.4.1982, Eda, Iwaki city, Fukushima pref., K. Akiyama lgt.”; 1 ♂ (EJCB): “Mt. Vaibosatsu, Yamanashi, 29.VI.1982, leg. K. Ikeda” (JENDEK, 1995 [#3768]); 1 ♂ (EJCB): “Eda, Iwaki shi, Fukushima Prefecture, 11.VII.1987, leg. S. Ohmomo” (JENDEK, 1995 [#3768]); 1 ♀ (EJCB): “Japan, Honshu, Osaka fu, Kongosan, 300 m, 2 km E of Amami, 12. May 2002, Bolm Leg.”. Kyushu: 1 (MNHN): “Kio Siou” (JENDEK, 1995 [#3768]); 1 (MNHN): “Japan, Kumamoto Prefecture, Kuruizawa, 12.VII.1908, leg. E. Gallois” (JENDEK, 1995 [#3768]); 1 ♂, 1 ♀ (EJCB): “Mt. Hikosan, Soeda machi, Fukuoka Pref., Kyushu, Japan, 15.iv.1990, Ryô Noda leg.”; 1 ♂ (EJCB): “Mt. Oboshi yama, Tsushima Isls., Nagasaki Pref., Kyushu, Japan, 4.v.1993, Ryô Noda leg.”. Tsushima: 1 (NMPC): “Japan, Tsushima is., (Sasuna) 5 7.6.1981, A. Torikata leg.”; 2 (EJCB): “Japan: Tsushima Is., Sasuna, 5 7.VI.1981, leg. A. Torikata” (JENDEK, 1995 [#3768]). RUSSIA: Khabarovskiy kray: 1 ♂ (EJCB): “USSR: Khabarovsk Province, Slavianka 20 km N of Troitskoye, Amur bank, 14 19.VI.1990, leg. W. Schawaller” (JENDEK, 1995 [#3768]). Primorskiy kray: 1 ♀ (EJCB): “SU Ussuri reg., Novovarvarovka, 6 10.7.1989, leg. Dunda” (JENDEK, 1995 [#3768]); 1 ♂ (JMCZ): “Ross. or., Prim. kr., Anisimovka, 3.viii.1989, V. Kuznecov leg.”; 1 ♀ (EJCB): “Sib. or. m., Primorje, Sichote Alin Mts., Sokolči, 1 15.7.1990, Kadlec + Voříšek lg.” (JENDEK, 1995 [#3768]); 1 ♀ (EJCB): “USSR, Primorskij kraj Arseneev env. 300 400 m 11. 19.VII.1990 B. Šiška & R. Čermák leg.”; 7 (EJCB): “USSR, Ussuri prov., Arsenev env., VII.1990, leg. B. Šiška & R. Čermák” (JENDEK, 1995 [#3768]); 1 ♀ (ZIN): “Primorsk. krai, Spassk, les, 13.06.1990, leg. Belokobyl’skij [in Russian]” (JENDEK, 1995 [#3768]); 1 ♀ (EJCB): “Russia, Primorski reg. Vladivostok, 22.06.1991, R. Filimonov leg.”; 1 ♀ (EJCB): “Sib. or., Primorje, Komarovka flum., Kamenushka env., 300 m, Vořišek lgt., VI.1992” (JENDEK, 1995 [#3768]); 1 ♀ (JMCZ): “Russia, Ussuri, Kajmanovka, VIII.1994, Snížek”; 1 ♂, 2 ♀ (EJCB): “Russia E., Primorskyi reg. Artyom town. env., 3 15.VII.1996, Plutenko leg.”; 1 ♀ (EJCB): “Russia Far East, 20 km SE Ussuriysk, Gornotaezhnoje vill. env., 27.06.2003, Lg. A. Napolov”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Tachingousa (Prosiolotchnaja) bay 2 6.07.04 leg. М. & L. Smirnov’s”; 1 ♂ (MNCA): “Russia, Ussuri, Primor’e region, Pedan Mts., 28 29.7.2006, leg. K. Hadulla”; 1 ♂ (MNCA): “Russia, Ussuri, Primor’e region, Pedan Mts., 11 17.7.2006, leg. K. Hadulla”; 1 ♂ (EJCB): “Russia, Ussuri Primor’e region, Pedan Mts., 28 29.7.2006, leg. K. Hadulla”.

Adult occurence: 4-5-6-7-8 (n=33). Altitude range: 3001300 m (n=6). Host plants: Carpinus: AKIYAMA & OHMOMO, 1997 [#4091]. – Machilus thunbergii: AKIYAMA & OHMOMO, 1997 [#4091]. – Quercus: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: CHINA: Hebei, Henan, Hunnan, Liaoning, Nei Mongol, Shaanxi. JAPAN: Hokkaido, Honshu, Kyushu, Ryukyu Islands, Shikoku, Tsushima. RUSSIA: Khabarovskiy kray, Primorskiy kray.

Agrilus terracotta Jendek, sp. nov. Fig. 26J (habitus of HT); Fig. 39K (aedeagus)

Description: 5.9-8.0 mm (Holotype 7.2 mm). BODY: Shape: subparallel, Color (dorsally): unicolored or bicolored, Color

205

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA (dominant): green. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: flat or moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): semispherical, Sculpture (density): dense. Eyes: Size: large, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: visually square, Sides: arcuate, rarely subangulate, Maximal width: at anterior margin or at middle, Anterior margin: wider than posterior or subequal to posterior. Anterior lobe: moderate or obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles or projecting beyond anterior pronotal angles. Posterior angles: Shape: obtuse, Apex: sharp. Disk: Convexity: flat, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial. Prehumerus: carinal, rarely tubercular, Shape: arcuate, Extent: to half of pronotal length, Posterior end: joining posterior pronotal angle or distant from pronotal angle or margin, Anterior end: close to pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: absent or present, Narrowest point: at posterior angles or at posterior 1/5-1/4 of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: bicolored, Alternative color (location): apical portion, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: conjoint, rarely separate, Width: wide, Shape: subtruncate, rarely arcuate, Truncation: transverse. Elytral pubescence: proximal and distal, Proximal: like V or upturned T, Distal: preapical, Color: bicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: flat, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): shallow, Emargination (width): moderate. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: rectangular, Angles (tips): sharp, Disk: impressed. ABDOMEN: Sexual modifications in male: not apparent, rarely medial sulcus on ventrite 1, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate. LEGS: Male sexual modifications: medial posterior angle of metacoxa protruding, metacoxa medially with cluster of setae and lower margin of metafemur with row of hairs. Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): apex of medial lobe markedly obtuse, Ovipositor: elongate. Diagnosis: body more elongate; pronotum visually quadrate; distal part of elytral pubescence wide; medial lobe of aedeagus long and narrow. Species group: Cyaneovirens. Relevant species: A. pterostigma Obenberger, 1927.

Type locality: China, Shaanxi province, Qinling mts., 6 km east of Xunyangba, altitude 1000-1300 m. Type specimens: Primary: Holotype (Fig. 26J), ♂, (EJCB): “China, 1000-1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.-13.vi.1998, I. H. Marshal leg.”. Secondary: CHINA: Shaanxi: 1 PT (EJCB): “China S, Shaanxi, 14-16.6.1995, Qinling Mts. N. slope, 33.56-59’/108.50’ 800-1200 m, Changan Co., L.+R. Businský lgt.”; 1 ♀ PT (EJCB): “C-China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000-1300m, 23.V.-13.VI. leg. C. Holzschuh 2000”; Zhejiang: 1 ♀ PT (MNHN): “11.vii.[19]36 O. Piel, coll., T’ienmu-Shan”; INDIA: Assam: 1 ♂ PT (EJCB): “NE India, Assam, 1999, 5 km N of Umrongso, 700m, 25°27’N, 92°43’E, 17.-25.v., Dembický & Pacholátko leg.”; West Bengal: 1 ♀ PT (EJCB): “Kalimpong, ix.1985, N. Dangal leg.”; VIETNAM: Lang Son: 6 PT (EJCB): “N Vietnam, 21°35N, 106°30E, 52 km SW of Lang Son, 27.iv.-6.v.1996, 370 m, Dembický & Pacholátko leg.”. Adult occurence: 4-5-6-7-9 (n=7). Altitude range: 3701300 m (n=5). Host plants: Unknown. Distribution: CHINA: Shaanxi, Zhejiang. INDIA: Assam, West Bengal. VIETNAM: Lang Son. Etymology: The specific name referes to the ceramic terracotta used as the embellishment of the Terracotta Army, one of the most remarkable historical attractions of the Shaanxi province.

Agrilus thibetanus Obenberger Fig. 19D (habitus); Fig. 36E (aedeagus); Fig. 53T (type) thibetanus Obenberger, 1928 (Agrilus) OBENBERGER, 1928: 70 [#2075] (description). OBENBERGER, 1936: 1054 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 359 [#2429] (checklist; China). JENDEK, 2000: 506 [#4851] (lectotype designation). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Taiwan). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2327 [#8576] (world catalog). JENDEK & GREBENNIKOV, 2009: 45, 52 [#8561] (characters in key; diagnosis).

Type material: Agrilus thibetanus Obenberger, 1928 Type locality: Vrionatong, Thibet. Type specimens: Primary: Lectotype (Fig. 53T), ♂, designated by JENDEK (2000: 506) [#4851], (NMPC): “Vrionatong Thibet LeMoult [p] \ Typus [p] [red label] \ Agrilus thibetanus m. Type [h] Det. Dr. Obenberger [p]”. Secondary: 2 PLT (NMPC). Described from unknown number of syntypes. Diagnosis: 6.5-7.3 mm; prosternal process in male enlarged; medial lobe of aedeagus with dentate sides and more acute apex. Species group: Cyanescens. Relevant species: A. cyanescens (Ratzeburg, 1837). Specimens examined: CHINA: Xizang: 1 ♂, 3 ♀ (MNHN): “Thibet, Vrionatong” (JENDEK & GREBENNIKOV, 2009 [#8561]).

Host plants: Unknown.

206 Distribution: CHINA: Xizang.

Agrilus tokyoensis Kurosawa Fig. 10A (habitus); Fig. 32P (aedeagus); Fig. 53V (type) tokyoensis Kurosawa, 1985 (Agrilus) KUROSAWA, 1985: 151 153 [#1640] (description). TÔYAMA, 1985: 24 [#3678] (iconography; Japan). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 41 [#4091] (checklist; Japan: Honshu, Kyushu). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2330 [#8576] (world catalog).

Type material: Agrilus tokyoensis Kurosawa, 1985 Type locality: Hinoharu, Yamanashi Pref., Honshu, Japan. Type specimens: Primary: Holotype (Fig. 53V), ♂, (NSMT): “Hinoharu Kitahoma-gun Yamanashi Pref. [p] 10 [h] - VII - 198 [p] I [h] A. Seki leg. [p] \ Holotype [p] Agrilus tokyoensis Y. Kurosawa, 1985 [h] [red label with black border]”. Secondary: 2 PT (NSMT). Described from 8 specimens (HT, AT, PT).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Type material: Agrilus tonkineus Kerremans, 1895 Type locality: Hanoï. Type specimens: Primary: Lectotype (Fig. 53W), ♀, designated by DESCARPENTRIES & VILLIERS (1963: 105, 109) [#700], (BMNH): “Type [p] [round label with red border] \ Hanoi, Donckier [h] \ tonkineus Kerr. Type [Kerremans’ MS] \ Kerremans 1903-59 [p] \ A. tonkineus Kerrem. Ton Kin [h]”. Described from unknown number of syntypes. Agrilus blatteiceps Bourgoin, 1925 Type locality: Laos: Vien Poukha. Type specimens: Holotype by monotypy (Fig. 43O), ♀, (MNHN): “Laos [h] Cochinchine [p] [strikethrough] Vien Pokha [h] le [p] 11-v-[h] 19 [p]18 [h] R. Vitalis de Salvaza [p] \ Agrilus blatteiceps Bourgoin [h] [black ink] Type ♂ [h] [red ink] \ tonkineus Kerr. comp. an Type [h] \ Muséum Paris Coll. Générale [yeallow label]”. Described from 1 specimen. Diagnosis: 7.5-9.5 mm; proximal part of elytral pubescence mosaic; elytral apices broadly arcuate. Species group: Occipitalis. Relevant species: A. auriventris Saunders, 1873

Diagnosis: 6.1-10.2 mm; head without medial impression; pronotum strongly convex with obvious anterior lobe. Species group: Spinipennis. Relevant species: A. ashlocki Baudon, 1968.

Specimens examined: CHINA: Yunnan: 1 ♀ (NMPC): “Yun’nan’, 30 km SW Czin’lina, 420 m, 26.IV.1956 Khuan Kèzhen’ i dr. [in Russian]”. VIETNAM: 2 ♂, 1 ♀ (MNHN): “Tonkin, Dap Can, Col. Sicard”.

Specimens examined: JAPAN: Honshu: 1 ♂ (EJCB): “1991.7.29., Hosaka, Nirasaki, Yamanashi, C Jpn, T. Kosaka leg.”. LAOS: Houaphan: 7 ♂, 3 ♀ (EJCB): “Laos NE, Houa Phan prov., 20°12 13.5’N, 103°59’.5 104°01’E, Ban Saluei → Phou Pane Mt., 1340 1870m, 15.iv. 15.v.2008, Lao collectors leg.”.

Adult occurence: 4-5-6 (n=3). Faunal records published by BAUDON, 1962 [#145] and 1963 [#148] are also included. Altitude range: ~420 m (n=1).

Adult occurence: 4-5-6-7-8 (n=5). Faunal records published by KUROSAWA, 1985 [#1640] are also included. Altitude range: 1340-1870 m (n=1).

Distribution: CHINA: Yunnan. LAOS. VIETNAM.

Host plants: Quercus acutissima: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: JAPAN: Honshu, Kyushu. LAOS.

Agrilus tonkineus Kerremans Fig. 19K (habitus); Fig. 36L (aedeagus); Figs. 43O, 53W (types) tonkineus Kerremans, 1895 (Agrilus) KERREMANS, 1895: 222 223 [#1416] (description). KERREMANS, 1903: 277 [#1441] (catalog). OBENBERGER, 1936: 1105 [#2191] (world catalog). BAUDON, 1963: 53 [#148] (faunal record; Laos). DESCARPENTRIES & VILLIERS, 1963: 105, 109 [#700] (lectotype designation: “Hanoi (type de tonkineus, British Museum)”; characters; faunal records; Tonkin; Laos). BAUDON, 1968: 130, 143 [#162] (note). KUROSAWA, 1974: 3 [#4575] (characters; notes). BELLAMY, 2008: 2330 [#8576] (world catalog). blatteiceps Bourgoin, 1925 (Agrilus) BOURGOIN, 1925: 131 [#373] (description). THÉRY, 1935: 15 [#2817] (synonym of tonkineus). OBENBERGER, 1936: 1105 [#2191] (synonym of tonkineus). BAUDON, 1962: 69 [#145] (faunal record; Laos). DESCARPENTRIES & VILLIERS, 1963: 109 [#700] (synonym of tonkineus). BELLAMY, 2008: 2321 [#8576] (synonym of tonkineus; world catalog).

Host plants: Unknown.

Agrilus trinotatus Saunders Fig. 23G (habitus); Fig. 38B (aedeagus); Fig. 54A (type) trinotatus Saunders, 1873 (Agrilus) SAUNDERS, 1873: 513 514 [#2554] (description). LEWIS, 1879: 15 [#3345] (catalog; Japan). KERREMANS, 1885: 154 [#1381] (catalog). SCHÖNFELDT, 1887: 113 [#8901] (Anambus; catalog; Japan). KERREMANS, 1892: 276 [#1397] (catalog). KERREMANS, 1903: 291 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 21 [#1809] (catalog; Japan). OBENBERGER, 1936: 1054 [#2191] (world catalog). KUROSAWA, 1949: 15 [#5028] (characters). CHÛJÔ & KUROSAWA, 1950: 9 [#501] (catalog; faunal records; Japan (Shikoku)). KUROSAWA, 1963: 153 [#1618] (characters; Japan). KUROSAWA, 1975: 2 [#4591] (note). TÔYAMA, 1985: 29 [#3678] (iconography; Japan). AKIYAMA, 1986: 25 [#3199] (faunal records; Japan: Kanagawa prefecture). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 232 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 186 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 41 42 [#4091] (checklist; Japan: Honshu, Shikoku, Kyushu). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2334 [#8576] (world catalog).

Type material: Agrilus trinotatus Saunders, 1873

207

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 54A) by present designation, ♀, (BMNH): “Type H. T. [p] [round label with red border] \ Japan, G. Lewis, 1910-320 [p] \ Agrilus trinotatus E. S Type [h]”. Described from unknown number of syntypes. Diagnosis: 4.-6.3 mm; body more flat dorsally; apical third of elytra more attenuate; distal part of pubescence very limited. Species group: Sulcicollis. Relevant species: A. sulcinotus Jendek, 2007. Specimens examined: JAPAN: Honshu: 1 (JMCZ): “Mt. Gragyu, Okayama pref. Japan, May 14, ‘67, T. Aono leg.”; 1 (EJCB): “Ishidojyuku, Kitamoto City, Saitama Pref., Japan, 13.v.1986”; 6 (EJCB): “1998.4.29, Mt. Atago, Iwama, Ibaraki, C Japan, S. Ohmomo”.

Adult occurence: 4-5 (n=3). Host plants: Quercus: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: JAPAN: Honshu, Kyushu, Shikoku.

Agrilus truncatus Jendek Fig. 24C (habitus of HT); Fig. 38J (aedeagus) truncatus Jendek, 2007 (Agrilus) JENDEK, 2007: 39 [#8006] (description). [#8576] (world catalog).

BELLAMY, 2008: 2335

Type material: Agrilus truncatus Jendek, 2007 Type locality: China, Shanxi, Lüliang Shan, 1000 m, road Fangshan-Jiaocheng, Hengjian env. Type specimens: Primary: Holotype (Fig. 24C), ♂, (EJCB): “China, Shanxi, 9.VI.2000, Lüliang Shan, 1000 m, road Fangshan-Jiaocheng, Hengjian env.”. Secondary: 1 ♂ PT (SOCI); 7 PT (EJCB). Described from 9 specimens (HT, PT). Diagnosis: 4.3-5.2 mm; pronotum with deep lateral impressions; elytral apices slender; aedeagus symmetric. Species group: Sulcicollis. Relevant species: A. fissus Obenberger, 1917. Specimens examined: CHINA: Beijing: 1 ♂ PT (EJCB): “China, Beijing distr., Yan Shan, Dongling Mts., 39°58’N, 115°26’E, 1400 m, Xiaolongmen, 15 16.VI.2001”; 1 ♂ PT (EJCB): “China: Beijing Reg., ca 1150 m, Xiaolongmen, 9 12.VI.2004, N39°58.074’, E115°25.882’, F.I.Trap, walnut / poplar woodland”. Shanxi: 3 PT (EJCB): “China, Shanxi, 9.VI.2000, Lüliang Shan, 1000 m, road Fangshan Jiaocheng, Hengjian env.” (JENDEK, 2007 [#8006]); 2 ♂ PT, 1 ♀ PT (EJCB): “China, W Shanxi, 9.VI.2000, 37.7N, 111.6E, Lüliang Shan, road Fangshan Jiaocheng, Hengjian env., ~1000m, Jaroslav Turna leg.”.

Adult occurence: 6 (n=4). Altitude range: 1000-1400 m (n=4). Host plants: Unknown. Distribution: CHINA: Beijing, Shanxi.

Agrilus tschitscherini Semenov Fig. 17B (habitus); Fig. 35G (aedeagus); Figs. 43C, 48O, 46Q, 48S, 54C, 54J, (types) tschitscherini Semenov, 1895 (Agrilus) SEMENOV, 1895: 135 137 [#3525] (description). HEYDEN, 1896: 57 [#5552] (catalog). ABEILLE DE PERRIN, 1897: 7 [#19] (characters in key). ESCHERICH, 1897: 26 [#8345] (faunal records; Turkey). SEMENOV, 1900: 601 [#2620] (note). KERREMANS, 1903: 292 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 658 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1053 [#2191] (world catalog). MAKHNOVSKIJ, 1958: 127 [#6090] ([Note: Misidentification according to the host plant cited]; notes; Uzbekistan). MAKHNOVSKIJ, 1959: 32 [#5829] ([Note: Misidentification of pistaciophagus according to MAKHNOVSKIJ, 1966: 80 [#7356]]; characters; biology; Uzbekistan). KULINITSH, 1963: 154 [#8648] (ecological notes; Tajikistan). KOSTIN, 1973: 108 [#3693] (characters in key; Kazakhstan). GURYEVA, 1974: 100 [#5428] (notes). KADYROV, 1976: 51 [#3461] (biology; Tajikistan). YAGDYEV, 1978: 90 [#4206] (faunal record; biology; Kazakhstan). JENDEK, 1993: 38 [#1354] (lectotype designation). ALEXEEV, 1998: 381 [#4322] (subgenus Agrilus; subgeneric classification). TLEPPAEVA, 1999: 184 [#6551] (cited as tschitsherini; notes; Kazakhstan). TLEPPAEVA & ISHKOV, 2004: 84 [#7211] (subgenus Agrilus; checklist; Kazakhstan). JENDEK, 2006: 389 [#7441] (subgenus Agrilus; Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2336 [#8576] (subgenus Agrilus; world catalog). attila Obenberger, 1930 (Agrilus) syn. nov. OBENBERGER, 1930: 111 [#2088] (description). ALEXEEV, 1989: 488 [#64] (cited as atilla; key to species; Chita; Buryatia; north Mongolia). ALEXEEV & VOLKOVITSH, 1989: 320, 349, 357 [#3241] (cited as atilla; characters in key; faunal records; notes; Mongolia). ALEXEEV, 1998: 381 [#4322] (subgenus Agrilus; cited as atilla; subgeneric classification). JENDEK, 2006: 388 [#7441] (subgenus Agrilus; Palaearctic catalog). BELLAMY, 2008: 1981 [#8576] (subgenus Agrilus; world catalog). lubischevi Stepanov, 1958 (Agrilus) syn. nov. STEPANOV, 1958: 165 166, 172 [#3664] (description). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2170 [#8576] (world catalog). uzbekistanus Stepanov, 1958 (Agrilus) syn. nov. STEPANOV, 1958: 166 168, 172 [#3664] (description). MAKHNOVSKIJ, 1958: 127 [#6090] (notes; Uzbekistan). MAKHNOVSKIJ, 1959: 32 [#5829] (notes; Uzbekistan). SINADSKIJ, 1963: 48 [#5038] (notes; biology). KOSTIN, 1973: 107 [#3693] (synonym of aurichalceus; characters in key; Kazakhstan). LINDEMAN & TURUNDAEVSKAYA, 1975: 327 [#3478] (bionomy; v Zapadnom Kazakhstane). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). LÖBL & SMETANA, 2007: 30 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2342 [#8576] (world catalog). gerschuni Stepanov, 1958 (Agrilus) syn. nov. STEPANOV, 1958: 168 170, 172 [#3664] (description). JENDEK, 2006: 398 [#7441] (Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2107 [#8576] (world catalog). machnovskii Stepanov, 1958 (Agrilus) syn. nov. STEPANOV, 1958: 170 171, 172 [#3664] (description). MAKHNOVSKIJ, 1958: 127 [#6090] (cited as machnovski; notes; Uzbekistan). MAKHNOVSKIJ, 1959: 32 [#5829] (notes; Uzbekistan). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2173 [#8576] (world catalog).

208 Type material: Agrilus tschitscherini Semenov Tian Shanskij, 1895 Type locality: Turkestan occid.: ad lac. Iskander-kul. Type specimens: Primary: Lectotype (Fig. 54C), ♀, designated by JENDEK (1993: 38) [#1354], (ZIN): “IskanderKul Glasunov 1892 [p] \ Agrilus Tschitscherini m Typ I. [18]95 [h] A. Semenov det. [p]”. Secondary: 1 PLT (ZIN). Described from 2 syntypes. Agrilus attila Obenberger, 1930 Type locality: Mongolia: Tunigol [Note: locality emended to “Tuin Gol river” by ALEXEEV & VOLKOVICH, 1989 [# 3241]]. Type specimens: Primary: Lectotype (Fig. 43C), ♂, by present designation, (NMPC): “Mongolia Tunigol [p] \ Typus [p] [red label] \ Agrilus Attila m. Type [h] Det. Dr. Obenberger [p]”. Secondary: 1 PLT (NMPC). Described from unknown number of syntypes. Agrilus lubischevi Stepanov, 1958 Type locality: v okresnostyakh g. Frunze Frunzenskoi oblasti Kirgizskoi SSR [in Russian]. Type specimens: Holotype by monotypy (Fig. 48O), (ZIN): “Frunzensk. ob. okr. g. Frunze 13-VI-46 A. Lyubishev leg. [h] [in Russian] \ Typus [h] [red labe] \ Agrilus lubischevi m. sp. n. 52 [h] V. Stepanov det. [p] \ Ris. 215 [h]”. Described from 1 specimen. Agrilus uzbekistanus Stepanov, 1958 Type locality: v Gorno-lesnom gosudarstvennom zapovednike Parkentskogo raiona Tashkentskoi oblasti, Uzbekskoi SSR, urochishche Shavazi-Kolon-saya [in Russian]. Type specimens: Primary: Lectotype (Fig. 54J) by present designation, ♂, (ZIN): “Uzbeksk. SSR. Tashkentsk. o. Parkentsk. r. 17-VI-51 g I. K. Makhnovskii [h] [in Russian] \ N9 Agrilus sp. na ive [h] [in Russian] Typus [h] [red labe] \ Agrilus uzbekistanus m. sp. n. 52 [h] V. Stepanov det. [p]”. Described from 2 syntypes. Agrilus gerschuni Stepanov, 1958 Type locality: v sovkhoze Pakhta-Aral (Golodnaya step’) Yuzhno Kazakhstanskoi oblasti Kazakhskoi SSR [in Russian]. Type specimens: Holotype by monotypy (Fig. 46Q), ♀, (ZIN): “Kazakhskaya SSR Golodnaya step’ Sovkhoz PakhtaAral 19-VI-1951 M. S. Gershun leg. [h] [in Russian] \ Typus [h] [red labe] \ Agrilus gerschunii [sic!] m. sp. n. 52 [h] V. Stepanov det. [p] \ Ris. 216 [h] \ Holotypus [p] [red label]”. Described from 1 specimen. Agrilus machnovskii Stepanov, 1958 Type locality: zapovednik Parkentskii raion Tashkentskoi oblasti, Uzbekskoi SSR [in Russian]. Type specimens: Primary: Lectotype (Fig. 48S) by present designation, ♀, (ZIN): “Uzbekskaya SSR Tashkentsk. ob. Parkentsk. r. n. 27-VI-51 g. I. K. Makhnovskii [h] [in Russian] Gorno-lesn. Z-k. na ive mertvye [h] [in Russian] [underside] \ Agrilus 2 èkz. na ive [h] [in Russian] \ Typus [h] [red labe] \ Agrilus machnovskii m. sp. n. 52 [h] V. Stepanov det. [p] \ Ris. 214 [h] \ Syntypus [p] [red label]”. Secondary: 1 PLT (ZIN). Described from 2 syntypes. Diagnosis: 5.7-9.6 mm; pronotum widest at anterior third; prehumerus absent; pronotum and elytra with distinct whitish pubescence; elytral apices obviously prolonged. Species group: Viridis. Relevant species: A. viridis (Linné, 1758).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Specimens examined: CHINA: 1 ♀ (MNHN): “Chine, Meyer D.”. KAZAKHSTAN: 1 (ZIN): “r. Temerlik, okr. pos. Temerlik, Uigursk. r n., Alma Atinsk. obl., Kazakh., Volkovitsh, 20.06.1988 [in Russian]”. 1 (NMPC): “Aulie Ata, Syr Darja”; 1 ♂, 1 ♀ (ZIN): “Yuzhno Kazazhstan. obl., Yu V kyzyl Kumy, Tugan, r. Syr Daria, vyveden iz ivy, 1967 [in Russian]”; 2 (EJCB): “USSR, Kazachstan, Čingan / Čatkal. chr., 1900m, 12.5.1977, Sv. Bílý lgt. \ reared from Salix”; 1 ♂, 1 ♀ (ZIN): “Yu.V.Kazakstan, khr. Toksandai, 1400 1500m, bl. Chuvara, 14.vii.1985, I. Lopatin [in Russian]”; 1 (ZIN): “ur. Kurtogai. r. Charyn, 35 km SZ pos. Kegen, Kazakhstan, Volkovitsh, 15 16.VII.1986 [in Russian]”; 1 (ZIN): “ur. Kurtogai. r. Charyn, 35 km SZ Kegena, Kazakhstan, Volkovitsh, 2 4.VII.1987 [in Russian]”; 1 (ZIN): “Koibynskoe ushch., g. Katutau, 25 km ZSZ Koktala, Kazakhst., Volkovitsh, 30.6 2.7.1988 [in Russian]”; 1 (MDCM): “27.4.1976 Chilik, Kazakhstan, iz vetok turangi, B. M. Mamaev”. KYRGYZSTAN: 1 (MKCY): “Kirg. SSR, r. Uu. Èolovi Zamki, 20.07.1978 [in Russian]”; 1 (ZIN): “r. Kolbor, 20 km YuZ Kazarmana, Kazarman sk. kotl. Naryn, Volkovitsh, 12 14.06.1989 \ v vetke Salix iz kolybel’ki [in Russian]”; 3 (MNCA): “USSR, Kirgizia mer., ca. 115 km E of Osch: Oy Tal, 1800 200m, NN, vii.1991, Bohne”. MONGOLIA: 1 (EJCB): “Mongoliam Chovd Ajmak, Ueng Gol, 30km N Ueng, 30.7.1975”; 2 (ZIN): “Mongoliya, Kobloski aimak, yr. Utyn Mod, 50km YuYuZ Uèncha, Kerzhner, 27.vi.1980 [in Russian]”; 2 (ZIN): “Mongoliya, Kobloski aimak, yr. Utyn Mod, 50km YuYuZ Uèncha, G. Medvedev, 27.vi.1980 [in Russian]”. TAJIKISTAN: 1 (NMPC): “Tadzhik. SSR, Ramit, 1950, Kulinich [in Russian]”; 2 (NMPC): “Kondara, Tadzh., s ivy, 13.vi.1956, Kulinich [in Russian]”; 1 (ZIN): “yshch. r. Sangvor, pr. r. Obikhkhingoi, Khr. Petra I, Tajz., Danilovich, 6.vii.1975, na Salix [in Russian]”; 4 (NMPC): “SSSR Tadžik., Dušanbe, Javroz, vi.1981, J. Hron lgt.”; 5 (EJCB): “USSR, Tajik.SSR, 30.6., Gissar mts., 1984, 40 km N of Dushanbe, Takob, E. Jendek leg.”; 2 (MKCY): “Tadzh. SSR, Ramit, poima r. Sardamliena, na ivakh, vi.1985 [in Russian]”. UZBEKISTAN: 1 (ZIN): “Tashkent, 30/iv 1916 [in Russian]”; 4 (NMPC): “Uzbekistan, 5.7.1976, Taškent, Čimgam, Sv. Bílý leg.”; 2 ♀ (EJCB): “USSR, Uzbekistan, Aktaš / Taškent, 1980, ex l., Vít Kubáň lgt.”; 11 (NMPC): “USSR, Uzbekistan, Aktaš / Taškent, 1980, ex l., Vít Kubáň lgt.”; 1 (EJCB): “Uzbekistan, Ferghana, Uzgen dist., Tuyuk riv., 29.06.1996, Tretyakov leg.”.

Adult occurence: 4-5-6-7 (n=21). Altitude range: 14002000 m (n=3). Host plants: Salix: STEPANOV, 1958 (as machnovskii, gerschuni, uzbekistanus) [#3664]; MAKHNOVSKIJ, 1958 (as machnovskii and uzbekistanus) [#6090]; MAKHNOVSKIJ, 1959 (as machnovskii and uzbekistanus) [#5829]; KULINITSH, 1963 [#8648]; SINADSKIJ, 1963 (as uzbekistanus) [#5038]; KADYROV, 1976 [#3461]; ALEXEEV, 1989 (as attila)[#64]; TLEPPAEVA & ISHKOV, 2004 [#7211]. – S. acutifolia: LINDEMAN & TURUNDAEVSKAYA, 1975 (as uzbekistanus) [#3478]. – S. fragilis: LINDEMAN & TURUNDAEVSKAYA, 1975 (as uzbekistanus) [#3478]. – S. triandra: LINDEMAN & TURUNDAEVSKAYA, 1975 (as uzbekistanus) [#3478]. Erroneous: Pistacia: MAKHNOVSKIJ, 1958 [#6090]; MAKHNOVSKIJ, 1959 [#5829]. – Populus: SINADSKIJ, 1963 (as uzbekistanus) [#5038]; TLEPPAEVA & ISHKOV, 2004 [#7211]. – P. diversifolia: YAGDYEV, 1978 [#4206]. – P. tremula: TLEPPAEVA, 1999 [#6551]. Distribution: CHINA: Nei Mongol. KAZAKHSTAN. KYRGYZSTAN. MONGOLIA. RUSSIA: Buryatiya Respublika, Zabaykalskyi kray. TAJIKISTAN. UZBEKISTAN.

209

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Remarks: Mongolian specimens are densely pubescent, on the ventral side almost tomentose.

Agrilus tsushimanus Kurosawa Fig. 27C (habitus); Fig. 39P (aedeagus) tsushimanus Kurosawa, 1963 (Agrilus; subspecies of acastus) KUROSAWA, 1963: 106 [#1617] (description). KUROSAWA, 1963: 154 [#1618] (subspecies of acastus; characters; Japan). KUROSAWA, 1974: 2 [#4575] (subspecies of acastus; characters; notes). AKIYAMA & OHMOMO, 1997: 4, 42 [#4091] (checklist; Japan: Tsushima). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2336 [#8576] (world catalog). JENDEK & GREBENNIKOV, 2009: 15 [#8563] (not belonging to muscarius species group).

Type material: Agrilus acastus tsushimanus Kurosawa, 1963 Type locality: Tsushima: Kamizaka - Shiratake. Type specimens: Holotype, ♂, not in NSMT (S. Nomura pers. com.). Described from 1 specimen (HT). Diagnosis: 4.4-6.1 mm; posterior part of elytral pubescence distinct; apices conjointly arcuate; medial lobe of aedeagus with obtuse apex. Species group: Cyaneovirens. Relevant species: A. euonymi Tôyama, 1985; A. sagittifer sp. nov. Specimens examined: CHINA: Guizhou: 2 ♂ (EJCB): “CH Guizhou NE, 27.V. 3.VI., 20 km NW of Jiangkou, 1995, Fanjing Shan Kuaichang, E. Jendek & O. Šauša leg.”. Henan: 4 ♂, 2 ♀ (EJCB): “China, N. Henan, 11 12.VI., Jiuligou, 550 m, 35º12’N, 112º26’E, Jaroslav Turna leg. 2007”. Shaanxi: 1 ♂, 1 ♀ (EJCB): “China, Shaanxi prov., 21 23.June 1998 \ Qing Ling Shan mts, 1000 m road Baoji Taibai vill., pass 40 km S Baoji, Zd. Jindra lgt.”; 24 (EJCB): “C China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000 1300m, 23.V. 13.VI. leg. C. Holzschuh 2000”; 1 ♂ (EJCB): “China Shaanxi, Lueang env., 15 km NW, 26.5. 31.5.2001, lgt. E. Kučera”. JAPAN: Kyushu: 2 ♂ (EJCB): “1997.6.15., Ohboshi, Tsushima, Nagasaki, Kyushu, S Jpn., T. Akashi leg.”.

Adult occurence: 5-6 (n=6). Altitude range: 550-1300 m (n=3). Host plants: Unknown. Distribution: CHINA: Guizhou, Henan, Shaanxi. JAPAN: Kyushu, Tsushima. Remarks: The concept of this taxon was judged from two Japanese specimens determined by S. Ohmomo.

Agrilus tsutsumiuchii Ohmomo Fig. 25J (habitus of PT); Fig. 39C (aedeagus) tsutsumiuchii Ohmomo, 2006 (Agrilus) OHMOMO, 2006: 247 248 [#7546] (description). 2336 (world catalog).

BELLAMY, 2008:

Type material: Agrilus tsutsumiuchii Ohmomo, 2006 Type locality: Ushuki-shi, Oita Pref. (Kyusuikei, Kokonoemachi, Oita Pref.). Type specimens: Primary: Holotype: ♂, (NSMT). Secondary: PT (SOCI); 1 PT (Fig. 25J)(EJCB). Described from 8 specimens (HT, PT).

Diagnosis: 6.85-7.05 mm; elytra except for pleural parts entirely pubescent; elytral apices evenly rounded with very fine serrulated margin; pygidium arcuate; tubercles on basal ventrite in male not transversely elongate. Species group: Tempestivus. Relevant species: A. asiaticus Kerremans, 1898. Specimens examined: Known to us only from the male PT.

Host plants: Zelkova serrata: OHMOMO, 2006 [#7546]. Distribution: JAPAN: Kyushu.

Agrilus tuber Jendek, sp. nov. Fig. 29A (habitus of HT); Fig. 4Q (aedeagus)

Description: Holotype 4.2 mm. BODY: Shape: subparallel, Color (dorsally): unicolored, Color (dominant): green. HEAD: Shape: markedly convex, Medial impression: present, Impression (extent): vertex, Impression (depth): shallow. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): irregular, Sculpture (density): dense. Eyes: Size: large, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: visually elongate, Sides: arcuate, Maximal width: at middle, Anterior margin: narrower than posterior. Anterior lobe: obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: rectangular, Apex: blunt. Disk: Convexity: moderately convex, Disk impressions: posterolateral. Prehumerus: carinal, Shape: bisinuate, Extent: to anterior pronotal angle, Posterior end: joining posterior pronotal angle, Anterior end: joining anterior pronotal angle, Arc: moderate. Marginal and submarginal carinae: Interspace: wide, Convergence: strongly convergent, Junction: present, Narrowest point: at posterior third of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: conjoint, Width: wide, Shape: subangulate. Elytral pubescence: entire, Color: unicolored, Character: homogenous. Tomentum: absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: prosternum with longer and denser pubescence. Prosternal lobe: large, Anterior margin: arcuate. Prosternal process: wide, Shape: dilated, Sides: straight, Angles: rectangular, Angles (tips): sharp, Disk: impressed with sides protruding. ABDOMEN: Sexual modifications in male: single medial tubercle on ventrite 1, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuately sinuate. LEGS: Male sexual modifications: metafemur markedly incrassate. Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical. Diagnosis: head strongly convex; eyes large; vertex very narrow; elytra without humeal carina; elytral apices

210 conjointly subangulate. Species group: Viridiobscurus. Relevant species: A. pliculipennis Obenberger, 1940. Type locality: China, Zheijang province. Type specimens: Primary: Holotype, ♂, (EJCB): “China, Zheijang”. Adult occurence: Host plants: Unknown. Distribution: CHINA: Zhejiang. Etymology: The specific name is the Latin noun tuber (swelling, bulb) in apposition. It refers to the single medial tubercle on the basal ventrite in the male of this species.

Agrilus tyndarus Kerremans Fig. 31J (habitus); Fig. 41V (aedeagus); Figs. 50J, 53S, 54D (types) tyndarus Kerremans, 1913 (Agrilus) KERREMANS, 1913: 115 [#1471] (description). MIWA, 1931: 125 [#6101] (catalog; Formosa). MIWA & CHŰJÔ, 1936: 21 [#1809] (catalog; Japan). OBENBERGER, 1936: 1106 [#2191] (world catalog). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Taiwan). MÜHLE, 2003: 47 [#6155] (checklist; Taiwan). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2338 [#8576] (world catalog). omissus Kerremans, 1914 ([preoccupied]) KERREMANS, 1914: 106 [#1481] (description). MIWA, 1931: 126 [#6101] (catalog; Formosa). OBENBERGER, 1935: 121 [#2189] (synonym of omissulus). MIWA & CHŰJÔ, 1936: 19 [#1809] (catalog; Japan). OBENBERGER, 1936: 1095 [#2191] (synonym of omissulus). JENDEK, 2000: 505 [#4851] (synonym of omissulus; lectotype designation). BELLAMY, 2008: 2218 [#8576] (synonym of omissulus; world catalog). omissulus Obenberger, 1935 (replacement name for omissus Kerremans, 1914 not Kerremans, 1903) syn. nov. OBENBERGER, 1935: 121 [#2189] (replacement name proposal). OBENBERGER, 1936: 1095 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 357 [#2429] (checklist; China). JENDEK, 2000: 505 [#4851] (synonymy). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Taiwan). MÜHLE, 2003: 47 [#6155] (checklist; Taiwan). JENDEK, 2006: 400 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2218 [#8576] (world catalog). therondi Descarpentries & Villiers, 1963 (Agrilus) syn. nov. DESCARPENTRIES & VILLIERS, 1963: 5, 19 [#701] (description). DESCARPENTRIES & VILLIERS, 1967: 148 [#710] (diagnosed with muong). BELLAMY, 2008: 2327 [#8576] (subgenus Sianagrilus; world catalog).

Type material: Agrilus tyndarus Kerremans, 1913 Type locality: [Formosa] Kankau (Koshun). Type specimens: Primary: Lectotype (Fig. 54D) by present designation , ♂, (MNHN): “Kankau (Koshun) [p] IV [h] Formosa [p] H. Sauter [p] IV [h] 1912 [p] \ Tyndarus Kerr. Type [h] \ Museum Paris Coll. Ch. Kerremans 1923 [p] [yellow label]”. Secondary: 2 PLT (ISNB); 2 PLT (NMPC); 2 PLT (MNHN); 5 PLT (DEI). Described from unknown number of syntypes. Agrilus omissus Kerremans, 1914

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Type locality: [Formosa] Kankau: Koshun. Type specimens: Primary: Lectotype (Fig. 50J), ♀, designated by JENDEK (2000: 505) [#4851], (DEI): “Kankau (Koshun) Formosa H. Sauter IV.1912 [p] \ Syntypus [p] [red label] \ Kerremanns [sic!] det. [p] \ Agrilus omissus Kerr. Type [h] [Kerremans’ MS] \ Coll. DEI Eberswalde [p]”. Described from unknown number of syntypes. Agrilus omissulus Obenberger, 1935 Type locality: See: Agrilus omissus Kerremans, 1914. Type specimens: See: A. omissus Kerremans, 1914. Agrilus therondi Descarpentries & Villiers, 1963 Type locality: Tonkin: Hoa Binh. Type specimens: Primary: Holotype (Fig. 53S), ♂, (MNHN): “Tonkin occ. Rég. de Hoa Binh 1919 [p] \ ♂ [h] Type [p] [red label] \ Agrilus therondi n. sp. Holotype ♂ nob[is]. A. Descarpentries et [h] A. Villiers det. 19 [p] 63 [h]”. Described from 2 specimens (HT, PT). Diagnosis: 4.6-5.3 mm; pronotal medial impression deeper and larger in basal part; elytra with perisutural stripes of white pubescence obscure in apical elytral third. Species group: Unassigned. Relevant species: A. perviridis Kerremans, 1894. Specimens examined: CHINA: Fujian: 1 ♂, 1 ♀ (EJCB): “China, prov. Fujian bor. occ., Shaowu env., 25km road Shaowu Taining, 13 16.6., M. Nikodým lgt., 1991”; 1 ♂ (EJCB): “China, Fujian prov., Shaowu env., 13 16.6.1991”; 1 (EJCB): “China, Fujian prov. Shaowu env., 5 10.VII.1991”. LAOS: Borikhamxai: 1 ♀ (EJCB): “Laos centr., Bolikhamsai prov., Ban Nape Kaew Nua Pass, 18.4. 1.5.1998, alt. 600±100 m, N18°22.3, E105°09.1 (GPS), E. Jendek & O. Šauša leg.”. THAILAND: Chiang Mai: 1 (EJCB): “Thailand N, 21.V. 4.VI.1995, Chiang Dao env., M. Snížek leg.”. VIETNAM: 1 ♂ (EJCB): “Tonkin, Phu Lang Thuong”; 1 (EJCB): “Vietnam, NW, 17.4.1963, Bai Tuong lang Tian, Kabakov leg.”; 3 ♂, 1♀ (EJCB): “N Vietnam, 21°35N, 106°30E, 52 km SW of Lang Son, 27.iv. 6.v.1996, 370 m, Dembický & Pacholátko leg.”.

Adult occurence: 4-5-6-7 (n=7). Altitude range: 370-700 m (n=2). Host plants: Unknown. Distribution: CHINA: Fujian. LAOS. THAILAND. VIETNAM.

Agrilus uenoi Kurosawa Fig. 28D (habitus); Fig. 40H (aedeagus); Fig. 54E (type) uenoi Kurosawa, 1963 (Agrilus) KUROSAWA, 1963: 102 104 [#1617] (description). KUROSAWA, 1975: 2 3 [#4591] (note). TÔYAMA, 1985: 20 [#3678] (iconography; Japan). TÔYAMA, 1985: 30 31 [#2924] (faunal records; Okinawa). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). AKIYAMA, 1993: 8 [#4875] (faunal record; Japan). MORIMOTO & TADAUCHI, 1995: 232 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 42 [#4091] (checklist; Japan: Kyushu, Tsushima; Yakushima I., Tokara nakanoshima I., Ryukyus (Amami oshima I., Tokuno shima I., Okinawa I., Kume jima I., Ishigaki jima I., Iriomote jima I.)). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2338 [#8576] (world catalog).

Type material: Agrilus uenoi Kurosawa, 1963

211

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Type locality: Tokara Is.: Nakano-shima I. Type specimens: Primary: Holotype (Fig. 54E), ♂, (NSMT): “Tokara Is. Nakanoshima 6. VI. 1953 S. Ueno [p] \ Holotype [p] Agrilus uenoi Y. Kurosawa, 196 [sic!] [h] [red label]”. Described from 9 specimens (HT, PT). Diagnosis: 3.3-4.7 mm; body slender; humeri feebly impressed. Species group: Densetomentosus. Relevant species: A. densetomentosus Obenberger, 1940. Specimens examined: JAPAN: Ryukyu isl. (Okinawa incl.): 1 ♀ (EJCB): “1996.10.24, Ishigaki Is., Ryukyu, S Jpn, K. Takahashi leg.”; 1 ♂ (EJCB): “1998.4.18, Ishigaki Is., Ryukyu, S Jpn, S Ohmomo leg.”.

Adult occurence: 4-5-6-7-8-10 (n=8). Faunal records published by KUROSAWA, 1963 [#1617] are also included. Host plants: Mallotus japonicus: TÔYAMA, 1985 [#2924]; AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: JAPAN: Kyushu, Ryukyu Islands, Tsushima.

Agrilus ulaangomiensis Cobos Fig. 31K (habitus); Fig. 41W (aedeagus); Figs. 45T, 54F (types)

Secondary: PT (MNMS); 1 PT (HNHM). Described from 3 specimens (HT, PT). Agrilus subelegans Cobos, 1972 Type locality: Mittelgobi aimak, Choot bulag, zw. Chuld und Delgerchangaj, 1.480 m. Type specimens: Not located. Described from 1 specimen. Agrilus emeljanovi Alexeev, 1975 Type locality: Mongolia ... Vostochno-Gobiiskii aimak: Nomt-Ula, 30 km YUYUV [= south-southestern] oz. Schokhoi-Nur [in Russian]. Type specimens: Primary: Holotype (Fig. 45T), sex not examined, (ZIN): “Mongoliya Vost.-Gob. aimak, Nomt-Ula, 30 km YUYUV oz. Shokhoi-Nur Kerzhner 26 VI 1971 [p] [in Russian] \ Holotypus [p] [red label] \ Agrilus emeljanovi Alex. [h]”. Secondary: 1 PT (ZIN). Described from 13 specimens (HT, PT). Diagnosis: 4.8-6.1 mm; pronotum conspicuously convex with distinct lateral impressions; anterior pronotal margin deeply bisinuate, with obvious medial lobe; prehumerus carinal and short; marginal and submarginal pronotal carinae convergent and conjoined posteriad; elytral apices narrow. Species group: Unassigned. Relevant species: Unknown.

ulaangomiensis Cobos, 1972 (Agrilus; subgenus Agrilus) COBOS, 1972: 47 49, 56 [#591] (description). ALEXEEV, 1975: 158 [#52] (cited as ulaangomensis; note; Mongolia). ALEXEEV & VOLKOVITSH, 1989: 319 [#3241] (cited as ulaangomensis; synonym of spessivcevi; type examination). JENDEK, 2005: 21 [#7217] (revalidated species; synonymy; notes). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2339 [#8576] (world catalog). subelegans Cobos, 1972 (Agrilus; subgenus Agrilus) COBOS, 1972: 49 51, 56 [#591] (description). ALEXEEV, 1975: 157 [#52] (note; close to A. solieri species group; Mongolia). ALEXEEV & VOLKOVITSH, 1989: 319 [#3241] (synonym of spessivcevi; type examination). JENDEK, 2005: 21 [#7217] (synonym of ulaangomiensis). JENDEK, 2006: 402 [#7441] (synonym of ulaangomiensis; Palaearctic catalog). BELLAMY, 2008: 2339 [#8576] (synonym of ulaangomiensis; world catalog). emeljanovi Alexeev, 1975 (Agrilus) ALEXEEV & ALEXEEV, 1975: 152 155 [#52] (description). VOLKOVITSH, 1989: 319 [#3241] (synonym of spessivcevi; type examination). JENDEK, 2005: 21 [#7217] (synonym of ulaangomiensis). JENDEK, 2006: 402 [#7441] (synonym of ulaangomiensis; Palaearctic catalog). BELLAMY, 2008: 2339 [#8576] (synonym of ulaangomiensis; world catalog).

Specimens examined: CHINA: Gansu: 1 ♂, 1 ♀ (EJCB): “China Gansu, Dingxi, 23.5. 30.5.1996, E. Kučera leg.”. Ningxia: 1 ♀ (EJCB): “China bor., 11.5.1996, Yanchi env., 107,6’ 37,4’, leg. J. Halada”. MONGOLIA: Bulgan: 1 ♀ (EJCB): “Mongolia, Bulgan, 16.6.1975, river Tuul, “Telengijn Bajtin [from Russian]”. RUSSIA: Tyva Respublika: 1 ♀ (ZIN): “Tuva, 25 km SZ Kyzyla, pos. Seserlig ( Germanovka), 12.vi.1979, Korotyaev [in Russian]”; 12 (EJCB): “Tuvinskaja ASSR, 15.6., 10 km NE Erzin, 1994, Z. Kletečka leg.”.

Type material: Agrilus ulaangomiensis Cobos, 1972 Type locality: Uvs aimak, SW. Rand des Sees Uvs nuur, 63 km W von der Stadt Ulaangom, 790 m. Type specimens: Primary: Holotype (Fig. 54F), ♀, (HNHM): “female sign [p] \ MONGOLIA: Uvs aimak SW Rand des Sees Uvs nuur, 63 km O von deer Stadt Ulaangom, 790 m, Ep. Dr. Z. Kaszab, 1968 [p] \ Nr. 1020 26. VI. 1968 [p] \ HOLOTYPUS A. Cobos [p] [red label] \ Agrilus ulaangomiensis nov. sp. Holotypus [h] A. Cobos det. 197 [p] 1 [h] \ Agrilus spesivcevi Obenb. [h] Alexeev det. [p] Cum typo comp. [h] \ (= Agrilus subelegans Cobos [h] Alexeev det. [p] (= emeljanovi Alex) [h] Alexeev det. [p]”.

Description: 4.1-5.2 mm (Holotype 4.5 mm). BODY: Shape: cuneiform, Build: robust, Color (dorsally): unicolored, Color (dominant): brown. HEAD: Medial impression: absent, rarely present, Impression (extent): vertex, Impression (depth): shallow, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (density): dense. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate, Sexual modification in male: not apparent. PRONOTUM: Shape: transverse, Sides: straight or arcuate, Maximal width: at anterior margin, Anterior margin: subequal

Adult occurence: 5-6-7 (n=13). Faunal records published by ALEXEEV, 1975 [#52] are also included. Altitude range: 1480 m (n=1). Host plants: Caragana: ALEXEEV, 1975 [#52] (as emeljanovi). – C. spinosa: ALEXEEV & VOLKOVITSH, 1989: [#3241] (as spessivcevi). Distribution: CHINA: Ningxia. MONGOLIA. RUSSIA: Tyva Respublika.

Agrilus ulmicola Jendek, sp. nov. Fig. 29G (habitus of HT); Fig. 40V (aedeagus)

212 to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles or projecting beyond anterior pronotal angles. Posterior angles: Shape: acute or rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: lateral only, Lateral impressions (deepness): shallow. Prehumerus: carinal, Shape: bisinuate, Extent: to anterior pronotal angle, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina or joining anterior pronotal angle, Arc: moderate. Marginal and submarginal carinae: Interspace: moderate, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior third of pronotal length, Modifications: submarginal carina divergent at posterior angles. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent, Apical part: feebly attenuate. Apices: Arrangement: separate, Shape: arcuate. Elytral pubescence: entire, Color: unicolored. Tomentum: absent. STERNUM: Metasternal projection: impressed, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate, Anterior margin: arcuately emarginate or angulately emarginate, Emargination (deepness): deep, Emargination (width): narrow. Prosternal process: narrow, Shape: narrowed, Sides: straight, Angles: absent, Disk: flat. ABDOMEN: Sexual modifications in male: impression at intercoxal projection, Apical margin of pygidium: arcuate, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Aedeagus (symmetry): symmetrical, Aedeagus (shape): markedly dorsoventrally bent, Ovipositor: elongate. Diagnosis: prosternal lobe markedly declivous and deeply emarginate; prosternal process narrow, vithout lateral angles, tapering to apex. Species group: Roscidus. Relevant species: A. eunuchus sp. nov. Type locality: China, Shanxi province, Yongji. Type specimens: Primary: Holotype (Fig. 29G), ♂, (EJCB): “China-Shanxi, Yongji, 23.5.-25.5.2001, lgt. E. Kučera”. Secondary: CHINA: Shanxi: 6 PT (EJCB): “China-Shanxi, Yongji, 23.5.-25.5.2001, lgt. E. Kučera”. Adult occurence: 5 (n=1). Host plants: Ulmus. Distribution: CHINA: Shanxi. Etymology: The specific name was derived from Ulmus (the Latin name of the host plant cited on the labels of the type specimens) and the Latin verb colo (to dwell, inhabit).

Agrilus unguiculosus Obenberger Fig. 22A (habitus); Fig. 37J (aedeagus); Figs. 46R, 54H (types) unguiculosus Obenberger, 1936 (Agrilus) OBENBERGER, 1936: 105 [#2205] (description). OBENBERGER, 1936: 1054 [#2191] (world catalog). ALEXEEV & VOLKOVITSH, 1989: 322 323, 351, 357 [#3241] (characters in key; faunal records; type

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA examination; remark on type locality; Mongolia). ALEXEEV, 1998: 382 [#4322] (subgenus Spiragrilus; subgeneric classification). JENDEK, 2006: 395 [#7441] (subgenus Spiragrilus; Palaearctic catalog). LÖBL & SMETANA, 2007: 29 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2340 [#8576] (subgenus Spiragrilus; world catalog). gobicola Cobos, 1968 (Agrilus; subgenus Agrilus) COBOS, 1968: 398 400 [#580] (description). COBOS, 1972: 56 [#591] (subgenus Agrilus; checklist; Mongolia). ALEXEEV, 1975: 159 [#52] (presumable variety of pseudolimoniastri; notes; faunal records; distributional summary; Mongolia; Tuva; Altay; Zaisan; Karaganda). ALEXEEV & VOLKOVITSH, 1989: 322 [#3241] (synonym of unguiculosus; type examination). JENDEK, 2006: 395 [#7441] (synonym of unguiculosus; Palaearctic catalog). BELLAMY, 2008: 2339 [#8576] (synonym of unguiculosus; world catalog).

Type material: Agrilus unguiculosus Obenberger, 1936 Type locality: Asia centralis: Fluvius Kenderlek. Type specimens: Primary: Lectotype (Fig. 54H) by present designation, ♀, (NMPC): ““Сауръ [Saur’]1060 m r. Кендерликь [Kenderlik’] 24.VI.[1]910 A. Якобсон [Yakobson] [h] \ Typus [p] [red label] \ Agrilus unguiculosus m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24959 [h] [orange label]”. Secondary: 1 ♀, PLT (NMPC). Described from unknown number of syntypes. Agrilus gobicola Cobos, 1968 Type locality: Chovd aimak, Mongol Altai Gebirge, Uljasutajan gol, 45 km. NNW. von somon Bulgan, 1400 m. alt. Type specimens: Primary: Holotype (Fig. 46R), ♂, (HNHM): “Mongolia, Chovd aimak, Mongol Altai Gebirge, Uljasutajan gol, 45 km. NNO [sic!]. von Somon Bulgan, 1400 m Exp. Dr. Z. Kaszab, 1966 [p] \ Nr. 637, .6 VII. 1966 [p] \ Holotypus A. Cobos [p] [red label] \ Agrilus gobicola nov. sp. Holotypus [h] A. Cobos det. 196 [p] 8 [p] \ Agrilus unguiculosus Obenb. [h] Alexeev det. [p] cum typo comp. [h] Alexeev det. [p]”. Secondary: 1 PT (MNMS); 1 PT (HNHM). Described from 4 specimens (HT, AT, PT). Diagnosis: 5.1-7 mm; posterior pronotal angles sharply acute; medial part of sternum in male with dense, long, erect pubescence. Species group: Hyperici. Relevant species: A. pseudolimoniastri Cobos, 1968. Specimens examined: MONGOLIA: 4 (EJCB): “Mongolia, Bayankhongor, 6.7.1975, 20 km SW lake Bon Cagan”; 2 ♂ (EJCB): “MGL Bayankhongor, 75 km S Bayankhongor, N45º20’, E100º48.5’, 1330m, 8.VII.2004, J. Halada leg.”. 7 (NMPC): “Mongolia, Khovd ajmak, Bajtag Bogd Nuru, 25.vii.1975”. 1 (EJCB): “Mongolia, Calgant, 10.vii.1975”.

Adult occurence: 6-7 (n=7). Faunal record published by COBOS, 1968 [#580] is also included. Altitude range: 10601550 m (n=5). Faunal record published by COBOS, 1968 [#580] is also included. Host plants: Caragana leucophloea: ALEXEEV & VOLKOVITSH, 1989 [#3241]. – Spirea: ALEXEEV, 1975 (as gobicola) [#52]; ALEXEEV & VOLKOVITSH, 1989 [#3241]. Distribution: KAZAKHSTAN. MONGOLIA. RUSSIA: Altay Kray, Tyva Respublika.

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA

Agrilus ussuricola Obenberger Fig. 24B (habitus); Fig. 38I (aedeagus); Figs. 48M, 54I (types) ussuricola Obenberger, 1924 (Agrilus) OBENBERGER, 1924: 46 [#2040] (description). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1055 [#2191] (world catalog). KUROSAWA, 1974: 2 [#4577] (related to A. lassiolus and A. angustulus). HOU, 1981: 395 396 [#4198] (cited as ussricota; notes; China: Xizang). COBOS, 1986: 231 [#609] (species with asymmetric aedeagus). HOU, 1987: 589 [#3490] (notes; Yunnan). PENG ZHONGLIANG, 1987: 359 [#2429] (checklist; China). ALEXEEV, 1989: 479 [#64] (key to species; Khabarovsk; Amursk; Primorsk; North Korea; northeastern China). ALEXEEV, 1998: 374 [#4322] (subgenus Quercuagrilus; subgeneric classification). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Heilongjiang, Yunnan, Xizang). JENDEK, 2005: 21 22 [#7217] (lectotype designation; type locality designation; synonymy; faunal records; notes; China (Shaanxi); Japan (Nagasaki); North Korea; Russia (Amursk, Primorsk)). JENDEK, 2006: 392 [#7441] (subgenus Quercuagrilus; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (checklist; Japan). BELLAMY, 2008: 234 2342 [#8576] (subgenus Queruagrilus; world catalog). VOLKOVITSH, 2009: 135 [#8850] (subgenus Quercuagrilus; faunal records; Russia: Primorskiy kray). lasiolus Obenberger, 1935 (Agrilus; variety of ussuricola) OBENBERGER, 1935: 170 [#2158] (description). OBENBERGER, 1936: 1055 [#2191] (variety of ussuricola; world catalog). KUROSAWA, 1963: 154 [#1618] (characters; Tsushima; Korea; East Siberia). KUROSAWA, 1974: 2 [#4577] (characters; notes; Tsushima; Korea; Ussuri). KUROSAWA, 1974: 2, 3 [#4575] (notes). AKIYAMA & OHMOMO, 1997: 35 [#4091] (checklist; Japan: Tsushima; Korea; Russia: East Siberia). JENDEK, 2005: 21 22 [#7217] (synonym of ussuricola; lectotype designation; type locality designation). JENDEK, 2006: 392 [#7441] (synonym of ussuricola; Palaearctic catalog). OHMOMO, 2006: 6 [#8002] (synonym of ussuricola). BELLAMY, 2008: 2342 [#8576] (synonym of ussuricola; world catalog). anomalus Théry, 1942 (Agrilus) THÉRY, 1942: 262 264 [#2875] (description). OBENBERGER, 1943: 10 [#2333] (synonym of ussuricola). JENDEK, 2006: 392 [#7441] (synonym of ussuricola; Palaearctic catalog). BELLAMY, 2008: 2342 [#8576] (synonym of ussuricola; world catalog).

Type material: Agrilus ussuricola Obenberger, 1924 Type locality: Ussuri (Vladivostok). Type specimens: Primary: Lectotype (Fig. 54I), ♂, designated by JENDEK (2005: 21-22) [#7217], (NMPC): “Amur [h] [in Russian] \ Typus [p] [red label] \ Agrilus ussuricola m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24261 [h] [orange label]”. Secondary: 2 PLT (NMPC); 1 PLT (NHMB). Described from unknown number of syntypes. Agrilus ussuricola variety lasiolus Obenberger, 1935 Type locality: Ussuri, Primorje. Type specimens: Primary: Lectotype (Fig. 48M), ♂, designated by JENDEK (2005: 21-22) [#7217], (NMPC): “s. Troickoye, oz. Khanka, Primors. Cherskij [p] 16.vi. [h] 09 [p] [in Russian] \ Typus [p] [red label] \ Agrilus ussuricola var. lasiolus m. ♂ Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24287 [h] [orange label]”. Secondary: 1 PLT (ZIN); 9 PLT (ZIN); 6 PLT (NMPC). Described from unknown number of syntypes.

213 Agrilus anomalus Théry, 1942 Type locality: Erzendjanzcy, Mandchourie. Type specimens: Not located. Described from 3 syntypes. Diagnosis: 3.3-5 mm; head smaller; elytral apices broadly separately arcuate. Species group: Sulcicollis. Relevant species: A. fissus Obenberger, 1917. Specimens examined: CHINA: Jilin: 2 (ZIN): “Hsiaoling, Kirin [ Jilin] Province, China 15.vi.1939 [in Russian]”. Shaanxi: 1 ♀ (EJCB): “China, Shaanxi, Foping env. 7.Jun., 120km SW Xian, 1992, M. Bok lgt.” (JENDEK, 2005 [#7217]); 1 ♂ (EKCS): “China Shaanxi, Lüeyang, 8.6. 14.6.1996, lgt. E. Kučera”; 1 ♂, 1 ♀ (EJCB): “China, 1150 1300 m, Shaanxi, Qinling mts., Foping (6 km N), 20 21.VI.1998, I. H. Marshal leg.” (JENDEK, 2005 [#7217]); 1 ♂ (EJCB): “China Shaanxi, Lüeang, 18.5. 21.5.2000, leg. E. Kučera” (JENDEK, 2005 [#7217]). Shanxi: 1 ♂ (JMCZ): “China c. 29 30.v., Jingangling, 111.7’ 50km W Linfen, 36.2’, leg. J. Halada 1996”; 2 ♂ (EJCB): “China c. 29 30.v., Jingangling, 111.7’ 50km W Linfen, 36.2’, leg. J. Halada 1996”. JAPAN: Tsushima: 1 ♀ (EJCB): “(Mt. Ohboshi) Tsushima Is., 17.vi.1997, coll. T. Akashi” (JENDEK, 2005 [#7217]). KOREA NORTH: 1 ♂ (EJCB): “D.P.R. Korea, Manmulsang, Kumgangsang Mts., lgt. M. Slovák, 25.5.1988” (JENDEK, 2005 [#7217]); 1 ♂, 2 ♀ (EJCB): “N. Korea, 19.7.1990, Toxong env. Omdouri” (JENDEK, 2005 [#7217]). RUSSIA: Amurskaya oblast’: 1 ♂ (EJCB): “USSR, Archara, Amursk. oblast, 7.89 Morozinski” (JENDEK, 2005 [#7217]). Primorskiy kray: 25 (EJCB): “USSR, Primorskij kraj Arseneev env. 300 400 m 11. 19.VII.1990 B. Šiška & R. Čermák leg.” (JENDEK, 2005 [#7217]); 15 (EJCB): “Sib.or. m., Primorje Sichote Alin Mts. Sokolči 1. 15.7.1990 Kadlec + Voříšek lg.” (JENDEK, 2005 [#7217]); cca 500 (EJCB): “Russia or, Primorskiy Kray, 44°10’00’’N, 133°15’00’’E, Arsen’yev env., June 1991, O. Šauša & M. Štrba leg.” (JENDEK, 2005 [#7217]); 2 ♂, 1 ♀ (EJCB): “Rossia, 2 10.VIII., Primorskij kraj, 1992, Ussurijsk, Kajmanovka, lgt. Smrž” (JENDEK, 2005 [#7217]); 3 ♀ (JMCZ): “Ross. or, Prim. kr., 5 km E Kraskino, 13 16.VII.1992, Boukal leg.”; 1 ♀ (EJCB): “Sib. or., Primorje, Komarovka flum., Kamenushka env., 300 m, Vořišek lgt., VI.1992” (JENDEK, 2005 [#7217]); 1 ♂ (EJCB): “Rossia, Primorskij k., Rjazanovka 4 8.vii.1993, Z. Kletečka leg.” (JENDEK, 2005 [#7217]); 1 ♂ (EJCB): “Russia, Primorski kray, Khankajski distr. 1995, lake Khanka, 17 19.vii., Kamen’ Rybolov” (JENDEK, 2005 [#7217]); 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Tachingousa (Prosiolotchnaja) bay 27.06.04 leg. М.& L. Smirnov’s”; 1 ♂, 1 ♀ (IMCM): “Yu Primor’e, Lazovskii raion, p. Preobrazhenie na usokhshikh Quercus mongolica, 9 14. vii.2006, leg. M. L. Smirnovy [in Russian]”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Preobrazhenie env. 9 17.07.06 leg. М. & L. Smirnov’s”; 1 (MSCI): “SE Maritime Prov., Lazovsky dist., Japan Sea coast, Preobrazhenie env. 14.07.06 leg. М.& L. Smirnov’s”; 1 ♂, 1 ♀ (ZIN): “Primorskii krai, s. Lazo, 1 22.VII.2007, V. Shokhrin [in Russian]”; 1 ♀ (ZIN): “Primorskii krai, s. Lazo, 9 14.07.2008, V. Shokhrin [in Russian]”; 1 ♂ (ZIN): “Primorskii krai, okr. s. Lazo, 06 10.VI.2008, V. Shokhrin [in Russian]”.

Adult occurence: 5-6-7-8 (n=25). Altitude range: 3001300 m (n=3). Host plants: Quercus: ALEXEEV, 1989 [#64]; AKIYAMA & OHMOMO, 1997 (as lasiolus) [#4091]. – Q. acutissima: KUROSAWA, 1974 (as lasiolus) [#4577]. – Q. serrata: KUROSAWA, 1974 (as lasiolus) [#4577]. Distribution: CHINA: Heilongjiang, Jilin, Shaanxi, Shanxi. JAPAN: Kyushu, Tsushima. KOREA NORTH. KOREA

214 SOUTH. RUSSIA: Amurskaya oblast’, Khabarovskiy kray, Primorskiy kray. Remarks: Distributional records without supporting data cited from Xizang and Yunnan (China) by HOU, 1981 [#4198] and HUA LI ZHONG, 2002 [#6696] were omitted.

Agrilus varius Kerremans Fig. 23K (habitus); Fig. 38F (aedeagus); Fig. 54K (type) varius Kerremans, 1895 (Agrilus) KERREMANS, 1895: 221 222 [#1416] (description). KERREMANS, 1903: 292 [#1441] (catalog). JAKOBSON, 1913: 796 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1055 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 359 [#2429] (checklist; China). JENDEK, 1995: 148 [#3768] (lectotype designation). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Anhui). JENDEK, 2006: 402 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2344 [#8576] (world catalog). VOLKOVITSH, 2009: 135 [#8850] (faunal records; Russia: Primorskiy kray).

Type material: Agrilus varius Kerremans, 1895 Type locality: Chine: Ho-Chan. Type specimens: Primary: Lectotype (Fig. 54K), ♀, designated by JENDEK (1995: 148) [#3768], (MNHN): “Chine ‘Ho - Chan R.P. Mouton [p] \ varius Kerr. Type [h] \ Kerremans Vidit 1895 [p]”. Described from unknown number of syntypes. Diagnosis: 4.2-6.2 mm; body less elongate; pronotum markedly enlarged anteriorly; elytral apices truncate with serrulate margin and projecting sutural angle. Species group: Sulcicollis. Relevant species: A. graminis Kiesenwetter, 1857. Specimens examined: CHINA: Henan: 1 ♂, 1 ♀ (EJCB): “China, N. Henan, 11 12.VI., Jiuligou, 550 m, 35º12’N, 112º26’E, Jaroslav Turna leg. 2007”. Hubei: 1 ♀ (EJCB): “P.R. China, Hubei, Dabieshan, N31˚07’06’’ E115˚48’57”, 11 21.vi.2008, 640 m, V. Grebennikov leg.”. Shaanxi: 1 ♀ (EJCB): “China S, Shaanxi, 14 16.6.1995, Qinling Mts. N. slope, 33.56 59’/108.50’ 800 1200 m, Changan Co., L.+R. Businský lgt.”; 1 ♂ (EJCB): “China Yunnan [sic!][should be Shaanxi], Lüyeang, 25.7. 30.7.1996, E. Kučera leg.”; 1 ♀ (EKCS): “China Shaanxi, Lüeyang, 8.6. 14.6.1996, lgt. E. Kučera”; 2 (EKCS): “China Shaanxi, Lüeang 33°07’N 106°05’E, 18.6. 24.6.[19]97 lgt. E. Kučera”; 1 ♀ (EJCB): “China, 1150 1300 m, Shaanxi, Qinling mts., Foping (6 km N), 20 21.VI.1998, I. H. Marshal leg.”; 5 (EJCB): “40 km SE of Taibai Shan Mts., Hou Zen Zi vill. env., Z. Jindra lgt. cca 1200m, China, Shaanxi prov., 11 July 1998, Qing Ling Shan mts.”; 1 ♀ (EKCS): “China Shaanxi, Lueyang, 18.5. 21.5.2000, lgt. E. Kučera”; 1 ♀ (EJCB): “C China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000 1300m, 23.V. 13.VI. leg. C. Holzschuh 2000”; 1 ♀ (EJCB): “C China, Shaanxi, Qinling, Shan, 6 km N of Foping, 1150 1300m, 20 21.VI.2000, leg. C. Holzschuh”; 1 ♀ (EJCB): “C China, Shaanxi, Qinling Shan, 1900 2250m, 12 km SW Xunyangba, 14. 18.VI. leg. C. Holzschuh 2000”; 1 ♀ (EJCB): “China Shaanxi, Lueang env., 15 km NW, 26.5. 31.5.2001, lgt. E. Kučera”; 1 ♀ (EKCS): “China Shaanxi, Lueyang, 29.5. 2.6.2005, lgt. E. Kučera”; 4 ♀ (EKCS): “China Shaanxi, Lueyang, 15.7. 22.7.2005, lgt. E. Kučera”. Sichuan: 1 ♂ (EJCB): “China Sichuan, Pingwu, 32°15’N, 104°16’E, 3 9.6.1997, lgt. E. Kučera”; 2 ♂, 2 ♀ (EKCS): “China N, Sechuan, Nanjiang,

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA 21 23.5.2002, lgt. E. Kučera”. KOREA SOUTH: 2 (JLCS): “Sammachi ri, Hongcheon eup, Hongcheon gun, Gangwon do, Korea, 5.vii.2003, coll. Jung gu Lee”. RUSSIA: Primorskiy kray: 1 ♀ (EJCB): “USSR, Ussuri prov. Arsenev env., VII.1990, leg. B. Šiška & R. Čermák”; 1 (ZIN): “Primorskii krai, Ussuriiskii r n., okr. p. Gornotaezhnyi, 41º43’23’’N, 132º09’19’’E, 19 27.vii.2000, leg. I. Mel’nik [in Russian]”; 1 ♀ (MNCA): “Russia, Ussuri Primor’e region, Arsenjev, 21 26.7.2006, leg. K. Hadulla”.

Adult occurence: 5-6-7 (n=21). Altitude range: 550-2250 m (n=8). Host plants: Unknown. Distribution: CHINA: Anhui, Henan, Hubei, Shaanxi, Shandong, Sichuan. KOREA SOUTH. RUSSIA: Primorskiy kray.

Agrilus ventricosus Fairmaire Fig. 9C (habitus); Fig. 32J (aedeagus); Figs. 47R, 53X, 54L (types) ventricosus Fairmaire, 1888 (Agrilus) FAIRMAIRE, 1888: 120 121 [#887] (description). KERREMANS, 1892: 277 [#1397] (catalog). KERREMANS, 1903: 292 [#1441] (catalog). JAKOBSON, 1913: 796 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 654 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 1055 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 359 [#2429] (checklist; China). JENDEK, 1995: 144 [#3768] (lectotype designation; synonymy; characters; northeastern China; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 42 [#4091] (cited as venticosus; checklist; Japan: Honshu; North China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Beijing). JENDEK, 2006: 403 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2345 [#8576] (world catalog). kalganicus Obenberger, 1935 (Agrilus) OBENBERGER, 1935: 163 164 [#2158] (description). OBENBERGER, 1936: 1004 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 356 [#2429] (checklist; China). JENDEK, 1995: 144 [#3768] (synonym of ventricosus; lectotype designation). AKIYAMA & OHMOMO, 1997: 42 [#4091] (synonym of ventricosus). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China (Kalgan)). JENDEK, 2006: 403 [#7441] (synonym of ventricosus; Palaearctic catalog). BELLAMY, 2008: 2346 [#8576] (synonym of ventricosus; world catalog). toyamai Kurosawa, 1985 (Agrilus) TÔYAMA, KUROSAWA, 1985: 153 156 [#1640] (description). 1985: 24 [#3678] (iconography; Japan). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). JENDEK, 1995: 144 [#3768] (synonym of ventricosus). MORIMOTO & TADAUCHI, 1995: 232 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 42 [#4091] (synonym of ventricosus). JENDEK, 2006: 403 [#7441] (synonym of ventricosus; Palaearctic catalog). BELLAMY, 2008: 2346 [#8576] (synonym of ventricosus; world catalog).

Type material: Agrilus ventricosus Fairmaire, 1888 Type locality: Pekin. Type specimens: Primary: Lectotype (Fig. 54L), ♀, designated by JENDEK (1995: 144) [#3768], (MNHN): “Chine A. David [p] \ Agrilus ventricosus Fairm. [h] [by Fairmaire hand]”. Described from unknown number of syntypes. Agrilus kalganicus Obenberger, 1935 Type locality: China: Inshan, Kalgan.

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Type specimens: Primary: Lectotype (Fig. 47R), ♀, designated by JENDEK (1995: 144) [#3768], (NMPC): “Inshan Kalgan [ = Zhangjiakou Shi, Hebei province] China [h] \ Typus [p] [red label] \ Agrilus kalganicus m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes. Agrilus toyamai Kurosawa, 1985 Type locality: Takahachi-yama, Tottori Pref., Honshu, Japan. Type specimens: Primary: Holotype (Fig. 53X), ♂, (NSMT): “[Japanese grapheme] 1979. 8. 16 [Japanese grapheme] leg. [h] \ Holotype [p] Agrilus toyamai Y. Kurosawa, 1985 [h] [red label with black border]”. Secondary: 2 PT (NSMT). Described from 11 specimens (HT, AT, PT). Diagnosis: 8.4-12 mm; prehumerus very short, carinate and distinctly arcuate; elytra with 1 or 2 pairs of vague pubescent spots; elytral apices widely subarcuate. Species group: Biguttatus. Relevant species: A. biguttatus (Fabricius, 1777); A. decoloratus Kerremans, 1892. Specimens examined: CHINA: 2 (MNHN): “Chine, A. David”; 1 (MNHN): “Tsienou 26.VIII.[19]34” (JENDEK, 1995 [#3768]). Nei Mongol: 1 (MNHN): “Chahar Yankiaping \ 3.VII.[19]31 O. Piel coll.” (JENDEK, 1995 [#3768]).

Adult occurence: 7-8 (n=5). Faunal records published by KUROSAWA, 1985 [#1640] (as toyamai) are also included. Host plants: Celtis bungeana var. jessoensis: KUROSAWA, 1985 (as toyamai) [#1640]; AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: CHINA: Beijing, Hebei, Nei Mongol. JAPAN: Honshu. Remarks: The holotype of A. toyamai Kurosawa, 1985 differs from A. ventricosus Fairmaire, 1888 by the deeper lateral pronotal impressions and the more sharply carinal prehumerus. More specimens are needed to resolve the status of these two nominal taxa.

Agrilus viduus Kerremans Fig. 14E (habitus); Fig. 34J (aedeagus); Figs. 44D, 44J, 47D, 54N, (types) viduus Kerremans, 1914 (Agrilus) KERREMANS, 1914: 105 [#1481] (description). MIWA, 1931: 125 [#6101] (catalog; Formosa). MIWA & CHŰJÔ, 1936: 21 [#1809] (catalog; Japan). OBENBERGER, 1936: 1107 [#2191] (world catalog). KUROSAWA, 1957: 192 [#1615] ( notes). KUROSAWA, 1963: 152 [#1618] (characters; Japan). KUROSAWA, 1975: 3 [#4591]. TÔYAMA, 1985: 30 [#3678] (iconography; Japan). ALEXEEV, 1989: 487 [#64] (key to species; distributional summary). LI JINGKE, 1992: 93 [#6755] (checklist; northeastern China). AKIYAMA & OHMOMO, 1997: 42 [#4091] (checklist; Taiwan; China). ALEXEEV, 1998: 381 [#4322] (subgenus Agrilus; subgeneric classification). JENDEK, 2000: 506 [#4851] (lectotype designation). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China: Taiwan). MÜHLE, 2003: 47 [#6155] (checklist; Taiwan). JENDEK, 2006: 389 [#7441] (subgenus Agrilus; Palaearctic catalog). BELLAMY, 2008: 2348 [#8576] (subgenus Agrilus; world catalog). hoscheki Obenberger, 1916 (Agrilus) syn. nov.

215 OBENBERGER, 1916: 273 [#1983] (description). OBENBERGER, 1924: 45 [#2040] (note). OBENBERGER, 1926: 655 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 997 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 356 [#2429] (checklist; China). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Guangdong). JENDEK, 2006: 399 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2127 [#8576] (world catalog). chinganicus Obenberger, 1922 (Agrilus) syn. nov. OBENBERGER, 1922: 27 28 [#2021] (description). OBENBERGER, 1926: 657 [#2057] (Palaearctic catalog). OBENBERGER, 1936: 969 [#2191] (world catalog). KUROSAWA, 1957: 192 [#1615] (subspecies of viduus; characters). KUROSAWA, 1975: 3 [#4591] (subspecies of viduus). ALEXEEV, 1979: 137 [#57] (subgenus Agrilus; subspecies of viduus; redescription; notes). PENG ZHONGLIANG, 1987: 355 [#2429] (checklist; China). ALEXEEV, 1989: 487 [#64] (subspecies of viduus; cited as chinensis rather than chinganicus; key to species; northeastern China; north Mongolia). ALEXEEV & VOLKOVITSH, 1989: 322, 351, 357 [#3241] (subspecies of viduus; characters in key; faunal records; Mongolia). ALEXEEV, 1998: 381 [#4322] (subgenus Agrilus; subgeneric classification). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Inner Mongolia (Chingan), Shandong). JENDEK, 2006: 389 [#7441] (subgenus Agrilus; subspecies of viduus; Palaearctic catalog). BELLAMY, 2008: 2348 [#8576] (subspecies of viduus; world catalog). VOLKOVITSH, 2009: 134 [#8850] (subgenus Agrilus; subspecies of viduus; faunal record; Russia: Primorskiy kray). cardaces Obenberger, 1936 (Agrilus) syn. nov. OBENBERGER, 1936: 38 [#2200] (description). MIWA & CHŰJÔ, 1936: 15 [#1809] (catalog; Japan). OBENBERGER, 1936: 1077 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 354 [#2429] (checklist; China). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Taiwan). MÜHLE, 2003: 46 [#6155] (checklist; Taiwan). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2014 [#8576] (world catalog). subviduus Kurosawa, 1957 (Agrilus; ssp. of viduus) syn. nov. KUROSAWA, KUROSAWA, 1957: 192 [#1615] (description). 1963: 152 [#1618] (subspecies of viduus; characters; Japan). KUROSAWA, 1975: 3 [#4591] (subspecies of viduus). TÔYAMA, 1985: 30 [#3678] (subspecies of viduus; iconography; Japan). HIRASHIMA, 1989: 324 [#8078] (subspecies of viduus; checklist; Japan). ANONYMOUS, 1991: 106 [#8552] (subspecies of viduus; faunal records; Japan: Aomori). MORIMOTO & TADAUCHI, 1995: 232 [#8080] (subspecies of viduus; checklist; Japan). AKIYAMA & AKIYAMA, 1996: 186 [#3962] (subspecies of viduus; faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 42 [#4091] (subspecies of viduus; checklist; Japan: Honshu; Kyushu, Tsushima). JENDEK, 2006: 389 [#7441] (subgenus Agrilus; subspecies of viduus; Palaearctic catalog). BELLAMY, 2008: 2348 [#8576] (subspecies of viduus; world catalog).

Type material: Agrilus viduus Kerremans, 1914 Type locality: [Formosa] Kankau: Koshun. Type specimens: Primary: Lectotype (Fig. 54N), ♀, designated by Jendek (2000: 506) [#4851], (DEI): “Kankau (Koshun) Formosa H. Sauter VII.1912 [p] \ Syntypus [p] [red label] \ Kerremanns [sic!] det. [p] \ Agrilus viduus Kerr. Type [h] [Kerremans’ MS] \ Coll. DEI Eberswalde [p]”. Secondary: 1 PLT (MNHN). Described from unknown number of syntypes. Agrilus hoscheki Obenberger, 1916 Type locality: Kiautschou (China).

216 Type specimens: Primary: Lectotype (Fig. 47D) by present designation, ♂ [aedeagus missing] (NMPC): “Kiau-Tchou, China [h] \ Typus [p] [red label] \ Agrilus Hoscheki m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24258 [h] [orange label]”. Described from unknown number of syntypes. Agrilus chinganicus Obenberger, 1922 Type locality: China bor. or.: Chingan. Type specimens: Primary: Lectotype (Fig. 44J) by present designation, ♂, (NMPC): “Linsihieng. Chingan mont. Chine [h] \ Typus [p] [red label] \ Agrilus chinganicus m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24780 [h] [orange label] \ Agrilus viduus Kerr. [h] Alexeev det. [p] ssp. chinganicus Obenb. [h]”. Described from unknown number of syntypes. Agrilus cardaces Obenberger, 1936 Type locality: Formosa. Type specimens: Primary: Lectotype (Fig. 44D) by present designation, ♀, (NMPC): “Formosa Dovnou [h] \ Typus [p] [red label] \ Agrilus cardaces m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 26860 [h] [orange label]”. Described from unknown number of syntypes. Agrilus viduus subviduus Kurosawa, 1957 Type locality: Nakatsugawa Valley, Niigata Pref. Type specimens: Holotype, ♂, (NSMT). Described from 12 specimens (HT, AT, PT). Types not examined. Diagnosis: 5.5-8.8 mm; prehumerus present, filamentar or finely carinate, elytra often with obvious perisutural stripes of pubescence; aedeagus as in Fig. 34J. Species group: Viduus. Relevant species: A. compacticornis sp. nov.; A. shan sp. nov. Specimens examined: CHINA: Beijing: 1 ♂, 1 ♀ (EJCB): “Peking, ex Ulmus, Kubáň leg.”. Gansu: 1 (EJCB): “China, 7 1989, Gansu prov., Linxia”. Nei Mongol: 1 ♂ (EJCB): “China, Nei Mongol, Wuhai, 8.5. 13.5.1996, E. Kučera leg.”; 9 (EKCS): “China Nei Mongol, Baotou, 27.7. 28.7.2007, lgt. E. Kučera [reared from Prunus persica, Kučera pers.com.]”. Shaanxi: 4 ♂, 2 ♀ (EJCB): “China S, Shaanxi, 14 16.6.1995, Qinling Mts. N. slope, 33.56 59’/108.50’ 800 1200 m, Changan Co., L.+R. Businský lgt.”; 2 (EKCS): “China Shaanxi, Lüeang 33°07’N 106°05’E, 18.6. 24.6.[19]97 lgt. E. Kučera”; 35 (EJCB): “C China, Shaanxi, Qinling Shan, 6 km E of Xunyangba, 1000 1300m, 23.V. 13.VI. leg. C. Holzschuh 2000”; 2 (EJCB): “China Shaanxi, Lüeang, 22.5. 25.5.2000, leg. E. Kučera”; 1 ♂ (EJCB): “China Shaanxi, 15 km N Lueyang, 2.7 8.7. 2007, lgt. E. Kučera”; 4 (EKCS): “China Shaanxi, 15 km N Lueyang, 2.7 8.7. 2007, lgt. E. Kučera”. Shanxi: 1 ♂, 1 ♀ (EJCB): “China, W Shanxi, 9.VI.2000, 37.7N, 111.6E, Lüliang Shan, road Fangshan Jiaocheng, Hengjian env., ~1000m, Jaroslav Turna leg.”. Sichuan: 1 (EJCB): “Sichuan, Xiangchen, 1982.vi.22”. Taiwan: 1 (EJCB): “Sengpei, Kaohsiung Hsien, Taiwan, 2.x.1980, Chen Wenlong leg.”; 2 (EJCB): “Mt. Dayuan shan, Kaohsiung Hsien, Taiwan, 2.viii.1988, Chen Wenlong leg.”; 1 (EJCB): “Mt. Dayuan shan, Kaohsiung Hsien, Taiwan, 22.viii.1988, Chen Wenlong leg.”; 1 (EJCB): “Chi Ai, Puli, Nantou Hsien, Rep. of China, V11 18 1972, F. J. Santana”; 5 ♂, 2 ♀ (EJCB): “Taiwan, Nantou, 1997, Lien Hua Chih, 4 km S Wuchen, 23 27 VI., B. Šiška & T. Spevár lgt.”. JAPAN: Honshu: 1 (JMCZ): “Japan, 4.7.[19]72, Takahashi, Okayama, T.Aono. lgt.”; 1 (EJCB): “Japan, 4.7.1972, Takahashi, Okayama, T. Aono lgt.”; 1 (EJCB): “Moniwa, Fukushima Pref., Japan, 29.v.1977, leg. S. Ohmomo”; 1 ♂, 1 ♀ (EJCB): “Moniwa, Iizaka Town, Fukushima City, Fukushima Pref., Japan, 25 June

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA 1995, Nobuo Sagiura leg.”; 2 ♀ (EJCB): “Japan, Yamagata, Myozawa, Oguni town, 20.vii.2005, N. Sugiura leg., Celtis sp. dead trunk”. Tsushima: 2 (EJCB): “Arisawa Rindō, Tsushima Is., vi.1984 em, Y. Matsumoto”; 1 (EJCB): “Arisawa Rindō, Tsushima Is., vii.1984 em, Y. Matsumoto”. KOREA NORTH: 1 (EJCB): “Korea, Onpho, ad Chongjin, 20.viii.1959, leg. B. Pisarski et Proszynski, J.”. KOREA SOUTH: 1 (JLCS): “Mt. Myeongjisan, Gyeonggi do, Korea, 7.vi.1992, coll. W. S. Kim”; 1 (JLCS): “Mt. Gayasan, Junggeun dong, Gwangyang si, Jeollanam do, Korea, 30 31.viii.2000, night, coll Tae ho An”. RUSSIA: Buryatiya Respublika: 1 ♂ (ZIN): “Buryatsk. ASSR, Gusinozersk. r n., c. Povorot (na Tchikoe), na Ulmus pumila (lyet), 25.vi.1976, G. Lindeman [in Russian]”. Primorskiy kray: 1 ♂, 2 ♀ (EJCB): “Sib. or. m., Primorje, Sichote Alin Mts., Sokolči, 1 15.7.1990, Kadlec + Voříšek lg.”; 2 ♂ (ZIN): “Primorskii krai, s. Lazo, 1 22.VII.2007, V. Shokhrin [in Russian]”; 3 (ZIN): “Russia, Far East, Lazovkiy reserve, 27.06 22.07.2007, V. Shokhrin”.

Adult occurence: 5-6-7-8-10 (n=36). Faunal records published by KUROSAWA, 1957 [#1615] (as viduus subviduus) are also included. Altitude range: 800-1300 m (n=3). Host plants: Celtis: see Specimens examined. – Prunus persica: see Specimens examined. – Ulmus: KUROSAWA, 1957 (as viduus subviduus) [#1615]; ALEXEEV, 1979 (as viduus chinganicus) [#57]; ALEXEEV, 1989 (as viduus chinganicus) [#64]. – U. pumila: ALEXEEV & VOLKOVITSH, 1989 (as viduus chinganicus) [#3241]. – U. wilsoniana: KUROSAWA, 1957 (as viduus chinganicus) [#1615]; KUROSAWA, 1975 (as viduus subviduus) [#4591]. – Zelkova serrata: KUROSAWA, 1957 (as viduus subviduus) [#1615]; AKIYAMA & OHMOMO, 1997 (as viduus subviduus) [#4091]. Distribution: CHINA: Beijing, Gansu, Guangdong, Hebei, Nei Mongol, Shaanxi, Shandong, Shanxi, Sichuan. JAPAN: Honshu, Kyushu, Tsushima. KOREA NORTH. KOREA SOUTH. MONGOLIA. RUSSIA: Primorskiy kray, Respublika Buryatiya, Respublika Tyva. Remarks: This species is very variable in size, color, extent of pubescence and tomentum, form and lenght of prehumerus, extent of prosternal lobe emargination as well as length and form of elytral apices. This variability does not seem to be geographically correlated.

Agrilus vigil Jendek, sp. nov. Fig. 18K (habitus of HT)

Description: Holotype 8.1 mm. BODY: Shape: subparallel, Build: robust, Color (dorsally): unicolored, Color (dominant): blue. HEAD: Medial impression: present, Impression (extent): vertex and frons, Impression (depth): deep, Epistoma: with raised upper margin. Frons: Shape: moderately convex. Vertex: Sculpture on vertex: rugae, Sculpture (density): dense. Eyes: Size: large, Shape: protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Shape: transverse, Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: moderate, Shape: arcuate, Position to anterior angles: not reaching level of anterior pronotal angles. Posterior angles: Shape: acute or rectangular, Apex: sharp. Disk: Convexity:

217

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA moderately convex, Disk impressions: medial only or medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Lateral impressions (deepness): shallow, Lateral impressions (width): narrow. Prehumerus: carinal, Shape: arcuate, Extent: to half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: close to pronotal marginal carina, Arc: obvious. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior 1/5-1/4 of pronotal length. Scutellum: Size: large, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Shape: cuspidate, Position of dominant cusp (spine): sutural margin. Tomentum: spots, Spots (locations): postmedial, rarely premedial. STERNUM: Metasternal projection: flat. Prosternal lobe: large, Anterior margin: arcuately emarginate, Emargination (deepness): moderate or deep, Emargination (width): moderate. Prosternal process: moderate, Shape: narrowed, Sides: straight or arcuate, Angles: absent or obtuse, Angles (tips): blunt. ABDOMEN: Sexual modifications in male: not apparent, Apical margin of pygidium: extended into long spine, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Ovipositor: elongate. Diagnosis: eyes markedly large, distinctly projecting beyond head outline; frons and vertex with deep medial carina. Species group: Laetecyanescens. Relevant species: A. yunnanus Obenberger, 1927. Type locality: China, Fo-kien [= Fujian] province. Type specimens: Holotype (Fig. 18K), ♀, (EJCB): Holotype, ♀, (EJCB): “China, prov. Fo-kien, G. Siemssen, vend. 4.ix.1913”. Adult occurence: 9 (n=1). Host plants: Unknown. Distribution: CHINA: Fujian. Etymology: The specific name is the Latin adjective vigil (awake, watchful). It refers to the large eyes of the species.

Agrilus violaceus Nonfried sp. inquirenda violaceus Nonfried, 1892 (Agrilus) NONFRIED, 1892: 85, 91 [#1924] (description). KERREMANS, 1900: 342 [#1434] (note). KERREMANS, 1903: 288 [#1441] (synonym of nonfriedi). OBENBERGER, 1923: 64 [#2026] (note). OBENBERGER, 1936: 1016 [#2191] (synonym of nonfriedi). JENDEK, 2006: 60, 403 [#7441] (valid name not homonym with violacellus Thomson, 1879; Palaearctic catalog). BELLAMY, 2008: 2350 [#8576] (world catalog). nonfriedi Kerremans, 1900 (Agrilus; unnecessary replacement name for violaceus Nonfried not violacellus Thomson) KERREMANS, 1900: 342 [#1434] (replacement name proposal). KERREMANS, 1903: 288 [#1441] (catalog). OBENBERGER, 1923: 64 [#2026] (note). OBENBERGER, 1936: 1016 [#2191] (world

catalog). PENG ZHONGLIANG, 1987: 357 [#2429] (checklist; China). HUA LI ZHONG, 2002: 90 [#6696] (checklist; China). JENDEK, 2006: 60, 403 [#7441] (synonym of violaceus; Palaearctic catalog). BELLAMY, 2008: 2350 [#8576] (synonym of violaceus; world catalog).

Type material: Agrilus violaceus Nonfried, 1892 Type locality: not given [“Südchina, Kwang-Si, Umgebung von Nienghali 24 nördl. Breite und 108 östl. Länge” is cited in the introduction]. Type specimens: Not located. Described from a single specimen. Agrilus nonfriedi Kerremans, 1900 Type locality and specimens: See: Agrilus violaceus Nonfried, 1892. Host plants: Unknown. Distribution: CHINA: Guangxi. Remarks: This taxon is species inquirenda and its name is nomen dubium.

Agrilus viridiobscurus Saunders Fig. 28F (habitus); Fig. 40J (aedeagus); Fig. 54O (type) viridiobscurus Saunders, 1873 (Agrilus) SAUNDERS, 1873: 515 516 [#2554] (description). LEWIS, 1879: 15 [#3345] (catalog; Japan). KERREMANS, 1885: 154 [#1381] (catalog). SCHÖNFELDT, 1887: 113 [#8901] (Anambus; catalog; Japan). KERREMANS, 1892: 277 [#1397] (catalog). KERREMANS, 1903: 292 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 21 [#1809] (catalog; Japan). OBENBERGER, 1936: 1056 [#2191] (world catalog). KUROSAWA, 1949: 15 [#5028] (characters). KUROSAWA, 1963: 154 [#1618] (characters; Japan). KUROSAWA, 1975: 3 [#4591] (cited as viridiobsculus). TÔYAMA, 1985: 20 [#3678] (iconography; Japan). JENDEK, HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). 1995: 148 [#3768] (lectotype designation; Japan). MORIMOTO & TADAUCHI, 1995: 232 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 187 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 42 43 [#4091] (cited as viridiobsculus; checklist; Japan: Hokkaido, Honshu; Shikoku, Kyushu). AKIYAMA & OHMOMO, 2000: 88 [#6240] (faunal records; Japan (Hiroshima)). FUKUTOMI & HORI, 2004: 6 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 403 [#7441] (Palaearctic catalog). ANONYMOUS, 2007: 32 [#8250] (cited as viridiobsculus; checklist; Japan: Honshu). BELLAMY, 2008: 2352 [#8576] (world catalog).

Type material: Agrilus viridiobscurus Saunders, 1873 Type locality: Japan. Type specimens: Primary: Lectotype (Fig. 54O), ♀, designated by JENDEK (1995: 148) [#3768], (BMNH): “Type H.T. [p] [round label with red border] \ Japan G. Lewis 1910320 [p] \ Agrilus viridiobscurus ES Type [h]”. Secondary: 1 PLT (BMNH). Described from unknown number of syntypes. Diagnosis: 3.8-5.4 mm; vertex protruding beyond head outline (posterodorsal view); medial pronotal impression visible only in proximal part. Species group: Viridiobscurus. Relevant

218 species: A. caudicula sp. nov.; A. foramenifer sp. nov. Specimens examined: CHINA: Henan: 1 ♀ (EJCB): “China, N Henan, 13.vi., Wangwushan, 1175 m, 35°11’N, 112°16’E, Jaroslav Turna leg., 2007”; JAPAN: Honshu: 1 (EJCB): “Mt. Iwawaki, Japan, 28.7.1963, Mizobe”; 1 ♀ (JMCZ): “Mt. Gragyu, Okayama pref. Japan, May 14, ‘67, T. Aono leg.”; 2 (EJCB): “Fussa City, Tokyo Met., 28.IV.1977, leg. A. Yamagami \ Host Plant Zelkova” (JENDEK, 1995 [#3768]); 1 ♀ (EJCB): “Japan, Honshu, Osaka fu, Iwawakiyama, 700 m, 6 km S of Kagata, 2 Jul 2002, Bolm lgt.”; 5 ♂, 2 ♀ (EJCB): “Japan, Honshu, Osaka fu, Kongosan, 300 m, 2 km E of Amami, 12. May 2002, Bolm Leg.”.

Adult occurence: 4-5-6-7 (n=6). Altitude range: 300-1175 m (n=3). Host plants: Alnus pendula: AKIYAMA & OHMOMO, 1997 [#4091]. – Carpinus tschonoskii: AKIYAMA & OHMOMO, 1997 [#4091]. – Hamamelis japonica: AKIYAMA & OHMOMO, 1997 [#4091]. – Juglans mandschurica var. sieboldiana: AKIYAMA & OHMOMO, 1997 [#4091]. – Parabenzoin praecox: AKIYAMA & OHMOMO, 1997 [#4091]. – Zelkova serrata: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: CHINA: Henan. JAPAN: Hokkaido, Honshu, Kyushu, Shikoku.

Agrilus viridis (Linné) Figs. 16C 16I (habitus); Fig. 35C (aedeagus); 43D, 43U, 43Y, 43X, 44C, 44E, 44P, 45M, 46P, 47F, 47W, 48B, 49G, 49O, 51D, 51O, 52H, 52N, 53Z, 54M, 54P, Figs. (types) viridis Linné, 1758 (Buprestis) LINNÉ, 1758: 410 [#1726] (description). LINNÉ, 1760: 214 [#1727] (Buprestis; notes; Suecia). FABRICIUS, 1781: 281 [#832] (Buprestis; characters; synonymy; Europe). GMELIN, 1790: 1937 [#1087] (Buprestis; characters; synonymy). FABRICIUS, 1792: 213 [#834] (Buprestis; characters; synonymy; references). PANZER, 1795: 232 [#4041] (Buprestis; characters; Germania). CEDERHIELM, 1798: 79 [#5314] (Buprestis; characters; references). FABRICIUS, 1801: 212 [#837] (Buprestis). MARSHAM, 1802: 397 [#6052] (Buprestis; characters; references). SCHÖNHERR, 1817: 244 245 [#3895] (Buprestis; synonymy; bibliography). DEJEAN, 1821: 30 [#4046] (Buprestis; catalog). FISCHER, 1829: 34 [#8133] (Buprestis; checklist). GEBLER, 1829: 79 [#3008] (notes; Altay). DEJEAN, 1833: 83 [#689] (Fabricius is cited as the author; catalog). DEJEAN, 1836: 94 [#692] (Fabricius is cited as the author; catalog). AUBÉ, 1837: 189 192 [#4034] ([Note: Species cited in this publication under the name viridis was described by GORY & LAPORTE, 1839 [#3738] as aubei]; larva; France: Boulogne). MANNERHEIM, 1837: 113 [#3018] (note). GORY & LAPORTE, 1839: 48 49 [#3738] ([Note: Misidentification of tenuis according to KIESENWETTER, 1857: 132 [#3015]]). LAPORTE, 1840: 220 [#4045] (characters). GEBLER, 1847: 121 [#4423] (Fabricius is cited as the author; faunal records; southwest Siberia). KIESENWETTER, 1857: 151 153 [#3015] (characters in key, redescription; notes; Germany). MARSEUL, 1857: 92 [#4371] (checklist; Europe). MOTSCHULSKY, 1859: 157, 275 [#1832] (catalog; bibliography on larval biology). MARSEUL, 1866: 436, 448 449 [#676] (characters in key; redescription; references; faunal records; distribution overview; larva; France). BOISDUVAL, 1867: 109 111 [#6844] (pest in horticulture; biology; France). GOUREAU, 1867: 24 26 [#7946] (Eschscholtz is cited as the author; life cycle). GEMMINGER & HAROLD, 1869: 1447 [#1057] (catalog). SAUNDERS, 1870: 35 [#2551] (catalog). SAUNDERS, 1871: 119 [#2552] (catalog). SOLSKY, 1871: 360 [#4756] (faunal

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA records; East Siberia). HOCHHUTH, 1873: 158 [#3453] (faunal records; Gouvernements Kiew und Volhynien). RUPERTSBERGER, 1880: 157, 275 [#8550] (commented larval bibliography, checklist). HEYDEN, 1881: 118 [#5548] (catalog). KEPPEN, 1882: 230 231 [#8636] (characters; biology; pest; notes). MARSEUL, 1887: 251 [#4707] (catalog). JUDEICH & NITSCHE, 1889: 319, 322 [#5686] (characters; biology; host plants overview). KERREMANS, 1892: 277 [#1397] (catalog). HEYDEN, 1893: 90 [#3450] (catalog). RUPERTSBERGER, 1894: 160, 287 [#4465] (larval bibliography; catalog). ABEILLE DE PERRIN, 1897: 4, 14 [#19] (characters in key; notes). KULIKOWSKI, 1897: 150 [#8666] (checklist; south Russia). KERREMANS, 1903: 275 [#1441] (catalog). ELLIOTT & MORLEY, 1907: 16 [#8725] (hymenopterous parasitoids). POMERANCEVA, 1908: 474 [#8667] (faunal record; Arkhangel’sk). CAILLOL, 1913: 509 510 [#4053] (biology). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). REH, 1913: 487 488 [#8153] (biology; references). BAU, 1914: 257 [#8668] (characters in key). OBENBERGER, 1916: 270 272 [#1983] (characters in key; infraspecific division). WEISS, 1916: 426 [#6742] ([Note: Misidentification of cuprescens]; faunal record; New Jersey). WEISS, 1916: 287 [#3913] ([Note: Misidentification of cuprescens]; faunal record; New Jersey). BEDEL, 1921: 200, 204, 212 213 [#170] (characters in key; references; faunal records; distribution; France). KROGERUS, 1922: 110, 111, 113 [#3916] (biology). ESCHERICH, 1923: 145, 149 [#6717] (characters in key; biology; larva; pest; control measures). OBENBERGER, 1926: 656 [#2057] (Palaearctic catalog). JAKOBSON, 1927: 179 [#8655] (characters in key). OBENBERGER, 1927: 197, 198, 203, 208, 209, 210 211, 213, 215, 225 227, 248 [#2064] (notes). DEKHTIAREV, 1928: 20 [#8663] ([Note: Misidentification of ribesi according to host plant cited]; pest; Ukraine). JAKOBSON & OGLOBLIN, 1931: 162 [#8314] (characters in key). OGLOBLIN & REICHARDT, 1932: 278 [#8654] (pest). OBENBERGER, 1936: 1056 1068 [#2191] (world catalog). POMERANCEV, 1938: 17 18 [#4977] (biology; notes). MIWA, 1940: 72 [#4586] (cited as “viridri Linné var.”; [Note: Misidentification of sinuatus yokoyamai according to KUROSAWA, 1975: 1 2 [#4578]]; checklist; Japan). MIWA & CHÛJÔ, 1940: 74 [#1810] (cited as viridri; faunal records; Sakhalin: Konuma). RIKHTER, 1945: 165 [#3507] (characters in key; European part of USSR, north to White Sea (Solovetskiye ostrova)). PALMÉN, 1946: 120 [#7730] (faunal records; Russia (Karelia)). IGA, 1947: 17 [#3273] (cited as “viridis Linnaeus var.”; [Note: Misidentification of sinuatus yokoyamai according to KUROSAWA, 1975: 1 2 [#4578]]). KANGAS, 1947: 108 113 [#3622] (notes on larval characters; Finnland). ILINSKIJ, 1948: 295 [#5670] (larval characters in key; notes). RIKHTER, 1948: 393 [#3879] (characters in key; European part of USSR). STEPANOV, 1948: 243, 244 [#3535] (checklist; faunal record; Chelyabinskaya oblast’). SCHAEFER, 1949: 355, 397 402, 491 [#2565] (characters; references; faunal records; biology; notes; ditributional overview; notes; France). RIKHTER, 1950: 158, 165 [#3881] (adult and larval characters in key). ALLEN, 1951: 29 30 [#6697] (notes on color variability). GRECHKIN, 1951: 51 66 [#8606] ([Note: Misidentification of suvorovi according to host plant cited]; larva; biology; life cycle; parasitoids; USSR: Moscow, Penza, Dnepropetrovsk). WOELFLE, 1951: 118 [#7086] (phytopatologic importance; Germany). ZAKHAROV & LEVKOVICH, 1951: 298, 299, 301, 302, 305 [#8638] (pest; Russia: Saratov; Volgograd regions). HUSSON et al., 1952: 250 255 [#7085] (phytopatologic importance; biology; Germany). MÜHLMANN, 1954: 82 [#8152] (notes on biology). HORION, 1955: 93 94 [#1285] (distribution overview; faunal records; biology; Deutschland; Österreich). STARK, 1955: 775 [#3533] (faunal record; USSR: Saratov). TER MINASYAN, 1955: 453 454 [#7893] (host plant; distribution; references). HEERING, 1956: 249 287 [#3716] (life cycle; biology; ecology; etology; parasitoids; eradication; Germany). HEERING, 1956: 76 114 [#1195] (life cycle; biology; ecology; ethology; parasitoids;

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA eradication; Germany). ALEXEEV, 1957: 152 [#3226] (characters in key). PLAVILSHCHIKOV, 1957: 210 [#8254] (characters; European part of Soviet Union). ZAGAIKEVICH, 1957: 22 [#4976] (biology; Ukraine). ZAKHARCHENKO, 1957: 64 [#8670] (larva; characters in key; Volga estuary). BASURMANOVA, 1958: 1039 1044 [#4974] (two biological forms on Acer; biology; larva; characters; Ukraine: Voroshilovgradskaya oblast’; Russia: Rostovskaya oblast’). SLIVKINA, 1958: 166 167 [#8660] ([Note: Misidentification of suvorovi according to host plant cited]; notes; north Kazakhstan). ALEXEEV, 1959: 13 [#3229] (characters in key). KHARAZISHVILI, 1960: 147 [#8630] ([Note: Misidentification of suvorovi according to host plant cited]; pest; Georgia). LOZOVOI, 1960: 134 [#8627] (notes; Transcaucasia). PADIJ, 1960: 141 [#8628] (pest; Ukraine). ZAGAIKEVICH, 1960: 257, 258 [#5615] (checklist; biology; Ukraine). ALEXEEV, 1961: 9 [#3243] (larval characters in key). BALACHOWSKI et al., 1962: 287, 292 294 [#96] (characters; biology; distribution; economic importance; bibliography). GRECHKIN & VORONCOV, 1962: 57 60 [#8242] ([Note: Misidentification of suvorovi according to host plant cited]; biology). ZAGAIKEVICH, 1962: 105 106 [#3249] (faunal records; biology; Ukraine; Krym). ZAGAIKEVICH, 1962: 60 [#5616] (checklist; Ukraine). SAMEDOV, 1963: 92, 100 [#6091] (characters; biology; host plants; Azerbaijan). KHRAMCOV & PADIJ, 1965: 80 82 [#8649] (characters; larva; biology; harms; eradication; USSR). RIKHTER & ALEXEEV, 1965: 299 [#2522] (characters in key; European part of USSR). ALEXEEV, 1971: 225 [#73] (phylogenetic origin). KOSTIN, 1973: 107 [#3693] (characters in key; Kazakhstan). GURYEVA, 1974: 101 [#5428] (pest of agriculture cultures; biology; references; USSR). KUROSAWA, 1974: 2 [#4577] (notes; Japan). ALEXEEV, 1975: 157 [#52] (notes; faunal records; distributional overview; Mongolia). ADELI & SOLEIMANI, 1976: 134 [#8300] (abundance; remarks; Iran). VOLKOVITSH, 1977: 25 [#2985] (faunal record; notes; USSR: Belgorod). LOMPE, 1979: 240 [#3628] (characters in key; Central Europe). ALEXEEV, 1981: 76 77 [#61] (larval characters in key). HOU, 1981: 395 [#4198] (notes; China: Xizang). SAVKOVSKII & RYBALOV, 1981: 82 86 [#6092] ([Note: Misidentification of ribesi according to host plant cited]; biology; Ukraine). BÍLÝ, 1982: 64, 68, 82 83 [#295] (characters in key; distribution overview; biology). ČAPEK et al., 1982: 350 [#6200] (hymenopterous parasitoids). ABAI & ADELI, 1984: 10, 15, 41, 89 [#5808] (pest; Iran). KRYZHANOVSKII, 1984: 20, 107 [#8665] (pest; USSR). TÔYAMA, 1985: 23 [#3678] (iconography; Japan). COBOS, 1986: 253 255, 317 [#609] (subgenus Agrilus; characters in key; distribution; synonymy; bionomy). KALASHIAN, 1986: 95 [#3865] (note on structure of ovipositor). KALASHIAN, 1986: 125 [#3866] (ovipositor structure). HOU, 1987: 588 589 [#3490] (notes; Yunnan). PENG ZHONGLIANG, 1987: 359 [#2429] (checklist; China). ALEXEEV, 1989: 488 [#64] (key to species, distribution overview). ALEXEEV & VOLKOVITSH, 1989: 350, 357 [#3241] (characters in key; Mongolia). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). LI JINGKE, 1992: 92 [#6755] (checklist; northeastern China). NIEHUIS & TEZCAN, 1993: 37, 46, 47 [#1910] (subgenus Agrilus; faunal records; distribution and host plants summary; Turkey). BÍLÝ, 1994: 112 [#8200] (larval characters in key). CURLETTI, 1994: 22, 173 175 [#658] (subgenus Agrilus; catalog; faunal records; Italia; Sicilia; Corsica; Sardegna). VOLKOVITSH & ALEXEEV, 1994: 437 [#2974] (checklist; Turkmenistan). MORIMOTO & TADAUCHI, 1995: 232 [#8080] (checklist; Japan). LAGUNOV & NOVOZHENOV, 1996: 47 [#7461] (checklist; Russia (Chelyabinsk)). OVTCHINNIKOV, 1996: 132 [#8621] (checklist; Kyrgyzstan). ALEXEEV, 1998: 380, 381 [#4322] (subgenus Agrilus; subgeneric classification). ALEXEEV & SLEPKO, 1998: 52 72 [#4887] (population persistence on riverine islands; North Caucasus). CURLETTI, 1998: 133 [#4357] (subgenus Agrilus; cited as type species of sg. Agrilus by original designation). TLEPPAEVA, 1999: 184 [#6551] (notes; Kazakhstan). BÍLÝ, 2002: 28 29 [#5877] (bionomy; host plants; distribution).

219 BRECHTEL & KOSTENBADER, 2002: 312 316, 326 328 [#5766] (biology; ecology; distribution). FRYDRYCHOVÁ & MAREC, 2002: 179 187 [#8240] (chromosomes). HUA LI ZHONG, 2002: 90 91 [#6696] (checklist; China: Heilongjiang, Jilin, Liaoning, Hebei, Henan, Shandong, Shaanxi, Xinjiang, Hubei, Fujian, Yunnan, Xizang). LIN GUI RUI, 2002: 234 [#7926] (checklist; pest). KADYRBEKOV et al., 2003: 40 [#7318] (notes; Kazakhstan). KENIS & HILSZCZANSKI, 2004: 489 493 [#8566] (natural enemies). TLEPPAEVA & ISHKOV, 2004: 84 [#7211] (subgenus Agrilus; checklist; Kazakhstan). BERNHARD et al., 2005: 601, 602 [#7261] (mitochondrial DNA; molecular systematics). DEDYUKHIN, NIKITSKY & SEMENOV, 2005: 303 [#8860] (checklist; Russia: Udmurtia). JENDEK, 2006: 389 [#7441] (subgenus Agrilus; Palaearctic catalog). JENDEK, 2007: 111 [#7967] (taxonomic notes; synonymy). BELLAMY, 2008: 2352 2358 [#8576] (subgenus Agrilus; world catalog). VOLKOVITSH, 2009: 134 [#8850] (subgenus Agrilus; faunal records; Russia: Primorskiy kray). rosaceus (Scopoli, 1763) (Mordella) SCOPOLI, 1763: 61 62 [#2606] (description). OLIVIER, 1790: 83 [#2409] (Buprestis; synonym of viridis). SCHÖNHERR, 1817: 244 [#3895] (Buprestis; synonym of viridis). GORY & LAPORTE, 1839: 49 [#3738] (synonym of viridis). GEMMINGER & HAROLD, 1869: 1447 [#1057] (synonym of viridis). SAUNDERS, 1870: 27, 35 [#2551] (synonym of viridis). SAUNDERS, 1871: 119 [#2552] (synonym of viridis). KERREMANS, 1892: 277 [#1397] (synonym of viridis; catalog). KERREMANS, 1903: 275 [#1441] (synonym of viridis). JAKOBSON, 1913: 797 [#4386] (synonym of viridis; catalog). OBENBERGER, 1935: 63 [#2178] (synonym of viridis). OBENBERGER, 1936: 1061 [#2191] (synonym of viridis). COBOS, 1986: 317 [#609] (synonym of viridis). CURLETTI, 1994: 173 [#658] (synonym of viridis). BÍLÝ, 2002: 28 [#5877] (synonym of viridis). BRECHTEL & KOSTENBADER, 2002: 312 [#5766] (synonym of viridis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2354 [#8576] (synonym of viridis; world catalog). serraticornis (Scopoli, 1763) (Mordella) SCOPOLI, 1763: 61 [#2606] (description). FABRICIUS, 1781: 281 [#832] (Buprestis; synonym of viridis). GMELIN, 1790: 1937 [#1087] (Buprestis; synonym of viridis). SCHÖNHERR, 1817: 246 [#3895] (Buprestis). SAUNDERS, 1870: 28, 35 [#2551] (synonym of viridis). SAUNDERS, 1871: 119 [#2552] (synonym of viridis). KERREMANS, 1892: 277 [#1397] (synonym of viridis; catalog). KERREMANS, 1903: 275 [#1441] (synonym of viridis). JAKOBSON, 1913: 797 [#4386] (synonym of viridis; catalog). BEDEL, 1921: 212 [#170] (synonym of viridis). OBENBERGER, 1935: 63 [#2178] (synonym of viridis). OBENBERGER, 1936: 1061 [#2191] (synonym of viridis). ZAGAIKEVICH, 1962: 105 [#3249] (synonym of viridis). CURLETTI, 1994: 173 [#658] (synonym of viridis). BÍLÝ, 2002: 28 [#5877] (synonym of viridis). BRECHTEL & KOSTENBADER, 2002: 312 [#5766] (synonym of viridis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2353 2354 [#8576] (synonym of viridis; world catalog). linearis (Schrank, 1781) (Buprestis; [preoccupied]) SCHRANK, 1781: 197 198 [#2601] (description). SCHRANK, 1782: 168 [#2602] (Buprestis; synonym of lineata). SCHÖNHERR, 1817: 244 [#3895] (Buprestis; synonym of viridis). GORY & LAPORTE, 1839: 49 [#3738] (synonym of viridis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). lineatus (Schrank, 1782) (Buprestis; replacement name for linearis Schrank von Paula not Linné; [preoccupied]) SCHRANK, 1782: 168 [#2602] (replacement name proposal). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). linearis (Fabricius, 1792) (Buprestis; [preoccupied])

220 FABRICIUS, 1792: 213 [#834] (description). PANZER, 1795: 233 [#4041] (Buprestis; characters; Austria). FABRICIUS, 1796: 26 [#6846] (Buprestis; register). STURM, 1796: 48 [#4850] (Buprestis). FABRICIUS, 1801: 213 [#837] (Buprestis). HERBST, 1801: 313 314 [#1216] (Buprestis; synonym of filiformis). FABRICIUS, 1803: 10 [#8134] (Buprestis; index). SCHÖNHERR, 1817: 258 [#3895] (Buprestis; bibliography). DEJEAN, 1821: 30 [#4046] (Buprestis; catalog). DEJEAN, 1833: 83 [#689] (catalog). DEJEAN, 1836: 94 [#692] (catalog). MANNERHEIM, 1837: 115 [#3018] (note). GORY & LAPORTE, 1839: 55 56 [#3738] (note). REDTENBACHER, 1847: 287 [#3912] (characters in key). KIESENWETTER, 1857: 147 [#3015] (synonym of viridis). BELKE, 1866: 502 [#8622] (checklist; Ukraine). SAUNDERS, 1870: 19, 35 [#2551] (synonym of viridis). SAUNDERS, 1871: 119 [#2552] (synonym of viridis). REITTER, 1911: 194 [#2518] (aberration of viridis; characters in key). CAILLOL, 1913: 510 [#4053] (variety of viridis; faunal records; France). JAKOBSON, 1913: 797 [#4386] (variety of viridis; catalog; Russia and Europe). OBENBERGER, 1916: 271, 272 [#1983] (aberration of viridis; characters in key). KROGERUS, 1922: 110, 113 [#3916] (aberration of viridis). ESCHERICH, 1923: 149 [#6717] (variety of viridis). OBENBERGER, 1926: 656 [#2057] (aberration of viridis; Palaearctic catalog). POMERANCEV, 1938: 17 [#4977] (variety of viridis; notes). RIKHTER, 1945: 165 [#3507] (aberration of viridis; Ratzeburg is cited as the author). ZIMSEN, 1964: 156 [#3548] (two type specimens in Copenhagen and two in Kiel). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). bicolor (Schrank, 1798) (Buprestis) SCHRANK, 1798: 603 [#5144] (description). FABRICIUS, 1803: 10 [#8134] (Buprestis; index). SAUNDERS, 1870: 5, 35 [#2551] (synonym of viridis). SAUNDERS, 1871: 119 [#2552] (synonym of viridis). KERREMANS, 1892: 277 [#1397] (synonym of viridis; catalog). KERREMANS, 1903: 275 [#1441] (synonym of viridis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2353 [#8576] (synonym of viridis; world catalog). filiformis (Herbst, 1801) (Buprestis; replacement name for linearis Fabricius not Linné) HERBST, 1801: 313 314 [#1216] (replacement name proposal). SCHÖNHERR, 1817: 245 [#3895] (Buprestis; synonym of olivacea Gyllenhal). RATZEBURG, 1837: 56 [#2498] (Buprestis). FALDERMANN, 1838: 75 [#928] (synonym of angustulus). KIESENWETTER, 1849: 40 [#8334] (catalog; Europe). GEMMINGER & HAROLD, 1869: 1447 [#1057] (synonym of viridis; catalog). SAUNDERS, 1870: 13, 35 [#2551] (synonym of viridis). SAUNDERS, 1871: 119 [#2552] (synonym of viridis). KERREMANS, 1892: 277 278 [#1397] (synonym of viridis; catalog). KERREMANS, 1903: 275 [#1441] (synonym of viridis). JAKOBSON, 1913: 797 [#4386] (synonym of viridis; catalog). OBENBERGER, 1935: 63 [#2178] (synonym of viridis). OBENBERGER, 1936: 1061 [#2191] (synonym of viridis). ZAGAIKEVICH, 1962: 105 [#3249] (synonym of viridis). COBOS, 1986: 317 [#609] (synonym of viridis). CURLETTI, 1994: 173 [#658] (synonym of viridis). BÍLÝ, 2002: 28 [#5877] (synonym of viridis). BRECHTEL & KOSTENBADER, 2002: 312 [#5766] (synonym of viridis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2354 [#8576] (synonym of viridis; world catalog). fagi (Ratzeburg, 1837) (Buprestis) RATZEBURG, 1837: 56 57, 59 [#2498] (description). KIESENWETTER, 1849: 40 [#8334] (catalog; Europe). LETZNER, 1855: 23 [#7215] (references to larval description). KIESENWETTER, 1857: 152 [#3015] (synonym of viridis). MARSEUL, 1857: 92 [#4371] (checklist; Europe). MOTSCHULSKY, 1859: 157, 275 [#1832] (variety of viridis; catalog; bibliography on larval biology). JACQUELIN DU VAL, 1860: 143 [#8086] (synonym of viridis).

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA MARSEUL, 1866: 448 [#676] (variety of viridis). GEMMINGER & HAROLD, 1869: 1447 [#1057] (variety of viridis; catalog). SAUNDERS, 1871: 119 [#2552] (synonym of viridis; catalog). RUPERTSBERGER, 1880: 275 [#8550] (variety of viridis; checklist). KEPPEN, 1882: 230 [#8636] (form of viridis; pest). MARSEUL, 1887: 251 [#4707] (variety of viridis). JUDEICH & NITSCHE, 1889: 319 [#5686] (variety of viridis). REITTER, 1891: 200 [#5278] (variety of viridis). KERREMANS, 1892: 278 [#1397] (synonym of viridis; catalog). RUPERTSBERGER, 1894: 161, 287 [#4465] (variety of viridis; larval bibliography; catalog). HEYDEN, 1897: 194 [#5286] (variety of viridis; characters). KERREMANS, 1903: 275 [#1441] (synonym of viridis). REITTER, 1911: 194 [#2518] (aberration of viridis; characters in key). CAILLOL, 1913: 510 [#4053] (variety of viridis; faunal records; France). JAKOBSON, 1913: 797 [#4386] (variety of viridis; catalog; Russia and Europe). BAU, 1914: 257 [#8668] (variety of viridis). KIZERITSKII, 1915: 175 [#8698] (variety of fagi; faunal record; Ukraine). WEISS, 1915: 134 [#5953] (variety of viridis; [Note: Misidentification of cuprescens according to host plant cited]). OBENBERGER, 1916: 271, 272 [#1983] (aberration of viridis; characters in key). WEISS, 1916: 213 [#5954] (variety of viridis; [Note: Misidentification of cuprescens according to host plant]; pest; New Jersey). BEDEL, 1921: 213 [#170] (variety of viridis). KROGERUS, 1922: 110, 113 [#3916] (aberration of viridis). ESCHERICH, 1923: 149 [#6717] (variety of viridis). FROST, 1924: 27 [#1030] (subspecies of viridis; [Note: Misidentification of cuprescens according to host plant cited]; collecting record; Massachusetts). BRITTON, 1926: 544 545 [#6727] (new pest; USA: Connecticut). CHAMBERLIN, 1926: 85 [#468] (variety of viridis; [Note: Misidentification of cuprescens according to host plant cited]). OBENBERGER, 1926: 656 [#2057] (aberration of viridis; Palaearctic catalog). LENG & MUTCHLER, 1927: 30 [#1703] (variety of viridis; [Note: Misidentification of cuprescens]; supplement to catalog; USA: Massachusetts). VASILEV, 1927: 147 149 [#5834] (variety of viridis; [Note: Misidentification of cuprescens according to host plant cited]; pest; biology; parasitoids, predators; notes; Ukraine). FISHER, 1928: 13, 209 213 [#980] (variety of viridis; [Note: Misidentification of cuprescens according to host plants cited]; characters; bibliography; notes; US: Connecticut, Massachusetts, Michigan, New Jersey, New York, Pennsylvania). OGLOBLIN & REICHARDT, 1932: 278 [#8654] (variety of viridis; pest). OBENBERGER, 1935: 64 [#2178] (synonym of viridis aberration aubei). OBENBERGER, 1936: 1062 1063 [#2191] (synonym of viridis aberration aubei ; world catalog). THÉRY, 1942: 164, 165 [#2873] (subgenus Agrilus; subspecies of viridis). SCHAEFER, 1949: 397, 399, 400, 491 [#2565] (variety of viridis; characters). MÜHLMANN, 1954: 82 83 [#8152] (variety of viridis; notes). HORION, 1955: 94 [#1285] (variety of viridis; notes). TER MINASYAN, 1955: 454 [#7893] ([Note: Misidentification of ribesi according to host plant cited]; host plant; distribution; references). BALACHOWSKI et al., 1962: 292 [#96] (synonym of viridis). ALEXEEV, 1969: 89 [#51] (aberration of viridis; characters in key). GURYEVA, 1974: 101 [#5428] (variety of viridis; pest of agriculture cultures; references; Soviet Union). LOMPE, 1979: 240 [#3628] (variety of viridis; characters in key; Central Europe). COBOS, 1986: 254 [#609] (subgenus Agrilus; form of viridis). ZIEGLER, 1992: 26 [#6505] (cited as valid species; faunal records; Germany). CURLETTI, 1994: 177 [#658] (synonym of aurichalceus). BÍLÝ, 2002: 28 [#5877] (synonym of viridis). BRECHTEL & KOSTENBADER, 2002: 312, 316 325, 328 330 [#5766] (cited as a subspecies of viridis on pages 316 325, 328 330 and as a synonym of viridis on page 312; biology; ecology; distribution; parasitoids; predators; forestry; Germany). BELLAMY & WESTCOTT, 2003: 16 [#6153] (subspecies of viridis; not in Nearctic region). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2355 [#8576]

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA (synonym of viridis; world catalog). nocivus (Ratzeburg, 1837) (Buprestis) REDTENBACHER, RATZEBURG, 1837: 56 [#2498] (description). 1847: 287 [#3912] (characters in key). KIESENWETTER, 1849: 40 [#8334] (catalog; Europe). JACQUELIN DU VAL, 1857: xii [#8203] (synonym of viridis). KIESENWETTER, 1857: 151 [#3015] (synonym of viridis). MARSEUL, 1857: 91 [#4371] (synonym of distinguendus; checklist). MOTSCHULSKY, 1859: 157, 275 [#1832] (variety of viridis; catalog; bibliography on larval biology). JACQUELIN DU VAL, 1860: 143 [#8086] (as a synonym of pseudocyaneus with Redtenbacher’s authorship and as a synonym of viridis with Ratzeburg’s authorship). MARSEUL, 1866: 448 [#676] (variety of viridis). GEMMINGER & HAROLD, 1869: 1447 [#1057] (variety of viridis; catalog). SAUNDERS, 1871: 119 [#2552] (synonym of viridis; catalog). HOCHHUTH, 1873: 158 [#3453] (variety of viridis; faunal records; Gouvernements Kiew und Volhynien). RUPERTSBERGER, 1880: 275 [#8550] (variety of viridis; checklist). KEPPEN, 1882: 230, 231 [#8636] (form of viridis; pest). MARSEUL, 1887: 251 [#4707] (variety of viridis). JUDEICH & NITSCHE, 1889: 319 [#5686] (variety of viridis). REITTER, 1891: 200 [#5278] (variety of viridis). KERREMANS, 1892: 278 [#1397] (synonym of viridis; catalog). HEYDEN, 1893: 90 [#3450] (variety of viridis; catalog). RUPERTSBERGER, 1894: 161, 287 [#4465] (variety of viridis; larval bibliography; catalog). HEYDEN, 1897: 194 [#5286] (variety of viridis; characters). KERREMANS, 1903: 275 [#1441] (synonym of viridis). REITTER, 1911: 194 [#2518] (aberration of viridis; characters in key). CAILLOL, 1913: 510 [#4053] (variety of viridis; faunal records; France). JAKOBSON, 1913: 797 [#4386] (variety of viridis; catalog; Russia and Europe). BAU, 1914: 257 [#8668] (variety of viridis). OBENBERGER, 1916: 271, 272 [#1983] (aberration of viridis; characters in key). BEDEL, 1921: 213 [#170] (variety of viridis). LABLER, 1921: 188 [#5928] (aberration of viridis; checklist; Germany). KROGERUS, 1922: 110, 113 [#3916] (aberration of viridis). ESCHERICH, 1923: 149 [#6717] (variety of viridis). OBENBERGER, 1926: 656 [#2057] (aberration of viridis; Palaearctic catalog). OBENBERGER, 1927: 211, 216, 229 [#2064] (variety of viridis). OBENBERGER, 1935: 64 [#2178] (variety of viridis; catalog). OBENBERGER, 1936: 1065 1067 [#2191] (aberration of viridis; world catalog). THÉRY, 1942: 164, 165 [#2873] (subgenus Agrilus; cited as subspecies of viridis on page 164 and as variety of viridis on page 165). RIKHTER, 1945: 165 [#3507] (aberration of viridis). SCHAEFER, 1949: 397, 399, 400, 491 [#2565] (variety of viridis; cited as nociva; characters). HORION, 1955: 94 [#1285] (variety of viridis; cited as nociva; notes). LOMPE, 1979: 240 [#3628] (variety of viridis; characters in key; Central Europe). COBOS, 1986: 254, 317 [#609] (synonym of viridis). CURLETTI, 1994: 173 [#658] (synonym of viridis). BÍLÝ, 2002: 28 [#5877] (synonym of viridis). BRECHTEL & KOSTENBADER, 2002: 312, 330 332 [#5766] (cited as a subspecies of viridis on pages 330 332 and as a synonym of viridis on page 312; biology; ecology; distribution; Germany). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2355 2356 [#8576] (synonym of viridis; world catalog). capreae (Chevrolat, 1838) (Buprestis; subgenus Agrilus) CHEVROLAT, 1838: 56 [#490] (description). KIESENWETTER, 1857: 151 [#3015] (synonym of viridis). MARSEUL, 1857: 92 [#4371] (checklist; Europe). JACQUELIN DU VAL, 1860: 143 [#8086] (synonym of viridis). GEMMINGER & HAROLD, 1869: 1447 [#1057] (synonym of viridis). SAUNDERS, 1871: 119 [#2552] (synonym of viridis). MARSEUL, 1887: 251 [#4707] (synonym of viridis). JUDEICH & NITSCHE, 1889: 319 [#5686] (variety of viridis). REITTER, 1891: 200 [#5278] (synonym of viridis). KERREMANS, 1892: 278 [#1397] (synonym of viridis; catalog). KERREMANS, 1903: 275 [#1441] (synonym of viridis). CAILLOL, 1913: 509 [#4053] (synonym of viridis). JAKOBSON, 1913: 797 [#4386]

221 (synonym of viridis; catalog). OBENBERGER, 1935: 63 [#2178] (synonym of viridis). OBENBERGER, 1936: 1061 [#2191] (synonym of viridis). BALACHOWSKI et al., 1962: 292 [#96] (synonym of viridis). ZAGAIKEVICH, 1962: 105 [#3249] (synonym of viridis). COBOS, 1986: 317 [#609] (synonym of viridis). CURLETTI, 1994: 173 [#658] (synonym of viridis). BÍLÝ, 2002: 28 [#5877] (cited as caprae; synonym of viridis). BRECHTEL & KOSTENBADER, 2002: 312 [#5766] (synonym of viridis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2355 [#8576] (synonym of viridis; world catalog). aubei Gory & Laporte, 1839 (Agrilus) GORY & LAPORTE, 1839: 44 45 [#3738] (description). AUBÉ, 1837: 189 192 [#4034] (cited as viridis; [Note: Species cited in this publication under the viridis was described by GORY & LAPORTE, 1839 [#3738] as aubei]; larva; France: Boulogne). KIESENWETTER, 1849: 40 [#8334] (catalog; Europe). JACQUELIN DU VAL, 1857: xii [#8203] (synonym of viridis). KIESENWETTER, 1857: 152 [#3015] (synonym of viridis). MARSEUL, 1857: 91 [#4371] (checklist; Europe). MOTSCHULSKY, 1859: 157 [#1832] (variety of viridis; catalog; bibliography on larval biology). JACQUELIN DU VAL, 1860: 143 [#8086] (synonym of viridis). MARSEUL, 1866: 448, 449 [#676] (synonym of viridis variety fagi). GEMMINGER & HAROLD, 1869: 1447 [#1057] (variety of viridis; catalog). SAUNDERS, 1871: 119 [#2552] (synonym of viridis; catalog). MARSEUL, 1887: 251 [#4707] (synonym of viridis variety fagi). JUDEICH & NITSCHE, 1889: 319 [#5686] (variety of viridis). REITTER, 1891: 200 [#5278] (synonym of viridis variety fagi). KERREMANS, 1892: 278 [#1397] (synonym of viridis; catalog). KERREMANS, 1903: 275 [#1441] (synonym of viridis). CAILLOL, 1913: 510 [#4053] (synonym of viridis variety fagi). JAKOBSON, 1913: 797 [#4386] (synonym of viridis; catalog). BEDEL, 1921: 213 [#170] (synonym of viridis variety fagi). OBENBERGER, 1927: 197, 210, 215, 227 228, 248 [#2064] (aberration of viridis). OBENBERGER, 1935: 64 [#2178] (aberration of viridis; catalog). OBENBERGER, 1936: 1062 [#2191] (aberration of viridis; world catalog). POMERANCEV, 1938: 17 [#4977] (variety of viridis; notes). SCHAEFER, 1949: 397, 491 [#2565] (synonym of viridis variety fagi). ALEXEEV, 1969: 89 [#51] (aberration of viridis; characters in key). VOLKOVITSH, 1977: 25 [#2985] (subspecies of viridis; [Note: Misidentification of ribesi according to host plant cited]; faunal record; notes; USSR: Belgorod). COBOS, 1986: 317 [#609] (synonym of viridis). NELSON & BELLAMY, 1993: 304 [#3167] (authorship and publication date). CURLETTI, 1994: 173 [#658] (synonym of viridis). JENDEK, 1998: 318 [#4481] (lectotype designation). BÍLÝ, 2002: 28 [#5877] (synonym of viridis). BRECHTEL & KOSTENBADER, 2002: 312 [#5766] (synonym of viridis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2354 2355 [#8576] (synonym of viridis; world catalog). distinguendus Gory & Laporte, 1839 (Agrilus) GORY & LAPORTE, 1839: 44 [#3738] (description). KIESENWETTER, 1849: 40 [#8334] (catalog; Europe). KIESENWETTER, 1857: 151 [#3015] (synonym of viridis). MARSEUL, 1857: 91 [#4371] (checklist; Europe). JACQUELIN DU VAL, 1860: 143 [#8086] (synonym of viridis). MARSEUL, 1866: 448 [#676] (synonym of viridis variety nocivus). GEMMINGER & HAROLD, 1869: 1447 [#1057] (variety of viridis; catalog). SAUNDERS, 1871: 119 [#2552] (synonym of viridis; catalog). MARSEUL, 1887: 251 [#4707] (synonym of viridis variety nocivus). JUDEICH & NITSCHE, 1889: 319 [#5686] (variety of viridis). REITTER, 1891: 200 [#5278] (synonym of viridis variety nocivus). KERREMANS, 1892: 278 [#1397] (synonym of viridis; catalog). KERREMANS, 1903: 275 [#1441] (synonym of viridis). CAILLOL, 1913: 510 [#4053] (synonym of viridis variety nocivus). JAKOBSON, 1913: 797 [#4386] (synonym of viridis; catalog). BEDEL, 1921: 213

222 [#170] (synonym of viridis variety nocivus). OBENBERGER, 1935: 64 [#2178] (synonym of viridis aberration nocivus). OBENBERGER, 1936: 1067 [#2191] (synonym of viridis variety nocivus; world catalog). SCHAEFER, 1949: 397, 491 [#2565] (synonym of viridis variety nocivus). COBOS, 1986: 317 [#609] (synonym of viridis). NELSON & BELLAMY, 1993: 304 [#3167] (authorship and publication date). CURLETTI, 1994: 174 [#658] (synonym of viridis). JENDEK, 1998: 321 [#4481] (synonym of viridis; lectotype designation). BÍLÝ, 2002: 28 [#5877] (synonym of viridis). BRECHTEL & KOSTENBADER, 2002: 312 [#5766] (synonym of viridis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2355 [#8576] (synonym of viridis; world catalog). viridipennis Gory & Laporte, 1839 (Agrilus) GORY & LAPORTE, 1839: 45 46 [#3738] (description). KIESENWETTER, 1849: 40 [#8334] (catalog; Europe). PERRIS, 1851: 111 113 [#3721] (larval and adult characters). LETZNER, 1855: 23 [#7215] (references to larval description). JACQUELIN DU VAL, 1857: ccii [#8203] (larva; references). KIESENWETTER, 1857: 151 [#3015] (synonym of viridis). MARSEUL, 1857: 92 [#4371] (synonym of capreae; checklist). JACQUELIN DU VAL, 1860: 143 [#8086] (synonym of viridis). GEMMINGER & HAROLD, 1869: 1447 [#1057] (synonym of viridis). SAUNDERS, 1871: 119 [#2552] (synonym of viridis). MARSEUL, 1887: 251 [#4707] (synonym of viridis). JUDEICH & NITSCHE, 1889: 319 [#5686] (variety of viridis). REITTER, 1891: 199 [#5278] (synonym of viridis). KERREMANS, 1892: 278 [#1397] (cited as irridipennis; synonym of viridis; catalog). KERREMANS, 1903: 275 [#1441] (cited as iridipennis; synonym of viridis). CAILLOL, 1913: 509 [#4053] (synonym of viridis). JAKOBSON, 1913: 797 [#4386] (synonym of viridis; catalog). BEDEL, 1921: 212 [#170] (synonym of viridis). OBENBERGER, 1935: 63 [#2178] (synonym of viridis). OBENBERGER, 1936: 1061 [#2191] (synonym of viridis). ALEXEEV, 1959: 13 [#3229] (synonym of viridis). BALACHOWSKI et al., 1962: 292 [#96] (synonym of viridis). ZAGAIKEVICH, 1962: 105 [#3249] (synonym of viridis). NELSON & BELLAMY, 1993: 304 [#3167] (authorship and publication date). CURLETTI, 1994: 173 [#658] (synonym of viridis). JENDEK, 1998: 332 [#4481] (lectotype designation). BÍLÝ, 2002: 28 [#5877] (synonym of viridis). BRECHTEL & KOSTENBADER, 2002: 312 [#5766] (synonym of viridis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2355 [#8576] (synonym of viridis; world catalog). littlei Curtis, 1840 (Agrilus; cited as littlii; [Note: Dedicated to W. Little. Incorrect original spelling (ICZN, Article 32.5.)]) CURTIS, 1840: 365 366 [#660] (description). GRAY, 1848: 37 [#5388] (cited as littlii with authorsip attributed to Shuckard; checklist of taxa in the collection of the British museum). KIESENWETTER, 1849: 40 [#8334] (cited as littlii; catalog; Europe). KIESENWETTER, 1857: 151 [#3015] (synonym of viridis; Shuckard is cited as the author). MARSEUL, 1857: 92 [#4371] (cited as littlii; checklist; Europe). JACQUELIN DU VAL, 1860: 143 [#8086] (Curtis is cited with the question mark; synonym of viridis). MARSEUL, 1866: 448 [#676] (synonym of viridis variety linearis). GEMMINGER & HAROLD, 1869: 1447 [#1057] (variety of viridis; catalog). SAUNDERS, 1871: 119 [#2552] (synonym of viridis; catalog). MARSEUL, 1887: 251 [#4707] (Laporte de Castelnau is cited as the author; synonym of viridis variety linearis). KERREMANS, 1892: 278 [#1397] (synonym of viridis; catalog). KERREMANS, 1903: 275 [#1441] (synonym of viridis). JAKOBSON, 1913: 797 [#4386] (synonym of viridis; catalog). OBENBERGER, 1935: 64 [#2178] (synonym of viridis aberration linearis). OBENBERGER, 1936: 1065 [#2191] (synonym of viridis variety linearis; world catalog). COBOS, 1986: 317 [#609] (synonym of viridis). CURLETTI, 1994: 174 [#658] (synonym of viridis). BÍLÝ, 2002: 28 [#5877] (synonym of viridis). BRECHTEL & KOSTENBADER, 2002: 312 [#5766] (synonym of viridis). JENDEK,

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2354 [#8576] (synonym of viridis; world catalog). bicolor Redtenbacher, 1847 (Agrilus) REDTENBACHER, 1847: 287 [#3912] (description). KIESENWETTER, 1849: 40 [#8334] (catalog; Europe). KIESENWETTER, 1857: 151 [#3015] (synonym of viridis). MARSEUL, 1857: 92 [#4371] (checklist; Europe). JACQUELIN DU VAL, 1860: 143 [#8086] (synonym of viridis). MARSEUL, 1866: 448 [#676] (synonym of viridis variety nocivus). GEMMINGER & HAROLD, 1869: 1447 [#1057] (variety of viridis; catalog). SAUNDERS, 1871: 119 [#2552] (synonym of viridis; catalog). MARSEUL, 1887: 251 [#4707] (synonym of viridis variety nocivus). JUDEICH & NITSCHE, 1889: 319 [#5686] (variety of viridis). REITTER, 1891: 200 [#5278] (synonym of viridis variety nocivus). KERREMANS, 1892: 278 [#1397] (synonym of viridis; catalog). KERREMANS, 1903: 275 [#1441] (synonym of viridis). CAILLOL, 1913: 510 [#4053] (synonym of viridis variety nocivus). JAKOBSON, 1913: 797 [#4386] (synonym of viridis; catalog). OBENBERGER, 1935: 64 [#2178] (synonym of viridis aberration nocivus). OBENBERGER, 1936: 1067 [#2191] (synonym of viridis variety nocivus; world catalog). SCHAEFER, 1949: 397, 491 [#2565] (synonym of viridis variety nocivus). COBOS, 1986: 317 [#609] (synonym of viridis). CURLETTI, 1994: 174 [#658] (synonym of viridis). BÍLÝ, 2002: 28 [#5877] (synonym of viridis). BRECHTEL & KOSTENBADER, 2002: 312 [#5766] (synonym of viridis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2356 [#8576] (misidentification of viridis; world catalog). quercinus Redtenbacher, 1847 (Agrilus) REDTENBACHER, 1847: 286 [#3912] (description). KIESENWETTER, 1849: 40 [#8334] (catalog; Europe). KIESENWETTER, 1857: 152 [#3015] (synonym of viridis). MARSEUL, 1857: 91 [#4371] (checklist; Europe). JACQUELIN DU VAL, 1860: 143 [#8086] (synonym of viridis). MARSEUL, 1866: 448 [#676] (synonym of viridis variety fagi). GEMMINGER & HAROLD, 1869: 1447 [#1057] (variety of viridis; catalog). SAUNDERS, 1871: 119 [#2552] (synonym of viridis; catalog). HOCHHUTH, 1873: 158 [#3453] (variety of viridis; faunal records; Gouvernements Kiew und Volhynien). MARSEUL, 1887: 251 [#4707] (synonym of viridis variety fagi). JUDEICH & NITSCHE, 1889: 319 [#5686] (variety of viridis). REITTER, 1891: 200 [#5278] (synonym of viridis variety fagi). KERREMANS, 1892: 278 [#1397] (synonym of viridis; catalog). KERREMANS, 1903: 275 [#1441] (synonym of viridis). CAILLOL, 1913: 510 [#4053] (synonym of viridis variety fagi). JAKOBSON, 1913: 797 [#4386] (synonym of viridis; catalog). OBENBERGER, 1935: 64 [#2178] (synonym of viridis aberration aubei). OBENBERGER, 1936: 1064 [#2191] (synonym of viridis variety aubei; world catalog). SCHAEFER, 1949: 397, 491 [#2565] (synonym of viridis variety fagi). COBOS, 1986: 317 [#609] (synonym of viridis). CURLETTI, 1994: 174 [#658] (synonym of viridis). BÍLÝ, 2002: 28 [#5877] (synonym of viridis). BRECHTEL & KOSTENBADER, 2002: 312 [#5766] (synonym of viridis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2356 [#8576] (synonym of viridis; world catalog). darwinii Wollaston, 1857 (Agrilus) WOLLASTON, 1857: 82, 214 [#3124] (description). WOLLASTON, 1865: 188 189 [#7784] (notes; Madeira). MARSEUL, 1866: 494 [#676] (cited as darwini; redescription). GEMMINGER & HAROLD, 1869: 1439 [#1057] (cited as darwini). SAUNDERS, 1871: 121 [#2552] (catalog). MARSEUL, 1887: 252 [#4707] (catalog). KERREMANS, 1892: 254 [#1397] (cited as darwini; catalog). FAUVEL, 1897: 58 [#8090] (cited as darwini; catalog; Madeira). KERREMANS, 1903: 284 [#1441] (catalog). JAKOBSON, 1913: 798 [#4386] (cited as darwini; catalog; Russia and Europe). BEDEL, 1921: 212 [#170] (cited as darwini synonym of viridis).

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA OBENBERGER, 1926: 659 [#2057] (cited as darwini; Palaearctic catalog). OBENBERGER, 1935: 63 [#2178] (synonym of viridis). OBENBERGER, 1936: 986 [#2191] (cited as darwini; world catalog). SCHAEFER, 1949: 398 [#2565] (cited as darwini; presumed synonym of viridis; notes). COBOS, 1986: 317 [#609] (synonym of viridis). CURLETTI, 1994: 174 [#658] (cited as darwini; synonym of viridis). BÍLÝ, 2002: 28 [#5877] (cited as darwini; synonym of viridis). BRECHTEL & KOSTENBADER, 2002: 312 [#5766] (cited as darwini; synonym of viridis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2356 [#8576] (synonym of viridis; world catalog). imbellis Kerremans, 1892 (Agrilus; [preoccupied]) KERREMANS, 1892: 215 [#1402] (description). Kerremans, 1903: 286 [#1441] (synonym of importunus; catalog). OBENBERGER, 1936: 1086 [#2191] (synonym of importunus). BELLAMY, 2008: 2136 [#8576] (synonym of importunus; world catalog). brevitarsis Lewis, 1893 (Agrilus) syn. nov. LEWIS, 1893: 336 [#1722] (description). KERREMANS, 1903: 287 [#1441] (synonym of brevitarsis; catalog). JAKOBSON, 1913: 798 [#4386] (synonym of lewisi; catalog). OBENBERGER, 1926: 659 [#2057] (synonym of lewisi). MIWA & CHÛJÔ, 1936: 17 [#1809] (cited as brevtarsis; synonym of lewisi; catalog; Japan). OBENBERGER, 1936: 1008 [#2191] (synonym of lewisi; world catalog). JENDEK, 1995: 147 [#3768] (valid not preoccupied name; lectotype designation; synonymy; characters; Japan). AKIYAMA & OHMOMO, 1997: 5, 30 31 [#4091] (checklist; Japan: central Honshu). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2008 [#8576] (world catalog). indulgens Kerremans, 1897 (Agrilus) KERREMANS, 1897: 76 [#1424] (description). OBENBERGER, 1960: 146 [#2394] (synonym of viridis aberration linearis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). rudis Abeille de Perrin, 1897 (Agrilus) ABEILLE DE PERRIN, 1897: 4, 13 14 [#19] (description). KERREMANS, 1903: 290 [#1441] (catalog). JAKOBSON, 1913: 797 [#4386] (catalog; Russia and Europe). OBENBERGER, 1916: 272 [#1983] (subspecies of viridis; characters in key). OBENBERGER, 1926: 656 [#2057] (subspecies of chrysoderes; Palaearctic catalog). OBENBERGER, 1927: 212, 221, 242 [#2064] (uncertain subspecies of communis). OBENBERGER, 1936: 1038 [#2191] (variety of rubicola; world catalog). THÉRY, 1942: 164 [#2873] (subgenus Agrilus; synonym of viridis). SCHAEFER, 1949: 403 [#2565] (subspecies of aurichalceus; notes). COBOS, 1986: 255, 318 [#609] (subgenus Agrilus; subspecies of aurichalceus). JENDEK, 2003: 9, 16 [#5878] (synonym of viridis; lectotype designation; notes). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2356 [#8576] (synonym of viridis; world catalog). importunus Kerremans, 1903 (Agrilus; replacement name for imbellis Kerremans not Crotch) syn. nov. KERREMANS, 1903: 286 [#1441] (replacement name proposal). OBENBERGER, 1936: 1086 [#2191] (world catalog). BELLAMY, 2008: 2135 [#8576] ( world catalog). lewisi Kerremans, 1903 (Agrilus; unnecessary replacement name for brevitarsis Lewis not Erichson, which is an unavailable name) KERREMANS, 1903: 287 [#1441] (replacement name proposal). JAKOBSON, 1913: 798 [#4386] (catalog; Russia and Europe). OBENBERGER, 1926: 659 [#2057] (Palaearctic catalog). MIWA & CHÛJÔ, 1936: 17 [#1809] (catalog; Japan). OBENBERGER, 1936: 1008 [#2191] (world catalog). KUROSAWA, 1974: 2 [#4577] (note). JENDEK, 1995: 147 [#3768] (synonym of brevitarsis). AKIYAMA & OHMOMO, 1997: 30 [#4091] (synonym of brevitarsis). JENDEK, 2006: 397 [#7441] (synonym of brevitarsis brevitarsis; Palaearctic catalog). BELLAMY, 2008: 2008 [#8576] (synonym of

223 brevitarsis; world catalog). montanellus Obenberger, 1916 (Agrilus; cited as a variety of viridis on page 270 and as a subspecies of viridis on p. 272) OBENBERGER, 1916: 270, 272 [#1983] (description; [Note: This name was treated as a senior homonym and thus is an available name (Article 45.6.4.1)]). OBENBERGER, 1926: 656 [#2057] (variety of chrysoderes; Palaearctic catalog). OBENBERGER, 1927: 211, 216, 217, 231 232 [#2064] (subspecies of viridis). OBENBERGER, 1936: 1068 [#2191] (subspecies of viridis; world catalog). SCHAEFER, 1949: 398 [#2565] (subspecies of viridis; notes). HORION, 1955: 94 [#1285] (variety of viridis; note). LOMPE, 1979: 240 [#3628] (variety of viridis; characters in key; Central Europe). COBOS, 1986: 253, 317 [#609] (subgenus Agrilus; subspecies of viridis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). JENDEK, 2006: 40 [#7359] (available name). BELLAMY, 2008: 2357 [#8576] (synonym of viridis; world catalog). gebleri Obenberger, 1924 (Agrilus) syn. nov. OBENBERGER, OBENBERGER, 1924: 42 [#2040] (description). 1926: 656 [#2057] (Palaearctic catalog). OBENBERGER, 1927: 224 [#2064] (note). OBENBERGER, 1936: 991 [#2191] (world catalog). IVLIEV & KONONOV, 1966: 109 [#8089](as gebleri aberration kamtschaticus [undescribed]; biology; faunal records; Russia: Kamchatka, Magadan, Yakutiya). COBOS, 1968: 389 [#580] (subgenus Agrilus; characters; faunal records; Mongolia). COBOS, 1972: 56 [#591] (subgenus Agrilus; checklist; Mongolia). ALEXEEV, 1975: 158 [#52] (presumably conspecific with transbaicalenesis; notes; faunal records; distributional overview; Mongolia). ALEXEEV, 1989: 489 [#64] (key to species; Kamchatka; Amursk; Chita; Buryatia; northeastern China; Mongolia). ALEXEEV & VOLKOVITSH, 1989: 321, 350, 357 [#3241] (characters in key; faunal records; Mongolia). ALEXEEV, 1998: 381 [#4322] (subgenus Agrilus; subgeneric classification). NONNAIZAB et al., 1999: 112 [#8247] (checklist; China: Inner Mongolia). JENDEK, 2006: 389 [#7441] (subgenus Agrilus; Palaearctic catalog). BELLAMY, 2008: 2106 [#8576] (subgenus Agrilus; world catalog). caenus Obenberger, 1924 (Agrilus; subspecies of viridis) OBENBERGER, OBENBERGER, 1924: 40 [#2040] (description). 1926: 656 [#2057] (subspecies of chrysoderes; Palaearctic catalog). OBENBERGER, 1927: 211, 220, 224, 241 [#2064] (subspecies of communis). OBENBERGER, 1936: 1038 [#2191] (variety of rubicola; world catalog). THÉRY, 1942: 166 [#2873] (subgenus Agrilus; aberration of aurichalceus). SCHAEFER, 1949: 403 [#2565] (subspecies of aurichalceus; notes). COBOS, 1986: 256 [#609] (subgenus Agrilus; subspecies of aurichalceus). CURLETTI, 1987: 152 [#653] (subspecies of aurichalceus). CURLETTI, 1994: 178 [#658] (subspecies of aurichalceus; catalog; Sicilia). JENDEK, 2003: 12 13, 16 [#5878] (synonym of viridis; lectotype designation; notes). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2357 [#8576] (synonym of viridis; world catalog). poppiusi Obenberger, 1924 (Agrilus; ssp. of viridis) syn. nov. OBENBERGER, 1924: 40 41 [#2040] (description). OBENBERGER, 1926: 656 [#2057] (subspecies of viridis; Palaearctic catalog). OBENBERGER, 1927: 211, 217, 233 [#2064] (subspecies of viridis). OBENBERGER, 1936: 1068 [#2191] (subspecies of viridis; world catalog). SCHAEFER, 1949: 398 [#2565] (subspecies of viridis; notes). COBOS, 1986: 254, 317 [#609] (subgenus Agrilus; subspecies of viridis). JENDEK, 2006: 389 [#7441] (subgenus Agrilus; subspecies of viridis; Palaearctic catalog). BELLAMY, 2008: 2358 [#8576] (subspecies of viridis; world catalog). krogerusi Obenberger, 1924 (Agrilus; subspecies of viridis) OBENBERGER, OBENBERGER, 1924: 41 [#2040] (description). 1926: 656 [#2057] (subspecies of viridis; Palaearctic catalog).

224 OBENBERGER, 1927: 211, 216, 232 [#2064] (subspecies of viridis). BRUNDIN, 1934: 361 [#8542] (subspecies of viridis; [Note: Misidentification]; faunal records; north Sweden). OBENBERGER, 1936: 1068 [#2191] (subspecies of viridis; world catalog). SCHAEFER, 1949: 398 [#2565] (subspecies of viridis; notes). COBOS, 1968: 389 390 [#580] (subgenus Agrilus; subspecies of viridis; characters; faunal records; Mongolia). COBOS, 1972: 46, 56 [#591] (subgenus Agrilus; subspecies of viridis; faunal records; Mongolia). ALEXEEV, 1975: 157 [#52] (synonym of viridis). COBOS, 1986: 253, 317 [#609] (subgenus Agrilus; subspecies of viridis). PENG ZHONGLIANG, 1994: 19 [#2432] (subspecies of viridis; faunal records; China). HUA LI ZHONG, 2002: 91 [#6696] (subspecies of viridis; checklist; China: Hunan). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2357 [#8576] (synonym of viridis; world catalog). melantatus Obenberger, 1924 (Agrilus; subspecies of viridis) OBENBERGER, OBENBERGER, 1924: 40 [#2040] (description). 1926: 656 [#2057] (subspecies of viridis; Palaearctic catalog). OBENBERGER, 1927: 211, 216, 232 233 [#2064] (subspecies of viridis). OBENBERGER, 1936: 1068 [#2191] (subspecies of viridis; world catalog). SCHAEFER, 1949: 398 [#2565] (subspecies of viridis; notes). POCHON, 1967: 133 [#2473] (subspecies of viridis; faunal record; Mongolia). ALEXEEV, 1975: 157 [#52] (synonym of viridis). COBOS, 1986: 254, 317 [#609] (subgenus Agrilus; subspecies of viridis). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2357 [#8576] (synonym of viridis; world catalog). brydli Obenberger, 1924 (Agrilus) syn. nov. OBENBERGER, OBENBERGER, 1924: 41 [#2040] (description). 1926: 656 [#2057] (Palaearctic catalog). OBENBERGER, 1927: 215 [#2064]. OBENBERGER, 1936: 968 [#2191] (world catalog). JENDEK, 2006: 397 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2009 [#8576] (world catalog). cernyi Obenberger, 1925 (Agrilus; subspecies of viridis) OBENBERGER, 1925: 59 [#2049] (description). OBENBERGER, 1926: 656 [#2057] (subspecies of chrysoderes; Palaearctic catalog). OBENBERGER, 1927: 211, 217, 224, 231 [#2064] (subspecies of viridis). ROUBAL, 1930: 115 [#5172] (notes). OBENBERGER, 1935: 64 [#2178] (subspecies of viridis; catalog). OBENBERGER, 1936: 1068 [#2191] (subspecies of viridis; world catalog). SCHAEFER, 1949: 398 [#2565] (subspecies of viridis; notes). HORION, 1955: 94 [#1285] (variety of viridis; note). LOMPE, 1979: 240 [#3628] (variety of viridis; characters in key; Central Europe). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2357 [#8576] (synonym of viridis; world catalog). vernadskii Obenberger, 1927 (Agrilus; cited as vernavskii) OBENBERGER, 1927: 15 [#2060] (description). OBENBERGER, 1936: 1055 [#2191] (world catalog). ALEXEEV, 1989: 488 [#64] (key to species; Khabarovsk; Amursk; Primorsk; Korean peninsula; northeastern China). ALEXEEV & VOLKOVITSH, 1989: 321, 350, 357 [#3241] (synonym of suvorovi; characters in key; faunal records; type examination; notes on type locality; Mongolia). ALEXEEV, 1998: 381 [#4322] (subgenus Agrilus; subgeneric classification). JENDEK, 2002: 2, 3, 4 [#5392] (synonym of viridis; taxonomic and nomenclatural notes). JENDEK, 2006: 389 [#7441] (synonym of viridis; Palaearctic catalog). BELLAMY, 2008: 2357 [#8576] (synonym of viridis; world catalog). kiricenkoi Obenberger, 1935 (Agrilus) transfer in syn. OBENBERGER, OBENBERGER, 1935: 166 [#2158] (description). 1936: 1004 [#2191] (world catalog). COBOS, 1972: 56 [#591] (subgenus Agrilus; checklist; Mongolia). ALEXEEV, 1975: 158 [#52] (notes; faunal records; Mongolia). PENG ZHONGLIANG, 1987: 356 [#2429] (checklist; China). ALEXEEV, 1989: 488 [#64] (synonym of attila). ALEXEEV & VOLKOVITSH, 1989: 320 [#3241]

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA (synonym of attila; lectotype designation). OVTCHINNIKOV, 1996: 132 [#8621] (cited as kiritschenkoi; checklist; Kyrgyzstan). HUA LI ZHONG, 2002: 89 [#6696] (checklist; China: Inner Mongolia). JENDEK, 2006: 388 [#7441] (synonym of attila; Palaearctic catalog). BELLAMY, 2008: 1982 [#8576] (synonym of attila; world catalog). transbaicalensis Obenberger, 1935 (Agrilus) syn. nov. OBENBERGER, OBENBERGER, 1935: 166 [#2158] (description). 1936: 1054 [#2191] (world catalog). MAKHNOVSKIJ, 1958: 127 [#6090] (cited as bajkalensis; bionomy; pest; Uzbekistan). MAKHNOVSKIJ, 1959: 31 32 [#5829] (cited as bajkalensis; characters; biology; Uzbekistan). COBOS, 1972: 46, 56 [#591] (subgenus Agrilus; faunal records; Mongolia). ALEXEEV, 1975: 158 [#52] (note; faunal records; Mongolia). ALEXEEV, 1989: 489 [#64] (key to species; Amursk; Chita; Buryatia; northeastern China; north Mongolia). ALEXEEV & VOLKOVITSH, 1989: 321, 350, 357 [#3241] (characters in key; faunal records; Mongolia). ALEXEEV, 1998: 381 [#4322] (subgenus Agrilus; subgeneric classification). NONNAIZAB et al., 1999: 113 [#8247] (checklist; China: Inner Mongolia). JENDEK, 2006: 389 [#7441] (subgenus Agrilus; Palaearctic catalog). BELLAMY, 2008: 2332 [#8576] (subgenus Agrilus; world catalog). pusztae Obenberger, 1936 (Agrilus) OBENBERGER, 1936: 108 109 [#2205] (description). OBENBERGER, 1936: 1026 [#2191] (world catalog). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). JENDEK, 2007: 111 [#7967] (synonym of viridis; lectotype designation). BELLAMY, 2008: 2358 [#8576] (synonym of viridis; world catalog). towadensis Miwa & Chûjô, 1940 (Agrilus) syn. nov. MIWA & CHÛJÔ, 1940: 54, 61 62 [#1810] (description). KUROSAWA, 1974: 2 [#4577] (subspecies of lewisi). AKIYAMA & OHMOMO, 1997: 5, 31 [#4091] (subspecies of brevitarsis; checklist; Japan: north Honshu). JENDEK, 2006: 397 [#7441] (subspecies of brevitarsis; Palaearctic catalog). BELLAMY, 2008: 2008 [#8576] (subspecies of brevitarsis; world catalog). salicivola Kurosawa, 1963 (Agrilus; subspecies of suvorovi) syn. nov. KUROSAWA, 1963: 106 107 [#1617] (description). KUROSAWA, 1963: 153 [#1618] (subspecies of suvorovi; characters; Japan). KUROSAWA, 1974: 1, 2 [#4575] (subspecies of suvorovi; cited as salicivora; characters; notes). KUROSAWA, 1975: 1 [#4591] (comment on correct specific name). AKIYAMA & OHMOMO, 1997: 4, 5, 38 [#4091] (checklist; Japan: Hokkaido, Honshu; Korea; Russia: Sachalin, East Siberia). FUKUTOMI & HORI, 2004: 7 8 [#6851] (checklist; faunal records; Japan (Hokkaido)). JENDEK, 2006: 401 [#7441] (Palaearctic catalog). ANONYMOUS, 2007: 32 [#8250] (checklist; Japan: Honshu). BELLAMY, 2008: 2277 [#8576] (world catalog). cojbalsanensis Cobos, 1968 (Agrilus; subgenus Agrilus) transfer in syn. COBOS, 1968: 404 406 [#580] (description). COBOS, 1972: 56 [#591] (subgenus Agrilus; checklist; Mongolia). ALEXEEV, 1975: 159 [#52] (presumably conspecific with pseudolimoniastri; note; Mongolia). ALEXEEV & VOLKOVITSH, 1989: 321 [#3241] (synonym of transbaicalensis; type examination). NONNAIZAB et al., 1999: 113 [#8247] (synonym of transbaicalensis). JENDEK, 2006: 389 [#7441] (synonym of transbaicalensis; Palaearctic catalog). BELLAMY, 2008: 2332 [#8576] (synonym of transbaicalensis; world catalog).

Type material: Buprestis viridis Linné, 1758 Type locality: Europa. Type specimens: Not located. Described from unknown number of syntypes.

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Mordella rosacea Scopoli, 1763 Type locality: not given [“Carniola” is cited in the title of the publication]. Type specimens: Not located. Described from unknown number of syntypes. Mordella serraticornis Scopoli, 1763 Type locality: not given [“Carniola” is cited in the title of the publication]. Type specimens: Not located. Described from unknown number of syntypes. Buprestis linearis Schrank von Paula, 1781 Type locality: in Kagran Austriae Passava. Type specimens: Not located. Described from unknown number of syntypes. Buprestis lineata Schrank von Paula, 1782 Type locality and specimens: See: Buprestis linearis Schrank, 1781. Buprestis linearis Fabricius, 1792 Type locality: Dania. Type specimens: Not located. Described from unknown number of syntypes. Buprestis bicolor Schrank von Paula, 1798 Type locality: not given. Type specimens: Not located. Described from unknown number of syntypes. Buprestis filiformis Herbst, 1801 Type locality and specimens: See: Buprestis linearis Fabricius, 1792. Buprestis fagi Ratzeburg, 1837 Type locality: not given [“Wäldern Preussens” are cited in the title of the publication]. Type specimens: Not located. Described from unknown number of syntypes. Buprestis nociva Ratzeburg, 1837 Type locality: not given [“Wäldern Preussens” are cited in the title of the publication]. Type specimens: Not located. Described from unknown number of syntypes. Buprestis (Agrilus) capreae Chevrolat, 1838 Type locality: Paris. Type specimens: Primary: Lectotype (Fig. 44C) by present designation, ♂, (BMNH): “SYN-Type [p] [round label with blue border] \ Env. Paris [Kerremans’ MS] \ coll. Chevrolat [p] \ capreae Chevrolat Type [Kerremans’ MS] \ Kerremans. 1903-59 [p]”. Secondary: 1 ♀ PLT (BMNH). Described from unknown number of syntypes. Agrilus aubei Gory & Laporte, 1839 Type locality: Paris, Boulogne. Type specimens: Primary: Lectotype (Fig. 43D), sex not examined, designated by JENDEK (1998: 318) [#4481], (MNHN): “Transcriptio [p] var. Aubei Gory Type Paris [h] \ Muséum Paris 1952 Coll. R. Oberthur [p] [yellow label]”. Secondary: 2 PLT (MNHN). Described from unknown number of syntypes. Agrilus distinguendus Gory & Laporte, 1839 Type locality: Paris (Boulogne). Type specimens: Primary: Lectotype (Fig. 45M), sex

225 not examined, designated by JENDEK (1998: 321) [#4481], (MNHN): “[male sign] [h] \ Transcriptio [p] var. Distinguendus Gory Type Paris France m. [h] \ Muséum Paris 1952 Coll. R. Oberthur [p] [yellow label]”. Secondary: 1 PLT (MNHN). Described from unknown number of syntypes. Agrilus viridipennis Gory & Laporte, 1839 Type locality: Paris. Type specimens: Primary: Lectotype (Fig. 54P), sex not examined, designated by JENDEK (1998: 332) [#4481], (MNHN): “Type Gory [h] \ Transcriptio [p] Viridis Fabr. Viridipennis Gory type Paris Europe [h] \ Muséum Paris 1952 Coll. R. Oberthur [p] [yellow label]”. Described from unknown number of syntypes. Agrilus littlei Curtis, 1840 Type locality: Rae Hills in Annandale, Dumfries-shire. Type specimens: Not located. Described from 20 syntypes. Agrilus bicolor Redtenbacher, 1847 Type locality: not given [Austria is cited in the title of the publication]. Type specimens: Not located. Described from unknown number of syntypes. Agrilus quercinus Redtenbacher, 1847 Type locality: not given [Austria is cited in the title of the publication]. Type specimens: Not located. Described from unknown number of syntypes. Agrilus darwinii Wollaston, 1857 Type locality: Ribeiro de São Jorge (in the north of Madeira proper). Type specimens: Not located. Described from unknown number of syntypes. Agrilus imbellis Kerremans, 1892 Type locality: Inde. [Note: Locality dubious] Type specimens: Primary: Lectotype (Fig. 47F) by present designation, ♀, (MNHN): “imbellis (Laferté) Inde [h] \ ExMusaeo Mniszech [p] \ Kerremans vidit 1892 [p] \ imbellis Kerr. Type [Kerremans’ MS]”. Secondary: 1 PLT (MNHN). Described from unknown number of syntypes. Agrilus brevitarsis Lewis, 1893 Type locality: Main island ... Chiuzenji. Type specimens: Primary: Lectotype (Fig. 43U), ♀, designated by JENDEK (1995: 147) [#3768], (BMNH): “Type H.T. [p] [round label with red border] \ 14.VI. 81 [h] \ Japan. G. Lewis 1910-320 [p] \ Agrilus brevitarsus [sic!] Lewis Type [by Lewis hand]”. Described from unknown number of syntypes. Agrilus indulgens Kerremans, 1897 Type locality: Bahia: St. Antonio da Barra [Note: Locality confused, see OBENBEREGER, 1960 [#2394]]. Type specimens: Not located. Described from unknown number of syntypes. Agrilus rudis Abeille de Perrin, 1897 Type locality: not given. Type specimens: Primary: Lectotype (Fig. 52H), ♀, designated by JENDEK (2003: 9, 16) [#5878], (MNHN): “Türkei [h] \ Type [p] [red ink] \ Muséum Paris Coll. A. de PERRIN 1919 [p] \ rudis Ab. [h]”. Described from unknown number of syntypes.

226 Agrilus importunus Kerremans, 1903 Type specimens and locality: See: Agrilus imbellis Kerremans, 1892. Agrilus lewisi Kerremans, 1903 Type locality and specimens: See: Agrilus brevitarsis Lewis, 1893. Agrilus viridis variety montanellus Obenberger, 1916 Type locality: Nordböhmen ... Trautenau. Type specimens: Holotype by monotypy (Fig. 49O), ♂, (NMPC): “Boh. bor., 1911 Trutnov, J. Obenberger [h] \ viridis var. montanellus m. Typus unicum [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24825 [h] [orange label]”. Described from 1 specimen. Agrilus gebleri Obenberger, 1924 Type locality: Transbaikalia (Čita). Type specimens: Primary: Lectotype (Fig. 46P) by present designation, ♂, (NMPC): “Tschita, Transbaikalien, Hermann Frieb [p] \ Typus [p] [red label] \ Agrilus gebleri m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24907 [h] [orange label]”. Secondary: 11 PLT (NMPC); 1 PLT (BMNH); 1 PLT (HNHM). Described from unknown number of syntypes. Agrilus viridis caenus Obenberger, 1924 Type locality: Sicilia. Type specimens: Primary: Lectotype (Fig. 43Y), ♂, designated by JENDEK (2003: 12-13, 16) [#5878], (NMPC): “Sicilia [h] \ Typus [p] [red label] \ A. viridis ssp. caeneus m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24357 [h] [orange label]”. Secondary: 1 PLT (NMPC). Described from unknown number of syntypes. Agrilus viridis poppiusi Obenberger, 1924 Type locality: Nikolskaja: Fluvius Lena. Type specimens: Primary: Lectotype (Fig. 51D) by present designation, ♀, (NMPC): “Nikolskaja [p] \ Fl. Lena [p] \ B. Poppius [p] \ Typus [p] [red label] \ viridis sp. Poppiusi m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24253 [h] [orange label]”. Described from unknown number of syntypes. Agrilus viridis krogerusi Obenberger, 1924 Type locality: Verchnyj Udukunsk (Sibiria) Odenval. Type specimens: Primary: Lectotype (Fig. 48B) by present designation, ♀, (NMPC): “V. Udukunsk [p] \ Odenvall [p] \ Typus [p] [red label] \ viridis ssp. Krogerusi m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24252 [h] [orange label]”. Described from unknown number of syntypes. Agrilus viridis melantatus Obenberger, 1924 Type locality: Transbaikalia. Type specimens: Holotype by monotypy (Fig. 49G), ♀, (NMPC): “Transbaikal. [h] \ Typus [p] [red label] \ viridis ssp. melantatus m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24254 [h] [orange label]”. Described from 1 specimen. Agrilus brydli Obenberger, 1924 Type locality: Caucasus borealis. Type specimens: Primary: Lectotype (Fig. 43X) by present designation, ♂, (NMPC): “Caucasus b[orealis] [h] \ Typus [p] [red label] \ Agrilus Brýdli m. Type [h] Det.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Dr. Obenberger [p]”. Described from unknown number of syntypes. Agrilus viridis cernyi Obenberger, 1925 Type locality: Austria inf.: Wien, Serbia (Brus). Type specimens: Primary: Lectotype (Fig. 44E) by present designation, ♂, (NMPC): “Wien: Fr. J. Land, 23.5.[19]16 Curti [p] \ Cernyi m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24826 [h] [orange label]”. Secondary: 1 PLT (NMPC). Described from unknown number of syntypes. Agrilus vernadskii Obenberger, 1927 Type locality: Sibiria orientalis, Chitaizski-Sterena ... verisimiliter: Kitajskaja storona [= Chinese side of Amur river near Radde, Jewish Autonomous Oblast, Russia. See BODEMEYER, 1915: 438, 439 [#8261] and BODEMEYER, 1928:72-73, 79 [#7705]]. Type specimens: Holotype by monotypy (Fig. 54M), sex not examined, (NMPC): “B. v. Bodemeyer [leg.] Sibiria orient. Chitaizki-Sterena [p] \ Type [p] [red label] \ Agrilus Vernadskyi m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24913 [h] [orange label]”. Described from 1 specimen. Agrilus kiricenkoi Obenberger, 1935 Type locality: Kuldja, Fluvius Tuni-gol, Chalcha, Mongolia. Type specimens: Primary: Lectotype, sex not examined, designated by ALEXEEV & VOLKOVITSH (1989: 320) [#3241], (ZIN): “r. Tyingol, sr. tech. r. Khalka, 26-30 VI 1926, Kirichenko [leg.]”. Secondary: 2 PLT (ZIN); 2 PLT (NMPC). Described from unknown number of syntypes. Paralectotype (Fig. 47W) from NMPC is illustrated. Agrilus transbaicalensis Obenberger, 1935 Type locality: Transbaicalia, fluvius Čikol; Šaragol. Type specimens: Primary: Lectotype (Fig. 53Z) by present designation, ♀, (NMPC): “Transbaicalia mr.occ. Fl. Tshikoj: Sharagol, 1.VII.1928 TH. Lukjanovitsh [p] \ Typus [p] [red label] \ Agrilus transbaicalensis m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24918 [h] [orange label]”. Secondary: 1 PLT (ISNB); 1 PLT (NHMB); 1 PLT (ZIN); cca 30 PLT (NMPC). Described from unknown number of syntypes. Agrilus pusztae Obenberger, 1936 Type locality: Hungaria: Szolnok. Type specimens: Primary: Lectotype (Fig. 51O), ♂, designated by JENDEK (2007: 111) [#7967], (NMPC): “Szolnok VI [19]28 [h] \ Typus [p] [red label] \ Agrilus Pusztae m. Type [h] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24 914 [h] [orange label]”. Described from unknown number of syntypes. Agrilus towadensis Miwa & Chûjô, 1940 Type locality: lake-side of Towada, Pref. Akita, Japan. Type specimens: Holotype by monotypy was not examined and should be preserved in TARI (AKIYAMA & OHMOMO, 1997 [#4091]). Described from 1 specimen. Agrilus suvorovi salicivola Kurosawa, 1963 Type locality: Fukushima Pref.: Mt. Aoki-yama, AizuWakamatsu City. Type specimens: Primary: Holotype (Fig. 52N), ♂,

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA (NSMT): “Odayama Mt. Monden V. Nr. Wakamatsu 27. V. 1948 Y. K[urosawa]. [h] \ Holotype [p] Agrilus suvorovi salicivora [sic!] Y. Kurosawa, 1963 [h] [red label, with black border]”. Secondary: 6 PT (NSMT). Described from unknown number of specimens (HT, AT, PT). Agrilus cojbalsanensis Cobos, 1968 Type locality: Cojbalsan aimak, Chamardavaa ul, 80 km SW. von somon Chalchingol. 600 m. alt. Type specimens: Primary: Holotype (Fig. 44P), ♀, (HNHM): “Mongolia, Čojbalsan aimak, Chamardavaa ul, 80 km SO [sic!] von Somon Chalchingol. 600 m. Exp. Dr. Z. KASZAB, 1965 [p] \ Nr. 401, .13. VIII. 1965 [p] \ Holotypus A. Cobos [p] [red label] \ Agrilus cojbalsanensis nov. sp. Holotypus [h] A. Cobos det. Described from 1 specimen. Diagnosis: 5.3-10.2 mm; body larger and more elongate; eyes moderate or large; pronotum widest at anterior half, rarely at middle; pronotal sides moderately arcuate; apical third of elytra more elongate. Species group: Viridis. Relevant species: A. cuprescens (Ménétriés, 1832); A. suvorovi Obenberger, 1935. Specimens examined: CHINA: Beijing: 1 ♂ (EJCB): “China, Beijing distr., Yan Shan, Dongling Mts., 39º58’N, 115º26’E, 1400 m, Xiaolongmen, 15 16.VI.2001”. Nei Mongol: 1 ♂ (EJCB): “Paoteou, 19.vii.1919”. Shaanxi: 1 ♂ (EJCB): “40 km SE of Taibai Shan Mts., Hou Zen Zi vill. env., Z. Jindra lgt. cca 1200m, China, Shaanxi prov., 11 July 1998, Qing Ling Shan mts.”. JAPAN: Honshu: 1 ♂ (EJCB): “Mt. Hanazono, Ibaraki Pref., Japan, 23. VII.1979, A. Nishiyama leg.”; 11 (EJCB): “Tochigi Pref., S. Ohmomo leg., 1995.6.17, Uwadaira, Shioya”; 5 (EJCB): “Fukushima Pref., NE JPN, S. Ohmomo leg., 1998.6.6 7, Tateiwa mura, S Aizu”; 1 ♂, 1 ♀ (EJCB): “Japan, Yamagata, Myozawa, Oguni town, 13.vii.2005, N. Sugiura leg.”. Kyushu: 1 ♂, 1 ♀ (EJCB): “Aizu, Wakamatsu, Japan, 27.v.1948”. KAZAKHSTAN: 3 (NMPC): “Kazch., 11 15.7.1977, Karkalinsk, Vasilenko leg.”; 1 (ZIN): “khr. Tarbagatai, 30km ZSZ p. Tarbagatai, Kazakhstan, Emelyanov, 19 20.vii.1983 [in Russian]”; 2 (EJCB): “r. Tainty, 15 km VSV Targyna, Altai, V Kazakh. obl. 1 2.vii.1983, Belokabyl’skii [in Russian]”; 3 (MKCY): “Aktyubinsk, 6.1985”; 1 (EJCB): “Kazakhstan NE, 100 km SEE Ust’ Kamenogorsk, 5 km NE Panteleymonovka, Kaiyndy forest, 16 20.6.1993, Napolov”; 1 ♂, 1 ♀ (EJCB): “W Kazakhstan, Aktyubinsk, Mugodzhany, Alinskol, 6.6.1995, leg. Dolin”. MONGOLIA: 1 (NMPC): “Heuve Tola, Urga, Mong.”; 1 (NMPC): “Mongolia, Songino ad., Ulan Bator, 13.vii.1963, leg. Burakowski et H. Szelegiewicz”; 3 ♀ (EJCB): “Songino u Ulan batoru, Nongolia [sic!], 11.vii.1965, Boruvka lgt.”; 2 (MKCY): “Ar Khangaiskii a., Tèvshrulèkh, 15.viii.1970 [in Russian]”; 1 (ZIN): “Mongoliya, Vost. aimak, g. Dèrkhin Cagan Obo, 60km VSV Byan Burda, G. Medvedev, 21.vii.1971 [in Russian]”; 1 (MKCY): “Khubs. a., r. Selenga, 30 km VYuV Ikh Ula, 19.vii.1972 [in Russian]”; 1 (MKCY): “Ar Khangaiskii a., r. Tamir y Cènkhèra, 7.vi.1972 [in Russian]”; 1 (EJCB): “Mongoliya, Ara Khangaickii aimak, 48 km Yu Tèvshrulèkha, Emelyanov, 2 3. vii.1975 [in Russian]”; 1 ♂ (ZIN): “Mongolia, Khubsugul’sk. aimak, r. Selenga, Ikh Ily, Emel’yanov 12.vii.1975 [in Russian],”; 4 (EJCB): “Mongolia, Arkhangal, 5 km S of lake Bulgana, 2.7.1975”; 1 (EJCB): “MNR, Centr. a., poima Toly, 22.8.1976, Lopov [in Russian]”; 4 (EJCB): “Mongolia, Bayan Ulgii ajmak, pass Shinedavg, 21.7.1976”; 1 (MKCY): “Selengiiski a., Shamor, 1.7.1978 [in Russian]”; 4 (EJCB): “Mongolia N, Selenga aymak env., Shamar, 01.07.1984, Kuzmin leg.”; 4 (ZIN): “MNR, Selenginsk. aimak, Shamar, 1.viii.1984, Kuzmin [in Russian]”; 1 ♂ (EJCB): “Mongolia or., Chentejský ajmak, dol. Mörön gol, B. Malec lgt. / 5 31.7.1985”; 1 (NMPC): “Mong., Chanuj via Bulgan,

227 23.8.1965, loc 34, Dlabola”; 6 (EJCB): “Mongolia, Selenga Aimak, Ero Gol Riv., 28 30.7.2002, 650m, 49º48’N, 106º35’E, M. Danilevsky leg.”; 55 (EJCB): “Mongolia, 20.vi.2003, 20km E Terelzh, 60km NE of Ulaanbaatar, Tuul riv.valley Terelzh N.P., Z. Jindra lgt., 1200 1400m”; 1 ♂ (EJCB): “Mongolia, 100 km E, Ulanbatar, 20 km NE, Tereltz, Tuul riv., 15 21.vii.2003, J. Halada leg.”; 4 ♂ (EJCB): “Mongolia, 14 20.7.2003, 20km E Terelzh, 1200 1400m, 60km NE of Ulaanbaatar, Tuul riv.valley Terelzh N.P., Z. Jindra lgt.”; 18 (EJCB): “Mongolia, 20. June 2003, 20km E Terelzh, 60km NE of Ulaanbaatar, Tuul riv.valley \ Terelzh N.P., cca 1200 1400m, Z. Jindra lgt.”; 31 (EJCB): “Mongolia, 20.June 2003, Ulaanbaatar, near Zaysan, Tuul riv.vall., Z Jindra leg.”; 28 (EJCB): “Mongolia, 50 km E Ulanbatar, Tuul riv. 22.vi., J. Halada leg. 2003”; 27 (EJCB): “Mongolia, 11.July 2003, Ulaanbaatar, near Zaysan, Tuul riv.vall., Z Jindra leg.”; 1 ♂ (EJCB): “Mongolia, 100 km E Ulanbatar, 20 km NE, Tereltz, Tuul riv., 15 21.vii.2003, J. Halada leg.”; 3 (EJCB): “Mongolia, 14 20. July 2003, 20km E Terelzh, 60km NE of Ulaanbaatar, Tuul riv.valley \ Terelzh N.P., cca 1200 1400m, Z. Jindra lgt.”; 1 ♂, 3 ♀ (EJCB): “Mongolia C, Mongol Els n.res., N17º24’, E103º39’, dunes, 1320 m, 31.7., Kadlecová lg., 2005”; 2 ♀ (EJCB): “Mongolia W, 40 km SW Uliastay, dunes, 18.vii.2005, J. Halada lg.”; 1 ♀ (EJCB): “Mongolia C, Mongol Els n.res., N17º24’, E103º39’, dunes, 1320 m, 31.7., J. Halada lg., 2005”. RUSSIA: Altay Respublika: 2 (MDCM): “18 km severno Gorno Altaiska, 15.7.1984, iva [in Russian]”. Amurskaya oblast’: 1 (EJCB): “USSR, 15.6.1975, Amurskaja Obl., Saskali, Bereja R., leg. V. Kuznetsov”. Buryatiya Respublika: 1 (ZIN): “Transbaicalia mr. occ. fl. Tshikoj: Sharagol 1.VII.1928 Th. Lukjanovitsh”; 1 ♀ (ZIN): “Buryatiya Verkh. r. Turki 20.8.1966 Kabakov [in Russian]”; 5 (FACF): “Russia Siberia, S. Buriatia reg., Novoselenginsk, 10.vi.2002”; 3 ♀ (EJCB): “Russia Siberia, S Burjatia reg., Ochetui, 16.vi.2002”. Chelyabinskaya oblast’: 9 (EJCB): “Russia, South Ural Mts., Cheliabinsk, 3.vii.1990”; 3 (EJCB): “Russia, Cheliabinsk reg., Chebarkal distr., Bishkil vill., 17.vii.2001”; 3 (FACF): “Russia Siberia, Cheliabinsk reg., Bishkil, 31.v.2001”. Irkutskaya oblast’: 1 (MDCM): “Irkutskaya obl. Slyudyanka 12.8.1971”; 1 (FACF): “Russia Siberia W Baikal lake, Sarma, 20.vi.1997”. Kamchatskiy kray: 1 ♂, 2 ♀ (EJCB): “Kamchatka, Shchapino, 24.vi.1960, Kononov leg. [in Russian]”. Leningradskaya oblast’: 1 (FACF): “Russia, St. Petersburg, Ivanovskoe, 2.vi.1999”. Lipetskaya oblast’: 1 (MCCD): “35км СВВ г. Елец, ур. Плющань, 7.6.1996 Цуриков”; 1 (MCCD): “Липецкая обл., Чаплыгинский р н, 7км Ю с. Дубовое 2.7.1997 Цуриков”; 1 (MCCD): “Липецкая обл., Добровский р н, окр. с. Гудово 1.7.1997 Цуриков”. Primorskiy kray: 1 ♂ (ZIN): “Russia, Primorsk, Krasnoarmeysk. r., 9.vii.1971, Mt. Armu, On Lonicera”; 1 (MKCY): “Prim. kr., Ussuriisk, 7 km vverkh ot s. Yakovka, 5.vii.1975, Kuznecov [in Russian]”; 1 (MDCM): “Ussuri, Sedanka, 26.7.1983 S. Seloborodov”; 1 ♂ (EJCB): “SU, Ussuri reg., Jasnoe, 12 19.vii.1989, R. Červenka leg.”; 1 ♀ (EJCB): “USSR Sibiria or., Ussuri region 11 14.7, Jasnoe, 450 m, 1989, leg. St. Jákl”; 1 ♂ (EJCB): “SU Ussuri reg. Novovarvarovka, 6 10.VII.1989, R. Červenka leg.”; 1 ♀ (EJCB): “SU Ussuri reg., Novovarvarovka, 6 10.7.1989, M. Nikodým lgt.”; 3 (EJCB): “Sib. or. m., Primorje, Sichote Alin Mts., Sokolči, 1 15.7.1990, Kadlec + Voříšek lg.”; 1 ♀ (EJCB): “USSR, Primorsk. kraj, Novocugujevka env., 16 18. VII.1990, leg. Snížek”; 1 ♂ (EJCB): “Sib. or. m., Primorje, Ussuri res., 20.7.1990, Kadlec + Voříšek lg.”; 1 ♀ (ZIN): “Prim. Khasansk., Bamburovo, na ive 20.8.[19]90 Napolov [in Russian]”; 1 ♀ (EJCB): “USSR, Primorsk. kraj, Novocugujevka env., 18.VII.1990, Kantner leg.”; 1 ♂ (EJCB): “Sib. or., Primorje, Komarovka flum., Kamenushka env., 300 m, Vořišek lgt., VI.1992”; 1 ♀ (EJCB): “Russia, Primorsk. reg., 18 21.7.1992, Rezanovka, E. Kučera leg., ex Lonicera”; 1 ♂ (EJCB): “SSSR, Primorskij k., Kajmanovka 10 22.vii.1993, Z. Kletečka leg.”; 2 (MKCY): “Russia, Far East, 20km SE Ussuriysk, Gornotaezhnoe vill. env., 30.06.2003, Napolov leg.”;

228 2 (EJCB): “Russia, Primorsky Kray, Kaymanovka vil., N43º37’, E132º14’, 7 11.vii.2007, 100m, V. Grebennikov”; 1 (EJCB): “Russia, Primorsky kray, Vladivostok, N43º12’, E131º59’, 6. vii.2007, 50m, Grebennikov”. Sakha Respublika: 4 (NMPC): “Yakutsk 1901 Olenin [in Russian]”; 1 (EJCB): “28.6.1987, Jakutia, Jakutsk, Dr. Sobota leg.”. Sakhalinskaya oblast’: 3 (MKCY): “Yuzhno Sakhalinsk, p. Novoalexandrovsk, po r. Krasnosel’skoi, M. Nesterov, 2.7.1975 [ in Russian]”; 1 ♂ (ZIN): “Russia, Kunashir isl., Goryachee lake, 12.ix.1976, broadleaf forest, Kuznetsov, V. leg.”; 1 ♂, 1 ♀ (ZIN): “S Sakhalin, Firsovo, 21.8.1977, Kuznetsov”; 1 (SGCB): “E Russia, Russland, J. Sachalin umg., Dolinsk / Starodubsk, 31.7.1977, leg. V. Kuznacov”; 1 (EJCB): “o. Sakhalin, Khopski r n., Pozharskoe, na Ive, 14.8.1977 g., V. Kuznetsov”; 1 ♀ (EJCB): “Sakhalin, Novoalexandrovsk, 15.III. [sic!].1989”; 1 ♀ (EJCB): “S Sakhalin, Ins. Nowoalersandrow [sic!], 4.1989, leg. Nesteron”; 1 ♂ (EJCB): “Russia or, Sakhalin, Okha env., 1 10. vii.1991, Babarukov leg.”; 1 (HMMG): “Russia, Sakhalin, Anira distr., 5km W Petropavlovkiy, tributary of Lyutoga riv., 20 21. vii.1993, Pütz & Wrase”; 1 ♀ (EJCB): “Russia, Sakhalin, Korsakov distr., 3 km W Kirilloco vill., Dryun river, 22/23.vii.1997, Pütz & Wrase”. Severnaya Osetiya Respublika: 3 (ZIN): “Kavkaz, okr. Ordzhonikidze 3.6.1949 Kuri [in Russian]”. Sverdlovskaya oblast’: 1 (MKCY): “11.07.1976, Sverdlovsk. obl., Sisertskii r n., Fomino, Kozarev, A. V. [in Russian]”. Tyva Respublika: 1 ♀ (ZIN): “Tuva, V. Zkamenka, 20.vii.1949 [in Russian]”; 1 (ZIN): “Tyva, c. Mugur Aksy, B. Korotyaev, 11 7 1970 [in Russian]”; 1 (ZIN): “Tyva, Dvyurskii r n, pos. Sagl[p]]y, B. Korotyaev, 7 7 1971 [in Russian]”; 1 (ZIN): “Tyva, pos. Èrzin, B. Korotyaev, Na ive, 1 7 1972 [in Russian]”; 1 ♂ (EJCB): “Tuvinskaja ASSR, Chadyn 29 30.94 Z. Kletečka lgt.”; 1 ♂, 1 ♀ (EJCB): “Russia, Tuva, Kara Chaak vill, 17.vi., Patrikeev V., leg. 1997”; 11 (EJCB): “Russia, Tuva, right bank of Biy Khem riv. 800 m, Ujukskiy Mts. 2 5.VII.1999, S. N. Vashchenko leg.”; 2 (EJCB): “Russia, Tuva, right bank of Biy Khem riv. 800 m, Ujukskiy Mts. 22.V. 6.VI.1999, S. N. Vashchenko leg.”; 11 (EJCB): “Russia, S Tuva, 2000, Kachemskiy distr., 1000 1400m, lower of Buren riv., 15 24.vii., A. Vashchenko leg.”; 33 (EJCB): “Siberia SW Tuva, Zapadniy Tannu Ola Mts., Khandagaity vill. river, 50°44’N, 92°02’E, 1200m, leg. Jan Hron, 6.vii.2002”. Voronezhskaya oblast’: 1 (MCCD): “25км NNO г. Воронежа, Усманский Бор, 9.6.1988 Цуриков”. Zabaykalskiy kray: 2 (PMCL): “USSR, Zabajkalí, Čitská o., Butevken jez. 3 6.7.1990 Moravec, P. lgt.”; 6 (PMCL): “USSR, Zabajkalí, Čitská o., Butevken jez. 3 6.7.1990 Moravec, P. lgt.”.

Adult occurence: 4-5-6-7-8-9 (n=109). Faunal records published by COBOS, 1972 [#591] (as viridis krogerusi) are also included. Altitude range: 50-1600 m (n=19). Faunal records published by COBOS, 1972 [#591] (as viridis krogerusi) are also included. Host plants: Acer: POMERANCEV, 1938 [#4977]; ILINSKIJ, 1948 [#5670]; SCHAEFER, 1949 [#2565]; TER MINASYAN, 1955 [#7893]; HEERING, 1956 [#3716]; BASURMANOVA, 1958 (as basurmanovae) [#4975]; KHRAMCOV & PADIJ, 1965 [#8649]; ALEXEEV, 1981 [#61]; ABAI & ADELI, 1984 [#5808]; VOLKOVITSH & ALEXEEV, 1994 [#2974]; CURLETTI, 1994 [#658]; BÍLÝ, 2002 [#5877]; HASTIR & GASPAR, 2002 [#6211]; HUA LI ZHONG, 2002 [#6696]. – A. campestre: BASURMANOVA, 1958 [#4974]. – A. ginnala: BASURMANOVA, 1958 [#4974]. – A. platanoides: ZAKHAROV & LEVKOVICH, 1951 [#8638]; BASURMANOVA, 1958 [#4974]. – A. pseudoplatanus: GUTOWSKI & MILKOWSKI, 2008 [#8244]. – A. tataricum: ZAKHAROV & LEVKOVICH, 1951 [#8638]; ALEXEEV, 1957 [#3226] (as aceris); BASURMANOVA, 1958 [#4974]. – Alnus: PANZER, 1806 (as linearis) [#4052]; JUDEICH & NITSCHE, 1889 [#5686];

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA ESCHERICH, 1923 [#6717]; ILINSKIJ, 1948 [#5670]; SCHAEFER, 1949 [#2565]; HORION, 1955 (also as nocivus) [#1285]; TER MINASYAN, 1955 [#7893]; HEERING, 1956 [#3716]; KHRAMCOV & PADIJ, 1965 [#8649]; IVLIEV & KONONOV, 1966 (as kamtschaticus (unavailable name)) [#8089]; ABAI & ADELI, 1984 [#5808]; NIEHUIS & TEZCAN, 1993 (uncertain) [#1910]; CURLETTI, 1994 [#658]; BÍLÝ, 2002 [#5877]; HASTIR & GASPAR, 2002 [#6211]. – A. glutinosa: ZAKHAROV & LEVKOVICH, 1951 [#8638]. – A. incana: ALEXEEV & SLEPKO, 1998 [#4887]. – A. viridis: FRANZ, 1974 (as nocivus) [#4334]. – Betula: PANZER, 1795 [#4041]; SAMOUELLE, 1819 [#7071]; AUBÉ, 1837 (also as aubei) [#4034]; GORY & LAPORTE, 1839 [#3738] (as aubei); MARSEUL, 1866 [#676]; JUDEICH & NITSCHE, 1889 [#5686]; ESCHERICH, 1923 [#6717]; SAINTE CLAIRE, 1935 (as aubei) [#8551]; POMERANCEV, 1938 [#4977]; ILINSKIJ, 1948 [#5670]; SCHAEFER, 1949 [#2565]; HORION, 1955 (also as fagi) [#1285]; KHRAMCOV & PADIJ, 1965 [#8649]; ALEXEEV, 1981 [#61]; NIEHUIS & TEZCAN, 1993 [#1910]; CURLETTI, 1994 [#658]; BÍLÝ, 2002 [#5877]; BRECHTEL & KOSTENBADER, 2002 (as fagi) [#5766]; HASTIR & GASPAR, 2002 [#6211]. – B. alba: LINNÉ, 1760 [#1727]; FABRICIUS, 1781 [#832]; GMELIN, 1790 [#1087]; FABRICIUS, 1792 [#834]; CEDERHIELM, 1798 [#5314]; MARSHAM, 1802 [#6052]; BEDEL, 1921 (also as fagi) [#170]; NIEHUIS, 1988 (as fagi) [#1912]. – B. nana: BRUNDIN, 1934 (as albisetulosus) [#8542]. – B. odorata: KROGERUS, 1922 [#3916]; KROGERUS, 1925 [#3472]; KANGAS, 1947 [#3622]. – B. tortuosa: BRUNDIN, 1934 (as krogerusi) [#8542]. – B. verrucosa: KROGERUS, 1922 [#3916]; KROGERUS, 1925 [#3472]; KANGAS, 1947 [#3622]; ZAKHAROV & LEVKOVICH, 1951 [#8638]. – Carpinus: PANZER, 1806 (as linearis) [#4052]; HORION, 1955 (as fagi) [#1285]; ZAGAIKEVICH, 1957 [#4976]; PADIJ, 1960 [#8628]; NIEHUIS & TEZCAN, 1993 (uncertain) [#1910]; CURLETTI, 1994 [#658]; BÍLÝ, 2002 [#5877]; HASTIR & GASPAR, 2002 [#6211]. – C. betulus: HUSSON et al. 1952 [#7085]. – C. orientalis: TER MINASYAN, 1955 [#7893]. – Corylus: PANZER, 1806 (as linearis) [#4052]; SAMOUELLE, 1819 [#7071]; HEERING, 1956 [#3716]; ALEXEEV, 1981 [#61]; CURLETTI, 1994 [#658]; BÍLÝ, 2002 [#5877]; HASTIR & GASPAR, 2002 [#6211]; MUSKOVITS & HEGYESSY, 2002 [#5806]. – Fagus: RATZEBURG, 1837 (as fagi and nocivus) [#2498]; KALTENBACH, 1874 (as fagi) [#8154]; KEPPEN, 1882 (as fagi and nocivus) [#8636]; JUDEICH & NITSCHE, 1889 [#5686]; BEDEL, 1921 (also as fagi) [#170]; ESCHERICH, 1923 [#6717]; SAINTE CLAIRE, 1935 (as aubei and nocivus) [#8551]; ILINSKIJ, 1948 [#5670]; SCHAEFER, 1949 [#2565]; WOELFLE, 1951 [#7086]; HORION, 1955 (also as nocivus and fagi) [#1285]; TER MINASYAN, 1955 [#7893]; KHRAMCOV & PADIJ, 1965 [#8649]; ALEXEEV, 1981 [#61]; NIEHUIS, 1988 [#1912]; NIEHUIS & TEZCAN, 1993 [#1910]; CURLETTI, 1994 [#658]; HASTIR & GASPAR, 2002 [#6211]. – F. orientalis: ADELI & SOLEIMANI, 1976 [#8300]; ABAI & ADELI, 1984 [#5808]. – F. sylvatica: HUSSON et al., 1952 [#7085]; HEERING, 1956 [#3716]; NOVAK, 1964 [#6567]; NIEHUIS, 1988 (as fagi) [#1912]; BRECHTEL & KOSTENBADER, 2002 (as fagi) [#5766]. – Salix: MARSEUL, 1866 [#676]; POMERANCEV, 1938 [#4977]; ILINSKIJ, 1948 [#5670]; SCHAEFER, 1949 [#2565]; HORION, 1955 [#1285]; TER MINASYAN, 1955 [#7893]; HEERING, 1956 [#3716]; MAKHNOVSKIJ, 1958 (as transbaicalensis) [#6090]; MAKHNOVSKIJ, 1959 (as transbaicalensis) [#5829]; KHRAMCOV & PADIJ, 1965 [#8649]; IVLIEV & KONONOV, 1966

229

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA (as kamtschaticus (unavailable name)) [#8089]; ALEXEEV, 1975 [#52]; ALEXEEV, 1981 [#61]; ALEXEEV, 1989 (as vernadskii, gebleri and transbaicalensis) [#64]; ALEXEEV & VOLKOVITSH, 1989 (as transbaicalensis) [#3241]; NIEHUIS & TEZCAN, 1993 [#1910]; CURLETTI, 1994 [#658]; KRÓLIK & MAJEWSKI, 1994 [#3471]; TLEPPAEVA, 1999 [#6551]; STUMPF et al., 2001 (as linearis) [#5788]; BÍLÝ, 2002 [#5877]; HASTIR & GASPAR, 2002 [#6211]; HUA LI ZHONG, 2002 [#6696]; TLEPPAEVA & ISHKOV, 2004 [#7211]. – S. acutifolia: ZAKHAROV & LEVKOVICH, 1951 [#8638]. – S. alba: ZAKHAROV & LEVKOVICH, 1951 [#8638]. – S. auriculata: CURTIS, 1840 (as littlei) [#660]. – S. aurita: BRECHTEL & KOSTENBADER, 2002 [#5766]. – S. caprea: CHEVROLAT, 1838 [#490] (as capreae); HUBENTHAL, 1902 (as linearis) [#8530]; DAEHNE, 1912 (as linearis) [#3609]; BEDEL, 1921 [#170]; KROGERUS, 1925 (as fagi) [#3472]; SAINTE CLAIRE, 1935 (as linearis) [#8551]; KANGAS, 1947 [#3622]; ZAKHAROV & LEVKOVICH, 1951 [#8638]; HORION, 1955 (as fagi) [#1285]; BRECHTEL & KOSTENBADER, 2002 [#5766]; GUTOWSKI & MILKOWSKI, 2008 [#8244]. – S. cinerea: SCHIÖDTE, 1864 [#2595]; MERKL, 1991 [#6564]; BRECHTEL & KOSTENBADER, 2002 [#5766]. – S. pentrandra: ZAKHAROV & LEVKOVICH, 1951 [#8638]. – S. purpurea: ZAKHAROV & LEVKOVICH, 1951 [#8638]; LISKENNE, 1994 (as ignotus) [#4279]. – S. sachalinensis: KUROSAWA, 1963 (as salicivola) [#1617]; AKIYAMA & OHMOMO, 1997 (as salicivola) [#4091]. –– S. triandra: ZAKHAROV & LEVKOVICH, 1951 [#8638]. – S. viminalis: SCOPOLI, 1763 (as serraticornis) [#2606]; BRECHTEL & KOSTENBADER, 2002 [#5766]. – S. yezoenis: KUROSAWA, 1963 (as salicivola) [#1617]; AKIYAMA & OHMOMO, 1997 (as salicivola) [#4091]. –Tilia: JUDEICH & NITSCHE, 1889 [#5686]; ESCHERICH, 1923 [#6717]; POMERANCEV, 1938 [#4977]; ILINSKIJ, 1948 [#5670]; HEERING, 1956 [#3716]; ALEXEEV, 1981 [#61]; CURLETTI, 1994 [#658]; BÍLÝ, 2002 [#5877]; HASTIR & GASPAR, 2002 [#6211]. – T. cordata: ZAKHAROV & LEVKOVICH, 1951 [#8638]. –Ulmus: HUA LI ZHONG, 2002 [#6696]. –U. minor: WEIDLICH, 1989 [#6165]. Dubious: Castanea: CURLETTI, 1994 [#658]; BÍLÝ, 2002 [#5877]. – Myrica: SAINTE CLAIRE, 1935 (as nocivus) [#8551]; HORION, 1955 (as nocivus) [#1285]; CURLETTI, 1994 [#658]. – M. gale: BEDEL, 1921 (as nocivus) [#170]; KROGERUS, 1925 (as nocivus) [#3472]. – Ostrya carpinifolia: NOVAK, 1964 (as aubei) [#6567]. – Rhamnus: CAILLOL, 1913 [#4053]. – Sorbus: MUSKOVITS & HEGYESSY, 2002 [#5806]. Erroneous: Populus (misidentification of A. suvorovi Obenberger): JUDEICH & NITSCHE, 1889 [#5686]; HEYDEN, 1897 (as nocivus) [#5286]; DAEHNE, 1912 [#3609]; BEDEL, 1921 (as nocivus) [#170]; POMERANCEV, 1938 [#4977]; ILINSKIJ, 1948 [#5670]; SCHAEFER, 1949 [#2565]; GRECHKIN, 1951 [#8606]; TER MINASYAN, 1955 [#7893]; ALEXEEV, 1957 (as erronesculus) [#3226]; SLIVKINA, 1958 [#8660]; KHARAZISHVILI, 1960 [#8630]; GRECHKIN & VORONCOV, 1962 [#8242]; KHRAMCOV & PADIJ, 1965 [#8649]; ALEXEEV, 1981 [#61]; ALEXEEV, 1989 (as vernadskii) [#64]; ALEXEEV & VOLKOVITSH, 1989 (as vernadskii) [#3241]; TLEPPAEVA, 1999 [#6551]; HUA LI ZHONG, 2002 [#6696]; TLEPPAEVA & ISHKOV, 2004 [#7211]. – Quercus: PANZER, 1806 [#4052]; JUDEICH & NITSCHE, 1889 [#5686]; DAEHNE, 1912 [#3609]; ESCHERICH, 1923 [#6717]; SCHAEFER, 1949 [#2565]; ABAI & ADELI, 1984 [#5808]. – Ribes (misidentification of A. ribesi Schaefer):

DEKHTIAREV, 1928 [#8663]; TER MINASYAN, 1955 (as fagi) [#7893]; VOLKOVITSH, 1977 (as aubei) [#2985]; ALEXEEV, 1981 [#61]; SAVKOVSKII & RYBALOV, 1981 [#6092]. – Rosa (misidentification of A. cuprescens (Ménétriés)): JUDEICH & NITSCHE, 1889 [#5686]; WEISS, 1916 [#6742]; WEISS, 1916 (as fagi) [#5954]; WEISS, 1916 [#3913]; ESCHERICH, 1923 [#6717]; FROST, 1924 (as fagi) [#1030]; BRITTON, 1926 (as fagi) [#6727]; FISHER, 1928 (as fagi) [#980]; CURLETTI, 1994 [#658]. – Rubus (misidentification of A. cuprescens (Ménétriés)): VASILEV, 1927 (as fagi) [#5834]; PJATAKOWA, 1930 [#8320]. Distribution: Local: CHINA: Beijing, Hebei, Heilongjiang, Henan, Hubei, Jilin, Liaoning, Nei Mongol, Shaanxi, Shandong, Shanxi, Xinjiang. JAPAN: Hokkaido, Honshu, Kyushu. MONGOLIA. RUSSIA: Amurskaya oblast’, Buryatiya Respublika, Kamchatskiy kray, Irkutskaya oblast’, Magadanskaya oblast’, Primorskiy kray, Sakhalinskaya oblast’, Tyva Respublika, Zabaykalskiy kray. Global: AFRICA: ALGERIA, MOROCCO, TUNISIA. ASIA: ARMENIA, AZERBAIJAN, CHINA, GEORGIA, IRAN, JAPAN, KAZAKHSTAN, KYRGYZSTAN, RUSSIA, TURKEY, TURKMENISTAN, UZBEKISTAN. EUROPE: ALBANIA, AUSTRIA, BELARUS, BELGIUM, BOSNIA & HERZEGOVINA, BULGARIA, CROATIA, CZECH REPUBLIC, DENMARK, ESTONIA, FINLAND, FRANCE, GERMANY, GREECE, HUNGARY, ITALY, LATVIA, LIECHTENSTEIN, LITHUANIA, LUXEMBOURG, MACEDONIA, MOLDOVA, NETHERLANDS, NORWAY, POLAND, PORTUGAL, ROMANIA, RUSSIA, SERBIA & MONTENEGRO, SLOVAKIA, SLOVENIA, SPAIN, SWEDEN, SWITZERLAND, TURKEY, UKRAINE, UNITED KINGDOM. Remarks: Distributional records from Fujian, Xizang and Yunnan (China) cited by HUA LI ZHONG, 2002 [#6696] need verification.

Agrilus volkovitshi Jendek Fig. 23F (habitus of HT); Fig. 38A (aedeagus) volkovitshi Jendek, 2007 (Agrilus) JENDEK, 2007: 33 34 [#8006] (description). [#8576] (world catalog).

BELLAMY, 2008: 2360

Type material: Agrilus volkovitshi Jendek, 2007 Type locality: China, Yunnan, 100 km W of Kunming, Diaolin Nat. Reservation. Type specimens: Primary: Holotype (Fig. 23F), ♂, (EJCB): “China, Yunnan, 22.V.-2.VI., 100km W of Kunming, 1993, Diaolin Nat. Reservation”. Secondary: 4 PT (CBCS); 4 PT (SOCI); 3 PT (NMPC); 3 PT (ZIN); 168 PT (EJCB). Described from 183 specimens (HT, PT). Diagnosis: 4.2-6.2 mm; frons strongly convex, without long erect pubescence; antennae slender and shorter, not reaching to posterior pronotal angles; pronotum without lateral impressions; elytra less prolonged; prosternal process flat, without sharply elevated angles; legs slender and aedeagus more asymmetric. Species group: Sulcicollis.

230 Relevant species: A. miwai Obenberger, 1936; A. reichardti Obenberger, 1935. Specimens examined: CHINA: Yunnan: 1 ♂ (EJCB): “P. R. China, Yunnan, Jizu Shan Mt., N25º57’, E100º24’, 2 5.vi.2007, 2000m, V. Grebennikov”.

Adult occurence: 6 (n=1). Altitude range: ~2000 m (n=1). Host plants: Quercus: JENDEK, 2007 [#8006]. Distribution: CHINA: Yunnan.

Agrilus voriseki Jendek Fig. 17G (habitus of HT); Fig. 35L (aedeagus) voriseki Jendek, 1995 (Agrilus) JENDEK, 1995: 175, 178 [#3767] (description). ALEXEEV, 1989: 487 ([Note: Misidentified as pratensis djukini]; characters in key; Chita; Buryatia). JENDEK, 2006: 403 [#7441] (Palaearctic catalog). LÖBL & SMETANA, 2007: 30 [#8558] (complement to distribution in Palaearctic catalog). BELLAMY, 2008: 2360 [#8576] (world catalog).

Type material: Agrilus voriseki Jendek, 1995 Type locality: Sib. or. Primorje, Komarovka flum, Kamenushka env. 300 m. Type specimens: Primary: Holotype (Fig. 17G), ♂, (EJCB): “Sib. or. Primorje, Komarovka flum, Kamenushka env. 300 m, Vorisek lgt. VI. 1992”. Secondary: 1 PT (HMMG); 2 PT (NMPC); 1 PT (MNCA); 9 PT (EJCB). Described from 14 specimens (HT, PT). Diagnosis: 4.7-6.5 mm; golden-green; golden-brown; rarely blue; lateral sides of pronotum less impressed, impressions with granulose sculpture; elytral apices subtruncate. Species group: Betuleti. Relevant species: A. pseudocyaneus Kiesenwetter, 1857. Specimens examined: CHINA: Heilongjiang: 1 ♀ (ZIN): “st. Gaolinczy, vostochnaya liniya Kvzh.d. Kitai 2.vii.940 [in Russian]”. KAZAKHSTAN: 1 ♂ (MNCA): “Kazakhstan, Semipalatinsk reg., Putintzevo, 20 km N Zyrijanovsk, 08 29.Juni, 2005, leg. K. Hadulla”; 1 ♀ (EJCB): “NE Kazakhstan, sS Altai, env. Zyranovsk, 700 m, 19 25.6.1994”. MONGOLIA: 1 (ZIN): “Mong. 12/vi 94 Tergildzhi 12 13.VI. [old style] [ 24 25.VI. new style] 1894 [V.A. Kashkarov] [in Russian]”; 1 ♂, 2 ♀ (ZIN): “Mong. 25/vii 94 Tolaken [Tolakei?] 22 27.VII., [old style] [ 3 8. VIII., new style] 1894 [V.A. Kashkarov] [in Russian]”. RUSSIA: Amurskaya oblast’: 1 ♂ (ZIN): “Priamur’e Khr. Myao Chan, r. Chalda 10.8.1956 Kabakov [in Russian]”; 1 ♂ (ZIN): “Amur. obl., Skovorodinovskii r n, st. Uldruch’i, 11.6.59 Kabakov [in Russian]”; 1 PT (EJCB): “Asia Amur r. Reg. Archara 1991 Oblucie env. 28.6. 15.7., Ing. Kadlec leg.” (JENDEK, 1995 [#3767]); 1 PT (EJCB): “USSR, Siberia, Amurska oblast, Kundur env., 1.7. 10.7.1991, J. Dalihod lgt.” (JENDEK, 1995 [#3767]); 2 (EJCB): “USSR, Siberia, Amurska oblast, Kundur env., 1.7. 10.7.1991, J. Dalihod lgt.”. Buryatiya Respublika: 1 ♂ (ZIN): “Khara Daban’ [in Russian] 1.vii.1916 Rodionoff”; 1 ♀ (ZIN): “Bass. r. Vitim, sr. tech. r. B. Amalat 17.8.1968 Kabakov [in Russian]”; 1 (MDCM): “Selenginskii raion 1.VII.1978”. Irkutskaya oblast’: 1 ♀ (ZIN): “Kultuk’ [in Russian] 5.vii.1928 Rodionoff”; 2 ♂ (ZIN): “Kultuk’ [in Russian] 9.vii.1928 Rodionoff”. Khabarovskiy kray: 1 ♀ (ZIN): “Bass. Ussuri, Bikinskii r n., Birskoe, r. Olen’ 4.7.1958

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA Kabakov [in Russian]”; 1 ♀ (ZIN): “Bass. Ussuri, Bikinskii r n., Birskoe 30.6.1958 Kabakov [in Russian]”; 3 ♂, 1 ♀ (ZIN): “Bass. Ussuri, Bikinskii r n., Birskoe 1.7.1958 Kabakov [in Russian]”; 1 ♂ (ZIN): “Bass. Ussuri, Bikinskii r n., Birskoe, r. Olen’ 5.7.1958 Kabakov [in Russian]”; 1 ♀ (ZIN): “Bass. Ussuri, Bikinskii r n., Birskoe, r. Olen’ 9.7.1958 Kabakov [in Russian]”; 1 ♂ (ZIN): “R. Ussuri, Bikinskii r n, Birskoe 1.VII.1968 Kabakov [in Russian]”; 1 ♀ (ZIN): “Yu Khabarovska, Korfovskaya 2 VII 1981 Kabakov [in Rusian]”. Primorskiy kray: 1 PT (NMPC): “Ussuri” (JENDEK, 1995 [#3767]); 1 ♂ (ZIN): “Primorsk. kr. Sikhotè Alin’ 25.vi.1953 / Sicy [in Russian]”; 2 ♂, 1 ♀ (ZIN): “Primorsk. kr. Sikhotè Alin’ 18.vii.1953 / Sicy [in Russian]”; 1 PT (EJCB): “Primorje Sokoltchi 20.6.79 Koshpaksiv” (JENDEK, 1995 [#3767]); 1 PT (EJCB): “SU Ussuri reg. Jasnoe 400 m 12 19.7.1989 M. Nikodým lgt.” (JENDEK, 1995 [#3767]); 1 ♂ (JMCZ): “USSR / Sibiria Primorskij Kraj. Novocugujevka 15 20.7.1990 leg. Boukal”; 2 PT (HMMG): “USSR / Sibiria Primorskij Kraj. Novocugujevka 15 20.7.1990 leg. Boukal” (JENDEK, 1995 [#3767]); 1 PT (EJCB): “USSR / Sibiria Primorskij Kraj. Novocugujevka 15 20.7.1990 leg. Boukal” (JENDEK, 1995 [#3767]); 2 ♂, 2 ♀ (EJCB): “Russia, Primorskij kraj, Černigovka env., Sinnij chrebet Mts., 14 16.vi.1997, leg. L. Roller, L. Vidlička”. Tomskaya oblast’: 1 ♂ (ZIN): “Semiluzhnoe Tomsk. u i gub. Emel’yanov 6.iv.08 [in Rusian]”; 1 ♂ (ZIN): “Tomsk. gub. po dol. r. Beloi 20 21.vi.1910 [in Russian]”. Yevreyskaya avtonomnaya oblast’: 2 PT (EJCB): “Asia Amur r. Reg. Archara Oblucie env. 28.6. 17.7.91 Ing. Lorenc” (JENDEK, 1995 [#3767]); 3 PT (MNCA): “Asia Amur r. Reg. Archara Oblucie env. 28.6. 17.7.91 Ing. Lorenc” (JENDEK, 1995 [#3767]). Zabaykalskiy kray: 1 ♂ (ZIN): “Urul’ga Zabaik. 8.vii.907 [in Russian]”; 3 (ZIN): “Chitinskaya obl., Krasnochikoiskii r n., B. Rechka (bas. r. Chikoi), 30 VII 59g na osinovom balagane v poime reki, G. V. Lindeman [in Russian]”; 1 (ZIN): “Chitinskaya Obl. Krasnochikoiskii r n. Dol. r. N. Elovki 14 vii 1959 g., na stvole usykhayushchei osiny G.V.Lindeman leg. [in Russian]”.

Adult occurence: 4-6-7-8 (n=37). Altitude range: 400-700 m (n=2). Host plants: Salix; Populus: ALEXEEV, 1989 [#64]. Distribution: CHINA: Heilongjiang. KAZAKHSTAN. MONGOLIA. RUSSIA: Amurskaya oblast’, Buryatiya Respublika, Irkutskaya oblast’, Khabarovskiy kray, Primorskiy kray, Tomskaya oblast’, Yevreyskaya avtonomnaya oblast’, Zabaykalskiy kray. Remarks: The record “Transbaikalia” published by JENDEK, 1995 [#3767] for A. supremus Obenberger, 1917 pertains to A. voriseki. Agrilus voriseki Jendek, 1995 was consistently misidentified in ZIN by Alexeev as A. pratensis djukini Obenberger, 1935, A. roberti djukini Obenberger, 1935, A. robertii Chevrolat, 1840; and by Stepanov as A. obrucevi Obenberger, 1936.

Agrilus yakushimensis Tôyama Fig. 24L (habitus of PT); Fig. 38S (aedeagus) yakushimensis Tôyama, 1985 (Agrilus) TÔYAMA, 1985: 34 35 [#2924] (description). TÔYAMA, 1985: 21 [#3678] (iconography; Japan). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 232 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 43 [#4091] (checklist; Japan: Kyushu, Yakushima I., Shimokoshiki I.). JENDEK, 2006: 403 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2365 [#8576] (world catalog).

231

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Type material: Agrilus yakushimensis Tôyama, 1985 Type locality: Mt. Miyanoura, Yakushima Is. Type specimens: Holotype, ♂, (NSMT). Secondary: 2 PT (EJCB). Described from 36 specimens (HT, AT, PT). Paratype (Fig. 24L) from EJCB is illustrated. Diagnosis: 4-5.8 mm; see remarks in TÔYAMA, 1985: 35 [#2924]. Species group: Sulcicollis. Relevant species: A. amamioshimanus Kurosawa, 1964. Specimens examined: JAPAN: Kyushu: 1 ♂ (EJCB): “1995.5.5., Nagata, Yaku Is., Kagoshima, Kyushu, S Japan, T. Akashi leg.”.

Adult occurence: 5 (n=1). Host plants: Unknown. Distribution: JAPAN: Kyushu, Ryukyu Islands. Remarks: A. yakushimensis might be conspecific with A. amamioshimanus Kurosawa. More specimens are needed to resolve their stratus.

Type locality: Nara Pref.: Mt. Ôdaigahara-san. Type specimens: Holotype not in NSMT (S. Nomura pers. com. 2001, 2003). Described from inexplicit number of specimens (HT, AT, PT). Agrilus ohmomoi Tôyama, 1985 Type locality: Akane Rindô, Haranomachi, Fukushima Pref. Type specimens: Primary: Holotype (Fig. 50H), ♂ (NSMT). Secondary: 1 PT (EJCB). Described from 14 specimens (HT, AT, PT). Diagnosis: 4-6.2 mm; prehumerus extending almost to anterior pronotal angles with anterior end joining lateral pronotal margin. Species group: Cyaneovirens. Relevant species: A. kucerai sp. nov. Specimens examined: JAPAN: Honshu: 1 ♀ (EJCB): “[Japanese grapheme], Kanagawa Pref., 4.vii.1988, E. Akiyama leg.”; 1 ♂ (EJCB): “2006.5.31, Kamisagano, Kawazu, Shizouka, C Japan, S. Tsuyuki leg.”.

Adult occurence: 5-7 (n=3). Faunal record published by KUROSAWA, 1963 [#1617] is also included.

Agrilus yamabusi Miwa & Chûjô

Host plants: Quercus: AKIYAMA & OHMOMO, 1997 [#4091].

Fig. 26K (habitus); Fig. 39L (aedeagus); Fig. 50H (type)

Distribution: JAPAN: Hokkaido, Honshu, Kyushu, Ryukyu Islands, Tsushima.

yamabusi Miwa & Chûjô, 1940 (Agrilus) MIWA & CHÛJÔ, 1940: 55, 63 64 [#1810] (description). CHÛJÔ & KUROSAWA, 1950: 11 [#501] (synonym of pilosovittatus; catalog). KUROSAWA, 1974: 4 [#4577] (synonym of pilosovittatus). AKIYAMA, 1993: 8 [#4875] (valid species). JENDEK, 1995: 138 139 [#3768] (synonym of pilosovittatus). AKIYAMA & OHMOMO, 1997: 43 [#4091] (checklist; Japan: Honshu, Kyushu, Tsushima, Yakushima). FUKUTOMI & HORI, 2004: 6 [#6851] (checklist; Japan (Hokkaido)). JENDEK, 2006: 403 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2365 [#8576] (world catalog). nakanei Kurosawa, 1963 (Agrilus; subspecies of acastus) KUROSAWA, KUROSAWA, 1963: 106 [#1617] (description). 1963: 154 [#1618] (subspecies of acastus; characters; Japan). KUROSAWA, 1974: 3 4 [#4577] (note). TÔYAMA, 1985: 21 [#3678] (iconography; Japan). TÔYAMA, 1985: 27 28 [#2924] (type series consist of two species, from which one is new and is described below as A. euonymi). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 43 [#4091] (synonym of yamabusi). JENDEK, 2006: 403 [#7441] (synonym of yamabusi; Palaearctic catalog). BELLAMY, 2008: 2365 [#8576] (synonym of yamabusi; world catalog). JENDEK & GREBENNIKOV, 2009: 15 [#8563] (not belonging to muscarius species group). ohmomoi Tôyama, 1985 (Agrilus) TÔYAMA, 1985: 31 32 [#2924] (description). TÔYAMA, 1985: 22 [#3678] (iconography; Japan). HIRASHIMA, 1989: 323 [#8078] (checklist; Japan). AKIYAMA, 1993: 8 [#4875] (synonym of yamabusi). MORIMOTO & TADAUCHI, 1995: 231 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 43 [#4091] (synonym of yamabusi). JENDEK, 2006: 403 [#7441] (synonym of yamabusi; Palaearctic catalog). BELLAMY, 2008: 2365 [#8576] (synonym of yamabusi; world catalog).

Type material: Agrilus yamabusi Miwa & Chûjô, 1940 Type locality: Mt. Hikosan, Kyusyu, Japan. Type specimens: Not located. Described from 1 specimen. Agrilus acastus nakanei Kurosawa, 1963

Agrilus yamajii Tôyama Fig. 31L (habitus); Fig. 41X (aedeagus); Fig. 54S (type) yamajii Tôyama, 1985 (Agrilus) TÔYAMA, 1985: 26 27 [#2924] (description). TÔYAMA, 1985: 30 [#3678] (iconography; Japan). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). MORIMOTO & TADAUCHI, 1995: 232 [#8080] (checklist; Japan). AKIYAMA & OHMOMO, 1997: 43 [#4091] (checklist; Japan: Honshu, Kyushu). JENDEK, 2006: 403 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2365 2366 [#8576] (world catalog).

Type material: Agrilus yamajii Tôyama, 1985 Type locality: Mt. Takahachi, Tottori Pref. Type specimens: Primary: Holotype (Fig. 54S), sex not examined, (NSMT): “Mt. Takahachi, Tottori Pref. 10. V. 1979 O. Yamaji leg. (host out) [h] \ Holotype [p] Agrilus yamajii M. Tôyama, 1985 [h] [red label]”. Described from 5 specimens (HT, AT, PT). Diagnosis: 4.2-4.7 mm; glabrous; head strongly convex; head and pronotum with very fine transverse sculpture; aedeagus with obvious chaetotaxy on parameres. Species group: Unassigned. Relevant species: Unknown. Specimens examined: JAPAN: Honshu: 1 ♂ (EJCB): “1991.6.8., Masuzawa, Tateiwa V., S Aizu, Fukushima, NE Japan, S. Ohmomo leg.”; 1 ♀ (EJCB): “Masuzawa, 800m alt., Tateiwa mura, Fukushima pref., 1991 vii 6, T. Shindou leg.”.

Adult occurence: 6-7 (n=2). Altitude range: ~800 m (n=1). Host plants: Celtis jessoensis: TÔYAMA, 1985 [#2924]; AKIYAMA & OHMOMO, 1997 [#4091].

232 Distribution: JAPAN: Honshu, Kyushu.

Agrilus yamawakii Kurosawa Fig. 20A (habitus); Fig. 36N (aedeagus); Fig. 54T (type) yamawakii Kurosawa, 1957 (Agrilus) KUROSAWA, 1957: 192 [#1615] (description). KUROSAWA, 1963: 152 [#1618] (characters; Japan). AKIYAMA, 1975: 10 [#3177] (faunal records; Izu Islands). KUROSAWA, 1975: 3 [#4591] (note). AKIYAMA, 1980: 85, 87 [#3194] (faunal record; Japan: Kanagawa). TÔYAMA, 1985: 33 [#2924] (faunal record; Ryukyu Islands). TÔYAMA, 1985: 24 [#3678] (iconography; Japan). HIRASHIMA, 1989: 324 [#8078] (checklist; Japan). LI JINGKE, 1992: 92 [#6755] (checklist; China: Liaoning). MORIMOTO & TADAUCHI, 1995: 232 [#8080] (checklist; Japan). AKIYAMA & AKIYAMA, 1996: 187 [#3962] (faunal records; Japan: Honshu). AKIYAMA & OHMOMO, 1997: 43 [#4091] (checklist; Japan: Honshu, Shikoku, Kyushu, Izu Island (Mikura), Tsushima, Yakushima Island, Tanegashima Island, Ryukyus Islands (Amami oshima); Korea; Taiwan). NONNAIZAB et al., 1999: 113 [#8247] (checklist; China: Inner Mongolia). HUA LI ZHONG, 2002: 91 [#6696] (checklist; China: Taiwan). MÜHLE, 2003: 47 [#6155] (checklist; Taiwan). JENDEK, 2006: 403 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2366 [#8576] (world catalog).

Type material: Agrilus yamawakii Kurosawa, 1957 Type locality: Mt. Fukuchiyama, Fukuoka Pref., Kyûshû, Japan. Type specimens: Primary: Holotype (Fig. 54T), sex not examined, (NSMT): “Fukuchiyama Mt. Fukuoka Pref. Kyushu 24.VII.1951, Y. Yamawaki [h] \ Holotype [p] Agrilus yamawakii Y. Kurosawa, 1957 [h] [red label]”. Secondary: 1 PT (MNHN). Described from 15 specimens (HT, AT, PT). Diagnosis: 6.9-11.7 mm; elytral pubescence obsolete at least in proximal part; pygidium with apical margin extended into long spine. Species group: Occipitalis. Relevant species: A. auriventris Saunders, 1873. Specimens examined: JAPAN: Honshu: 1 ♀ (EJCB): “Yamanaka cho, Yokosuka shi, Kanagawa pref., 24 26.vii.1996, K. Ishida et al. leg.”. Kyushu: 1 ♀ (EJCB): “Mt. Kumada, Chikugo, 16.vii.1961”. Ryukyu isl. (Okinawa incl.): 2 (EJCB): “Tanegashima Is., 10.v.1984, Fayara allanthoides”. KOREA NORTH: 1 (EJCB): “Korea, Tesôngsan bei Pjôngiang, 23.7.1974, Josifov leg.”. KOREA SOUTH: 1 (JLCS): “Korea, Prov. Gyeonggido Namiangju, Mt. Chukryeongsan, 24.vi.2000, leg. J.C. Sohn”; 1 (JLCS): “Seodun dong, Suwon si, Gyeonggi do, Korea, vi.2002, coll. Hae chul Park”.

Adult occurence: 5-6-7 (n=9). Faunal records published by KUROSAWA, 1957 [#1615] are also included. Host plants: Fagara ailanthoides: AKIYAMA & OHMOMO, 1997 [#4091]. – F. mantchurica: AKIYAMA & OHMOMO, 1997 [#4091]. Distribution: CHINA: Liaoning, Nei Mongol, Taiwan. JAPAN: Honshu, Kyushu, Ryukyu Islands, Shikoku, Tsushima. KOREA NORTH. KOREA SOUTH.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA yunnanicola Obenberger, 1936 (Agrilus) OBENBERGER, 1936: 116 [#2205] (description). OBENBERGER, 1936: 1069 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 359 [#2429] (checklist; China). JENDEK, 2000: 507 [#4851] (lectotype designation). HUA LI ZHONG, 2002: 91 [#6696] (checklist; China: Yunnan). JENDEK, 2006: 403 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2366 [#8576] (world catalog).

Type material: Agrilus yunnanicola Obenberger, 1936 Type locality: Yunnan-sen. Type specimens: Primary: Lectotype (Fig. 54U), ♀, designated by JENDEK (2000: 507) [#4851], (NMPC): “China: Yunnan sen [h] \ typus [p] [red label] \ Agrilus Yunnanicola m. Type [h] [Obenberger’s MS] Det. Dr. Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24991 [h] [orange label]”. Described from unknown number of syntypes. Diagnosis: 4.2-8.4 mm; vertex with spiral sculpture; body more elongate; elytral apices distinctly expanded. Species group: Subrobustus. Relevant species: A. bonadonai Descarpentries & Villiers, 1963 Specimens examined: CHINA: Yunnan: 1 ♀ (EJCB): “Junnam [sic!], Yunnan Sen”; 2 ♂ (EJCB): “Yunnan 1800 2000m, 25.04N 101.55E, Yipinglang 17 20.6.1994,Vít Kubáň leg.”; 60 (EJCB): “CH Yunnan, 13 14.VI.1995, 100 km W of Kunming, Diaolin Nat. Reservation, E. Jendek & O. Šauša leg.”.

Adult occurence: 6 (n=2). Altitude range: 1800-2000 m (n=1). Host plants: Unknown. Distribution: CHINA: Yunnan.

Agrilus yunnanus Obenberger Fig. 18J (habitus); Fig. 35Z (aedeagus); Fig. 54V (type) yunnanus Obenberger, 1927 (Agrilus) OBENBERGER, 1927: 74 75 [#2062] (description). OBENBERGER, 1936: 1069 [#2191] (world catalog). PENG ZHONGLIANG, 1987: 359 [#2429] (checklist; China). HUA LI ZHONG, 2002: 91 [#6696] (checklist; China: Yunnan). JENDEK, 2006: 403 [#7441] (Palaearctic catalog). BELLAMY, 2008: 2366 [#8576] (world catalog).

Type material: Agrilus yunnanus Obenberger, 1927 Type locality: Yunnan borealis occidentalis: Djo-Kou-La, alt. 1200 m. Type specimens: Primary: Lectotype (Fig. 54V) by present designation, ♀, (NMPC): “Djo-Kou-La, alt. 1200 m Nord Ouest Yunnan [p] \ Typus [p] [red label] \ Agrilus yunnanus m. n. sp. Type [h] Det. Dr Obenberger [p]”. Secondary: 1 PLT (RMNH). Described from unknown number of syntypes. Diagnosis: 7.1-8 mm; elytra with adsutural pubescence in apical third; elytral apices spinose. Species group: Laetecyanescens. Relevant species: A. laetecyanescens Obenberger, 1940. Specimens examined: CHINA: Yunnan: 2 ♂ (MNHN): “Djo Kou La, Yunnan”.

Agrilus yunnanicola Obenberger Fig. 14J (habitus); Fig. 34N (aedeagus); Fig. 54U (type)

Host plants: Unknown.

233

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA Distribution: CHINA: Yunnan.

Agrilus zanthoxylumi Li Meng Lou Fig. 20C (habitus); Fig. 36P (aedeagus)

Type specimens: Primary: Holotype (Fig. 19H), ♂, (EJCB): “China, Yunnan 1994 Zhongdian, 16–20.vi. leg. Šiška T. Spevár”. Secondary: 8 PT (EJCB). Described from 9 specimens (HT, PT). Diagnosis: 5.8-7.9 mm; pronotal disk flat; legs longer; prosternal lobe markedly protruding and spinose, prosternal process subparallel and narrow with sharp angles; median lobe of aedeagus as in Fig. 36I. Species group: Cyanescens. Relevant species: A. cyanescens (Ratzeburg, 1837).

zanthoxylumi Li Meng Lou, 1989 (Agrilus) LI MENG LOU, 1989: 60 63 [#8117] (description; [Note: Li Meng Lou attributed the species name to Hou et Feng but we did not find their use of it and Li Menglou presented characters]). ZHANG RUNZI, 1988: 16 17 [#6695] ([Note: no scientific name is cited]; biology). LI MENG LOU et al., 1990: 34 38 [#8082] (Hou & Feng are cited as the authors; biology; damage character; spatial distribution; China: Shaanxi). ZHANG RUNG KE & WANG TONG MU, 1992: 402 403 [#6193] (Hou is cited as the author; characters; biology). JENDEK, 2006: 403 [#7441] (Zhang & Wang are cited as the authors; Palaearctic catalog). WU HAI, 2006: 236 239 [#7654] (Hou and Feng are cited as the authors; biology; control methods; China: Shandong). LÖBL & SMETANA, 2007: 29 [#8558] (authorship of species corrected to Zhang R. & Wang). BELLAMY, 2008: 2367 [#8576] (Zhang & Wang are cited as the authors; world catalog).

Specimens examined: CHINA: Yunnan: 2 ♂ PT, 4 ♀ PT (EJCB): “China, Yunnan, 1994, Zhongdian, 16 20.vi., leg. Šiška T. Spevár” (JENDEK & GREBENNIKOV, 2009 [#8561]); 2 ♂ PT (EJCB): “China NW, Yunnan, cca 3600 m, road Deqen Yanjing, 10 km W of Deqen, 1997, 28°28’N 98°53’E, 21 22.vi., M. Trýzna et O. Šafránek lgt.” (JENDEK & GREBENNIKOV, 2009 [#8561]).

Type material: Agrilus zanthoxylumi Li Meng Lou, 1989 Type locality: Shaanxi (Baoji, Weinan). Type specimens: Not located. Described from unknown number of syntypes.

Agrilus zhuang Jendek, sp. nov.

Diagnosis: 7.8-10.3 mm; frons and vertex with narrow, deep, medial impression; vertex distinctly pubescent; preapical transverse pubescent band on elytra narrow, undulated; elytral apices very finely denticulated. Species group: Occipitalis. Relevant species: A. perroti Descarpentries & Villiers, 1963. Specimens examined: CHINA: Gansu: 4 ♂, 2 ♀ (EJCB): “China Gansu, Wudu, 1 7.6.1996, E. Kučera leg.”; 1 ♂ (EJCB): “China S, Gansu, Wenxian, 28.vi.2002, Gorodinski”. Hubei: 1 ♀ (EJCB): “Hubei”. Shaanxi: 1 ♂ (EJCB): “China, Shaanxi, 1992, 120 km E Xi an, Hua Shan, 3 4.vi., Jaroslav Turna”. Yunnan: 1 ♂ (MNHN): “Chine, Yunnan, recu de Lou nan, 1931”. Zhejiang: 1 ♀ (EJCB): “Zheijiang, 1984 V 24”.

Adult occurence: 5-6 (n=4). Host plants: Zanthoxylum: LI MENG LOU, 1989 [#8117]. – Z. bungeanum: WU HAI, 2006 [#7654]. Distribution: CHINA: Gansu, Hubei, Shaanxi, Shandong, Yunnan, Zhejiang. Remarks: The concept of this taxon was judged from the picture kindly sent by Dr. Xiao-Yi Wang (Beijing) who helped in resolving the correct authorship of this species.

Agrilus zhongdian Jendek Fig. 19H (habitus of HT); Fig. 36I (aedeagus) zhongdian Jendek, 2009 (Agrilus) JENDEK in JENDEK & GREBENNIKOV, 2009: 45, 55 [#8561] (description).

Type material: Agrilus zhongdian Jendek, 2009 Type locality: China, Yunnan province, Zhongdian 25°48’N, 99°42’E.

Adult occurence: 6 (n=2). Altitude range: ~3600 m (n=1). Host plants: Unknown. Distribution: CHINA: Yunnan.

Fig. 26B (habitus of HT)

Description: 6.4-6.9 mm (Holotype 6.9 mm). BODY: Shape: fusiform, Build: robust, Color (dorsally): unicolored, Color (dominant): black or green. HEAD: Medial impression: present, Impression (extent): vertex, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: protruding from head outline, Sculpture on vertex: rugae, Sculpture (density): dense. Eyes: Size: moderate, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate or long. PRONOTUM: Shape: visually elongate or visually square, Sides: arcuate, Maximal width: at anterior margin. Anterior lobe: moderate or obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles or projecting beyond anterior pronotal angles. Posterior angles: Shape: rectangular or obtuse, Apex: sharp. Disk: Convexity: moderately convex, Disk impressions: medial and lateral, Medial impression (shape): anteromedial and posteromedial, Medial impression (deepness): deep. Prehumerus: carinal, Shape: arcuate, Extent: beyond half of pronotal length, Posterior end: joining posterior pronotal angle, Anterior end: joining pronotal marginal carina, Arc: obvious. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior third of pronotal length. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Shape: subtruncate, Truncation: transverse. Elytral pubescence: proximal only, Proximal: quadrangular, Color: unicolored, Character: homogenous. Tomentum: absent. Prosternal lobe: moderate, Anterior margin: arcuately emarginate, Emargination (deepness): moderate. Prosternal process: moderate, Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Apical margin of pygidium: arcuate, Apical margin of last

234 ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): somewhat longer than mesotarsus. Tarsomere 1 (relative size): subequal or longer than 2-4 combined. GENITALIA: Ovipositor: elongate. Diagnosis: elytral apices transversely truncate; elytra with wide stripe of perisutural pubescence in anterior half. Species group: Tempestivus. Relevant species: A. tempestivus Lewis, 1893. Type locality: China, west Jiangxi province, Jinggang Shan, Ciping. Type specimens: Primary: Holotype (Fig. 26B), ♀, (EJCB): “China, Jiangxi W, Jinggang Shan, Ciping, 2-14.VI.1994, E. Jendek & O. Šauša leg.”. Secondary: CHINA: Jiangxi: 1 ♀ PT (EJCB): “China, Jiangxi W, Jinggang Shan, Ciping, 2-14. VI.1994, E. Jendek & O. Šauša leg.”. Adult occurence: 6 (n=1). Host plants: Unknown. Distribution: CHINA: Jiangxi. Etymology: Noun in apposition; the specific name refers to the Chinese ethnic group of people Zhuang living in southern China.

Agrilus zhui Jendek, sp. nov. Fig. 9I (habitus of HT)

Description: Holotype 9.6 mm. BODY: Shape: cuneiform, Build: robust, Color (dorsally): bicolored, Color (dominant): brown. HEAD: Medial impression: present, Impression (extent): vertex, Impression (depth): deep, Epistoma: with raised upper margin. Frons: Shape: moderately convex, Outline: not protruding from head outline. Vertex: Outline: not protruding from head outline, Sculpture on vertex: rugae, Sculpture (aspect): semispherical, Sculpture (density): dense, Sculpture (intensity): rough. Eyes: Size: large, Shape: not protruding from head outline, Lower margin: below antennal socket. Antennae: Length: moderate. PRONOTUM: Sides: arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: obvious, Shape: arcuate, Position to anterior angles: at level with anterior pronotal angles. Posterior angles: Shape: rectangular, Apex: sharp. Disk: Convexity: moderately convex, Disk modifications: with obvious tomentose spots, Disk impressions: medial and lateral, Medial impression

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA (shape): anteromedial and posteromedial, Medial impression (deepness): shallow, Anteromedial and posteromedial impressions: smaller anteromedial and larger posteromedial, Lateral impressions (deepness): deep, Lateral impressions (width): narrow. Prehumerus: carinal, Shape: bisinuate, Extent: beyond half of pronotal length, Posterior end: distant from pronotal angle or margin, Anterior end: joining pronotal marginal carina, Arc: moderate. Marginal and submarginal carinae: Interspace: narrow, Convergence: moderately convergent, Junction: absent, Narrowest point: at posterior 1/5-1/4 of pronotal length, Modifications: submarginal carina posteriorly obliterate. Scutellum: Size: moderate, Disk: not impressed, Scutellar carina: present. ELYTRA: Color: unicolored, Humeral carina: absent. Apices: Arrangement: separate, Shape: arcuate or subangulate. Elytral pubescence: proximal only, Proximal: triangular, Color: unicolored, Character: homogenous. Tomentum: spots, Spots (locations): humeral, premedial or postmedial. STERNUM: Metasternal projection: flat, Sexual modification in male: without obvious sexual modifications. Prosternal lobe: moderate, Anterior margin: arcuate. Prosternal process: moderate, Shape: narrowed or subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disk: flat. ABDOMEN: Apical margin of pygidium: extended into long spine, Apical margin of last ventrite: arcuate. Sternal groove: Shape on apex of last ventrite: arcuate. LEGS: Metatarsus (size to mesotarsus): about as long as mesotarsus. Tarsomere 1 (relative size): longer than 2-3 combined but shorter than 2-4 combined. GENITALIA: Ovipositor: elongate. Diagnosis: pronotum with obvious pronotal lobe; prehumerus extending beyond half of pronotal length with anterior end joining marginal carina. Species group: Spinipennis. Relevant species: A. fleischeri Obenberger, 1925. Type locality: China, Shaanxi province, Qinling mts., 6 km east of Xunyangba, altitude 1000-1300 m. Type specimens: Holotype (Fig. 9I), ♀, (EJCB): “China, 1000-1300 m, Shaanxi, Qinling mts., Xunyangba (6 km E), 23.v.-13.vi.1998, I. H. Marshal leg.”. Adult occurence: 5-6 (n=1). Altitude range: 1000-1300 m (n=1). Host plants: Unknown. Distribution: CHINA: Shaanxi. Etymology: The specific name is the Chinese noun zhui (spine) in apposition. It refers to the pygidium of this species, which is projecting into a long spine.

TAXONOMIC INVENTORY OF AGRILUS TAXA OF EAST ASIA

235

AGRILUS TAXA OUTSIDE THE TARGET AREA Taxa cited in this part, although not known from the target region of East Asia, were examined in order to settle some taxonomic issues pertaining to the present publication. Only the resulting taxonomic-nomenclatural acts and necessary data are cited.

Agrilus albogularis albogularis Gory albogularis Gory, 1841 (Agrilus) GORY, 1841: 235 236 [#3737] (description). sieversi Abeille de Perrin, 1897 (Agrilus; variety of zigzag) transfer in syn. ABEILLE DE PERRIN, 1897: 7, 28 29 [#19] (description).

Type material: Agrilus sieversi Abeille de Perrin, 1897 Type locality: Russie méridionale. Type specimens: Primary: Lectotype by present designation, ♀, (MNHN): “Samara [h] \ A. Sieversi Gangl. an convexifrons Kiesw? [h] \ Type [p] [red ink] \ Muséum Paris 1919 Coll. A. DE PERRIN [p]”. Described from unknown number of syntypes.

Agrilus sericans Kiesenwetter sericans Kiesenwetter, 1857 (Agrilus) KIESENWETTER, 1857: 124, 127 [#3015] (description). sericarius Abeille de Perrin, 1897 (Agrilus) syn. nov. ABEILLE DE PERRIN, 1897: 7, 25 26 [#19] (description). daghestanicus Obenberger, 1930 (Agrilus) syn. nov. OBENBERGER, 1930: 109 110 [#2088] (description).

Type material: Agrilus sericans Kiesenwetter, 1857 Type locality: Caucasus. Type specimens: Primary: Lectotype, sex not examined, designated by JENDEK (2003: 186) [#5823], (NMPC): “Transcriptio [p] Cuprescens Ménétr. Chalconotus (Megerle) Caucase Typ Gory [h] \ Muséum Paris 1952 Coll. R. Oberthur [p] [yellow label]”. Described from unknown number of syntypes. Agrilus sericarius Abeille de Perrin, 1897 Type locality: Daghestan (provinces Causasiques). Type specimens: Holotype by monotypy, ♀, (MNHN): “Caucasus Daghestan E. Koenig [p] \ sericarius Ab. [h] \ Muséum Paris 1919 Coll. A. DE PERRIN [p] \ Holotype [p] [red label] \ sericarius Abeille [h] Type [p] [red ink] \ Agrilus sericans Kiesw. [h] Alexeev det. [p]”. Described from 1 specimen. Agrilus daghestanicus Obenberger, 1930 Type locality: Daghestan. Type specimens: Primary: Lectotype by present designation, ♀, (NMPC): “Daghestan [h] \ Typus [p] [red label] \ Agrilus daghestanicus m. Type [h] Det. Dr Obenberger [p]”. Described from unknown number of syntypes.

Agrilus zigzag Marseul zigzag Marseul, 1866 (Agrilus) MARSEUL, 1866: 439, 478 479 [#676] (description).

scutellatus Abeille de Perrin, 1897 (Agrilus) transfer in syn. ABEILLE DE PERRIN, 1897: 8 [#19] rossicus Obenberger, 1916 (Agrilus) (syn. by ALEXEEV, 1959 [#3229] OBENBERGER, 1916: 276 [#1983] (description). simbirensis Obenberger, 1925 (Agrilus) syn. nov. OBENBERGER, 1925: 102 103 [#2047] (description). semirjeciae Obenberger, 1936 (Agrilus) syn. nov. OBENBERGER, 1936: 105 [#2205] (description). kusakievici Obenberger, 1936 (Agrilus) syn. nov. OBENBERGER, 1936: 107 [#2205] (description).

Type material: Agrilus zigzag Marseul, 1866 Type locality: Russie méridionale. Type specimens: Primary: Lectotype by present designation, ♂, MNHN: “Agrilus [ilegible word] zigzag Rus. m. Krb 6H [h] [round label] \ zigzag Mars. Rus. mer. Ab. 66.47 [h] \ Museum Paris Coll. De Marseul 284290 [p] \ Museum Paris Coll. Générale [p] [yellow label]”. Secondary: 1 PLT (BMNH). Described from unknown number of syntypes. Agrilus scutellatus Abeille de Perrin, 1897 Type locality: not given (See Lectotype locality data). Type specimens: Primary: Lectotype by present designation, ♂, MNHN: “Saratow [p] \ Typ Ballion [h] \ Agrilus scutellatus Ball. [h] \ Type [p] [red ink] \ Scutellatus Ball. [h] \ Muséum Paris 1919 Coll. A. DE PERRIN [p]”. Described from unknown number of syntypes. Agrilus rossicus Obenberger, 1916 Type locality: Sarepta in Südrußland. Type specimens: Primary: Lectotype by present designation, ♀, NMPC: “Sarepta [h] \ Typus [p] [red label] \ Agrilus russicus [sic!] m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24948 [h] [orange label]”. Secondary: 2 ♀ PLT (NMPC), 3 PLT (DEI), 1 PLT (NMBS). Described from unknown number of syntypes. Agrilus simbirensis Obenberger, 1925 Type locality: Russia occ. - Simbirsk. Gub. Type specimens: Holotype by monotypy, ♀, NMPC: “Simbirsk-Gub. Rossia oc. Dr. Jureček 1917 [p] \ Typus [p] [red label] \ Agrilus simbirensis m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24946 [h] [orange label]”. Described from 1 specimen. Agrilus semirjeciae Obenberger, 1936 Type locality: Semirječia: Semirječensk: Lepsa. Type specimens: Primary: Lectotype by present designation, ♀, NMPC: “Семиръченск. oбл. окр. Лепсинска 10.V.1908 Силагаевъ / Typus [p] [red label] \ Agrilus semirječiae m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24324 [h] [orange label]”. Secondary: 1 PLT (NMPC). Described from unknown number of syntypes.

236 Agrilus kusakievici Obenberger, 1936 Type locality: Semirječia. Type specimens: Primary: Lectotype (Fig. 48H) by present designation, ♀, (NMPC): “Лепса, Семиръчье Кушакевичь 72 [p] \ Typus [p] [red label] \ Agrilus kušakieviči m. Type [h] Det. Dr Obenberger [p] \ Mus. Nat. Pragae Inv. [p] 24323 [h] [orange label]”. Described from unknown number of syntypes.

MISIDENTIFICATIONS, NOMINA NUDA, AND OTHER UNAVAILABLE NAMES

Misidentifications. A. angustulus (Illiger, 1803) is one of the most common West Palaearctic species distributed eastwards to the Respublika Tatarstan in Russia, i.e., not reaching the target area of East Asia. Surprisingly, 1 ♂ has been detected from Russian Far East labeled: “Russia, Primorski reg. Khasan distr. Ryazanovka 05.06.1991 Filimonov leg.”. This record needs further confirmation. NONNAIZAB, QI BAOYING & LI YABAI, 1999 [#8247] and partly also HUA LI ZHONG, 2002 [#6696] and HUANG RENXIN, 2005 [#8248] cited in their checklists the following Agrilus

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA from Inner Mongolia, Hebei or Xinjiang: A. albogularis Gory, 1841; A. biguttatus (Fabricius, 1777); A. hyperici (Creutzer, 1799); A. macroderus Abeille de Perrin, 1897; A. pseudocyaneus Kiesenwetter, 1857; A. roscidus Kiesenwetter, 1857; A. sericans Kiesenwetter, 1857. The occurence of these West Palaearctic species in China is highly unlikely and they should be excluded from the list of East Asian Agrilus. The faunal record from “Transbaikalia” cited by JENDEK, 1995 [#3767] as A. supremus was misidentified and should by assigned to A. voriseki Jendek, 1995. Unavailable names. The following unavailable names which have never been proposed as the name for taxon have been cited in the following publications: A. tsonglokianus – cited by PENG ZHONGLIANG, 1992 [#2431] and LIN GUI RUI, 2002 [#7926] in the synonymy of Agrilus auriventris Saunders, 1873; A. citri – cited as valid name of species by QUAYLE, 1938 [#7340], FANG ZHIGANG & WU HONG, 2001 [#6624], LIN GUI RUI, 2002 [#7926] and HUA LI ZHONG, 2002 [#6696]; A. marchali – cited as a valid name of species by PENG ZHONGLIANG, 2002 [#6628]; A. kamtschaticus Stepanov – cited as a valid name of species by IVLIEV & KONONOV, 1966 [#8089].

237

ANALYSIS In this chapter we attempt to take advantage of the fact that the jewel-beetle Agrilus data reported in this book originate from a highly structured and extensive relational database. Such data might be variously classified, filtered and analyzed with relative ease, as compared, for example, to the same amount of data stored as plain text. Various graphical and statistical outputs resulting from the analysis (Figs. 55A-57B) are discussed in this chapter. Additionally, we exported locality data into a mapping program and generated various spatial outputs visualizing different distributional patterns of East Asia Agrilus (Figs. 58A-68H). In most cases the referred graphs and maps are self-explanatory and easily interpreted even without references to the text.

TAXONOMIC-NOMENCLATURAL ANALYSIS From 641 Agrilus specific names relevant to the target area of East Asia, 278 names are valid and the rest (363) are invalid (259) or unavailable (104). More than one quarter of the valid names (74) are those proposed herein as new. The high rate of unavailable names is caused by numerous infrasubspecific names, unpublished names, incorrect subsequent spellings, misidentifications, names in collections, mistyping, etc. The rate of invalid names relevant to east-Asian Agrilus versus those valid is remarkably high when analyzed individually by relevant author (Fig. 55A). It should be noted that even Jan Obenberger (1892-1964), a careful and experienced entomologist, introduced 95 invalid or unavailable names compared to 48 valid. The most taxonomically ancient and widely distributed A. viridis (Linné, 1758) has, not surprisingly, the highest number of synonyms (64). Among ten Agrilus taxa with the highest number of synonyms (Fig. 55B), eight are widely distributed Eurosiberian species. The most productive period for describing new eastAsian Agrilus taxa was the 20th century with the absolute maximum between the two World Wars (Fig. 55C). Figure 55D shows the rate of new taxa descriptions in 15-year intervals for particular countries of East Asia, the most noticeable is China with the highest, most recent spike. The first described Agrilus taxon endemic to East Asia was A. confinis Faldermann, 1835, a species native to China. The Agrilus species native to East Asia and most often cited in publications is A. viridis (Linné, 1758). This species is distributed throughout the entire Palaearctic region and has 729 subsequent references pertaining to the target area. Other widely distributed Agrilus taxa follow the list, even though their bibliography has been markedly reduced by omitting numerous extraterritorial local references.

EXAMINED MATERIAL Type specimens. Type specimens (mostly primary) were examined for 459 of the 513 available type bearing names of Agrilus pertaining to East Asia. The remaining types of 55 names of nominal taxa were not found, including those of 16 nominal taxa with valid names. The following is a list of available names, where types were not located. The current status of each name is cited in the round brackets after the name, where necessary: ainu Miwa & Chûjô, 1940 (syn. of friebi Obenberger, 1922); albizziae Kurosawa, 1963 (syn. of subrobustus Saunders, 1873);

anomalus Théry, 1942 (syn. of ussuricola Obenberger, 1924); aurichalceus Redtenbacher, 1847 (syn. of cuprescens cuprescens (Ménétriés, 1832)); betuleti (Ratzeburg, 1837); bicolor (Schrank, 1798) (syn. of viridis (Linné, 1758)); bicolor Redtenbacher, 1847 (syn. of viridis (Linné, 1758)); caeruleus (Rossi, 1792) (syn. of cyanescens cyanescens (Ratzeburg, 1837)); confinis Faldermann, 1835; cyaneomicans Nonfried, 1892; cyanescens (Ratzeburg, 1837); darwinii Wollaston, 1857 (syn. of viridis (Linné, 1758)); enisus Théry, 1942 (syn. of friebi Obenberger, 1922); fagi (Ratzeburg, 1837) (syn. of viridis (Linné, 1758)); foveola Rey, 1891 (syn. of obscuricollis Kiesenwetter, 1857); igai Kurosawa, 1963 (syn. of asiaticus Kerremans, 1898); imasakai Tôyama, 1985; indulgens Kerremans, 1897 (syn. of viridis (Linné, 1758)); kanohi Kurosawa, 1954 (syn. of auropictus Kerremans, 1912); kochi Théry, 1942 (syn. of fleischeri Obenberger, 1925); kuangcenensis Obenberger, 1940; laticornis (Illiger, 1803); linearis (Fabricius, 1792) (syn. of viridis (Linné, 1758)); linearis (Schrank, 1781) (syn. of viridis (Linné, 1758)); littlei Curtis, 1840 (syn. of viridis (Linné, 1758)); lubricus Théry, 1942 (syn. of fissus Obenberger, 1917); mali Matsumura, 1924; nakanei Kurosawa, 1963 (syn. of yamabusi Miwa & Chûjô, 1940); nipponicola Kurosawa, 1963 (syn. of fleischeri Obenberger, 1925); nocivus (Ratzeburg, 1837) (syn. of viridis (Linné, 1758)); obscuripennis Fairmaire, 1888 (syn. of sinensis sinensis Thomson, 1879); obscurus Delherm de Larcenne, 1885 (syn. of obscuricollis Kiesenwetter, 1857); panphilovi Alexeev, 1959 (syn. of asiaticus Kerremans, 1898); praeclarus Krogerus, 1925 (syn. of pratensis (Ratzeburg, 1837)); pratensis (Ratzeburg, 1837); proximus Rey, 1891 (syn. of cuprescens cuprescens (Ménétriés, 1832)); pseudimitans Miwa & Chûjô, 1940 (syn. of sinensis sinensis Thomson, 1879); pseudoconstantini Alexeev, 1981; quercinus Redtenbacher, 1847 (syn. of viridis (Linné, 1758)); rosaceus (Scopoli, 1763) (syn. of viridis (Linné, 1758)); rozhkovi Alexeev, 1964 (syn. of alashanensis Obenberger, 1936); sachalinicola Obenberger, 1940; scaberrimus (Ratzeburg, 1837) (syn. of laticornis (Illiger, 1803)); serraticornis (Scopoli, 1763) (syn. of viridis (Linné, 1758)); subelegans Cobos, 1972 (syn. of ulaangomiensis Cobos, 1972); subviduus Kurosawa, 1957 (syn. of viduus Kerremans, 1914); tamanukii Stepanov, 1954 (syn. of sachalinicola Obenberger, 1940); tsushimanus Kurosawa, 1963; turdus Théry, 1942 (syn. of sibiricus sibiricus Obenberger, 1912); violaceus Nonfried, 1892; viridis (Linné, 1758); yagii Tôyama, 1985 (syn. of euonymi Tôyama, 1985); yamabusi Miwa & Chûjô, 1940; yokoyamai Iga, 1955 (syn. of sinuatus sachalinensis Obenberger, 1935); zanthoxylumi Li Meng Lou, 1989.

Among the16 nominal taxa with valid names and missing primary types, 11 taxa have their identity adequately known: A. betuleti (Ratzeburg, 1837); A. cyanescens (Ratzeburg, 1837); A. imasakai Tôyama, 1985; A. laticornis (Illiger, 1803); A. mali Matsumura, 1924; A. pratensis (Ratzeburg, 1837); A. pseudoconstantini Alexeev, 1981; A. sachalinicola

238 Obenberger, 1940; A. tsushimanus Kurosawa, 1963; A. viridis (Linné, 1758); A. yamabusi Miwa & Chûjô, 1940.

Four other taxa are species inquirendae with unknown identity due to missing type and insufficient description: A. confinis Faldermann, 1835; A. cyaneomicans Nonfried, 1892; A. kuangcenensis Obenberger, 1940; A. violaceus Nonfried, 1892).

One taxon most likely has no primary type fixed: A. zanthoxylumi Li Meng Lou, 1989.

Other material examined. While the vast majority of Agrilus records cited in this work originate from Asia (8,905 specimens and 2,521 collecting records), some of the more widely distributed species have their faunal records also cited for Europe (177 specimens and 91 records) and North America (30 specimens and 15 records). Figure 58B shows Agrilus collecting sites with known coordinates from the target area of East Asia. The number of collecting sites for each sub-region is shown in Fig. 59B. The majority of east-Asian Agrilus specimens (Fig. 55E) originate from China (2,797 specimens and 846 collecting records) and Russia (2,721 specimens and 669 records). The least amount of material (94 specimens and 70 collecting records) examined was from the Korean peninsula. Most Chinese specimens originate from Yunnan, Shaanxi, Sichuan and Shanxi (Fig. 55F), while the vast majority of Russian Agrilus are from Primorskiy kray (Fig. 56E). More than 50 faunal records are cited from both Mongolia and the island of Honshu, Japan. No specimens were available from the Chinese provinces of Guangdong, Jiangsu, Liaoning, Macau and Shandong and from the Russian northeastern administrative unit Chukotskiy avtonomnyy okrug. The majority of the examined Agrilus specimens was collected during the last 30 years (Fig. 56B), with the absolute 10-year maximum between 1990 and 1999. The altitudinal distribution of Agrilus by country is shown in Fig. 56C. For records with the altitude originally given as an interval the calculated value is counted as their mean. The lowest recorded altitude is 10 m, the highest is 4,210 m. Most altitudinal records fall into the 700-1500 m interval. All Agrilus records collected above 2,200 m originate from China. The highest altitude, 3800-4600 m, is recorded for A. reichardti Obenberger, 1935. The largest vertical distribution from 400 to 4500 m is recorded for A. ecarinatus Marseul, 1866. The following Agrilus taxa were recorded from above 3,000 m: A. adelphinus Kerremans, 1895; A. asiaticus Kerremans, 1898; A. compacticornis sp. nov.; A. dali Jendek, 2009; A. ecarinatus Marseul, 1866; A. fissus Obenberger, 1917; A. foramenifer sp. nov.; A. klapperichianus krupkai Alexeev & Bílý, 1980; A. kostali Jendek, 2007; A. laetecyanescens Obenberger, 1940; A. lugu sp. nov.; A. mali Matsumura, 1924; A. mandala sp. nov.; A. miao sp. nov.; A. morawitzi Obenberger, 1936; A. myopic Jendek, 2000; A. nivosus Abeille de Perrin, 1900; A. paralleloides sp. nov.; A. pekinensis accolus Jendek, 1995; A. prosternalis sp. nov.; A. reichardti Obenberger, 1935; A. souliei Jendek, 1994; A. zhongdian Jendek, 2009.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

HOST PLANTS Among 278 east-Asian Agrilus, 106 taxa have at least one larval host plant record, while 67 taxa have two and more host plants (Fig. 56D). The absolute maximum,14 host plants, is recorded for A. viridis (Linné, 1758), followed by A. viridiobscurus Saunders, 1873 (6 hosts), A. laticornis (Illiger, 1803) and A. obscuricollis Kiesenwetter, 1857 (each with 5 hosts; Fig. 56D). At present, 69 host plant species are known for east-Asian Agrilus (Fig. 56F). The most commonly recorded host plant genera are Quercus (host of 23 Agrilus taxa), Ulmus (12 taxa), Salix (9 taxa) and Zelkova (8 taxa).

TAXA DIVERSITY Eight Agrilus species cited in this book are not yet confirmed to occur in East Asia, although they are likely to occur there: A. adspersus (Kerremans, 1890); A. chlorophyllus Abeille de Perrin, 1904; A. laticornis (Illiger, 1803); A. macrotatopus Obenberger, 1930; A. nepalensis Tôyama, 1988; A. pecirkai Obenberger, 1916; A. priamus Kerremans, 1912; A. salicis Frivaldszky, 1877. Among 270 species-group Agrilus taxa known from East Asia, seven are subspecies (A. komareki sapporoensis Obenberger, 1935; A. pekinensis accolus Jendek, 1995; A. sibiricus fukushimenis Jendek, 1994; A. sinensis splendidicollis Fairmaire, 1889; A. sinuatus sachalinensis Obenberger, 1935; A. smaragdinus habros ssp. nov.; A. subauratus amurensis Obenberger, 1922). Two species, A. confinis Faldermann, 1835, and A. proditor Obenberger, 1917, have their distribution known only from the vaguely given type locality (“China boreali” or “China”, respectively), and, therefore, are not included in calculations of sub-regional Agrilus diversity. Within East Asia, China has the highest diversity of Agrilus taxa (229, including 4 species inquirenda), followed by Japan (72), Russia (57), North Korea (34), Mongolia (31) and South Korea (29). The distributional overview of all concerned Agrilus taxa on the sub-regional level is given in APPENDIX IV. A map depicting sub-regional Agrilus diversity is shown in Fig. 59B. Among the sub-regions, Yunnan (79), Shaanxi (73), Honshu (57), Sichuan (48) and Primorskiy kray (47) have the highest number of Agrilus, while sub-regions like Macau and Chukotskskiy okrug have no records. The sub-regional diversity of Agrilus usually correlates with the number of records and/or number of examined specimens (Fig. 60A). The relatively low Agrilus diversity of some coastal Chinese provinces such as Guangdong (5), Guangxi (10), Jiangsu (4), Liaoning (13), Shandong (9) compared to the comparably sized province of Fujian (27), or even to the more northern Russian Primorskiy kray (47) most likely reflect the acute lack of data, rather than a depopulate Agrilus fauna. Japanese islands, due to the lasting history and tradition of Japanese entomology, are most likely the best explored and the most completely known Agrilus fauna. Current data for Honshu (57 taxa) might well be approaching the real Agrilus biodiversity, even though new Agrilus taxa continue to be discovered there.

ANALYSIS

DISTRIBUTION General distribution. The distributional valuation of Agrilus native to East Asia (Figs. 57A and 57B) is based on the sum of sub-regions occupied by each species either inside or outside the target area. Subspecies data are not provided separately, but merged for respective species. Calculated east-Asian sub-regional units are those consistently used throughout this work (see “Geographical coverage”). Additional distributional records included in the calculation are those of the east-Asian species known to occur outside the target area, either naturally or through introduction. Such extra-territorial records are counted by countries (such as, for example, Laos) or by the first level administrative unit (e.g. province or equivalent) for larger countries such as Canada, India, the non east-Asian part of Russia and the United States. Figure 57A shows the 50 most widely distributed species of Agrilus native to East Asia. The first seven most widely distributed species (A. cuprescens (Ménétriés, 1832); A. viridis (Linné, 1758); A. cyanescens (Ratzeburg, 1837); A. pratensis (Ratzeburg, 1837); A. sinuatus (Olivier, 1790); A. suvorovi Obenberger, 1935) and A. betuleti (Ratzeburg, 1837) are native to the entire Eurasian region. Invasive species that were also introduced to the American continent are indicated in blue (Fig. 57A). Three of them (A. cuprescens (Ménétriés, 1832), A. cyanescens (Ratzeburg, 1837), A. sinuatus (Olivier, 1790)) score among the five most widely distributed taxa. All six east-Asian Agrilus known to be invasive outside their natural distribution are scored among the first 37 widely distributed species. Figure 57B shows the 50 most widely distributed Agrilus species native to east-Asian. The numbers of sub-regions they inhabit are grouped by continent (Asia, Europe, Africa and North America). Among these species five are known to occur on three continents: A. cuprescens (Ménétriés, 1832); A. cyanescens (Ratzeburg, 1837); A. obscuricollis Kiesenwetter, 1857; A. sinuatus (Olivier, 1790) and A. viridis (Linné, 1758). Species endemic to East Asia. Among 278 Agrilus taxa recorded for East Asia, 135 are not known to occur outside the target region and, therefore, are presently considered to be endemic. These species can be grouped by the country they inhabit: China (105 endemic taxa): A. aerosus Ganglbauer, 1889; A. aurinotus sp. nov.; A. auristernum Obenberger, 1924; A. bai sp. nov.; A. bicuspidatus sp. nov.; A. bidentulus Ganglbauer, 1889; A. bituberculatus Jendek, 2007; A. bocaki sp. nov.; A. brevinotus sp. nov.; A. businskorum sp. nov.; A. caudicula sp. nov.; A. chaetocoxalis sp. nov.; A. charismaticus Jendek, 2000; A. ciping sp. nov.; A. conspersus sp. nov.; A. cordicollis sp. nov.; A. cyaneomicans Nonfried, 1892; A. dali Jendek, 2009; A. daur sp. nov.; A. densetomentosus Obenberger, 1940; A. diao sp. nov.; A. diaolin Jendek, 2001; A. emmerichi Obenberger, 1935; A. eunuchus sp. nov.; A. foliatus Jendek, 2007; A. foramenifer sp. nov.; A. fornicatus sp. nov.; A. fusciapex sp. nov.; A. fusiformis sp. nov.; A. gaoligong Jendek, 2000; A. han sp. nov.; A. hani sp. nov.; A. holzschuhi Jendek, 1994; A. huashanus Jendek, 2001; A. hui sp. nov.; A. hunanus Jendek, 2009; A. improcerus sp. nov.; A. jindrai sp. nov.; A. kostali Jendek, 2007; A. kuangcenensis Obenberger, 1940; A. kucerai sp. nov.; A. lacrima sp. nov.; A. lahu

239 sp. nov.; A. latipalpis Jendek, 2007; A. lei sp. nov.; A. lisu sp. nov.; A. lobatus sp. nov.; A. lugu sp. nov.; A. majzlani Jendek, 2007; A. mandala sp. nov.; A. miao sp. nov.; A. mio sp. nov.; A. myopic Jendek, 2000; A. naxi sp. nov.; A. ningpoensis Obenberger, 1927; A. notoclavus Jendek, 2000; A. obscurosericeus Obenberger, 1940; A. ornatoides sp. nov.; A. paralleloides sp. nov.; A. pekinensis accolus Jendek, 1995; A. pleuralis sp. nov.; A. pleurostigmus sp. nov.; A. pliculipennis Obenberger, 1940; A. presbys sp. nov.; A. prolatus sp. nov.; A. prosternalis sp. nov.; A. pseudobscuricollis Jendek, 2007; A. pterostigma Obenberger, 1927; A. pumi sp. nov.; A. pusillesculptus Obenberger, 1940; A. qiang sp. nov.; A. qinling Jendek, 2000; A. qinlingcola sp. nov.; A. recticollis sp. nov.; A. sagittifer sp. nov.; A. schillhammeri sp. nov.; A. sentis sp. nov.; A. shan sp. nov.; A. shaowensis Jendek, 1994; A. she sp. nov.; A. sichuanus sp. nov.; A. siskai sp. nov.; A. smaragdifrons Ganglbauer, 1889; A. smaragdinus habros ssp. nov.; A. smetanai sp. nov.; A. souliei Jendek, 1994; A. spathulifer sp. nov.; A. spiculum sp. nov.; A. stigma sp. nov.; A. subtilipilis Obenberger, 1940; A. suturicuspidatus Jendek, 2007; A. tai sp. nov.; A. thibetanus Obenberger, 1928; A. truncatus Jendek, 2007; A. tuber sp. nov.; A. ulmicola sp. nov.; A. vigil sp. nov.; A. violaceus Nonfried, 1892; A. volkovitshi Jendek, 2007; A. yunnanicola Obenberger, 1936; A. yunnanus Obenberger, 1927; A. zanthoxylumi Li Meng Lou, 1989; A. zhongdian Jendek, 2009; A. zhuang sp. nov.; A. zhui sp. nov. Japan (23 endemic taxa): A. alesi Obenberger, 1935; A. esakii Kurosawa, 1964; A. fortunatus Lewis, 1893; A. hattorii Nakane, 1983; A. imasakai Tôyama, 1985; A. imitans Lewis, 1893; A. japanocarinatus Ohmomo, 2002; A. komareki sapporoensis Obenberger, 1935; A. koyoi Ohmomo, 2002; A. mallotiellus Kurosawa, 1985; A. ogatai Ohmomo, 2002; A. shashamboe Kurosawa, 1963; A. shibatai Kurosawa, 1964; A. sudai Kurosawa, 1985; A. suginoi Tôyama, 1985; A. sugiurai Jendek, 2007; A. takahashii Tôyama, 1988; A. trinotatus Saunders, 1873; A. tsutsumiuchii Ohmomo, 2006; A. uenoi Kurosawa, 1963; A. yakushimensis Tôyama, 1985; A. yamabusi Miwa & Chûjô, 1940; A. yamajii Tôyama, 1985. South Korea (1 endemic taxon): A. coreanus Obenberger, 1935. Mongolia (2 endemic taxa): A. mongolorum Obenberger, 1936; A. plavilstshikovi Alexeev, 1989. Russia (4 endemic taxa): A. arsenevi Jendek, 2009; A. diversornatus sp. nov.; A. pseudoconstantini Alexeev, 1981; A. rudicollis Alexeev, 1979.

Regionally endemic taxa. Agrilus taxa presently known from a single east-Asian sub-region (as defined above) are considered endemic to them (Figs. 60B, 61A). Many of such cases derive, most likely, from the scarcity of available data, although in other instances it may indicate sub-regions with the higher rate of endemic Agrilus species. If all sub-regions with poorly known Agrilus faunas are removed from the analysis, the number of endemic taxa increases from north to south, and, less conclusively, from west to east. The highest number of endemic Agrilus has been recorded from Yunnan (28) followed by Shaanxi (11) and Sichuan (7). When compared to the respective sub-regional faunas, a remarkably high proportion of endemic Agrilus taxa is noticeable for the south-western provinces of China and particularly for Yunnan (35.4%) and Guizhou (26.1%). Neighbouring Guangxi and Guangdong may fall in the same category, if not being negatively affected by their faunal data deficiency (Fig. 60A). APPENDIX IV provides a list of all Agrilus species separately for each east-Asian sub-region.

240 Distributional data on east-Asian Agrilus do not indicate a strong correlation between sub-regional species diversity and the rate of endemic taxa. The Japanese island of Honshu, for example, is home to a relatively high number of Agrilus (57), but only two (3.5 %) are endemic. This ratio is considerably less than that found in Sichuan with 48 Agrilus taxa, and seven (14.6%) endemics. Another way to compare the data is to note that the main Japanese island has fewer endemic Agrilus taxa than that of Primorskiy kray (4 endemic taxa, 8.5%,) located on the mainland right across the strait.

BIOGEOGRAPHIC ANALYSIS Agrilus fauna of East Asia (Fig. 57C) belongs to two biogeographic realms: Palaearctic and Indo-Malayan (Fig. 61B). These two centres of biogeographic origin are discussed below. Palaearctic fauna Within the Palaearctic realm, five distributional centers are distinguished, namely Eurosiberian, West Palaearctic, East Palaearctic Central Asian and East-Asian. The first three, Eurosiberian, West Palaearctic and East Palaearctic correspond approximately to the biomes of Boreal Forest (Taiga). Central Asian matches the biomes of Deserts and Xeric Shrublands, and Temperate Grasslands, Savannas and Shrublands (Fig. 58A). Eurosiberian distributional pattern (Figs. 62A, 62B) is characteristic to Agrilus distributed throughout Europe and Siberia (Figs. 68A, 68B) and developing primarily in pioneer plants such as Alnus, Betula, Corylus, Lonicera, Populus, Rhamnus, Rosa, Rubus, Salix and Sorbus which occur continuously across the entire northern Eurasia. In East Asia, Eurosiberian Agrilus fauna is predominantly found in the northern regions of Russia where they constitute 100% of the Agrilus fauna of Magadan and Kamchatka. An example of a Eurosiberian species is Agrilus viridis (Linné, 1758) (Fig. 68A). The highest number of Eurosiberian Agrilus species (7-8) was recorded in Respublika Buryatiya, Zabaykalskiy and Primorskiy kray (Fig. 62B). 8 taxa: A. betuleti (Ratzeburg, 1837); A. cuprescens cuprescens (Ménétriés, 1832) (Fig. 68B); A. cyanescens cyanescens (Ratzeburg, 1837); A. delphinensis Abeille de Perrin, 1897; A. pratensis (Ratzeburg, 1837); A. ribesi Schaefer, 1946; A. suvorovi Obenberger, 1935; A. viridis (Linné, 1758) (Fig. 68A).

West Palaearctic distributional pattern (Fig. 62A) is characteristic to Agrilus taxa with a distribution predominantly in the West Palaearctic region which extends eastward to the river Yenisey. Only one east-Asian taxon, A. obscuricollis Kiesenwetter, 1857, falls into this category, having its eastern limit in Respublika Tyva (Fig. 68C). Its principal larval host plant is Quercus, but can probably also develop in substitute hosts such as Acer, Betula and Carpinus, which may explain its occurrence outside the native range of Quercus. 1 taxon: A. obscuricollis Kiesenwetter, 1857 (Fig. 68C).

East Palaearctic distributional pattern (Figs. 62A, 63A) is characteristic to Agrilus taxa having a predominantly East Palaearctic region distribution, with its western border

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA along the river Yenisey. In East Asia, East Palaearctic taxa are distributed predominantly in the southern Russian subregions, Mongolia and neighboring Chinese provinces. Among the sub-regions, only Primorskiy kray has all four taxa from this group, although their faunal proportion on subregional fauna is rather low (Fig. 63A). Known host plants include Artemisia, Corylus, Nitraria, Populus, and Salix. 4 taxa: A. ecarinatus Marseul, 1866; A. kaluganus Obenberger, 1940; A. nicolanus Obenberger, 1924; A. voriseki Jendek, 1995 (Fig. 68D)

Central Asian distributional pattern (Figs. 62A, 63B) is characteristic to Agrilus distributed predominantly in Central Asia which is defined as the area bordered by the Caspian Sea on the west, China on the east, Afghanistan to the south, and Russia to the north (Fig. 68E). In East Asia, Agrilus species with Central Asian distributional pattern are present predominantly in the western part of the region and compose 25% of the faunas of Xinjiang and Respublika Tyva. The highest number of Central Asian Agrilus species is seen in Mongolia (7), followed by Respublika Tyva (5) (Fig. 63B). The northern limit for the Central Asian taxa lies in Zabaykalskiy kray, the eastern one is in Nei Mongol. The larval hosts of Central Asian Agrilus are plants growing in arid and semiarid conditions such as Artemisia, Caragana and Ceratoides and pioneer trees such as Populus and Salix. 7 taxa: A. ganglbaueri Semenov, 1891 (Fig. 68E); A. morawitzi Obenberger, 1936; A. nivosus Abeille de Perrin, 1900; A. pseudolimoniastri Cobos, 1968; A. pseudosericans Obenberger, 1936; A. tschitscherini Semenov, 1895; A. unguiculosus Obenberger, 1936;

East-Asian distributional pattern (Fig. 64A) includes species occurring exclusively in the target area of East Asia, or those with the predominant distribution in East Asia but occasionally extend to the neighboring regions of the IndoMalayan realm (Figs. 68F, 68G). Most of Agrilus with eastAsian distributional pattern (41-61) are found in Yunnan, Sichuan, Shaanxi, Honshu and Kyushu (Fig. 64A). Their highest proportion in sub-regional faunas is achieved in the central-east Chinese provinces, South Korea and Japan (Fig. 64A). 200 taxa: A. adelphinus Kerremans, 1895 (Fig. 68G); A. aerosus Ganglbauer, 1889; A. alashanensis Obenberger, 1936; A. alesi Obenberger, 1935; A. alutaceicollis Obenberger, 1930; A. andrusi Baudon, 1968; A. arsenevi Jendek, 2009; A. asahinai Kurosawa, 1956; A. asiaticus Kerremans, 1898 (Fig. 68F); A. aurinotus sp. nov.; A. auristernum Obenberger, 1924; A. aurithorax sp. nov.; A. bai sp. nov.; A. bicuspidatus sp. nov.; A. bidentulus Ganglbauer, 1889; A. bituberculatus Jendek, 2007; A. bocaki sp. nov.; A. brevinotus sp. nov.; A. businskorum sp. nov.; A. caudicula sp. nov.; A. chaetocoxalis sp. nov.; A. charismaticus Jendek, 2000; A. chekiangensis Gebhardt, 1929; A. chujoi Kurosawa, 1985; A. ciping sp. nov.; A. compacticornis sp. nov.; A. confinis Faldermann, 1835; A. conspersus sp. nov.; A. cordicollis sp. nov.; A. coreanus Obenberger, 1935; A. cupes Lewis, 1893; A. cyaneomicans Nonfried, 1892; A. cyaneoniger Saunders, 1873; A. daimio Obenberger, 1936; A. dali Jendek, 2009; A. daur sp. nov.; A. densetomentosus Obenberger, 1940; A. diao sp. nov.; A. diaolin Jendek, 2001; A. discalis Saunders, 1873; A. diversornatus sp. nov.; A. dureli sp. nov.; A. emmerichi Obenberger, 1935; A. esakii Kurosawa, 1964; A. eunuchus sp. nov.; A. euonymi Tôyama, 1985; A. fareastensis Jendek, 1995;

241

ANALYSIS A. fissus Obenberger, 1917; A. fleischeri Obenberger, 1925; A. foliatus Jendek, 2007; A. foramenifer sp. nov.; A. fornicatus sp. nov.; A. fortunatus Lewis, 1893; A. friebi Obenberger, 1922; A. fusciapex sp. nov.; A. fusiformis sp. nov.; A. gaoligong Jendek, 2000; A. gussakovskiji Alexeev, 1981; A. han sp. nov.; A. hani sp. nov.; A. hattorii Nakane, 1983; A. holzschuhi Jendek, 1994; A. huashanus Jendek, 2001; A. hui sp. nov.; A. hunanus Jendek, 2009; A. imasakai Tôyama, 1985; A. imitans Lewis, 1893; A. improcerus sp. nov.; A. insulicola Kerremans, 1912; A. japanocarinatus Ohmomo, 2002; A. jindrai sp. nov.; A. kawarai Kurosawa, 1963; A. komareki komareki Obenberger, 1926; A. komareki sapporoensis Obenberger, 1935; A. kostali Jendek, 2007; A. koyoi Ohmomo, 2002; A. kuangcenensis Obenberger, 1940; A. kucerai sp. nov.; A. kurumi Kurosawa, 1957; A. lacrima sp. nov.; A. lahu sp. nov.; A. latipalpis Jendek, 2007; A. lei sp. nov.; A. lisu sp. nov.; A. lobatus sp. nov.; A. lugu sp. nov.; A. majzlani Jendek, 2007; A. mali Matsumura, 1924; A. mallotiellus Kurosawa, 1985; A. manchu sp. nov.; A. mandala sp. nov.; A. marginicollis Saunders, 1873; A. miao sp. nov.; A. mio sp. nov.; A. miwai Obenberger, 1936; A. moerens Saunders, 1873; A. mongolorum Obenberger, 1936; A. myopic Jendek, 2000; A. naxi sp. nov.; A. nelsonodaimio Ohmomo, 2006; A. ningpoensis Obenberger, 1927; A. nipponigena Obenberger, 1935; A. notoclavus Jendek, 2000; A. obscurosericeus Obenberger, 1940; A. ogatai Ohmomo, 2002; A. ornatoides sp. nov.; A. paralleloides sp. nov.; A. pekinensis accolus Jendek, 1995; A. pekinensis pekinensis Obenberger, 1924; A. peregrinus Kiesenwetter, 1879; A. pilosovittatus Saunders, 1873; A. plagiatus Ganglbauer, 1889; A. planipennis Fairmaire, 1888; A. plavilstshikovi Alexeev, 1989; A. pleuralis sp. nov.; A. pleurostigmus sp. nov.; A. pliculipennis Obenberger, 1940; A. presbys sp. nov.; A. proditor Obenberger, 1917; A. prolatus sp. nov.; A. prosternalis sp. nov.; A. przewalskii Obenberger, 1936; A. pseudobscuricollis Jendek, 2007; A. pseudoconstantini Alexeev, 1981; A. pterostigma Obenberger, 1927; A. pumi sp. nov.; A. pusillesculptus Obenberger, 1940; A. qiang sp. nov.; A. qinling Jendek, 2000; A. qinlingcola sp. nov.; A. quadrisignatus Marseul, 1866; A. recticollis sp. nov.; A. ribbei Kiesenwetter, 1879; A. rudicollis Alexeev, 1979; A. sachalinicola Obenberger, 1940; A. sagittifer sp. nov.; A. satoi Kurosawa, 1954; A. schillhammeri sp. nov.; A. sentis sp. nov.; A. shan sp. nov.; A. shaowensis Jendek, 1994; A. shashamboe Kurosawa, 1963; A. she sp. nov.; A. shibatai Kurosawa, 1964; A. sibiricus fukushimenis Jendek, 1994; A. sibiricus sibiricus Obenberger, 1912; A. sichuanus sp. nov.; A. sinensis sinensis Thomson, 1879; A. sinuatus sachalinensis Obenberger, 1935; A. siskai sp. nov.; A. smaragdifrons Ganglbauer, 1889; A. smaragdinus habros ssp. nov.; A. smaragdinus smaragdinus Solsky, 1876; A. smetanai sp. nov.; A. sospes Lewis, 1893; A. soudeki Obenberger, 1926; A. souliei Jendek, 1994; A. spathulifer sp. nov.; A. spiculum sp. nov.; A. spinipennis Lewis, 1893; A. stigma sp. nov.; A. subauratus amurensis Obenberger, 1922; A. subrobustus Saunders, 1873; A. subtilipilis Obenberger, 1940; A. sudai Kurosawa, 1985; A. suginoi Tôyama, 1985; A. sugiurai Jendek, 2007; A. suturicuspidatus Jendek, 2007; A. tai sp. nov.; A. takahashii Tôyama, 1988; A. tazoei Kurosawa, 1985; A. tempestivus Lewis, 1893; A. thibetanus Obenberger, 1928; A. tokyoensis Kurosawa, 1985; A. trinotatus Saunders, 1873; A. truncatus Jendek, 2007; A. tsushimanus Kurosawa, 1963; A. tsutsumiuchii Ohmomo, 2006; A. tuber sp. nov.; A. uenoi Kurosawa, 1963; A. ulaangomiensis Cobos, 1972; A. ulmicola sp. nov.; A. ussuricola Obenberger, 1924; A. varius Kerremans, 1895; A. ventricosus Fairmaire, 1888; A. viduus Kerremans, 1914; A. vigil sp. nov.; A. violaceus Nonfried, 1892; A. viridiobscurus Saunders, 1873; A. volkovitshi Jendek, 2007; A. yakushimensis Tôyama, 1985; A. yamabusi Miwa & Chûjô,

1940; A. yamajii Tôyama, 1985; A. yamawakii Kurosawa, 1957; A. yunnanicola Obenberger, 1936; A. yunnanus Obenberger, 1927; A. zanthoxylumi Li Meng Lou, 1989; A. zhongdian Jendek, 2009; A. zhuang sp. nov.; A. zhui sp. nov. The following 31 Agrilus taxa with east Asian distributional pattern are known to occur in the northernmost parts of the Indo Malayan region: A. adelphinus Kerremans, 1895; A. alesi Obenberger, 1935; A. andrusi Baudon, 1968; A. asiaticus Kerremans, 1898; A. aurithorax sp. nov.; A. cyaneoniger Saunders, 1873; A. daimio Obenberger, 1936; A. discalis Saunders, 1873; A. dureli sp. nov.; A. esakii Kurosawa, 1964; A. insulicola Kerremans, 1912; A. mallotiellus Kurosawa, 1985; A. manchu sp. nov.; A. miwai Obenberger, 1936; A. moerens Saunders, 1873; A. nelsonodaimio Ohmomo, 2006; A. plagiatus Ganglbauer, 1889; A. planipennis Fairmaire, 1888; A. satoi Kurosawa, 1954; A. shibatai Kurosawa, 1964; A. sinensis sinensis Thomson, 1879; A. spinipennis Lewis, 1893; A. subrobustus Saunders, 1873; A. suginoi Tôyama, 1985; A. tempestivus Lewis, 1893; A. tokyoensis Kurosawa, 1985; A. uenoi Kurosawa, 1963; A. viduus Kerremans, 1914; A. yakushimensis Tôyama, 1985; A. yamabusi Miwa & Chûjô, 1940; A. yamawakii Kurosawa, 1957.

Indo-Malayan fauna Indo-Malayan aspect (Figs. 61B, 64B) relates to the biome of Tropical and Subtropical Moist Broadleaf Forests (Fig. 58A). It comprises Agrilus occurring predominantly in the Indo-Malayan realm but extending into the neighboring regions of the target area (Fig. 68H). Most Agrilus of IndoMalayan origin occur in Yunnan (34) and other nearby provinces of southeast China. The highest proportion of IndoMalayan fauna on the regional fauna is present in southeast China (Fig. 64B). North Korea is the northernmost limit for Indo-Malayan species. 50 Taxa: A. agnatus Kerremans, 1892; A. ashlocki Baudon, 1968; A. aureofasciatus sp. nov.; A. aurigaster sp. nov.; A. auriventris Saunders, 1873; A. auropictus Kerremans, 1912; A. baroni Gory & Laporte, 1839; A. barrati Descarpentries & Villiers, 1963; A. blatteicollis Bourgoin, 1922; A. bonadonai Descarpentries & Villiers, 1963; A. bonnottei Bourgoin, 1925; A. caligans Bourgoin, 1925; A. convictor Descarpentries & Villiers, 1963; A. coomani Bourgoin, 1925; A. cyaneovirens Bourgoin, 1922; A. cyanipennis Gory & Laporte, 1839; A. decoloratus Kerremans, 1892; A. dianthus Kerremans, 1892 (Fig. 68H); A. erythrostictus Bourgoin, 1922; A. fevreae Descarpentries & Villiers, 1963; A. grisator Kerremans, 1893; A. hmong sp. nov.; A. inamoenus Kerremans, 1892; A. ineptus Kerremans, 1892; A. jaminae Baudon, 1968; A. laetecyanescens Obenberger, 1940; A. liscapia Jendek, 2003; A. loosdregti Baudon, 1965; A. lubopetri Jendek, 2000; A. maculifer Saunders, 1873; A. madeci Baudon, 1968; A. magnus sp. nov.; A. malloti Théry, 1930; A. muongoides sp. nov.; A. muscarius Kerremans, 1895; A. niveoguttatus Kerremans, 1892; A. odetteae Baudon, 1968; A. olivaceidorsis Obenberger, 1917; A. perroti Descarpentries & Villiers, 1963; A. plasoni plasoni Obenberger, 1917; A. quentini Descarpentries & Villiers, 1963; A. reichardti Obenberger, 1935; A. rivalieri Descarpentries & Villiers, 1963; A. rougeoti Descarpentries & Villiers, 1963; A. sikkimensis Obenberger, 1928; A. sinensis splendidicollis Fairmaire, 1889; A. sulcinotus Jendek, 2007; A. terracotta sp. nov.; A. tonkineus Kerremans, 1895; A. tyndarus Kerremans, 1913.

242 Agrilus taxa occurring in East Asia and neighboring regions along southern border Agrilus taxa occurring in the neighboring southern regions come from two distributional centers. The first includes Agrilus taxa with a predominant distribution in East Asia but extending into the Indo-Malayan region. The second includes taxa with a predominant distribution in the Indo-Malayan region but with limited overlap in East Asia. In Fig. 65A, the taxa from both categories are combined to illustrate their regional affinities and proportions in East Asia. The highest numbers of Agrilus taxa from both categories are present in Yunnan and islands of Japan. Yunnan has the higher proportion of Indo-Malayan, while the Japanese islands have dominance of East-Asian Agrilus taxa.

SIMILARITY OF SUB-REGIONAL FAUNAS Four East Asian sub-regions (Mongolia, Primorskyi kray, Japan and Yunnan) were selected as standard reference points for comparison with other sub-regional Agrilus faunas. The results of these comparisons are graphically depicted on four maps (Figs. 65B, 66A, 66B, 67A). Each map illustrates the degree of the regional similarity for every east-Asian subregion to the one chosen as a reference point. Among the noticeable results is the fact that the Mongolian Agrilus fauna (Fig. 65B) constitutes more than 50% of local diversity for all bordering sub-regions, as well as all sub-regions north of Mongolia. Faunal similarity between Primorskyi kray and Mongolia is rather remarkable, with the former sub-region having 18 Agrilus species (38.29% of its fauna) also recorded for Mongolia, which also represent 58.1% of all Mongolian Agrilus taxa. All sub-regions neighboring Primorskyi kray and all subregions northward of it have more than 75% of their Agrilus species also represented in Primorskyi kray (Fig. 66A). The nearby Japanese islands, however, share less than 50% of their Agrilus with Primorskyi kray. Remarkable is the high faunal affinity between Primorskyi kray and the relatively remote Chinese provinces of Sichuan and Shaanxi with 29.8% and 53.2% of all their Agrilus identical with those occurring in Primorskyi kray. Except for Yunnan, each sub-region of East Asia with more than 10 known Agrilus species has at least one quarter of its regional Agrilus diversity shared with Japan. The fauna of Japan (Fig. 66B) has the strongest affinity to the geographically closest mainland regions and also, rather surprisingly, to the relatively remote Chinese provinces of Shaanxi (31.5%) and Sichuan (21.9%). Only one widely distributed species, Agrilus cyaneoniger Saunders, 1873, originally described from Yunnan, reaches its northernmost limit in Amurskaya oblast’ of Russia. This example is rather exceptional, since more than 25% of all Yunnan Agrilus are shared only by its geographically most closest southern Chinese sub-regions (Fig. 67A), while among them, the neighboring Sichuan has the highest number of identical species.

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

CONCLUSIONS Estimated diversity of Asian Agrilus. The current revision reports 278 Agrilus taxa from the east-Asian region. How comprehensive is our present knowledge and what might be the real numbers of Agrilus species diversity? Figure 67B is an attempt to estimate the east-Asian Agrilus diversity separately for various countries. The pie charts illustrate the rate between known species and the estimated total diversity by a country. This assessment is based on the reasonably accurate information on Agrilus species known to inhabit these countries. The “estimated” part of each circle is a preliminary assessment derived primarily from a large number of undescribed Agrilus species for each respective country available in the collection of the first author, further enhanced by considering the country’s climate, diversity of regional flora and fauna, as well as the overall degree of entomological exploration (Fig. 59A). The Russian Far East and Japan with their relatively poor Agrilus diversity and long-standing entomological tradition are the two best explored regions of East Asia. Nonetheless, new Agrilus species and new regional faunal records are still regularly reported from both regions. The real diversity of the main Japanese islands, particularly owing to the subtropical climate of Kyushu, may well reach 100 species. The recent entomological exploration of China, conducted mostly by amateur entomologists, presents the very first insight to the real richness of this large and biologically diverse country. Local Agrilus diversity remains only partly, or not at all, explored for many Chinese sub-regions (Fig. 59A), including southern-most provinces like Hainan adjacent with the subtropical regions of Indochina, which may well double the total of 229 currently known Chinese Agrilus species. Our assessment of the Agrilus diversity for the Oriental region, i.e. immediately south of the area targeted in this work, is the most transparent when based on the data assembled for Laos, a country best explored in terms of Agrilus. Presently 160 Agrilus species have been known to occur in Laos, while 400 more new species recently collected in that country are pending descriptions. These new species were collected by a handful of European amateur entomologists during a number of somewhat sporadic expeditions. Considering these some 560 species already known from Laos, most of them unknown, a prediction of 700 Laotian Agrilus species can hardly be an overestimation. Similarly, the rich Agrilus diversity may also be expected for other tropical countries of the Oriental region. Malaysia and Indonesia, each having a warmer climate and a remarkably rich and endemic flora and fauna, are expected to be even more diverse with about one thousand species existing in these countries. Subgeneric subdivision. As already mentioned, the current Agrilus subgeneric classification is artificial, highly confusing and in acute need of comprehensive re-evaluation. It has been predominantly based on a local concept of Agrilus diversity, mainly referring to Palaearctic species, and therefore hardly serves its purpose. In our work we entirely avoid employing any subgeneric names and use instead informal species-groups. We consider the introduction of additional subgeneric names as highly undesirable at present and would urge others to join us in such approach. The adequate natural

ANALYSIS subgeneric Agrilus classification should reflect the historic pattern of species diversification and follow a well-supported phylogenetic hypothesis. Phylogenetic attempts within Agrilus, however, seem presently hardly possible, mainly due to the fact that many already described species need taxonomic re-examination, let alone the need to describe numerous others. Once the vast majority of Agrilus species become known and unambiguously linked with their correct names, then it will be time to start detecting monophyletic species-groups and linking them into a phylogenetically meaningful network. Species groups. This revision provides the first consistently implemented attempt to group Agrilus species in informal species groups. The main goal is to simplify species identification, and, therefore, these species groups are not necessarily monophyletic. Proposed species grouping, even when performed for Agrilus fauna of a relatively large territory of East Asia, still faces the same issue as any regionally-based classification, i.e. lack of relevant data. To group east-Asian species better we need to consider the Oriental Agrilus species, which goes well beyond our present scope. We hope, however, that at least some of the presently

243 defined species-groups might be later shown to represent monophyletic species clusters, while much more effort will go eventually on direction. Single polymorphic species versus a complex of closely related species. Some Agrilus species treated herein are highly morphologically and biologically polymorphic and widely distributed taxa. In many cases, species variability correlates with spatial distribution, time of adult occurrence and host plants, which might suggest that a multitude of closely related species be lumped together under a single name. Examples of such highly polymorphic species might be found among the Albogularis and Viridis species-groups. One particular species, A. viridis, appears to consist of the most intricate, diversified and taxonomically unsettled cluster of most closely related forms gradually interchanging potential morpho-ecospecies of uncertain rank. In such cases it is almost impossible to arrive to an adequate taxonomic species concept by using morphological characters alone, particularly with often limited comparative material. Employment of molecular techniques supplemented by fine biological data seems to be the most adequate approach in such cases.

244

LITERATURE

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

CITED

ABAI, M. & ADELI, E., 1984: List of pests of forest trees & shrubs of Iran. Tehran, Ministry of agriculture and rural development, Agricultural and Natural Resources research organisation, 147 pp, map. [#5808] ABEILLE DE PERRIN, E., 1869: Nouveaux coléoptères Français. Annales de la Société Entomologique de France, (4) 9: 79 90. [#4] ABEILLE DE PERRIN, E., 1869: Petites nouvelles. Agrilus sulcaticeps. Petites Nouvelles Entomologiques, 1 (11): 42. [#4493] ABEILLE DE PERRIN, E., 1895: Notes coléoptérologiques. Annales de la Société Entomologique de France, Bulletin, 64: CCLXXIV CCLXXV. [#3905] ABEILLE DE PERRIN, E., 1895: Buprestides paléarctiques réputés nouveaux. Annales de la Société Entomologique de France, Bulletin, 64: CXVI CXXVI. [#15] ABEILLE DE PERRIN, E., 1897: Notes sur les Buprestides paléarctiques (Suite). Revue d’Entomologie, 16: 1 33. [#19] ABEILLE DE PERRIN, E., 1900: Diagnoses de Coléoptères présumés nouveaux. Bulletin de l’Académie de Marseille, 1900: 3 23. [#21] ABEILLE DE PERRIN, E., 1904: Buprestides. Boletín de la Real Sociedad Española de Historia Natural, 4: 206 224. [#22] ABIVARDI, C., 2001: Iranian entomology. An introduction. Applied entomology. Volume 2. Berlin, Heidelberg, New York, Springer, p. 445 1033. [#8093] ADELI, E., 1972: Beitrag zur Kenntnis der im Forst schädlichen Insekten des Iran. 1. Coleoptera. Zeitschrift für angewandte Entomologie, 70 (1): 8 14. [#4871] ADELI, E. & SOLEIMANI, P., 1976: Insects on oriental beech (Fagus orientalis ssp. macrophylla) in Iran and their importance for forestry practices and wood utilization. Zeitschrift für angewandte Entomologie, 80: 132 138. [#8300] AHMAD, S., HAQUE, A., FARUQUZZAMAN, A. K. M., HUSSAIN, M. & HOSSAIN, M. A., 2001: Field evaluation of kenaf cultivars and their hybrids for their reactions to spiral borer. Online Journal of Biological Sciences, 1 (12): 1158 1160. [#7043] AIBASOV, C. A., 1974: Zhuki vrediteli turangi na yugo vostoke Kazakhstana. In: Fauna, sistematika i biologiya nasekomykh Kazakhastana. Trudy instituta zoologii Kazakhskoi SSR, 35: 143 154. (in Russian) [#7212] AKIYAMA, K., 1974: [New record of Agrilus sachalinicola Obenberger, from Shizuoka prefecture central Japan (Coleoptera, Buprestidae)]. Coleopterists’ News, Tokyo, 23 24: 8. (in Japanese) [#3176] AKIYAMA, K., 1975: [The buprestids fauna of Izu Islands (Coleoptera, Buprestidae)]. Coleopterists’ News, Tokyo, 25/26: 10. (in Japanese) [#3177] AKIYAMA, K., 1975: [The buprestids fauna of the Ryûkyû Archipelago (Coleoptera, Buprestidae)]. Coleopterists’ News, Tokyo, 29/30: 4. (in Japanese) [#3178] AKIYAMA, K., 1980: [Four new records of Buprestids beetles from Kanagawa Prefecture, with notes on the genus Nipponobuprestis and its allied genera (Coleoptera, Buprestidae)]. Natural History Report of Kanagawa, 1: 85 87. (in Japanese) [#3194] AKIYAMA, K., 1986: [New records of buprestid beetles from Kanagawa prefecture (Coleoptera, Buprestidae)]. Kanagawa Chûhô, 79: 24 25. (in Japanese) [#3199] AKIYAMA, K., 1993: [Records of five buprestid beetles from Japan]. Gekkan Mushi, 274: 8. (in Japanese) [#4875]

AKIYAMA, Y. & AKIYAMA, K., 1996: The Buprestid beetles (Coleoptera: Buprestidae) from Hiroshima Prefecture, southwestern Japan. Miscellaneous Reports of the Hiwa Museum for Natural History, 34: 181 192. (in Japanese with English subtitle and summary) [#3962] AKIYAMA, K. & OHMOMO, S., 1997: A check list of the Japanese Buprestidae. Gekkan Mushi, (Supplement 1), 67 pp. [#4091] AKIYAMA, K. & OHMOMO, S., 2000: The buprestid beetles of the world. Mushi Sha’s iconographic series of insects 4. Fujita, H. (ed.), Tokyo, Shizawa printing Co. Ltd. 341 pp. [#5370] AKIYAMA, K. & OHMOMO, S., 2000: Distributional notes of the buprestid beetles (Buprestidae, Coleoptera) from Hiroshima Prefecture, Southwestern Japan (3). Miscellaneous Reports of the Hiwa Museum for Natural History, 39 (3): 87 90. (in Japanese with English subtitle and summary) [#6240] ALEXEEV, A. V., 1957: Zlatki Stalingradskoi oblasti (Coleoptera, Buprestidae). Uchenye zapiski Orekhovo Zuevskogo pedagogicheskogo instituta, 10: 115 158. (in Russian) [#3226] ALEXEEV, A. V., 1958: Zlatki Orekhovo Zuevskogo raiona Moskovskoi oblasti (Coleoptera, Buprestidae). Uchenye zapiski Orekhovo Zuevskogo pedagogicheskogo instituta, 11: 183 191. (in Russian) [#3982] ALEXEEV, A. V., 1959: Novye dannye po sinonimike zlatok roda Agrilus Curtis SSSR i Kitaiskoi narodnoi respubliki (Coleoptera, Buprestidae). Sbornik rabot po ekologii i sistematike zhivotnykh, Orekhovo Zuevskii Pedagogicheskii Institut, Moskva, 1: 26 27. (in Russian) [#3230] ALEXEEV, A. V., 1959: Zlatki novye dlya fauny SSSR (Coleoptera, Buprestidae). Sbornik rabot po ekologii i sistematike zhivotnykh, Orekhovo Zuevskii Pedagogicheskii Institut, Moskva, 1: 28 30. (in Russian) [#3231] ALEXEEV, A. V., 1959: Opredelitel’ zlatok roda Agrilus Curtis Evropeiskoi chasti SSSR (Coleoptera, Buprestidae). Sbornik rabot po ekologii i sistematike zhivotnykh, Orekhovo Zuevskii Pedagogicheskii Institut, Moskva, 1: 3 25. (in Russian) [#3229] ALEXEEV, A. V., 1960: K morfologii i sistematike lichinok nekotorykh vidov zlatok roda Agrilus Curt. Evropeiskoi chasti SSSR (Coleoptera, Buprestidae). Zoologicheskii Zhurnal, (1959) 39 (10): 1497 1510. (in Russian with English summary) [#45] ALEXEEV, A. V., 1961: Opredelitel’ zlatok roda Agrilus Curtis Evropeiskoi chasti SSSR. II. Opredelitel’ lichinok. Sbornik rabot po ekologii i sistematike zhivotnykh, Orekhovo Zuevskii Pedagogicheskii Institut, Moskva, p. 3 10. (in Russian) [#3243] ALEXEEV, A. V., 1964: Novye dannye po morfologii i sistematike lichinok uzkotelykh zlatok roda Agrilus Curtis Evropeiskoi chasti SSSR. Sbornik rabot po ekologii i sistematike zhivotnykh, Orekhovo Zuevskii Pedagogicheskii Institut, Moskva, 3: 18 27. (in Russian) [#3847] ALEXEEV, A. V., 1964: Novye vidy zlatok (Coleoptera, Buprestidae) s territorii SSSR. Sbornik rabot po ekologii i sistematike zhivotnykh, Orekhovo Zuevskii Pedagogicheskii Institut, Moskva, 3: 5 17. (in Russian) [#3846] ALEXEEV, A. V., 1968: Pereopisanie nekotorykh vidov zlatok (Coleoptera, Buprestidae) fauny SSSR. Èntomologicheskoe Obozrenie, 47 (4): 878 884. (in Russian with English summary) [#50]

LITERATURE CITED ALEXEEV, A. V., 1969: O pishchevykh formakh uzkoteloi zlatki Agrilus viridis (Coleoptera, Buprestidae). Zoologicheskii Zhurnal, 48 (1): 85 92. (in Russian with English summary) [#51] ALEXEEV, A. V., 1971: Proiskhozhdenie i vremya formirovaniya fauny zlatok roda Agrilus Curtis Evropeiskoi chasti SSSR. Trudy XIII mezdhunarodnogo èntomologicheskogo kongressa, Moskva, 2 9 avgusta 1968, Leningrad, Nauka, 1: 225 226. (in Russian) [#73] ALEXEEV, A. V., 1975: K faune zhukov zlatok (Coleoptera, Buprestidae) Mongolskoi narodnoi respubliki. Nasekomye Mongolii, Leningrad, Nauka, 3: 132 164. (in Russian with English subtitle) [#52] ALEXEEV, A. V., 1979: Novye, ranee neizvestnye s territorii SSSR i maloizvestnye vidy zhukov zlatok (Coleoptera, Buprestidae) vostochnoi Sibiri i Dal’nego Vostoka, p. 123 139. (in Russian) In: KRIVOLUTSKAYA, G. O. (ed), Zhuki Dalnego Vostoka i vostochnoi Sibirii (novye dannye po faune i sistematike), Vladivostok, 157 pp. [#57] ALEXEEV, A. V., 1981: Dva novykh vida zhukov zlatok Agrilus Curt. (Coleoptera, Buprestidae) iz Primorskogo kraya, p. 3 9. (in Russian) In: KULIKOVA, L. S., AREFIN, V. S. & SIMAKOVA, T. P. (eds), Fauna i èkologiya nasekomykh Primorskogo kraya i Kamchatki. Vladivostok, 149 pp. [#62] ALEXEEV, A. V., 1981: Dva novykh vida zlatok roda Agrilus Curtis (Coleoptera, Buprestidae) s territorii SSSR. Sbornik trudov zoologicheskovo muzeya MGU, 19: 53 58. (in Russian) [#59] ALEXEEV, A. V., 1981: Opredelitel’ lichinok zlatok roda Agrilus Curtis (Coleoptera, Buprestidae) Evropeiskoi chasti SSSR. Sbornik trudov zoologicheskovo muzeya MGU, 19: 65 84. (in Russian) [#61] ALEXEEV, A. V., 1989: Buprestidae Zlatki, p. 463 489. (in Russian) In: LER, P. A. (ed), Opredelitel’ nasekomykh Dal’nego Vostoka SSSR, Tom 3, Zhestkokrylye, ili zhuki, Chast’ 1, Leningrad, Nauka, 572 pp. [#64] ALEXEEV, A. V., 1990: Nekotorye aspekty skhodstva faun zlatok roda Agrilus Curtis (Coleoptera, Buprestidae) Palearktiki i Nearktiki, p. 8 9. (in Russian) In: MEDVEDEV, G. S. & KOROTYAEV, B. A. (eds), Uspekhi èntomologii v SSSR: Zhestkokrylye nasekomye. Materialy X s’ezda Vsesoyuznogo èntomologicheskogo obshchestva, 11 15 sentyabrya 1989, Leningrad. [#65] ALEXEEV, A. V., 1998: K podrodovoi klassifikacii zlatok roda Agrilus Curtis (Coleoptera, Buprestidae) fauny palearktiki. Èntomologicheskoe Obozrenie, LXXVII: 367 383. (in Russian with English summary and subtitle) [#4322] ALEXEEV, A. V. & BÍLÝ, S., 1980: Novye zlatki roda Agrilus Curtis (Coleoptera, Buprestidae) iz Evrazii. Èntomologicheskoe Obozrenie, 59 (3): 600 606. (in Russian with English summary) [#44] ALEXEEV, A. V. & KULINITSH, P. N., 1962: Novyi vid uzkoteloi zlatki povrezhdayushchei fistashku v Tadzhikistane i drugikh respublikakh Srednei Azii (Coleoptera, Buprestidae). Izvestiya otdeleniya biologicheskikh nauk AN Tadzhitskoi SSR, Dushanbe, 2 (9): 92 95. (in Russian) [#3235] ALEXEEV, A. V. & SLEPKO, N. G., 1998: Khronicheskie ochagi zelenoi uzkoteloi zlatki v dolinakh rek severnogo Kavkaza. Chronic Agrilus viridis loci in river valleys of the north Caucasus. Lesovedenie, 4: 52 57, figs. (in Russian with English summary) [#4887] ALEXEEV, A. V. & VOLKOVITSH, M. G., 1989: Obzor zhukov zlatok (Coleoptera, Buprestidae) Mongol’skoi narodnoi respubliki, Nasekomye Mongolii, Leningrad, Nauka, Vypusk 10: 301 368. (in Russian) [#3241]

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250 FRIVALDSZKY, J. VON, 1877: Adatok Temes és Krassó megyék faunájához. Data ad faunam Hungariae meridionalis comitatum Temes et Krassó. p. 285 378. (in Hungarian) In: SZABÓ, J. (ed), Mathematikai és Természettudományi Közlemények. Vonatkozólag a hazai viszonyokra. Kiadja a Magyar Tudományos Akadémia Mathematikai és Természettudományi Állandó Bizottsága, Budapest, XIII. Kötet, 1875/6: 378 pp. [#3933] FROST, C. A., 1924: Agrilus viridis in Massachusetts. Bulletin of the Brooklyn Entomological Society, 19: 27. [#1030] FROST, S. W., 1922: Occurence of Agrilus coeruleus Rossi in America. Canadian Entomologist, 54: 96. [#4495] FRYDRYCHOVÁ, R. & MAREC, F., 2002: Repeated losses of TTAGG telomere repeats in evolution of beetles (Coleoptera). Genetica, 115: 179 187. [#8240] FUKUTOMI, K. & HORI, S., 2004: The buprestid beetles of Hokkaido in color (Coleoptera, Buprestidae). Jezoensis, Sapporo, 30: 1 25. (in Japanese) [#6851] GANGLBAUER, L., 1889: Insecta, a Cl. G. N. Potanin in China et in Mongolia novissime lecta. VII. Buprestidae, Oedemeridae, Cerambycidae. Trudy Russkago Èntomologicheskogo obshchestva, 24: 21 85. [#3427] GARLICK, W. G., 1940: Notes on the rose stem girdler, Agrilus communis rubicola Abeille. Canadian Entomologist, 72: 21 23 [Abstract in: Revue of applied Entomology, 1940, A, 28: 543]. [#8683] GATES, M., 2004: Notes on Henryana magnifica Yoshimoto and Oodera sp. Chalcid Forum, 26: 3. [#7136] GATES, M. & GIBSON, G., 2004: Host association of Mymarommatidae. Chalcid Forum, 26: 12. [#7132] GEBHARDT, A. VON, 1929: Neue paläarktische Buprestiden (Coleoptera) aus China. Coleopterologisches Centralblatt, (1928) 3 (1 2): 20 34. [#1051] GEBHARDT, A. VON, 1929: Neue paläarktische Buprestiden (Col.) aus China. Coleopterologisches Centralblatt, (1928) 3 (3 4): 94 104. [#3349] GEBLER, F. A. VON, 1829: III. Bemerkungen über die Insekten Sibiriens, vorzüglich des Altai. p. 1 228. In: LEDEBOUR, C. F. von (ed), Reise durch das Altai Gebirge und die soongorische Kirgisen Steppe. Zweiter Theil, Berlin, G. Reimer, 427 pp. [#3008] GEBLER, F. A. VON, 1847: Verzeichniss der im Kolywano Woskresenskischem Hüttenbezirke Süd West Sibiriens. Beobachten Kaefer mit Bemerkungen und Beschreibungen. Bulletin de la Société Impériale des Naturalistes de Moscou, 20 (2): 391 512. [#4423] GEISER, R., 1984: Rote liste der Käfer (Coleoptera). p. 75 114. In: BLAB, J., NOWAK, E., TRAUTMANN, W. & SUKOPP, H. (eds), Rote Liste der gefährdeten Tiere und Pflanzen in der Bundesrepublik Deutschland. 4. Auflage, Grewen, Kilda. [#6841] GEMMINGER, M. & HAROLD, E. VON, 1869: Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. V, Buprestidae, Trixagidae, Monommidae, Eucnemidae, Elateridae, Cebrionidae. Monachii, Sumptu E. H. Gummi, p. 1347 1608. [#1057] GENTRY, J. W., 1965: Crop insects of Northeast Africa Southwest Asia. United States Department of Agriculture, Agricultural Handbook No. 273: 1 210. [#8590] GEORGIEV, G., LJUBOMIROV, T., RAIKOVA, M., IVANOV, K. & SAKALIAN, V. P., 2004: Insect inhabitants of old larval galleries of Saperda populnea (L.) (Coleoptera: Cerambycidae) in Bulgaria. Journal of Pest Science, 77: 235 243. [#6736] GERINI, F., 1953: Note sui Buprestidi italiani. 1. Bolletino della Società entomologica italiana, 82 (9 10): 85 92. [#1060]

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA GHAHARI, H., BELLAMY, C. L., SAKENIN, H. & PETTERSON, P., 2008: A contribution to new records of Iranian Buprestidae (Coleoptera). Munis Entomology & Zoology, 3 (2): 636 642. [#8160] GIBSON, G. A. P., 2005: The world species of Balcha Walker (Hymenoptera: Chalcidoidea: Eupelmidae), parasitoids of wood boring beetles. Zootaxa, 1033: 1 62. [#7138] GMELIN, J. F., 1790: Caroli a Linné systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima tertia, aucta, reformata. Tom. I. Pars IV. Classis V. Insecta, Lipsiae, Georg Emanuel Beer, p. 1517 2224. [#1087] GOBBI, G., 1973: Su alcuni Buprestidi del Lazio (Coleoptera Buprestidae). Bolletino dell’Associazione Romana di Entomologia, 28 (3 4): 51 54. [#1093] GOBBI, G., 1983: Interessanti reperti di Buprestidi italiani e diagnosi di Anthaxia liae n. sp. (Coleoptera, Buprestidae). Bolletino dell’Associazione Romana di Entomologia, (1981) 36: 33 41. [#1098] GOBBI, G., 2002: Su alcuni interessanti coleotteri xilofagi dell’Appennino centrale (Coleoptera, Buprestidae, Cerambycidae). Bolletino dell’Associazione Romana di Entomologia, 57 (1 4): 27 31. [#6898] GORY, H. L., 1841: Histoire naturelle et iconographie des insectes coléoptères, Supplément aux buprestides. Tome IV, Livraisons 43 52, Paris, P. Duménil. [#3737] GORY, H. L. & LAPORTE DE CASTELNAU, F. L., 1839: Histoire naturelle et iconographie des insectes coléoptères, publiée par monographies séparées. Suite aux buprestides. Tome II, Paris, P. Duménil, genera paged separately. [#3738] GOTTWALD, S. & HORNBURG, M., 2004: Ein Wiederfund von Buprestis haemorrhoidalis Herbst, 1780 sowie weitere bemerkenswerte Prachtkäferfunde in Brandeburg und Berlin (Coleoptera: Buprestidae). Märkische Entomologische Nachrichten, Potsdam, 6 (2): 47 54. [#7007] GOULD, J., RODA, A. & REARDON, D., 2004: Biocontrol of the emerald ash borer, p. 35. In: MASTRO, V., & REARDON, R. (eds), Emerald ash borer research and technology development meeting. Port Huron, Michigan, September 30 October 1, 2003. Forest health technology enterprise team. Abstracts. West Virginia, Morgantown, VI + 44 pp. [#6793] GOUREAU, C. C., 1867: Les insectes nuisibles aux forêts et aux arbres d’avenues. Bulletin de la Société des sciences historiques et naturelles de l’Yonne, Extrait, 1er trimestre: 375 pp. [#7946] GRAY, J. E., 1848: Nomenclature of coleopterous insects in the collection of the British museum. Part III. Buprestidae. London, E. Newman, 52 pp. [#5388] GRECHKIN, V. P., 1951: Zelenaya uzkotelaya zlatka i zabolevanie topolei. In: Ocherki po biologii vreditelei lesa. Materialy k poznaniyu fauny i flory SSSR, Moskva, Novaya seriya, Otdel zoologicheskii, 31: 51 67. (in Russian) [#8606] GRECHKIN, V. P. & VORONCOV, A. I., 1962: Vrediteli i bolezni topolei i mery bor’by s nimi. Moskva, Goslesbumizdat, 150 pp. (in Russian) [#8242] GROOT, P. DE, BIGGS, W. D., LYONS, D. B., SCARR, T., CZERWINSKI, E., EVANS, H. J., INGRAM, W. & MARCHANT, K., 2006: A visual guide to detecting emerald ash borer damage. Ontario, Ontario ministry of natural resources, 16 pp. [#8068] GURYEVA, E. L., 1974: Buprestidae Zlatki, p. 96 112. In: KRYZHANOVSKII, O. L. (ed), Nasekomye i kleshchi vrediteli sel’skokhozyaistvennykh kul’tur. Tom II, zhestkokrylye. Leningrad, Nauka, 334 pp. (in Russian) [#5428] GUSSAKOVKII, V. V., 1949: Otryad Coleoptera. Zhestkokrylye ili zhuki. p. 57 109, 205 219, 281 297, 330 342. (in Russian) In: PAVLOVSKII, E. N. (ed), Vrednye zhivotnye Srednei Azii. Izdatel’stvo AN SSSR, Moskva Leningrad, 404 pp. [#7805]

LITERATURE CITED GUTOWSKI, J. M., 1993: Agrilus bialowiezaensis sp. n. from Poland (Coleoptera: Buprestidae). Genus, Wroclaw, 4 (4): 295 302. [#3638] GUTOWSKI, J. M. & MILKOWSKI, M., 2008: Bogatkowate (Coleoptera: Buprestidae) Puszczy Kozienickiej. Buprestidae (Coleoptera) of the Koziencka forest. Parki narodowe i Rezerwaty Przyrody, 27 (2): 49 85. (in Pollish with English subtitle and abstract) [#8244] HAACK, R. A., 2006: Exotic bark and wood boring Coleoptera in the United States: recent establishments and interceptions. Canadian journal of forest research, 36: 269 288. [#8288] HAACK, R. A., JENDEK, E., LIU HOUPING., MARCHANT, K. R., PETRICE, T. R., POLAND, T. M. & YE, H., 2002: The emerald ash borer: a new exotic pest in North America. Newsletter of the Michigan Entomological Society, 47 (3 4): 1 5. [#5632] HAACK, R. A. & PETRICE, T. R., 2004: Emerald ash borer adult dispersal, p. 10. In: MASTRO, V., & REARDON, R. (eds), Emerald ash borer research and technology development meeting. Port Huron, Michigan, September 30 October 1, 2003. Forest health technology enterprise team. Abstracts. West Virginia, Morgantown, VI + 44 pp. [#6774] HAACK, R. A. & PETRICE, T. R., 2004: Emerald ash borer attraction to trap logs of different lengths and species, p. 14. In: MASTRO, V., & REARDON, R. (eds), Emerald ash borer research and technology development meeting. Port Huron, Michigan, September 30 October 1, 2003. Forest health technology enterprise team. Abstracts. West Virginia, Morgantown, VI + 44 pp. [#6778] HAACK, R. A. & PETRICE, T. R., 2004: Emerald ash borer survival in firewood, p. 23. In: MASTRO, V., & REARDON, R. (eds), Emerald ash borer research and technology development meeting. Port Huron, Michigan, September 30 October 1, 2003. Forest health technology enterprise team. Abstracts. West Virginia, Morgantown, VI + 44 pp. [#6785] HAACK, R. A. & PETRICE, T. R., 2004: Evaluation of perma guard d 20 and imidacloprid to control emerald ash borer, p. 26. In: MASTRO, V., & REARDON, R. (eds), Emerald ash borer research and technology development meeting. Port Huron, Michigan, September 30 October 1, 2003. Forest health technology enterprise team. Abstracts. West Virginia, Morgantown, VI + 44 pp. [#6787] HAACK, R. A., PETRICE, T. R., MILLER, D. L., BAUER, L. S. & SCHIFF, N. M., 2004: Host range of emerald ash borer, p. 38. In: MASTRO, V., & REARDON, R. (eds), Emerald ash borer research and technology development meeting. Port Huron, Michigan, September 30 October 1, 2003. Forest health technology enterprise team. Abstracts. West Virginia, Morgantown, VI + 44 pp. [#6795] HANSEN, J., MOULTON, J. K. & KLINGEMAN, W. E., 2010: Range expansion and adult flight activity of Agrilus subrobustus (Coleoptera: Buprestidae) in Tennessee. Florida Entomologist, 93: 444 445. [#9057] HARIRRI, G., 1971: A list of recorded insect fauna of Syria. Part 2, Syria, University of Aleppo, Faculty of agriculture, 306 pp. [#8105] HAROLD, E. VON, 1869: Abänderungen vergebener Namen. Coleopterogische Hefte, 5: 122 125. [#3943] HARRIS, T. W., 1829: Contributions to entomology, No. vii. New England Farmer, and Horticulture Journal, 8 (1): 2 3. [#1166] HASTIR, P. & GASPAR, C., 2002: Les «richards» (Coleoptera, Buprestidae) de la faune de Belgique: éthologie, phénologie, classification et systematique. Notes faunistiques de Gembloux, 47: 3 39. [#6211]

251 HATTORI, T., 2010: Records on some Buprestidae from Island of Akuseki, the Tokara archipelago. Coleopterists’ News, Tokyo, 169: 4. [#8899] HATTORI, T., 2010: Records of some Buprestidae from island of Kuchinoshima, the Tokara archipelago. Coleopterists’ News, Tokyo, 170: 21. [#8921] HEERING, H., 1956: Zur Biologie, Ökologie und zum Massenwechsel des Buchenprachtkäfers (Agrilus viridis L.). I. Teil. Zeitschrift für angewandte Entomologie, 38 (3): 249 287. [#3716] HEERING, H., 1956: Zur Biologie,Ökologie und zum Massenwechsel des Buchenpracht Käfers (Agrilus viridis L.). II. Teil. Zeitschrift für angewandte Entomologie, 39 (1): 76 114. [#1195] HELLRIGL, K. G., 1978: Ökologie und Brutpflanzen europäischer Prachtkäfer (Col., Buprestidae). Teil 2. Zeitschrift für angewandte Entomologie, 85 (3): 253 275. [#3442] HELLWIG, D. I. C., 1795: Fauna Etrusca. Sistens Insecta quae in provinciis Florentina et Pisana praesertim collegit Petrus Rossius in regio Pisano athenaeo publ. prof. et soc. Ital. Tomus primus. Mantissae priore parte adiecta, iterum edita et annotatis perpetuis aucta. Helmstadii, Litteris C. G. Fleckeisen, XXVIII + 475 pp. [#3508] HERBST, J. F. W., 1801: Natursystem aller bekannten in und ausländischen Insekten, als eine Fortsetzung der Büffonschen Naturgeschichte. Der Käfer. Neunter Theil. Berlin, Buchhandlung des Geh. Commerzien Raths Bauti, 344 pp. [#1216] HERMS, D. A., STONE, A. K. & CHATFIELD, J. A., 2003: Emerald ash borer: The beginning of the end of ash in North America? p. 2 10. In: BENNETT, P., BLOETSCHER, B., BOGGS, J., CHATFIELD, J., DRAPER, E., DYKE, D., GAO, G., GOERIG, D., MALINICH, T., MARTIN. J., SHERRATT, A. S. & YOUNG, C. (eds), Special emerald ash borer issue. Buckeye yard and garden line (BYGL) 2003. The Ohio State University, Extension. [#6801] HERMS, D., SMITLEY, D. R., MCCULLOUGH, D. G. & BONELLO, P., 2004: Evaluation of resistance of Asian and North American ashes to emerald ash borer, p. 41. In: MASTRO, V., & REARDON, R. (eds), Emerald ash borer research and technology development meeting. Port Huron, Michigan, September 30 October 1, 2003. Forest health technology enterprise team. Abstracts. West Virginia, Morgantown, VI + 44 pp. [#6797] HESPENHEIDE, H. A., 1968: An immigrant Agrilus (Coleoptera: Buprestidae). Entomological News, 79 (3): 76 80. [#1233] HESPENHEIDE, H. A., 1974: Nomenclatural notes on the Agrilinae (Coleoptera, Buprestidae): II Agrilus. Entomological News, 85 (3): 48 53. [#1221] HEYDEN, L. F. J. D. VON, 1863: Beitrag zur Coleopterenfauna des Ober Engadins, insbesondere der Umgegend von St. Moriz. Jahresbericht der Naturforschenden Gesellschaft Graubünden, (N. F.), 8: 1 52. [#8478] HEYDEN, L. F. J. D. VON, 1881: Catalog der Coleopteren von Sibirien, mit Einschluss derjenigen der Turanischen Länder, Turkestans und der chinesischen Grenzgebiete. Mit specieller Angabe der einzelnen Fundorte in Siberien und genauer Citirung der darauf bezüglichen einzelnen Arbeiten nach eigenem Vergleich, sowie mit besonderer Rücksicht auf die geographische Verbreitung der einzelnen Arten über die Grenzländer, namentlich Europa und Deutschland. Berlin. A. W. Schade’s Buchdruckerei (L. Schade), 1880 1881, XXIV + 224 pp. [#5548] HEYDEN, L. F. J. D. VON, 1884: Beitrag zur Coleopterenfauna der Insel Askold und anderer Theile des Amurgebietes. Deutsche Entomologische Zeitschrift, 28: 273 300. [#8482]

252 HEYDEN, L. F. J. D. VON, 1893: Catalog der Coleopteren von Sibirien, mit Einschluss derjenigen des östlichen Caspi Gebietes, von Turcmenien, Turkestan, Nord Thibet und des Amur Gebietes. Mit specieller Angabe der einzelnen Fundorte und genauer Citirung der darauf bezüglichen Literatur. Nachtrag I. Berlin, A. W. Schade Buchdruckerei (L. Schade), 217 pp. [#3450] HEYDEN, L. F. J. D. VON, 1896: Catalog der Coleopteren von Sibirien, mit Einschluss derjenigen des östlichen Caspi Gebietes, von Turcmenien, Turkestan, Nord Thibet und des Amur Gebietes. Mit specieller Angabe der einzelnen Fundorte und genauer Citirung der darauf bezüglichen Literatur. Nachtrag II. Berlin, A. W. Schade’s Buchdruckerei (L. Schade), 84 pp. [#5552] HEYDEN, L. F. J. D. VON, 1897: Die deutschen und österreichischen Arten der Buprestiden Gattung Agrilus (nach Abeille de Perrin’s Tabelle). Deutsche Entomologische Zeitschrift, 1897: 193 196. [#5286] HEYDEN, L. F. J. D. VON, 1898: Catalog der Coleopteren von Sibirien, mit Einschluss derjenigen des östlichen Caspi Gebietes, von Turcmenien, Turkestan, Nord Thibet und des Amur Gebietes. Mit specieller Angabe der einzelnen Fundorte und genauer Citirung der darauf bezüglichen Literatur. Nachtrag III. Berlin, A. W. Schade’s Buchdruckerei (L. Schade), 1 24 pp. [#5553] HIRASHIMA, Y., 1989: Buprestidae, p. 320 327. In: MORIMOTO, K. & TADAUCHI, O. (eds), A check list of Japanese insects. I, Fukoka, Entomological laboratory, Kyushu University Express, XI + 540 pp. [#8078] HOCHHUTH, I. H., 1873: Enumeration der in den russischen Gouvernements Kiew und Volhynien bisher aufgefundenen Käfer. Bulletin de la Société Impériale des Naturalistes de Moscou, 46: 124 164. [#3453] HOLYNSKI, R. B., 2003: New subgenus and five new species of Agrilinae Cast. (Coleoptera: Buprestidae) from the Indo Pacific and Australian regions. Annals of the Upper Silesian Museum, 12: 15 27. [#7129] HORION, A., 1955: Faunistik der mitteleuropäischen Käfer. Sternoxia (Buprestidae), Fossipedes, Macrodactylia, Brachymera. Entomologische Arbeiten aus dem Museum G. Frey, Band 4: XXI + 280 pp, 7 pl. [#1285] HOU TAOQIAN, 1981: Coleoptera: Buprestidae, 395 396. (in Chinese with English subtitle and summary) In: Zhongguo Kexueyuan Qingzang Gaoyuan Zonghe Kexue Kaocha Dui. Xizang kunchong. [Insects Of Xizang (Tibet). The Series of the Comprehensive Scientific Expedition to the Qinghai Xizang Plateau]. Volume 1, Peking, Science Press, I XI, 600 pp. [#4198] HOU TAOQIAN, 1987: Buprestidae, p. 583 599. In: HUANG FUSHENG, HOU TAOQIAN, YIN HUIFEN, GUO ZHILI & WANG YONGXIAN (eds), Forest insects of Yunnan. Yunnan science and technology press. 1622 pp, 16 color plates. [#3490] HOU TAOQIAN, 1988: Coleoptera: Buprestidae, 279 280. (in Chinese with English summary) In: HUANG FUSHENG, WANG, P., YIN, W., YU, P., LEE, T., YANG, C. & WANG, X. (eds), Insects of Mt. Namjagbarwa region of Xizang. Series of mountaineering and scientific expedition. The mountaineering and scientific expedition, Academica Sinica. Beijing, Science Press, XII + 621 pp. [#4900] HOU TAOQIAN & ZENG RUI, 2002: Coleoptera: Buprestidae, Trictenotomidae, p. 356 359. (in Chinese with English subtitle and summary) In: HUANG FUSHENG (ed), Forest insects of Hainan. Beijing, Ke xue chu ban she, 1064 pp + 31 pls. [#6626] HUA LI ZHONG, 2002: List of Chinese Insects. Vol. II. Guangzhou, Zhongshan (Sun Yat sen) University Press, 612 pp. [#6696] HUANG RENXIN, 2005: The fauna of the desert insects of Xinjiang and its formation and evolution. Xinjiang science and technology publishing house, 232 pp + col. pls. [#8248]

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA HUANGFU WEI GUO, WEI SHU JUN, ZHENG HONG HAI, LIU PENG CHENG, HUANG WEI, SHI ZU HUA & CHEN XUE XIN, 2007: Ovarian development of Agrilus auriventris Saunders (Coleoptera: Buprestidae). Acta Entomologica Sinica, 50 (7): 682 688. (in Chinese with English subtitle) [#8169] HUSSON, R., STAUDER, F. & DAUM, J., 1952: Sur la présence d’Agrilus Viridis L. (Coleoptera, Buprestidae) dans les forêts de la Sarre. Annales univ. Saraviensis Wissenschaft. Jg, 1952: 250 255. [#7085] ICZN (International Commission on Zoological Nomenclature), 1999: International Code of Zoological Nomenclature. Fourth Edition, adopted by the International Union of Biological Sciences. London, International Trust for Zoological Nomenclature, XXIX + 306 pp. [#4793] IGA, M., 1947: Fauna of buprestid beetles from Mt. Tsurugi, (1955 m.), Awa, Shikoku, with description of a new variety. Transactions of the Kinki Coleopterological Society, 2 (2): 9 18. [#3273] IGA, M., 1955: Buprestidae, p. 73 82. (in Japanese with English subtitle) In: NAKANE, T. (ed), Coloured illustrations of the insects of Japan. Coleoptera. Enlarged and revised edition. Osaka, Hoikusha, 274 pp. [#3828] IGA, M., 1962: Buprestidae, p. 73 82. (in Japanese with English subtitle) In: NAKANE, T. (ed), Coloured illustration of the insects of Japan (Coleoptera). Osaka, Hoikusha, 274 pp. [#3351] ILINSKIJ, A. I., 1948: Opredelitel’ yaicekladok lichinok i kukolok nasekomykh, vrednykh v lesnom khozyaistve. Moskva Leningrad, Gosudarstvennoe lesotechnicheskoe izdatel’stvo, 336 pp. (in Russian) [#5670] ISRIGAYA, T., 1963: [On the occurence and control of citrus flat headed borer [Agrilus auriventris] in Wakayama Prefecture]. Plant Protection, 17 (9): 351 355. (in Japanese) [#7016] IVLIEV, L. A. & KONONOV, D. G., 1966: Zlatki (Buprestidae) Magadanskoi oblasti i Kamchatki. p. 97 111. In: SOBOLEVA, R. G. & IVLIEV, L. A. (eds), 1966: Èntomofauna lesov Kuril’skikh ostrovov, poluostrova Kamchatki, Magadanskoi oblasti, Moskva Leningrad, Nauka, 138 + 2 pp.(in Russian) [# 8089] JACQUELIN DU VAL, P. N. C., 1857: Manuel entomologique. Genera des coléoptères d’Europe comprenant leur classification en familles naturelles, la description des tous les genres, des tableaux dichotomiques destinés à faciliter l’étude, le catalogue de toutes les espèces de nombreux dessins au trait de caractères. CCLXXVI pp + 15 pls. [#8203] JACQUELIN DU VAL, P. N. C., 1860: Catalogue de la famille des buprestides. p. 139 143. In: JACQUELIN DU VAL, P. N. C. & FAIRMAIRE, L., Manuel entomologique. Genera des coléoptères d’Europe comprenant leur classification en familles naturelles, la description de tous les genres, des tableaux synoptiques destinés à faciliter l’étude, le catalogue de toutes les espèces, de nombreux dessins au trait de caractères et plus de treize cents types représentant un ou plusieurs insectes de chaque genre dessinés et peints d’après nature avec le plus grand soin par M. Jules Migneaux. Tome troiséme (1859 1863), Paris, A. Deyrolle, 463 pp + p. 126 200 [Catalogue] + 100 col. pls [Issued in parts, Catalogue appeared separately]. [#8086] JAKOBSON, G. G., 1913: Zhuki Rossii i zapadnoi Evropy. Rukovodstvo k’’ opredeleniyu zhukov. Vypusk’’ X yi. A. F. Devrien’’, S. Peterburg’’, (2) + p. 721 864, pls 76 83 [Buprestidae: 770 800]. (in Russian) [#4386] JAKOBSON, G. G., 1927: Opredelitel’ zhukov. Izuchenie nasekomykh SSSR. Moskva Leningrad, Gosudarstvennoe izdatel’stvo, 522 pp. (in Russian) [#8655]

LITERATURE CITED JAKOBSON, G. G. & OGLOBLIN, D. A., 1931: Opredelitel’ zhukov. Izdanie 2 e., Moskva Leningrad, Gosudarstvennoe izdatel’stvo sel’skokhozyastvennoi i kolkhozno kooperativnoi literatury, 472 pp. [#8314] JANICKI, T. P., 1998: Buprestidae (Coleoptera) of Roztocze. Fragmenta Faunistica, Warszawa, 41 (4): 29 32, figs. [#4786] JENDEK, E., 1993: Lectotype designations in the Buprestidae collection of the Zoological Institute Petersburg. Entomological Problems, Bratislava, 24 (2): 38. [#1354] JENDEK, E., 1994: New Agrilus Dahl, 1823 species from Asia (Coleoptera: Buprestidae). Entomological Problems, Bratislava, 25 (1): 27 36. [#1356] JENDEK, E., 1994: Studies in the East Palaearctic species of the genus Agrilus Dahl, 1823 (Coleoptera: Buprestidae). Part I. Entomological Problems, Bratislava, 25 (1): 9 24. [#1355] JENDEK, E., 1995: Studies in the East Palaearctic species of the genus Agrilus (Coleoptera: Buprestidae). Part II. Entomological Problems, Bratislava, 26 (2): 137 150. [#3768] JENDEK, E., 1995: Taxonomical notes on the Agrilus betuleti species group with description of two new species (Coleoptera: Buprestidae). Koleopterologische Rundschau, 65: 171 178. [#3767] JENDEK, E., 1998: Lectotype designations in the Palaearctic and Oriental Agrilus species (Coleoptera: Buprestidae) of the Oberthür’s collection in the Muséum national d’Histoire naturelle, Paris. Acta Societatis Zoologicae Bohemicae, 62: 315 333. [#4481] JENDEK, E., 2000: Studies in the Palaearctic and Oriental Agrilus (Coleoptera, Buprestidae). I. Biológia, Bratislava, 55 (5): 501 508. [#4851] JENDEK, E., 2000: Eight new species of the genus Agrilus from Asia (Coleoptera: Buprestidae). Entomological Problems, Bratislava, 31 (2): 149 154. [#4853] JENDEK, E., 2000: Revision of the Agrilus cyaneoniger species group (Coleoptera: Buprestidae). Entomological Problems, Bratislava, 31 (2): 187 193. [#4852] JENDEK, E., 2001: Studies in the Palaearctic and Oriental Agrilus (Coleoptera, Buprestidae). II. Biológia, Bratislava, 56 (2): 171 174. [#5035] JENDEK, E., 2001: Revision of the Agrilus plasoni species group (Coleoptera: Buprestidae). Zootaxa, 13: 1 11. [#5211] JENDEK, E., 2002: Nomenclatural and taxonomic notes on Agrilus cyanescens (Ratzeburg, 1837), A. pratensis (Ratzeburg, 1837) and A. convexicollis Redtenbacher, 1849 (Coleoptera: Buprestidae: Agrilinae). Zootaxa, 77: 1 11. [#5709] JENDEK, E., 2002: Nomenclatural and taxonomic notes on Agrilus ater (Linné), A. biguttatus (Fabricius) and A. subauratus Gebler (Coleoptera: Buprestidae: Agrilinae). Zootaxa, 120: 1 12. [#5744] JENDEK, E., 2002: Taxonomic and nomenclatural notes on Agrilus salicis Frivaldszky (Coleoptera: Buprestidae: Agrilinae). Zootaxa, 24: 1 6. [#5348] JENDEK, E., 2002: Taxonomic and nomenclatural notes on Agrilus suvorovi Obenberger (Coleoptera: Buprestidae: Agrilinae). Zootaxa, 52: 1 8. [#5392] JENDEK, E., 2003: Studies in the Palaearctic and Oriental Agrilus (Coleoptera, Buprestidae). III. Biológia, Bratislava, 58 (2): 179 190. [#5823] JENDEK, E., 2003: Revision of Agrilus cuprescens (Ménétriés, 1832) and related species (Coleoptera: Buprestidae). Zootaxa, 317: 1 22. [#5878] JENDEK, E., 2004: Revision of Agrilus acutus (Thunberg, 1787) and related species (Coleoptera: Buprestidae). Zootaxa, 507: 1 19. [#6675]

253 JENDEK, E., 2005: Taxonomic and nomenclatural notes on the genus Agrilus Curtis (Coleoptera: Buprestidae: Agrilini). Zootaxa, 1073: 1 29. [#7217] JENDEK, E., 2006: New nomeclatorial and taxonomic acts, and comments. Buprestidae: Agrilus p. 60. Catalog: genus Agrilus Curtis, 1825, p. 388 403. In: LÖBL, I. & SMETANA, A. (eds), Catalogue of the Palaearctic Coleoptera, Vol. 3, Stenstrup, Apollo Books, 690 pp. [#7441] JENDEK, E., 2006: Taxonomic and nomenclatural notes on the genera Sarawakita Obenberger and Agrilus Curtis (Coleoptera: Buprestidae: Agrilinae). Zootaxa, 1153: 33 42. [#7359] JENDEK, E., 2007: Nineteen new Agrilus species from the Palaearctic and Oriental regions (Coleoptera: Buprestidae: Agrilinae). Entomological Problems, Bratislava, 37 (1 2): 31 43. [#8006] JENDEK, E., 2007: Taxonomic and nomenclatural clarifications on the genus Agrilus (Coleoptera: Buprestidae: Agrilini). Entomological Problems, Bratislava, 37 (1 2): 87 95. [#8008] JENDEK, E., 2007: Taxonomic notes on the European species of the genus Agrilus (Coleoptera: Buprestidae: Agrilini). Folia Heyrovskyana, Series A, 14 (3): 109 112. [#7967] JENDEK, E. & GREBENNIKOV, V., 2009: Agrilus sulcicollis (Coleoptera: Buprestidae) a new alien species in North America. Canadian Entomologist, 141: 236 245. [#8724] JENDEK, E. & GREBENNIKOV, V., 2009: Revision of the Agrilus cyanescens species group (Coleoptera: Buprestidae) with description of three new species from the east Palaearctic region. Zootaxa, 2139: 43 60. [#8561] JENDEK, E. & GREBENNIKOV, V., 2009: Revision of the Agrilus muscarius species group (Coleoptera: Buprestidae) with description of thirteen new species from Palaearctic and Oriental regions. Zootaxa, 2168: 1 33. [#8563] JI YING, JI RONG & HUANG REN XIN, 2004: Invasive Species Agrilus Mali Matsumura and Damage in Xinjiang. Xinjiang Agricultural Sciences, 41 (1): 31 33. (in Chinese with English subtitle) [#7326] JUDEICH, J. F. & NITSCHE, H., 1889: Lehrbuch der Mitteleuropäischen Forstinsektenkunde mit einem Anhange : Die forstschadlichen Wirbelthiere. Als achte Aulage von von Dr. J. T. C. Ratzeburg: Die Waldverderber und ihre Feinde. II. Abtheilung. Wien, Eduard Hölzel, p. 265 623. [#5686] KABIR, A. K. M. F., 1995: Flatheaded borer Agrilus acutus Thumb. In: Juta pest in Bangladesh, BJRI, Dhaka, p. 59. [#7048] KADYRBEKOV, R. H., TLEPPAEVA, A. M. & CHILDEBAEV, M. K., 2003: K faune zhukov drovosekov (Cerambycidae) i zlatok (Buprestidae) nacional’nogo prirodnogo parka “Burabai”. Izvestiya Ministerstva Obrazovaniya i Nauki Respubliki Kazakhstan, Seriya Biologicheskaya i Meditsinskaya, 6: 34 42. (in Russian with English summary) [#7318] KADYROV, A. K., 1973: Materialy k izucheniyu vrednoi èntomofauny zelenykh nasazhdenii g. Dushanbe i poselkov yugo zapadnogo Tadjikistana. In: Fauna i èkologiya chlenistonogikh Tadzhikistana. Dushanbe, Tadzhikistanskii gosuniversitet, p. 50 65. (in Russian) [#8631] KADYROV, A. K., 1973: Puti i sposoby formirovanya vrednoi èntomofauny zelenykh nasazhdenii v Tadzhikistane. In: Fauna i èkologiya chlenistonogikh Tadzhikistana. Dushanbe, Tadzhikistanskii gosuniversitet, p. 65 75. (in Russian) [#8632] KADYROV, A. K., 1976: Materialy k faune zhukov zlatok (Coleoptera, Buprestidae) yugo zapadnogo Tadzhikistana. Izvestiya akademii nauk Tadzhikskoi SSR, Otdelenie biologicheskikh nauk, No. 2 (63): 45 52. (in Russian) [#3461]

254 KALASHIAN, M. Y., 1985: Redkie, maloizvestnye i novye dlya Armenii vidy zhukov zlatok (Coleoptera, Buprestidae). Biologicheskii zhurnal Armenii, Erevan, 38 (8): 718 720. (in Russian) [#1366] KALASHIAN, M. Y., 1986: Lozhnyi yaiceklad zhukov zlatok podsemeistva Agrilinae (Coleoptera, Buprestidae) i ego sistematicheskoye znacheniye. In: Tezisy dokladov 5 nauchnotekhnicheskoi konferentsii molodykh uchenykh i specialistov raiona 26 Bakinskikh Komissarov, Yerevan, p. 125. (in Russian) [#3866] KALASHIAN, M. Y., 1986: K klassifikacii lozhnykh yaitsekladov zhukov zlatok (Coleoptera, Buprestidae). In: Pervaya Zakavkazskaya konferentsia po èntomologii, Tezisy dokladov,Yerevan, p. 94 96. (in Russian) [#3865] KALSHOVEN, L. G. E., 1951: De plagen van de cultuurgewassen in Indonesië. Deel. II, Van Hoeve’s Bandoeng, 1 p + 513 1065 pp + 8 col. pls, 281 figs. (in Dutch with English translation of the text for illustrations) [#3718] KALTENBACH, J. H., 1874: Die Pflanzenfeinde aus der Klasse der Insekten. Stuttgart, J. Hoffmann, 848 pp. [#8154] KANGAS, E., 1947: Studien über die Agrilus larven (Col. Buprestidae). Annales Entomologici Fennici, 13: 104 113. [#3622] KAPLIN, V. G., 1981: Kompleksy chlenistonogykh zhivotnykh, obimayushchikh v tkanyakh rastenii peschanykh pustyn’. Ashkhabad, Ylym, 376 pp. (in Russian) [#5853] KARAS, V. & KLETEČKA, Z., 2002: Nálezy vzácnych a nových druhů brouků (Coleoptera) pro faunu jižních Čech. Records of rare and new beetle species (Coleoptera) from the fauna of South Bohemia. Sborník Jihočeského Muzea v Českých Budejovicích, Přírodní vědy, 42: 89 98. (in Czech with English subtitle and summary) [#5869] KENIS, M. & HILSZCZANSKI, J., 2004: Natural enemies of Cerambycidae and Buprestidae infesting living trees, p. 475 498. In: LIEUTIER, F., DAY, K. R., BATTISTI, A., GRÉGOIRE, J. C. & EVANS, H. F. (eds), Bark and Wood Boring Insects in Living Trees in Europe: A Synthesis. Kluwer Academic publishers, 569 pp. [#8566] KEPPEN, F., 1882: Vrednyya nas’komyya. Tom vtoroi, Special’naya chast’ I, Pryamokrylyya, zhuki i pereponchatokrylyya. Sanktpeterburg’, Tipografiya imperatorskoi akademii nauk’, 585 pp. (in Russian) [#8636] KERREMANS, C., 1885: Énumération des Buprestides décrits postérieurement au Catalogue de MM. Gemminger & de Harold. 1870 1883. Annales de la Société Entomologique de Belgique, 29: 119 157. [#1381] KERREMANS, C., 1890: Note sur les Buprestides du Chota Nagpore. Bulletin ou Comptes Rendus des Séances de la Société Entomologique de Belgique, 34: CC CCIX. [#1389] KERREMANS, C., 1892: Coléoptères du Bengale occidental. 19me mémoire. Buprestides de l’Inde. Annales de la Société Entomologique de Belgique, 36: 171 226. [#1402] KERREMANS, C., 1892: Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIX. Buprestides. Annali del Museo Civico di Storia Naturale di Genova, Serie 2, 12 (32): 809 832. [#1398] KERREMANS, C., 1892: Catalogue synonymique des Buprestides decrits de 1758 à 1890. Séance du 5 septembre 1891. Mémoires de la Société Entomologique de Belgique, 1: 1 304. [#1397] KERREMANS, C., 1893: Additions aux Buprestides des Indes Orientales. Annales de la Société Entomologique de Belgique, 37: 326 357. [#1406] KERREMANS, C., 1894: Buprestides Indo Malais. Annales de la Société Entomologique de Belgique, 38: 468 480. [#1412] KERREMANS, C., 1895: Buprestides Indo Malais. Deuxième partie. Annales de la Société Entomologique de Belgique, 39: 192 224. [#1416]

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA KERREMANS, C., 1897: Contribution a l’étude de la faune intertropicale Américaine. Voyages de M. E. Gounelle au Brésil. Buprestides. Mémoires de la Société Entomologique de Belgique, 6: 1 146. [#1424] KERREMANS, C., 1898: Buprestides nouveaux de l’Australie et des régions voisines. Annales de la Société Entomologique de Belgique, 42: 113 182. [#1427] KERREMANS, C., 1900: Buprestides nouveaux et remarques synonymiques. Annales de la Société Entomologique de Belgique, 44: 282 351. [#1434] KERREMANS, C., 1900: Contribution a l’etude de la faune entomologique de Sumatra (Côte ouest Vice résidence de Païnan). Chasses de M. J. L. Weyers VI. Buprestides. Mémoires de la Société Entomologique de Belgique, 7: 1 60. [#1435] KERREMANS, C., 1900: Buprestides Indo Malais. Troisiéme partie. Mémoires de la Société Entomologique de Belgique, 7: 61 93. [#1436] KERREMANS, C., 1903: Coleoptera Serricornia. Fam. Buprestidae, Fasc. 12b, 12c, 12d, p. 49 338. In: WYTSMAN, P. (ed), Genera Insectorum. Tome II, Fascicules XII XIV. Bruxelles, Verteneuil & Desmet. [#1441] KERREMANS, C., 1912: H. Sauter’s Formosa Ausbeute. Buprestiden. Archiv für Naturgeschichte, 78, Abteilung A, Heft 7: 203 209. [#1377] KERREMANS, C., 1913: H. Sauter’s Formosa Ausbeute. Buprestiden. 2. Teil. Archiv für Naturgeschichte, (1912) 78, Abteilung A, Heft 11: 110 116. [#1471] KERREMANS, C., 1914: H. Sauter’s Formosa Ausbeute. Buprestiden. 3. Teil. Archiv für Naturgeschichte, (1913) 79, Abteilung A, Heft 8: 103 107. [#1481] KETTERING, H., 1978: Agrilus pseudocyaneus Kiesw. in der Pfalz (Coleoptera, Buprestidae). Nachrichtenblatt der Bayerischen Entomologen, 27 (5): 81 83. [#3623] KHARAZISHVILI, K. V., 1960: K izycheniyu skrytnostvolovykh vreditelei topolya v Gruzii. In: 4. S’ezd vsesoyuznogo èntomologicheskogo obshchestva. Leningrad, 29.I. 3.II. 1960 g., Tezisy dokladov, Moskva, Leningrad, Izdatel’stvo akademii nauk SSSR, p. 146 147. (in Russian) [#8630] KHRAMCOV, N. N. & PADIJ, N. N., 1965: Stvolovye vrediteli lesa i bor’ba s nimi. Moskva, Izdatel’stvo Lesnaya promyshlennost’, 159 pp. (in Russian) [#8649] KIESENWETTER, H. VON, 1849: Verzeichnis europ. Käfer. Catalogus Coleopterorum Europae. Herausgeben vom entomologischen Vereine in Stettin. 3. Auflage, Bautzen, E. M. Monse [mit genus register p. 1 7 von C. A. Dohrn]. 82 pp. [#8334] KIESENWETTER, H. VON, 1857: Buprestidae, Eucnemidae, Bogen 1 11, Naturgeschichte der Insecten Deutschlands begonnen von Dr. W. F. Erichson, fortgesetzt von Prof. Dr. H. Schaum, Dr. G. Kraatz und H. v. Kiesenwetter. Erste Abtheilung Coleoptera. Vierter Band. Berlin, Nicolaische Verlagsbuchhandlung (G. Parthey), p. 1 178. [#3015] KIESENWETTER, H. VON, 1879: Neue Amur Käfer. Deutsche Entomologische Zeitschrift, 23 (1): 145 146. [#3347] KIZERITSKII, V. A., 1915: K’ faun’ zhukov Poltavskoi gubernii. Contribution à la faune des Coléoptères du governement de Poltava. Revue Russe d’Entomologie, 15: 167 184. (in Ukrainian with French subtitle) [#8698] KLETEČKA, Z., 1998: To the bionomy of Dicerca amphibia Mars. and of accompanying species of jewel beetles (Coleoptera, Buprestidae) on poplars Populus sp. div. in eastern Ukraine. Jewel Beetles, No. 7: 19 29, figs. [#4416]

LITERATURE CITED KOROTYAEV, B. A., KONSTANTINOV, A. S., LINGAFELTER, S. W., MANDELSHTAM, M. Y. & VOLKOVITSH, M. G., 2005: Gall inducing Coleoptera, 239 271. In: RAMAN, A., SCHAEFER, C. W. & WITHERS, T. M. (eds), Biology, ecology and evolution of gall inducing arthropods: Volume 2. Enfield, Science Publishers, Inc., I XXI, 431 817. [#7284] KOSTIN, I. A., 1973: Zhuki dendrofagi Kazakhstana (Koroedy, drovoseki, zlatki). The dendrophagous beetles of Kazakhstan (Buprestidae, Cerambycidae, Ipidae), Alma Ata, Nauka, 286 pp. (in Russian with English subtitle and summary) [#3693] KRAATZ, G. & KIESENWETTER, H. VON, 1879: Ueber Buprestiden vom Amur. Deutsche Entomologische Zeitschrift, 23 (2): 253 256. [#1595] KRIVOLUTSKAYA, G. O., 1973: Èntomofauna Kuril’skikh ostrovov. Osnovnye cherty i proiskhozhdenie. Entomofauna of the Kuril islands. Principal features and origin. Leningrad, Nauka, 315 pp. (in Russian with English subtitle and summary) [#8102] KRIVOSHEINA, N. P., 1976: Kompleksy nasekomykh rizobiontov i ikh znachenie v pustynnykh biogeocenozakh. In: Tezisy dokladov vsesoyuznoi nauchnoi konferencii po kompleksnomu izucheniyu i osvoeniyu pustynnykh territorii SSSR. Sekciya IV, Ashkhabad, Ilym, p. 22 24. (in Russian) [#8641] KROGERUS, R., 1922: Studien über Agrilus Arten. II. Zur Biologie des Agrilus ater L. ( sexguttatus auct.). Notulae Entomologicae, 2: 109 113. [#3916] KROGERUS, R., 1925: Zur Kenntnis der Agriliden Finnlands. Meddelanden af Societas pro Fauna et Flora Fennica, Helsingfors, 49: 70 76. [#3472] KRÓLIK, R. & MAJEWSKI, T., 1994: Agrilus ribesi Schaeffer [sic!], 1946 (Coleoptera, Buprestidae) nowy dla fauny Polski gatunek chrzaszcza. Agrilus ribesi Schaeffer [sic!], 1946 (Coleoptera, Buprestidae) species of beetle new for the fauna of Poland. Wiadomosci Entomologiczne, Poznan, 13 (2): 107 113. (in Polish with English sumary) [#3471] KRYUKOVA, E. A. & TEREKHOVA, N. A., 1974: Perenoschiki trakheomikoza duba na yugo vostoke Èvropeiskoi chasti RSFSR. In: Materialy sed’mogo c’ezda vsesoyuznogo èntomologicheskogo obshchestva, chast’ vtoraya, Sel’skokhozyaistvennaya èntomologiya, Lesnaya èntomologiya, Leningrad, Akademiya nauk SSSR, p. 225. (in Russian) [#8643] KRYZHANOVSKII, O. L., 1984: Semeistvo Zlatki Buprestidae, p. 20, 105 108. (in Russian) In: KOPANEVA, L. M. (ed), Opredelitel’ vrednykh i poleznykh nasekomykh i kleskchei plodovykh i yagodnykh kul’tur v SSSR. Leningrad, Kolos, 288 pp. [#8665] KUBÁŇ, V., 1978: Příspěvek k faunistice krasců Moravy a Slovenska s ekologickými a bionomickými poznámkami. Beitrag zur Faunistik Prachtkäfer Mährens und der Slowakei mit den ekologische und bionomische Bemerkungen. (Coleoptera Buprestidae). Zprávy Československé společnosti entomologické při ČSAV, Praha, 14: 21 24. (in Czech with German summary) [#3474] KULIKOWSKI, E., 1897: Materialy dlya fauny Coleoptera Yuzhnoi Rossii. Die Käferfauna v. Süd Russlands. Mémoires de la sociéte des naturalistes de la Nouvelle Russie, T. XXI, P. I: 1 270. (in Russian with German subtitle) [#8666] KULINITSH, P. N., 1963: Materialy k izucheniyu zhestkokrylykh stvolovykh vreditelei gornykh lesov Tadzhikistana. In: Pyatoe soveshchanie vsesoyuznogo èntomologicheskogo obshchestva. Tashkent, sentyabr 1963 g., Tezisy dokladov, Moskva, Leningrad, Akademiya nauk SSSR, p. 153 154. (in Russian) [#8648]

255 KUROLI, G., 1999: A málna karcsúdíszbogár (Agrilus aurichalceus Redtenbacher). Rose stem girdler (Agrilus aurichalceus Redt.). Növényvédelem, 35 (6): 257 263. (in Hungarian with English summary) [#5916] KUROSAWA, Y., 1947: [Note on the distribution of Agrilus cyanenoniger E. Saunders]. Transactions of the Kinki Coleopterological Society, 2 (1): 1 3. (in Japanese) [#3339] KUROSAWA, Y., 1949: [The illustration on the family Buprestidae of Japan (2)]. Shin Konchu, Tokyo, 3 (1): 13 15, 28. (in Japanese) [#5028] KUROSAWA, Y., 1950: Iconographia Insectorum Japonicorum. [pars Buprestidae p. 3183 3216]. p. 1031 1186. (in Japanese) [#3274] KUROSAWA, Y., 1954: Buprestid fauna of Eastern Asia, (Coleoptera) (I). Bulletin of the National Science Museum, Tokyo, N. S. Vol. 1, No 1 (No. 34): 29 39. [#1612] KUROSAWA, Y., 1954: Buprestid fauna of Eastern Asia (2). Bulletin of the National Science Museum, Tokyo, N. S. Vol. 1, No 2 (No. 35): 82 93. [#1613] KUROSAWA, Y., 1956: Buprestidae, p. 1111 1122. In: ESAKI, T., ISHII, T., KAWAMURA, T. & KINOSHITA, S. (eds), Iconographia insectorum Japonicorum, Editio secunda, reformata, Tokyo, Hokuryukan, 1738 + 203 pp. [#8107] KUROSAWA, Y., 1956: Buprestid fauna of Eastern Asia (3). Bulletin of the National Science Museum, Tokyo, N. S. Vol. 3, No. 1 (No. 38): 33 41. [#1614] KUROSAWA, Y., 1957: Buprestid fauna of Eastern Asia (4). (Coleoptera). Bulletin of the National Science Museum, Tokyo, Vol. 3, No. 3 (No. 40): 183 194. [#1615] KUROSAWA, Y., 1963: Buprestidae. 147 156 [Buprestidae], 167 168, 187 188. (in Japanese) In: NAKANE, T., OHBAYASHI, K., NOMURA, S. & KUROSAWA, Y. (eds), Iconographia Insectorum Japonicorum. Colore naturali edita. Volumen II (Coleoptera). Tokyo, Hokuryukan, 443 pp, 192 pls. [#1618] KUROSAWA, Y., 1963: Buprestid fauna of Eastern Asia (5). (Coleoptera). Bulletin of the National Science Museum, Tokyo, (1962) Vol. 6, No. 2 (No. 51): 90 111. [#1617] KUROSAWA, Y., 1964: Descriptions of several new species and subspecies of the genus Agrilus Curtis from Amami Oshima of the Loo Choo Archipelago. Kontyû, Tokyo, 32 (2): 323 334. [#1619] KUROSAWA, Y., 1974: [A comment to the Japanese Buprestidae (10)]. Coleopterists’ News, Tokyo, 17/18: 1 4. (in Japanese) [#4575] KUROSAWA, Y., 1974: [A comment to the Japanese Buprestidae (11)]. Coleopterists’ News, Tokyo, 21/22: 1 2. (in Japanese) [#4576] KUROSAWA, Y., 1974: [A comment to the Japanese Buprestidae (12)]. Coleopterists’ News, Tokyo, 23/24: 1 4. (in Japanese) [#4577] KUROSAWA, Y., 1975: [A comment to the Japanese Buprestidae (13)]. Coleopterists’ News, Tokyo, 25/26: 1 2. (in Japanese) [#4578] KUROSAWA, Y., 1975: [A comment to the Japanese Buprestidae (14)]. Coleopterists’ News, Tokyo, 27/28: 1 4. (in Japanese) [#4591] KUROSAWA, Y., 1976: Notes on the Oriental species of the Coleopterous family Buprestidae (I). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 2 (1): 51 58. [#1627] KUROSAWA, Y., 1976: Notes on the Oriental species of the Coleopterous family Buprestidae (II). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 2 (2): 129 136. [#1628] KUROSAWA, Y., 1981: A new buprestid beetle of the genus Agrilus from Northern Thailand, with description of a new subgenus (Coleoptera, Buprestidae). Entomological Review of Japan, 36 (1): 5 8. [#1637]

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266 WOLLASTON, T. V., 1865: Coleoptera Atlantidum, being an enumeration of the coleopterous insects of the Madeiras, Salvages, and Canaries. London, John van Voorst, XLVII + 526 + 140 pp. [#7784] WOO SHEH MING, 1964: Studies on the citrus buprestid beetle, Agrilus auriventris Saunders. Acta Phytophylacica Sinica, 3 (1): 61 71. (in Chinese with English subtitle and summary) [#7020] WU HAI., 2006: Bionomics and control of Agrilus zanthoxylumi. Chinese Bulletin of entomology, 43 (2): 236 239. (in Chinese with English summary) [#7654] XU GONGTIAN, 2003: Agrilus marcopoli Obenberger, p. 321 322. (in Chinese) In: XU GONGTIAN (ed), Atlas of ornamental pests and diseases. Beijing, China Agriculture Press. [#6836] YABLOKOV KHNZORYAN, S. M., 1957: Zhestkokrylye duba v Armyanskoi SSR. In: Materialy po izuchenyu fauny Armyanskoi SSR. Zoologicheskii sbornik, Akademya nauk Armyanskoi SSR, Erevan, Vypusk 10, (3): 59 152. (in Russian) [#5612] YAGDYEV, A., 1974: Vrediteli stvolov i vetvei turangi. In: Materialy sed’mogo c’ezda vsesoyuznogo èntomologicheskogo obshchestva, chast’ vtoraya, Sel’skokhozyaistvennaya èntomologiya, Lesnaya èntomologiya, Leningrad, Akademiya nauk SSSR, p. 257. (in Russian) [#8644] YAGDYEV, A., 1975: Stvolovye vrediteli turangy v Turkmenii. Izvestiya akademii nauk Turkmenskoi SSR, Seriya biologicheskikh nauk, 6: 60 64. (in Russian) [#7775] YAGDYEV, A., 1978: Obnaruzhenie zlatki Agrilus tschitscherini Sem. na turange v doline reki Ili. Izvestiya akademii nauk Turkmenskoi SSR, Seriya biologicheskikh nauk, 2: 90. (in Russian) [#4206] YAGDYEV, A., 1984: Kompleksy zhukov ksilophagov v tugainykh lesakh yuga Srednei Azii. Izvestiya akademii nauk Turkmenskoi SSR, Seriya biologicheskikh nauk, 4: 69 71. (in Russian) [#6196] YANG XINGKE, SUN HONGGUO & JIANG GUOMEI, 1991: Catalogue of the type specimens in the insect collection of the Institute of Zoology Academica Sinica, Beijing, China. Beijing, Agricultural Publishing House, 163 pp. (in Chinese) [#6756] YANG ZHONGQI, STRAZANAC, J. S., MARSH, P. M., ACHTERBERG, C. VAN & CHOI WONYOUNG, 2005: First recorded parasitoid from China of Agrilus planipennis: A new species of Spathius (Hymenoptera: Braconidae: Doryctinae). Annals of the Entomological Society of America, 98 (5): 636 642. [#7263] YU CHENGMIN, 1992: Agrilus marcopoli Obenberger, p. 400 401. In: XIAO, GANGROU (ed), Forest Insects of China. The second edition (Revised & enlarged). Beijing, China forestry publishing house, 1362 pp. [#5982] YUASA, H., 1932: Family Buprestidae, p. 652 660. In: ESAKI, T., HORI, H. & YASUMATSU, K., Nippon Konchu Zukan. Iconographia insectorum Japonicorum, Editio prima, Tokyo, Hokuryukan, 2241 pp. [#3369] YUASA, H., 1933: On the structure of some Japanese buprestid larvae, with notes on their life history. Journal of the Imperial Agricultural Experiment Station, Tokyo, 2 (2): 263 282. (in Russian with English summary) [#3132] YUASA, H., 1949: Buprestidae p. 126 128. In: Illustrated pocket book of Japanese insects. Tokyo, Hokuryukan, 432 + 33 pp. (in Japanese with English subtitle) [#8108] YURCHENKO, G. I., TUROVA, G. I. & KUZMIN, E. A., 2007: To distribution and ecology of emerald ash borer (Agrilus planipennis Fairmaire) in the Russian Far East. A.I. Kurentsov’s Annual Memorial Meetings, XVIII: 94 98. (in Russian with English subtitle and summary) [#8892]

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267

APPENDICES APPENDIX I. NEW TAXONOMIC AND NOMENCLATURAL ACTS CHANGES ON GENERIC LEVEL 1. Teres Harris, 1829 available junior synonym of Agrilus Curtis, 1825 2. Buprestis granulata Say, 1823 designated as the type species of Teres Harris, 1829 (fixation by subsequent designation) 3. Paradomorphus albicollis Waterhouse, 1887 t type species of Paradomorphus 4. Agrilus pulchellus Bland, 1865 type species of Engyaulus

NEW AGRILUS TAXA All newly proposed names are attributed to senior author. 1. A. aureofasciatus sp. nov. 2. A. aurigaster sp. nov. 3. A. aurinotus sp. nov. 4. A. aurithorax sp. nov. 5. A. bai sp. nov. 6. A. bicuspidatus sp. nov. 7. A. bocaki sp. nov. 8. A. brevinotus sp. nov. 9. A. businskorum sp. nov. 10. A. caudicula sp. nov. 11. A. chaetocoxalis sp. nov. 12. A. ciping sp. nov. 13. A. compacticornis sp. nov. 14. A. conspersus sp. nov. 15. A. cordicollis sp. nov. 16. A. daur sp. nov. 17. A. diao sp. nov. 18. A. diversornatus sp. nov. 19. A. dureli sp. nov. 20. A. eunuchus sp. nov. 21. A. foramenifer sp. nov. 22. A. fornicatus sp. nov. 23. A. fusciapex sp. nov. 24. A. fusiformis sp. nov. 25. A. han sp. nov. 26. A. hani sp. nov. 27. A. hmong sp. nov. 28. A. hui sp. nov. 29. A. improcerus sp. nov. 30. A. jindrai sp. nov. 31. A. kucerai sp. nov. 32. A. lacrima sp. nov. 33. A. lahu sp. nov. 34. A. lei sp. nov. 35. A. lisu sp. nov. 36. A. lobatus sp. nov. 37. A. lugu sp. nov. 38. A. magnus sp. nov. 39. A. manchu sp. nov. 40. A. mandala sp. nov. 41. A. miao sp. nov.

42. 43. 44. 45. 46. 47. 48. 49. 50. 51. 52. 53. 54. 55. 56. 57. 58. 59. 60. 61. 62. 63. 64. 65. 66. 67. 68. 69. 70. 71. 72. 73. 74.

A. mio sp. nov. A. muongoides sp. nov. A. naxi sp. nov. A. ornatoides sp. nov. A. paralleloides sp. nov. A. pleuralis sp. nov. A. pleurostigmus sp. nov. A. presbys sp. nov. A. prolatus sp. nov. A. prosternalis sp. nov. A. pumi sp. nov. A. qiang sp. nov. A. qinlingcola sp. nov. A. recticollis sp. nov. A. sagittifer sp. nov. A. schillhammeri sp. nov. A. sentis sp. nov. A. shan sp. nov. A. she sp. nov. A. sichuanus sp. nov. A. siskai sp. nov. A. smaragdinus habros ssp. nov. A. smetanai sp. nov. A. spathulifer sp. nov. A. spiculum sp. nov. A. stigma sp. nov. A. tai sp. nov. A. terracotta sp. nov. A. tuber sp. nov. A. ulmicola sp. nov. A. vigil sp. nov. A. zhuang sp. nov. A. zhui sp. nov.

REVALIDATED NAMES 1. olivaceidorsis Obenberger, 1917 nom. rev. resurrected from the synonymy of A. auriventris Saunders, 1873 as the valid specific name 2. sapporoensis Obenberger, 1935 nom. rev. resurrected from the synonymy of A. komareki Obenberger, 1926 as the valid name of subspecies A. komareki sapporoensis Obenberger, 1935.

CHANGE IN RANK 1. sachalinensis Obenberger, 1935 downgraded to subspecies, as A. sinuatus sachalinensis Obenberger, 1935 stat. nov.

TRANSFERS IN SYNONYMY 1. cojbalsanensis Cobos, 1968 from synonymy of A. gebleri Obenberger, 1924 to synonymy of A. viridis (Linné, 1758) 2. kiricenkoi Obenberger, 1935 from synonymy of A. attila Obenberger, 1930 to synonymy of A. viridis (Linné, 1758)

268 3. scutellatus Abeille de Perrin, 1897 from synonymy of A. ecarinatus Marseul, 1866 to synonymy of A. zigzag Marseul, 1866 4. sieversi Abeille de Perrin, 1897 from synonymy of A. zigzag Marseul, 1866 to synonymy of A. albogularis albogularis Gory, 1841

NEW SYNONYMS 1. alazon Lewis, 1893 syn. nov. of A. decoloratus Kerremans, 1892 2. aridosus Obenberger, 1936 syn. nov. of A. macrotatopus Obenberger, 1936 3. aritai Tôyama, 1985 syn. nov. of A. alesi Obenberger, 1935 4. assamensis Obenberger, 1936 syn. nov. of A. caligans Bourgoin, 1925 5. attila Obenberger, 1930 syn. nov. of A. tschitscherini Semenov, 1895 6. brevitarsis Lewis, 1893 syn. nov. syn. nov. of A. viridis (Linné, 1758) 7. brydli Obenberger, 1924 syn. nov. of A. viridis (Linné, 1758) 8. candidiventris Obenberger, 1936 syn. nov. of A. nivosus Abeille de Perrin, 1900 9. cardaces Obenberger, 1936 syn. nov. of A. viduus Kerremans, 1914 10. chinganicus Obenberger, 1922 syn. nov. of A. viduus Kerremans, 1914 11. chopardi Descarpentries & Villiers, 1963 syn. nov. of A. auropictus Kerremans, 1912 12. daghestanicus Obenberger, 1930 syn. nov. of A. sericans Kiesenwetter, 1857 13. diolaus Obenberger, 1958 syn. nov. of A. malloti Théry, 1930 14. djukini Obenberger, 1935 syn. nov. of A. pratensis (Ratzeburg, 1837) 15. duporti Bourgoin, 1922 syn. nov. of A. olivaceidorsis Obenberger, 1917 16. emisi Ohmomo, 2005 syn. nov. of A. komareki sapporoensis Obenberger, 1935 17. fairmairei Kerremans, 1892 syn. nov. of A. sinensis sinensis Thomson, 1879 18. filigranus Obenberger, 1916 syn. nov. of A. salicis Frivaldszky, 1877 19. fleutiauxi Bourgoin, 1922 syn. nov. of A. auriventris Saunders, 1873 20. gebleri Obenberger, 1924 syn. nov. of A. viridis (Linné, 1758) 21. gerschuni Stepanov, 1958 syn. nov. of A. tschitscherini Semenov, 1895 22. graptelytrus Obenberger, 1914 syn. nov. of A. auriventris Saunders, 1873 23. hoscheki Obenberger, 1916 syn. nov. of A. viduus Kerremans, 1914 24. imbellis Kerremans, 1892 syn. nov. of A. viridis (Linné, 1758) 25. importunus Kerremans, 1903 syn. nov. of A. viridis (Linné, 1758) 26. kanohi Kurosawa, 1954 syn. nov. of A. auropictus Kerremans, 1912 27. kanssuanus Obenberger, 1930 syn. nov. of A. smaragdifrons Ganglbauer, 1889 28. kuldjensis Obenberger, 1936 syn. nov. of A. nivosus Abeille de Perrin, 1900 29. kusakievici Obenberger, 1936 syn. nov. of A. zigzag Marseul, 1866

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA 30. lubischevi Stepanov, 1958 syn. nov. of A. tschitscherini Semenov, 1895 31. lugovoi Stepanov, 1958 syn. nov. of A. salicis Frivaldszky, 1877 32. machnovskii Stepanov, 1958 syn. nov. of A. tschitscherini Semenov, 1895 33. malyi Obenberger, 1936 syn. nov. of A. subrobustus Saunders, 1873 34. melanolucidus Ohmomo, 2005 syn. nov. of A. nipponigena Obenberger, 1935 35. mentor Kerremans, 1912 syn. nov. of A. plagiatus Ganglbauer, 1889 36. milletae Baudon, 1968 syn. nov. of A. coomani Bourgoin, 1925 37. mushanus Kurosawa, 1976 syn. nov. of A. madeci Baudon, 1968 38. nagaoi Nakane, 1983 syn. nov. of A. maculifer Saunders, 1873 39. nalajchanus Cobos, 1968 syn. nov. of A. friebi Obenberger, 1922 40. nipponicola Kurosawa, 1963 syn. nov. of A. fleischeri Obenberger, 1925 41. omissulus Obenberger, 1935 syn. nov. of A. tyndarus Kerremans, 1913 42. ornamentifer Obenberger, 1940 syn. nov. of A. olivaceidorsis Obenberger, 1917 43. panhensis Baudon, 1968 syn. nov. of A. adelphinus Kerremans, 1895 44. pidjinus Obenberger, 1924 syn. nov. of A. auriventris Saunders, 1873 45. pistaciophagus Alexeev & Kulinitsh, 1962 syn. nov. of A. chlorophyllus Abeille de Perrin, 1904 46. poppiusi Obenberger, 1924 syn. nov. of A. viridis (Linné, 1758) 47. praeclarus Krogerus, 1925 syn. nov. of A. pratensis (Ratzeburg, 1837) 48. pseudograminis Jendek, 2007 syn. nov. of A. andrusi Baudon, 1968 49 pseudoussuricola Alexeev, 1979 syn. nov. of A. kaluganus Obenberger, 1940 50. richteri Alexeev, 1975 syn. nov. of A. morawitzi Obenberger, 1936 51. rokuyai Kurosawa, 1976 syn. nov. of A. madeci Baudon, 1968 52. ryukyuensis Tôyama, 1985 syn. nov. of A. auropictus Kerremans, 1912 53. sacchari Obenberger, 1940 syn. nov. of A. alesi Obenberger, 1935 54. salicivola Kurosawa, 1963 syn. nov. of A. viridis (Linné, 1758) 55. semirjeciae Obenberger, 1936 syn. nov. of A. zigzag Marseul, 1866 56. sericarius Abeille de Perrin, 1897 syn. nov. of A. sericans Kiesenwetter, 1857 57. sibiricola Obenberger, 1924 syn. nov. of A. laticornis (Illiger, 1803) 58. simbirensis Obenberger, 1925 syn. nov. of A. zigzag Marseul, 1866 59. spesivcevi Obenberger, 1927 syn. nov. of A. pilosovittatus Saunders, 1873 60. subviduus Kurosawa, 1957 syn. nov. of A. viduus Kerremans, 1914 61. suensoni Obenberger, 1940 syn. nov. of A. insulicola Kerremans, 1912 62. therondi Descarpentries & Villiers, 1963 syn. nov. of A. tyndarus Kerremans, 1913

269

NEW TAXONOMIC AND NOMENCLATURAL ACTS 63. towadensis Miwa & Chûjô, 1940 syn. nov. syn. nov. of A. viridis (Linné, 1758) 64. toyoshimai Tôyama, 1988 syn. nov. of A. esakii Kurosawa, 1964 65. transbaicalensis Obenberger, 1935 syn. nov. of A. viridis (Linné, 1758) 66. uzbekistanus Stepanov, 1958 syn. nov. of A. tschitscherini Semenov, 1895 67. wilsoni Baudon, 1968 syn. nov. of A. quentini Descarpentries & Villiers, 1963 68. yokoyamai Iga, 1955 syn. nov. of A. sinuatus sachalinensis Obenberger, 1935 69. zhelochovtsevi Alexeev, 1979 syn. nov. of A. sinuatus sachalinensis Obenberger, 1935

LECTOTYPE DESIGNATIONS (lectotypes are cited in the original combination and rank) 1. 2. 3. 4. 5. 6. 7. 8. 9. 10. 11. 12. 13. 14. 15. 16. 17. 18. 19. 20. 21. 22. 23. 24. 25. 26. 27.

Agrilus aerosus Ganglbauer, 1889 Agrilus alesi Obenberger, 1935 Agrilus aridosus Obenberger, 1936 Agrilus assamensis Obenberger, 1936 Agrilus attila Obenberger, 1930 Agrilus auriventris Saunders, 1873 Agrilus brodjaga Obenberger, 1914 Agrilus brydli Obenberger, 1924 Agrilus caligans Bourgoin, 1925 Agrilus candidiventris Obenberger, 1936 Agrilus cardaces Obenberger, 1936 Agrilus chinganicus Obenberger, 1922 Agrilus chlorophyllus Abeille de Perrin, 1904 Agrilus daghestanicus Obenberger, 1930 Agrilus dentiapex Obenberger, 1940 Agrilus discalis Saunders, 1873 Agrilus duporti Bourgoin, 1922 Agrilus egorovi Alexeev, 1989 Agrilus emmerichi Obenberger, 1935 Agrilus fleutiauxi Bourgoin, 1922 Agrilus fortunatus Lewis, 1893 Agrilus gebleri Obenberger, 1924 Agrilus graptelytrus Obenberger, 1914 Agrilus hoscheki Obenberger, 1916 Agrilus imbellis Kerremans, 1892 Agrilus imitans Lewis, 1893 Agrilus insulicola Kerremans, 1912

28. 29. 30. 31. 32. 33. 34. 35. 36. 37. 38. 39. 40. 41. 42. 43. 44. 45. 46. 47. 48. 49. 50. 51. 52. 53. 54. 55. 56. 57. 58. 59. 60. 61. 62. 63. 64. 65. 66. 67. 68. 69. 70. 71. 72.

Agrilus jenissejensis Obenberger, 1924 Agrilus kanssuanus Obenberger, 1930 Agrilus kuldjensis Obenberger, 1936 Agrilus kusakievici Obenberger, 1936 Agrilus lugovoi Stepanov, 1958 Agrilus machnovskii Stepanov, 1958 Agrilus macrotatopus Obenberger, 1930 Agrilus malloti Théry, 1930 Agrilus malyi Obenberger, 1936 Agrilus mentor Kerremans, 1912 Agrilus mongolorum Obenberger, 1936 Agrilus morawitzi Obenberger, 1936 Agrilus ningpoensis Obenberger, 1927 Agrilus obscurosericeus Obenberger, 1940 Agrilus olivaceidorsis Obenberger, 1917 Agrilus ornamentifer Obenberger, 1940 Agrilus peregrinus prasinulus Obenberger, 1924 Agrilus pidjinus Obenberger, 1924 Agrilus pliculipennis Obenberger, 1940 Agrilus priamus Kerremans, 1912 Agrilus przewalskii Obenberger, 1936 Agrilus pusillesculptus Obenberger, 1940 Agrilus robertii Chevrolat, 1840 Agrilus rossicus Obenberger, 1916 Agrilus sacchari Obenberger, 1940 Agrilus scutellatus Abeille de Perrin, 1897 Agrilus semirjeciae Obenberger, 1936 Agrilus sieversi Abeille de Perrin, 1897 Agrilus sikkimensis Obenberger, 1928 Agrilus smaragdifrons Ganglbauer, 1889 Agrilus sospes Lewis, 1893 Agrilus spinipennis Lewis, 1893 Agrilus suensoni Obenberger, 1940 Agrilus transbaicalensis Obenberger, 1935 Agrilus trinotatus Saunders, 1873 Agrilus tyndarus Kerremans, 1913 Agrilus unguiculosus Obenberger, 1936 Agrilus uzbekistanus Stepanov, 1958 Agrilus viridicaerulans esfandiarinus Davatchi, 1958 Agrilus viridis cernyi Obenberger, 1925 Agrilus viridis krogerusi Obenberger, 1924 Agrilus viridis poppiusi Obenberger, 1924 Agrilus yunnanus Obenberger, 1927 Agrilus zigzag Marseul, 1866 Buprestis (Agrilus) capreae Chevrolat, 1838

270

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

APPENDIX II. SYNOPTIC CATALOG OF EAST ASIAN AGRILUS All names are listed in the alphabetical order. The new nomeclatural and taxonomic acts are highlighted in bold. adelphinus Kerremans, 1895 egorovi Alexeev, 1989 nigrocoerulans Obenberger, 1924 nonfriedanus Obenberger, 1923 nonfriedi Obenberger, 1914 panhensis Baudon, 1968 syn. nov.

adspersus (Kerremans, 1890) aerosus Ganglbauer, 1889 agnatus Kerremans, 1892 alashanensis Obenberger, 1936 rozhkovi Alexeev, 1964 steppensis Cobos, 1968

alesi Obenberger, 1935 aritai Tôyama, 1985 syn. nov. sacchari Obenberger, 1940 syn. nov.

alutaceicollis Obenberger, 1930 andrusi Baudon, 1968 pseudograminis Jendek, 2007 syn. nov.

arsenevi Jendek, 2009 asahinai Kurosawa, 1956 pseudorotundicollis Kurosawa, 1956 stepanovi Alexeev, 1979

ashlocki Baudon, 1968 asiaticus Kerremans, 1898 hauserellus Obenberger, 1935 igai Kurosawa, 1963 panphilovi Alexeev, 1959 planefasciatus Obenberger, 1936

aureofasciatus Jendek sp. nov. aurigaster Jendek sp. nov. aurinotus Jendek sp. nov. auristernum Obenberger, 1924 aurithorax Jendek sp. nov. auriventris Saunders, 1873 fleutiauxi Bourgoin, 1922 syn. nov. graptelytrus Obenberger, 1914 syn. nov. pidjinus Obenberger, 1924 syn. nov. Unavailable: citri Matsumura tsonglokianus Kerremans

auropictus Kerremans, 1912 chopardi Descarpentries & Villiers, 1963 syn. nov. kanohi Kurosawa, 1954 syn. nov. ryukyuensis Tôyama, 1985 syn. nov. Unavailable: marchali Obenberger

bai Jendek sp. nov. baroni Gory & Laporte, 1839 mopsus Théry, 1930

barrati Descarpentries & Villiers, 1963 betuleti (Ratzeburg, 1837) foveicollis Marseul, 1869 impressicollis Marseul, 1866 saliceti Obenberger, 1924 Unavailable: cyaneoatratus Obenberger, 1935 fuegneri Obenberger, 1917 schuleri Schaefer, 1936

tristis Fügner, 1891

bicuspidatus Jendek sp. nov. bidentulus Ganglbauer, 1889 bituberculatus Jendek, 2007 blatteicollis Bourgoin, 1922 bocaki Jendek sp. nov. bonadonai Descarpentries & Villiers, 1963 bonnottei Bourgoin, 1925 brevinotus Jendek sp. nov. businskorum Jendek sp. nov. caligans Bourgoin, 1925 assamensis Obenberger, 1936 syn. nov.

caudicula Jendek sp. nov. chaetocoxalis Jendek sp. nov. charismaticus Jendek, 2000 chekiangensis Gebhardt, 1929 semivittatus Tôyama, 1985

chlorophyllus Abeille de Perrin, 1904 esfandiarinus Davatchi, 1958 pistaciophagus Alexeev & Kulinitsh, 1962 syn. nov.

chujoi Kurosawa, 1985 ciping Jendek sp. nov. compacticornis Jendek sp. nov. confinis Faldermann, 1835 conspersus Jendek sp. nov. convictor Descarpentries & Villiers, 1963 coomani Bourgoin, 1925 milletae Baudon, 1968 syn. nov.

cordicollis Jendek sp. nov. coreanus Obenberger, 1935 cupes Lewis, 1893 cuprescens cuprescens (Ménétriés, 1832) altaicola Obenberger, 1935 aurichalceus Redtenbacher, 1847 calcicola Obenberger, 1916 communis Obenberger, 1924 egenus Abeille de Perrin, 1895 epistomalis Abeille de Perrin, 1897 foveolatus Abeille de Perrin, 1897 krasai Obenberger, 1924 kuznecovi Obenberger, 1935 kuznecovinus Obenberger, 1936 lacrymans Abeille de Perrin, 1895 mokrzeckii Obenberger, 1927 obtusifera Schaefer, 1949 obtusus Abeille de Perrin, 1897 proximus Rey, 1891 rubicola Abeille de Perrin, 1897 Unavailable: antiquus Gavoy chalconatus Megerle chrysoderes Bedel chrysoderes Bétis fagi Weiss fagi Glover, 1878 indigacellus Obenberger, 1916 politus Weiss salicis Csiki

271

SYNOPTIC CATALOG OF EAST ASIAN AGRILUS viridis Seidlitz viridis Weiss

cyaneomicans Nonfried, 1892 cyaneoniger Saunders, 1873 ataman Obenberger, 1924 cyaneoniger Thomson, 1879 impressifrons Kiesenwetter, 1879 jamesi Jakobson, 1913 marquardti Obenberger, 1914 melanopterus Solsky, 1876 mikado Obenberger, 1924 Unavailable: cupreoviridis Lewis, 1893

cyaneovirens Bourgoin, 1922 cyanescens cyanescens (Ratzeburg, 1837) acuticornis Abeille de Perrin, 1897 amabilis Gory & Laporte, 1839 caeruleus (Rossi, 1792) cockerelli Fisher, 1925 fissifrons Abeille de Perrin, 1897 italicus Obenberger, 1920 kyselyi Obenberger, 1924 pooli Théry, 1936 sulcaticeps Abeille de Perrin, 1869 teriolensis Obenberger, 1916 Unavailable: amethystinus Schönherr combinator Obenberger, 1925 cuprescens Fleischer, 1934 cyaneus Gory & Laporte cyaneus Lacordaire elisae Gerini, 1953 labronicus Gerini, 1953 pseudroberti Fleischer, 1934 subcarpathicus Roubal, 1929 tyrolensis Porta, 1929 virens Schilsky, 1888

cyanipennis Gory & Laporte, 1839 agilis Kerremans, 1894 amabilicolor Obenberger, 1936 amabilis Kerremans, 1893 dalbergiae Théry, 1930 insignis Saunders, 1871 orientalis Harold, 1869 ornativentris Saunders, 1866

daimio Obenberger, 1936 dali Jendek, 2009 daur Jendek sp. nov. decoloratus Kerremans, 1892 afghanistanicus Alexeev, 1992 ajax Kerremans, 1912 alazon Lewis, 1893 syn. nov. lunlingensis Obenberger, 1958 octostictus Bourgoin, 1922 pyrrhus Kerremans, 1913

delphinensis Abeille de Perrin, 1897 eogenes Obenberger, 1930

densetomentosus Obenberger, 1940 dianthus Kerremans, 1892 coelicus Bourgoin, 1922 dorri Bourgoin, 1925 nodosus Kerremans, 1894

diao Jendek sp. nov. diaolin Jendek, 2001

discalis Saunders, 1873 diversornatus Jendek sp. nov. dureli Jendek sp. nov. ecarinatus Marseul, 1866 gracilicornis Ganglbauer, 1889 grassator Kerremans, 1903 Unavailable: daemonius Obenberger, 1920 wulffiusi Obenberger, 1936

emmerichi Obenberger, 1935 erythrostictus Bourgoin, 1922 barmensis Obenberger, 1936

esakii Kurosawa, 1964 toyoshimai Tôyama, 1988 syn. nov.

eunuchus Jendek sp. nov. euonymi Tôyama, 1985 yagii Tôyama, 1985

fareastensis Jendek, 1995 fevreae Descarpentries & Villiers, 1963 fissus Obenberger, 1917 lubricus Théry, 1942

fleischeri Obenberger, 1925 kochi Théry, 1942 kurosawai Obenberger, 1940 nipponicola Kurosawa, 1963 syn. nov. tscherepanovi Stepanov, 1954 Unavailable: coreicus Kurosawa, 1954

foliatus Jendek, 2007 foramenifer Jendek sp. nov. fornicatus Jendek sp. nov. fortunatus Lewis, 1893 Unavailable: tsurugianus Iga, 1947

friebi Obenberger, 1922 ainu Miwa & Chûjô, 1940 enisus Théry, 1942 kiangsuanus Obenberger, 1935 nalajchanus Cobos, 1968 syn. nov. vodaki Obenberger, 1936

fusciapex Jendek sp. nov. fusiformis Jendek sp. nov. ganglbaueri Semenov, 1891 albifrons Reitter, 1897

gaoligong Jendek, 2000 grisator Kerremans, 1893 gussakovskiji Alexeev, 1981 han Jendek sp. nov. hani Jendek sp. nov. hattorii Nakane, 1983 hmong Jendek sp. nov. holzschuhi Jendek, 1994 huashanus Jendek, 2001 hui Jendek sp. nov. hunanus Jendek, 2009 imasakai Tôyama, 1985 imitans Lewis, 1893 improcerus Jendek sp. nov. inamoenus Kerremans, 1892 ineptus Kerremans, 1892 insulicola Kerremans, 1912 suensoni Obenberger, 1940 syn. nov.

jaminae Baudon, 1968 japanocarinatus Ohmomo, 2002

272

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

jindrai Jendek sp. nov. kaluganus Obenberger, 1940 pseudoussuricola Alexeev, 1979 syn. nov.

kawarai Kurosawa, 1963 komareki komareki Obenberger, 1926 ronino Obenberger, 1935

komareki sapporoensis Obenberger, 1935 nom. rev. emisi Ohmomo, 2005 syn. nov.

kostali Jendek, 2007 koyoi Ohmomo, 2002 kuangcenensis Obenberger, 1940 kucerai Jendek sp. nov. kurumi Kurosawa, 1957 lacrima Jendek sp. nov. laetecyanescens Obenberger, 1940 lahu Jendek sp. nov. laticornis (Illiger, 1803) asperrimus Marseul, 1866 scaberrimus (Ratzeburg, 1837) sibiricola Obenberger, 1924 syn. nov. Unavailable: aceris Chevrolat angustulus Kerremans coerulescens Schilsky, 1888 diabolicus Obenberger, 1913 laticollis Kiesenwetter obscurocupreus Roubal, 1936 schilskyi Pic, 1909 subscaber Rey, 1891 vultaviensis Obenberger, 1917

latipalpis Jendek, 2007 lei Jendek sp. nov. liscapia Jendek, 2003 apicalis (Bourgoin, 1923)

lisu Jendek sp. nov. lobatus Jendek sp. nov. loosdregti Baudon, 1965 lubopetri Jendek, 2000 lugu Jendek sp. nov. macrotatopus Obenberger, 1930 aridosus Obenberger, 1936 syn. nov.

maculifer Saunders, 1873 nagaoi Nakane, 1983 syn. nov.

madeci Baudon, 1968 mushanus Kurosawa, 1976 syn. nov. rokuyai Kurosawa, 1976 syn. nov.

magnus Jendek sp. nov. majzlani Jendek, 2007 mali Matsumura, 1924 jenissejensis Obenberger, 1924

malloti Théry, 1930 diolaus Obenberger, 1958 syn. nov.

mallotiellus Kurosawa, 1985 malloti Kurosawa, 1957

manchu Jendek sp. nov. mandala Jendek sp. nov. marginicollis Saunders, 1873 beppuensis Obenberger, 1936 hastuliferoides Gebhardt, 1929

miao Jendek sp. nov. mio Jendek sp. nov. miwai Obenberger, 1936 moerens Saunders, 1873 araxicola Abeille de Perrin, 1897 mandjuricus Obenberger, 1922

marinus Obenberger, 1922 paphius Théry, 1942 rotundicollis Saunders, 1873 Unavailable: vladivostokanus Obenberger, 1922

mongolorum Obenberger, 1936 morawitzi Obenberger, 1936 richteri Alexeev, 1975 syn. nov. Unavailable: pseudalbogularis Alexeev

muongoides Jendek sp. nov. muscarius Kerremans, 1895 komiyai Tôyama, 1987 seladon Obenberger, 1940 Unavailable: muscarinus Baudon

myopic Jendek, 2000 naxi Jendek sp. nov. nelsonodaimio Ohmomo, 2006 nepalensis Tôyama, 1988 nicolanus Obenberger, 1924 piscatorius Obenberger, 1935

ningpoensis Obenberger, 1927 nipponigena Obenberger, 1935 melanolucidus Ohmomo, 2005 syn. nov.

niveoguttatus Kerremans, 1892 nivosus Abeille de Perrin, 1900 candidiventris Obenberger, 1936 syn. nov. kuldjensis Obenberger, 1936 syn. nov.

notoclavus Jendek, 2000 obscuricollis Kiesenwetter, 1857 foveola Rey, 1891 obscurus Delherm de Larcenne, 1885 reyi Bauduer, 1878 savalae Stark, 1953 Unavailable: chlorodius Megerle coelestinus Schaefer, 1936 jirnensis Obenberger, 1937 mequignoni Obenberger, 1930 subauraticolor Obenberger, 1937

obscurosericeus Obenberger, 1940 odetteae Baudon, 1968 ogatai Ohmomo, 2002 olivaceidorsis Obenberger, 1917 nom. rev. duporti Bourgoin, 1922 syn. nov. ornamentifer Obenberger, 1940 syn. nov.

ornatoides Jendek sp. nov. paralleloides Jendek sp. nov. pecirkai Obenberger, 1916 pekinensis pekinensis Obenberger, 1924 boreooccidentalis Obenberger, 1936 cerskii Obenberger, 1936 charbinensis Théry, 1942 klapperichi Obenberger, 1940 Unavailable: aureosplendidus Obenberger, 1936

pekinensis accolus Jendek, 1995 peregrinus Kiesenwetter, 1879 brodjaga Obenberger, 1914

perroti Descarpentries & Villiers, 1963 pilosovittatus Saunders, 1873 pusillipubis Gebhardt, 1929 spesivcevi Obenberger, 1927 syn. nov.

273

SYNOPTIC CATALOG OF EAST ASIAN AGRILUS zemani Obenberger, 1925

plagiatus Ganglbauer, 1889 mentor Kerremans, 1912 syn. nov.

planipennis Fairmaire, 1888 feretrius Obenberger, 1936 marcopoli Obenberger, 1930 ulmi Kurosawa, 1956

plasoni plasoni Obenberger, 1917 jeanvoinei Descarpentries & Villiers, 1963

plavilstshikovi Alexeev, 1989 pleuralis Jendek sp. nov. pleurostigmus Jendek sp. nov. pliculipennis Obenberger, 1940 pratensis (Ratzeburg, 1837) chankae Obenberger, 1935 djukini Obenberger, 1935 syn. nov. fennicus Obenberger, 1936 praeclarus Krogerus, 1925 syn. nov. pseudocoeruleus Obenberger, 1930 robertii Chevrolat, 1840 Unavailable: aeneicolor Schaefer, 1946 auricollis Sturm, 1843 confusinellus Schaefer, 1949 confusus Schaefer, 1936 linearis Gory & Laporte linearis Paykull paracyaneus Schaefer, 1937 strigicollis Rey, 1891

presbys Jendek sp. nov. priamus Kerremans, 1912 proditor Obenberger, 1917 prolatus Jendek sp. nov. prosternalis Jendek sp. nov. przewalskii Obenberger, 1936 pseudobscuricollis Jendek, 2007 pseudoconstantini Alexeev, 1981 pseudolimoniastri Cobos, 1968 pseudosericans Obenberger, 1936 angulicollis Cobos, 1968

pterostigma Obenberger, 1927 pumi Jendek sp. nov. pusillesculptus Obenberger, 1940 qiang Jendek sp. nov. qinling Jendek, 2000 qinlingcola Jendek sp. nov. quadrisignatus Marseul, 1866 lama Obenberger, 1935 mongoliae Obenberger, 1922

quentini Descarpentries & Villiers, 1963 wilsoni Baudon, 1968 syn. nov.

recticollis Jendek sp. nov. reichardti Obenberger, 1935 ribbei Kiesenwetter, 1879 bakinensis Obenberger, 1935 corax Obenberger, 1917 freyi Théry, 1939 gracilipes Lewis, 1893 ignoratus Obenberger, 1924 iturupicus Alexeev, 1979 japonicus Kerremans, 1898 lewisiellus Kerremans, 1903 prinadai Fisher, 1925 taigicola Obenberger, 1924 tibialis Lewis, 1893

Unavailable: ingnoratus Obenberger, 1924 maderianus Obenberger, 1943

ribesi Schaefer, 1946 rivalieri Descarpentries & Villiers, 1963 rougeoti Descarpentries & Villiers, 1963 rudicollis Alexeev, 1979 sachalinicola Obenberger, 1940 tamanukii Stepanov, 1954

sagittifer Jendek sp. nov. salicis Frivaldszky, 1877 acutangulus Schaefer, 1949 alutaceus Obenberger, 1916 filigranus Obenberger, 1916 syn. nov. lugovoi Stepanov, 1958 syn. nov. Unavailable: acutangulus Théry, 1942 albisensis Obenberger, 1937 audrasi Schaefer, 1946 aurichalceus Abeille de Perrin boleslavensis Obenberger, 1920

satoi Kurosawa, 1954 schillhammeri Jendek sp. nov. sentis Jendek sp. nov. shan Jendek sp. nov. shaowensis Jendek, 1994 shashamboe Kurosawa, 1963 she Jendek sp. nov. shibatai Kurosawa, 1964 hokkaidoensis Tôyama, 1985

sibiricus sibiricus Obenberger, 1912 insuspectus Obenberger, 1924 turdus Théry, 1942

sibiricus fukushimenis Jendek, 1994 sichuanus Jendek sp. nov. sikkimensis Obenberger, 1928 sinensis sinensis Thomson, 1879 fairmairei Kerremans, 1892 syn. nov. obscuripennis Fairmaire, 1888 pseudimitans Miwa & Chûjô, 1940 Unavailable: chinensis Kerremans, 1903

sinensis splendidicollis Fairmaire, 1889 bayeri Obenberger, 1935 lameyi Obenberger, 1924

sinuatus sachalinensis Obenberger, 1935 stat. nov. yokoyamai Iga, 1955 syn. nov. zhelochovtsevi Alexeev, 1979 syn. nov.

siskai Jendek sp. nov. smaragdifrons Ganglbauer, 1889 kanssuanus Obenberger, 1930 syn. nov.

smaragdinus smaragdinus Solsky, 1876 prasinulus Obenberger, 1924 Unavailable: epicyanus Obenberger, 1934

smaragdinus habros Jendek ssp. nov. smetanai Jendek sp. nov. sospes Lewis, 1893 kinoshitae Obenberger, 1936 quadristictulus Obenberger, 1940

soudeki Obenberger, 1926 sericeus Théry, 1942

souliei Jendek, 1994 spathulifer Jendek sp. nov.

274

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

spiculum Jendek sp. nov. spinipennis Lewis, 1893 dentiapex Obenberger, 1940

stigma Jendek sp. nov. subauratus amurensis Obenberger, 1922 demuthi Théry, 1942 Unavailable: burjaticus Obenberger, 1924 euchryseus Obenberger, 1924 linearioides Obenberger, 1922 lineimargo Obenberger, 1924

subrobustus Saunders, 1873 albizziae Kurosawa, 1963 boisreymondi Alexeev & Bílý, 1980 kumamotoensis Obenberger, 1935 kumamotoi Tôyama, 1987 kurosawus Bellamy, 1998 malyi Obenberger, 1936 syn. nov. sorocinus Kerremans, 1914

subtilipilis Obenberger, 1940 sudai Kurosawa, 1985 suginoi Tôyama, 1985 sugiurai Jendek, 2007 sulcinotus Jendek, 2007 suturicuspidatus Jendek, 2007 suvorovi Obenberger, 1935 brussae Obenberger, 1956 populneus Schaefer, 1946 Unavailable: charbinensis Obenberger, 1935 cyanophilus Schaefer, 1946 pseudofagi Obenberger, 1956 raddei Obenberger, 1935 simoni Schaefer, 1961

tai Jendek sp. nov. takahashii Tôyama, 1988 tazoei Kurosawa, 1985 tempestivus Lewis, 1893 terracotta Jendek sp. nov. thibetanus Obenberger, 1928 tokyoensis Kurosawa, 1985 tonkineus Kerremans, 1895 blatteiceps Bourgoin, 1925

trinotatus Saunders, 1873 truncatus Jendek, 2007 tschitscherini Semenov, 1895 attila Obenberger, 1930 syn. nov. gerschuni Stepanov, 1958 syn. nov. lubischevi Stepanov, 1958 syn. nov. machnovskii Stepanov, 1958 syn. nov. uzbekistanus Stepanov, 1958 syn. nov.

tsushimanus Kurosawa, 1963 tsutsumiuchii Ohmomo, 2006 tuber Jendek sp. nov. tyndarus Kerremans, 1913 omissulus Obenberger, 1935 syn. nov. omissus Kerremans, 1914 therondi Descarpentries & Villiers, 1963 syn. nov.

uenoi Kurosawa, 1963 ulaangomiensis Cobos, 1972 emeljanovi Alexeev, 1975 subelegans Cobos, 1972

ulmicola Jendek sp. nov. unguiculosus Obenberger, 1936 gobicola Cobos, 1968

ussuricola Obenberger, 1924 anomalus Théry, 1942 lasiolus Obenberger, 1935

varius Kerremans, 1895 ventricosus Fairmaire, 1888 kalganicus Obenberger, 1935 toyamai Kurosawa, 1985

viduus Kerremans, 1914 cardaces Obenberger, 1936 syn. nov. chinganicus Obenberger, 1922 syn. nov. hoscheki Obenberger, 1916 syn. nov. subviduus Kurosawa, 1957 syn. nov.

vigil Jendek sp. nov. violaceus Nonfried, 1892 nonfriedi Kerremans, 1900

viridiobscurus Saunders, 1873 viridis (Linné, 1758) aubei Gory & Laporte, 1839 bicolor (Schrank, 1798) bicolor Redtenbacher, 1847 brevitarsis Lewis, 1893 syn. nov. brydli Obenberger, 1924 syn. nov. caenus Obenberger, 1924 capreae (Chevrolat, 1838) cernyi Obenberger, 1925 cojbalsanensis Cobos, 1968 transfer in syn. darwinii Wollaston, 1857 distinguendus Gory & Laporte, 1839 fagi (Ratzeburg, 1837) filiformis (Herbst, 1801) gebleri Obenberger, 1924 syn. nov. imbellis Kerremans, 1892 syn. nov. importunus Kerremans, 1903 syn. nov. indulgens Kerremans, 1897 kiricenkoi Obenberger, 1935 transfer in syn. krogerusi Obenberger, 1924 lewisi Kerremans, 1903 linearis (Schrank, 1781) linearis (Fabricius, 1792) lineatus (Schrank, 1782) littlei Curtis, 1840 melantatus Obenberger, 1924 montanellus Obenberger, 1916 nocivus (Ratzeburg, 1837) poppiusi Obenberger, 1924 syn. nov. pusztae Obenberger, 1936 quercinus Redtenbacher, 1847 rosaceus (Scopoli, 1763) rudis Abeille de Perrin, 1897 salicivola Kurosawa, 1963 syn. nov. serraticornis (Scopoli, 1763) towadensis Miwa & Chûjô, 1940 syn. nov. transbaicalensis Obenberger, 1935 syn. nov. vernadskii Obenberger, 1927 viridipennis Gory & Laporte, 1839 Unavailable: aceris Alexeev, 1957 albisetulosus Brundin androgynus Théry, 1942 anthraculus Obenberger, 1935 ater Fabricius ater Kiesenwetter basurmanovae Stepanov belial Obenberger, 1917

275

SYNOPTIC CATALOG OF EAST ASIAN AGRILUS cepelaki Obenberger coeruleus Herbst cyaneus Olivier erronesculus Obenberger, 1935 fagiformis Obenberger, 1935 ignotus Schaefer, 1946 kamtschaticus Stepanov linearis Panzer mixtulus Obenberger, 1924 novicus Westhoff, 1881 obtusus Obenberger praeclarus Rey, 1891 purpureicolor Pic, 1918 ragusanus Obenberger, 1922 rhodanensis Schaefer, 1934 salicicola Roubal, 1936

virescens Besser viridicupreus (Geoffroy, 1764)

volkovitshi Jendek, 2007 voriseki Jendek, 1995 yakushimensis Tôyama, 1985 yamabusi Miwa & Chûjô, 1940 nakanei Kurosawa, 1963 ohmomoi Tôyama, 1985

yamajii Tôyama, 1985 yamawakii Kurosawa, 1957 yunnanicola Obenberger, 1936 yunnanus Obenberger, 1927 zanthoxylumi Li Meng Lou, 1989 zhongdian Jendek, 2009 zhuang Jendek sp. nov. zhui Jendek sp. nov.

276

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

APPENDIX III. CHECKLIST OF SPECIES-GROUP NAMES (AVAILABLE AND UNAVAILABLE) aceris Chevrolat unavailable syn. of laticornis (Illiger, 1803)

aritai Tôyama, 1985 syn. of alesi Obenberger, 1935

aceris Alexeev, 1957 unavailable syn. of viridis (Linné, 1758)

arsenevi Jendek, 2009

acutangulus Théry, 1942 unavailable syn. of salicis Frivaldszky, 1877

asahinai Kurosawa, 1956

acutangulus Schaefer, 1949 syn. of salicis Frivaldszky, 1877

asiaticus Kerremans, 1898

acuticornis Abeille de Perrin, 1897 syn. of cyanescens cyanescens (Ratzeburg, 1837)

asperrimus Marseul, 1866 syn. of laticornis (Illiger, 1803)

adelphinus Kerremans, 1895 adspersus (Kerremans, 1890) aeneicolor Schaefer, 1946 unavailable syn. of pratensis (Ratzeburg, 1837) aerosus Ganglbauer, 1889 afghanistanicus Alexeev, 1992 syn. of decoloratus Kerremans, 1892 agilis Kerremans, 1894 syn. of cyanipennis Gory & Laporte, 1839 agnatus Kerremans, 1892 ainu Miwa & Chûjô, 1940 syn. of friebi Obenberger, 1922 ajax Kerremans, 1912 syn. of decoloratus Kerremans, 1892 alashanensis Obenberger, 1936 alazon Lewis, 1893 syn. of decoloratus Kerremans, 1892

ashlocki Baudon, 1968

assamensis Obenberger, 1936 syn. of caligans Bourgoin, 1925 ataman Obenberger, 1924 syn. of cyaneoniger Saunders, 1873 ater Fabricius (unavailable syn. of viridis (Linné, 1758)) ater Kiesenwetter (unavailable syn. of viridis (Linné, 1758)) attila Obenberger, 1930 syn. of tschitscherini Semenov, 1895 aubei Gory & Laporte, 1839 syn. of viridis (Linné, 1758) audrasi Schaefer, 1946 unavailable syn. of salicis Frivaldszky, 1877 aureofasciatus Jendek sp. nov. aureosplendidus Obenberger, 1936 unavailable syn. of pekinensis pekinensis Obenberger, 1924 aurichalceus Abeille de Perrin unavailable syn. of salicis Frivaldszky, 1877

albifrons Reitter, 1897 syn. of ganglbaueri Semenov, 1891

aurichalceus Redtenbacher, 1847 syn. of cuprescens cuprescens (Ménétriés, 1832)

albisensis Obenberger, 1937 unavailable syn. of salicis Frivaldszky, 1877

auricollis Sturm, 1843 unavailable syn. of pratensis (Ratzeburg, 1837)

albisetulosus Brundin unavailable syn. of viridis (Linné, 1758)

aurigaster Jendek sp. nov.

albizziae Kurosawa, 1963 syn. of subrobustus Saunders, 1873

aurinotus Jendek sp. nov.

alesi Obenberger, 1935

auristernum Obenberger, 1924

altaicola Obenberger, 1935 syn. of cuprescens cuprescens (Ménétriés, 1832)

aurithorax Jendek sp. nov.

alutaceicollis Obenberger, 1930

auropictus Kerremans, 1912

alutaceus Obenberger, 1916 syn. of salicis Frivaldszky, 1877

bai Jendek sp. nov.

amabilicolor Obenberger, 1936 syn. of cyanipennis Gory & Laporte, 1839

bakinensis Obenberger, 1935 syn. of ribbei Kiesenwetter, 1879

amabilis Gory & Laporte, 1839 syn. of cyanescens cyanescens (Ratzeburg, 1837)

baroni Gory & Laporte, 1839

amabilis Kerremans, 1893 syn. of cyanipennis Gory & Laporte, 1839

auriventris Saunders, 1873

barmensis Obenberger, 1936 syn. of erythrostictus Bourgoin, 1922 barrati Descarpentries & Villiers, 1963 basurmanovae Stepanov unavailable syn. of viridis (Linné, 1758)

amethystinus Schönherr unavailable syn. of cyanescens cyanescens (Ratzeburg, 1837)

bayeri Obenberger, 1935 syn. of sinensis splendidicollis Fairmaire, 1889

androgynus Théry, 1942 unavailable syn. of viridis (Linné, 1758)

belial Obenberger, 1917 unavailable syn. of viridis (Linné, 1758)

andrusi Baudon, 1968

beppuensis Obenberger, 1936 syn. of marginicollis Saunders, 1873

angulicollis Cobos, 1968 syn. of pseudosericans Obenberger, 1936

betuleti (Ratzeburg, 1837)

angustulus Kerremans unavailable syn. of laticornis (Illiger, 1803)

bicolor Redtenbacher, 1847 syn. of viridis (Linné, 1758)

anomalus Théry, 1942 syn. of ussuricola Obenberger, 1924

bicuspidatus Jendek sp. nov.

anthraculus Obenberger, 1935 unavailable syn. of viridis (Linné, 1758)

bidentulus Ganglbauer, 1889

antiquus Gavoy unavailable syn. of cuprescens cuprescens (Ménétriés, 1832)

blatteiceps Bourgoin, 1925 syn. of tonkineus Kerremans, 1895

apicalis (Bourgoin, 1923) syn. of liscapia Jendek, 2003 araxicola Abeille de Perrin, 1897 syn. of moerens Saunders, 1873 aridosus Obenberger, 1936 syn. of macrotatopus Obenberger, 1930

bicolor (Schrank, 1798) syn. of viridis (Linné, 1758)

bituberculatus Jendek, 2007 blatteicollis Bourgoin, 1922 bocaki Jendek sp. nov. boisreymondi Alexeev & Bílý, 1980 syn. of subrobustus Saunders, 1873

277

CHECKLIST OF SPECIES-GROUP NAMES boleslavensis Obenberger, 1920 unavailable syn. of salicis Frivaldszky, 1877

coelestinus Schaefer, 1936 unavailable syn. of obscuricollis Kiesenwetter, 1857

bonadonai Descarpentries & Villiers, 1963

coelicus Bourgoin, 1922 syn. of dianthus Kerremans, 1892

bonnottei Bourgoin, 1925

coerulescens Schilsky, 1888 unavailable syn. of laticornis (Illiger, 1803)

boreooccidentalis Obenberger, 1936 syn. of pekinensis pekinensis Obenberger, 1924

coeruleus Herbst unavailable syn. of viridis (Linné, 1758)

brevinotus Jendek sp. nov.

cojbalsanensis Cobos, 1968 syn. of viridis (Linné, 1758)

brevitarsis Lewis, 1893 syn. of viridis (Linné, 1758)

combinator Obenberger, 1925 unavailable syn. of cyanescens cyanescens (Ratzeburg, 1837)

brodjaga Obenberger, 1914 syn. of peregrinus Kiesenwetter, 1879 brussae Obenberger, 1956 syn. of suvorovi Obenberger, 1935 brydli Obenberger, 1924 syn. of viridis (Linné, 1758) burjaticus Obenberger, 1924 unavailable syn. of subauratus amurensis Obenberger, 1922 businskorum Jendek sp. nov. caenus Obenberger, 1924 syn. of viridis (Linné, 1758) caeruleus (Rossi, 1792) syn. of cyanescens cyanescens (Ratzeburg, 1837) calcicola Obenberger, 1916 syn. of cuprescens cuprescens (Ménétriés, 1832)

communis Obenberger, 1924 syn. of cuprescens cuprescens (Ménétriés, 1832) compacticornis Jendek sp. nov. confinis Faldermann, 1835 confusinellus Schaefer, 1949 unavailable syn. of pratensis (Ratzeburg, 1837) confusus Schaefer, 1936 unavailable syn. of pratensis (Ratzeburg, 1837) conspersus Jendek sp. nov. convictor Descarpentries & Villiers, 1963

caligans Bourgoin, 1925

coomani Bourgoin, 1925

candidiventris Obenberger, 1936 syn. of nivosus Abeille de Perrin, 1900

corax Obenberger, 1917 syn. of ribbei Kiesenwetter, 1879

capreae (Chevrolat, 1838) syn. of viridis (Linné, 1758)

coreanus Obenberger, 1935

cardaces Obenberger, 1936 syn. of viduus Kerremans, 1914 caudicula Jendek sp. nov.

coreicus Kurosawa, 1954 unavailable syn. of fleischeri Obenberger, 1925

cepelaki Obenberger unavailable syn. of viridis (Linné, 1758)

cupes Lewis, 1893

cernyi Obenberger, 1925 syn. of viridis (Linné, 1758)

cupreoviridis Lewis, 1893 unavailable syn. of cyaneoniger Saunders, 1873

cerskii Obenberger, 1936 syn. of pekinensis pekinensis Obenberger, 1924 chaetocoxalis Jendek sp. nov. chalconatus Megerle unavailable syn. of cuprescens cuprescens (Ménétriés, 1832) chankae Obenberger, 1935 syn. of pratensis (Ratzeburg, 1837) charbinensis Obenberger, 1935 unavailable syn. of suvorovi Obenberger, 1935

cordicollis Jendek sp. nov.

cuprescens (Ménétriés, 1832) cuprescens Fleischer, 1934 unavailable syn. of cyanescens cyanescens (Ratzeburg, 1837) cyaneoatratus Obenberger, 1935 unavailable syn. of betuleti (Ratzeburg, 1837) cyaneomicans Nonfried, 1892 cyaneoniger Thomson, 1879 syn. of cyaneoniger Saunders, 1873

charbinensis Théry, 1942 syn. of pekinensis pekinensis Obenberger, 1924

cyaneoniger Saunders, 1873

charismaticus Jendek, 2000

cyanescens (Ratzeburg, 1837)

chekiangensis Gebhardt, 1929

cyaneus Lacordaire unavailable syn. of cyanescens cyanescens (Ratzeburg, 1837)

chinensis Kerremans, 1903 unavailable syn. of sinensis sinensis Thomson, 1879 chinganicus Obenberger, 1922 syn. of viduus Kerremans, 1914

cyaneovirens Bourgoin, 1922

cyaneus Gory & Laporte unavailable syn. of cyanescens cyanescens (Ratzeburg, 1837)

chlorodius Megerle unavailable syn. of obscuricollis Kiesenwetter, 1857

cyaneus Olivier unavailable syn. of viridis (Linné, 1758)

chlorophyllus Abeille de Perrin, 1904

cyanophilus Schaefer, 1946 unavailable syn. of suvorovi Obenberger, 1935

chopardi Descarpentries & Villiers, 1963 syn. of auropictus Kerremans, 1912 chrysoderes Bedel unavailable syn. of cuprescens cuprescens (Ménétriés, 1832)

cyanipennis Gory & Laporte, 1839

daemonius Obenberger, 1920 unavailable syn. of ecarinatus Marseul, 1866 daimio Obenberger, 1936

chrysoderes Bétis unavailable syn. of cuprescens cuprescens (Ménétriés, 1832)

dalbergiae Théry, 1930 syn. of cyanipennis Gory & Laporte, 1839

chujoi Kurosawa, 1985

darwinii Wollaston, 1857 syn. of viridis (Linné, 1758)

ciping Jendek sp. nov.

daur Jendek sp. nov.

citri Matsumura unavailable syn. of auriventris Saunders, 1873

decoloratus Kerremans, 1892

cockerelli Fisher, 1925 syn. of cyanescens cyanescens (Ratzeburg, 1837)

delphinensis Abeille de Perrin, 1897

dali Jendek, 2009

278

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

demuthi Théry, 1942 syn. of subauratus amurensis Obenberger, 1922

feretrius Obenberger, 1936 syn. of planipennis Fairmaire, 1888

densetomentosus Obenberger, 1940

filiformis (Herbst, 1801) syn. of viridis (Linné, 1758)

dentiapex Obenberger, 1940 syn. of spinipennis Lewis, 1893

filigranus Obenberger, 1916 syn. of salicis Frivaldszky, 1877

diabolicus Obenberger, 1913 unavailable syn. of laticornis (Illiger, 1803)

fissifrons Abeille de Perrin, 1897 syn. of cyanescens cyanescens (Ratzeburg, 1837)

dianthus Kerremans, 1892

fissus Obenberger, 1917

diao Jendek sp. nov.

fleischeri Obenberger, 1925

fevreae Descarpentries & Villiers, 1963

diaolin Jendek, 2001

fleutiauxi Bourgoin, 1922 syn. of auriventris Saunders, 1873

diolaus Obenberger, 1958 syn. of malloti Théry, 1930

foliatus Jendek, 2007

discalis Saunders, 1873

foramenifer Jendek sp. nov.

distinguendus Gory & Laporte, 1839 syn. of viridis (Linné, 1758)

fornicatus Jendek sp. nov.

diversornatus Jendek sp. nov.

fortunatus Lewis, 1893

djukini Obenberger, 1935 syn. of pratensis (Ratzeburg, 1837)

foveicollis Marseul, 1869 syn. of betuleti (Ratzeburg, 1837

dorri Bourgoin, 1925 syn. of dianthus Kerremans, 1892

foveola Rey, 1891 syn. of obscuricollis Kiesenwetter, 1857

duporti Bourgoin, 1922 syn. of olivaceidorsis Obenberger, 1917 dureli Jendek sp. nov.

foveolatus Abeille de Perrin, 1897 syn. of cuprescens cuprescens (Ménétriés, 1832)

ecarinatus Marseul, 1866

freyi Théry, 1939 syn. of ribbei Kiesenwetter, 1879

egenus Abeille de Perrin, 1895 syn. of cuprescens cuprescens (Ménétriés, 1832)

friebi Obenberger, 1922

egorovi Alexeev, 1989 syn. of adelphinus Kerremans, 1895

fuegneri Obenberger, 1917 unavailable syn. of betuleti (Ratzeburg, 1837)

elisae Gerini, 1953 unavailable syn. of cyanescens cyanescens (Ratzeburg, 1837)

fusciapex Jendek sp. nov.

emeljanovi Alexeev, 1975 syn. of ulaangomiensis Cobos, 1972

ganglbaueri Semenov, 1891

emisi Ohmomo, 2005 syn. of komareki sapporoensis Obenberger, 1935

gaoligong Jendek, 2000

emmerichi Obenberger, 1935 enisus Théry, 1942 syn. of friebi Obenberger, 1922 eogenes Obenberger, 1930 syn. of delphinensis Abeille de Perrin, 1897 epicyanus Obenberger, 1934 unavailable syn. of smaragdinus smaragdinus Solsky, 1876

fusiformis Jendek sp. nov.

gebleri Obenberger, 1924 syn. of viridis (Linné, 1758) gerschuni Stepanov, 1958 syn. of tschitscherini Semenov, 1895 gobicola Cobos, 1968 syn. of unguiculosus Obenberger, 1936 gracilicornis Ganglbauer, 1889 syn. of ecarinatus Marseul, 1866 gracilipes Lewis, 1893 syn. of ribbei Kiesenwetter, 1879 graptelytrus Obenberger, 1914 syn. of auriventris Saunders, 1873

epistomalis Abeille de Perrin, 1897 syn. of cuprescens cuprescens (Ménétriés, 1832)

grassator Kerremans, 1903 syn. of ecarinatus Marseul, 1866

erronesculus Obenberger, 1935 unavailable syn. of viridis (Linné, 1758)

gussakovskiji Alexeev, 1981

erythrostictus Bourgoin, 1922

hani Jendek sp. nov.

esakii Kurosawa, 1964

hastuliferoides Gebhardt, 1929 (syn. of marginicollis Saunders, 1873)

esfandiarinus Davatchi, 1958 syn. of chlorophyllus Abeille de Perrin, 1904 euchryseus Obenberger, 1924 unavailable syn. of subauratus amurensis Obenberger, 1922 eunuchus Jendek sp. nov. euonymi Tôyama, 1985 fagi (Ratzeburg, 1837) syn. of viridis (Linné, 1758) fagi Weiss unavailable syn. of cuprescens cuprescens (Ménétriés, 1832) fagi Glover, 1878 unavailable syn. of cuprescens cuprescens (Ménétriés, 1832)

grisator Kerremans, 1893 han Jendek sp. nov.

hattorii Nakane, 1983 hauserellus Obenberger, 1935 syn. of asiaticus Kerremans, 1898 hmong Jendek sp. nov. hokkaidoensis Tôyama, 1985 syn. of shibatai Kurosawa, 1964 holzschuhi Jendek, 1994 hoscheki Obenberger, 1916 syn. of viduus Kerremans, 1914 huashanus Jendek, 2001 hui Jendek sp. nov. hunanus Jendek, 2009

fagiformis Obenberger, 1935 unavailable syn. of viridis (Linné, 1758)

igai Kurosawa, 1963 syn. of asiaticus Kerremans, 1898

fairmairei Kerremans, 1892 syn. of sinensis sinensis Thomson, 1879

ignotus Schaefer, 1946 unavailable syn. of viridis (Linné, 1758)

fareastensis Jendek, 1995

imbellis Kerremans, 1892 syn. of viridis (Linné, 1758)

fennicus Obenberger, 1936 syn. of pratensis (Ratzeburg, 1837)

imitans Lewis, 1893

ignoratus Obenberger, 1924 syn. of ribbei Kiesenwetter, 1879 imasakai Tôyama, 1985

279

CHECKLIST OF SPECIES-GROUP NAMES importunus Kerremans, 1903 syn. of viridis (Linné, 1758) impressicollis Marseul, 1866 syn. of betuleti (Ratzeburg, 1837)

kumamotoensis Obenberger, 1935 syn. of subrobustus Saunders, 1873

impressifrons Kiesenwetter, 1879 syn. of cyaneoniger Saunders, 1873

kumamotoi Tôyama, 1987 syn. of subrobustus Saunders, 1873

improcerus Jendek sp. nov.

kurosawus Bellamy, 1998 syn. of subrobustus Saunders, 1873

inamoenus Kerremans, 1892

kurumi Kurosawa, 1957

indigacellus Obenberger, 1916 unavailable syn. of cuprescens cuprescens (Ménétriés, 1832)

kusakievici Obenberger, 1936 syn. of zigzag Marseul, 1866

indulgens Kerremans, 1897 syn. of viridis (Linné, 1758) ineptus Kerremans, 1892

kurosawai Obenberger, 1940 syn. of fleischeri Obenberger, 1925

kuznecovi Obenberger, 1935 syn. of cuprescens cuprescens (Ménétriés, 1832)

ribbei

kuznecovinus Obenberger, 1936 syn. of cuprescens cuprescens (Ménétriés, 1832)

insignis Saunders, 1871 syn. of cyanipennis Gory & Laporte, 1839

kyselyi Obenberger, 1924 syn. of cyanescens cyanescens (Ratzeburg, 1837)

ingnoratus Obenberger, Kiesenwetter, 1879

1924

unavailable

syn.

of

insulicola Kerremans, 1912

labronicus Gerini, 1953 unavailable syn. of cyanescens cyanescens (Ratzeburg, 1837)

insuspectus Obenberger, 1924 syn. of sibiricus sibiricus Obenberger, 1912

lacrima Jendek sp. nov.

italicus Obenberger, 1920 syn. of cyanescens cyanescens (Ratzeburg, 1837)

lacrymans Abeille de Perrin, 1895 syn. of cuprescens cuprescens (Ménétriés, 1832)

iturupicus Alexeev, 1979 syn. of ribbei Kiesenwetter, 1879

laetecyanescens Obenberger, 1940

jamesi Jakobson, 1913 syn. of cyaneoniger Saunders, 1873

lahu Jendek sp. nov.

jaminae Baudon, 1968

lama Obenberger, 1935 syn. of quadrisignatus Marseul, 1866

japanocarinatus Ohmomo, 2002

lameyi Obenberger, 1924 syn. of sinensis splendidicollis Fairmaire, 1889

japonicus Kerremans, 1898(syn. of ribbei Kiesenwetter, 1879 jeanvoinei Descarpentries & Villiers, 1963 syn. of plasoni plasoni Obenberger, 1917

lasiolus Obenberger, 1935 syn. of ussuricola Obenberger, 1924) laticollis Kiesenwetter unavailable syn. of laticornis (Illiger, 1803)

jenissejensis Obenberger, 1924 syn. of mali Matsumura, 1924

laticornis (Illiger, 1803)

jindrai Jendek sp. nov.

latipalpis Jendek, 2007

jirnensis Obenberger, 1937 unavailable syn. of obscuricollis Kiesenwetter, 1857

lei Jendek sp. nov.

kalganicus Obenberger, 1935 syn. of ventricosus Fairmaire, 1888

lewisiellus Kerremans, 1903 syn. of ribbei Kiesenwetter, 1879

kaluganus Obenberger, 1940 kamtschaticus Stepanov unavailable syn. of viridis (Linné, 1758)

linearioides Obenberger, 1922 unavailable syn. of subauratus amurensis Obenberger, 1922

kanohi Kurosawa, 1954 syn. of auropictus Kerremans, 1912

linearis (Fabricius, 1792) syn. of viridis (Linné, 1758)

kanssuanus Obenberger, 1930 syn. of smaragdifrons Ganglbauer, 1889

linearis Paykull unavailable syn. of pratensis (Ratzeburg, 1837)

lewisi Kerremans, 1903 syn. of viridis (Linné, 1758)

kawarai Kurosawa, 1963

linearis Gory & Laporte unavailable syn. of pratensis (Ratzeburg, 1837)

kiangsuanus Obenberger, 1935 syn. of friebi Obenberger, 1922

linearis Panzer unavailable syn. of viridis (Linné, 1758)

kinoshitae Obenberger, 1936 syn. of sospes Lewis, 1893

linearis (Schrank, 1781) syn. of viridis (Linné, 1758)

kiricenkoi Obenberger, 1935 syn. of viridis (Linné, 1758)

lineatus (Schrank, 1782) syn. of viridis (Linné, 1758)

klapperichi Obenberger, 1940 syn. of pekinensis pekinensis Obenberger, 1924

lineimargo Obenberger, 1924 unavailable syn. of subauratus amurensis Obenberger, 1922

kochi Théry, 1942 syn. of fleischeri Obenberger, 1925

liscapia Jendek, 2003

komareki Obenberger, 1926

lisu Jendek sp. nov.

komareki sapporoensis Obenberger, 1935 ssp. of komareki Obenberger, 1926

littlei Curtis, 1840 syn. of viridis (Linné, 1758)

komiyai Tôyama, 1987 syn. of muscarius Kerremans, 1895

loosdregti Baudon, 1965

kostali Jendek, 2007

lubischevi Stepanov, 1958 syn. of tschitscherini Semenov, 1895

koyoi Ohmomo, 2002

lubopetri Jendek, 2000

krasai Obenberger, 1924 syn. of cuprescens cuprescens (Ménétriés, 1832)

lubricus Théry, 1942 syn. of fissus Obenberger, 1917

krogerusi Obenberger, 1924 syn. of viridis (Linné, 1758)

lugu Jendek sp. nov.

kuangcenensis Obenberger, 1940

lunlingensis Obenberger, 1958 syn. of decoloratus Kerremans, 1892

kucerai Jendek sp. nov. kuldjensis Obenberger, 1936 syn. of nivosus Abeille de Perrin, 1900

lobatus Jendek sp. nov.

lugovoi Stepanov, 1958 syn. of salicis Frivaldszky, 1877

machnovskii Stepanov, 1958 syn. of tschitscherini Semenov, 1895 macrotatopus Obenberger, 1930

280

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

maculifer Saunders, 1873

nepalensis Tôyama, 1988

madeci Baudon, 1968

nicolanus Obenberger, 1924

maderianus Obenberger, 1943 unavailable syn. of ribbei Kiesenwetter, 1879

nigrocoerulans Obenberger, 1924 syn. of adelphinus Kerremans, 1895

magnus Jendek sp. nov.

ningpoensis Obenberger, 1927

majzlani Jendek, 2007

nipponicola Kurosawa, 1963 syn. of fleischeri Obenberger, 1925

mali Matsumura, 1924

nipponigena Obenberger, 1935

malloti Théry, 1930

niveoguttatus Kerremans, 1892

malloti Kurosawa, 1957 syn. of mallotiellus Kurosawa, 1985

nivosus Abeille de Perrin, 1900

mallotiellus Kurosawa, 1985

nocivus (Ratzeburg, 1837) syn. of viridis (Linné, 1758)

malyi Obenberger, 1936 syn. of subrobustus Saunders, 1873

nodosus Kerremans, 1894 syn. of dianthus Kerremans, 1892

manchu Jendek sp. nov. mandala Jendek sp. nov.

nonfriedanus Obenberger, 1923 syn. of adelphinus Kerremans, 1895

mandjuricus Obenberger, 1922 syn. of moerens Saunders, 1873

nonfriedi Kerremans, 1900 syn. of violaceus Nonfried, 1892

marchali Obenberger unavailable syn. of auropictus Kerremans, 1912

nonfriedi Obenberger, 1914 syn. of adelphinus Kerremans, 1895

marcopoli Obenberger, 1930 syn. of planipennis Fairmaire, 1888

novicus Westhoff, 1881 unavailable syn. of viridis (Linné, 1758)

marginicollis Saunders, 1873

obscuricollis Kiesenwetter, 1857

marinus Obenberger, 1922 syn. of moerens Saunders, 1873 marquardti Obenberger, 1914 syn. of cyaneoniger Saunders, 1873

obscuripennis Fairmaire, 1888 syn. of sinensis sinensis Thomson, 1879

melanolucidus Ohmomo, 2005 syn. of nipponigena Obenberger, 1935

obscurocupreus Roubal, 1936 unavailable syn. of laticornis (Illiger, 1803)

melanopterus Solsky, 1876 syn. of cyaneoniger Saunders, 1873

obscurosericeus Obenberger, 1940

melantatus Obenberger, 1924 syn. of viridis (Linné, 1758)

obscurus Delherm de Larcenne, 1885 syn. of obscuricollis Kiesenwetter, 1857

mentor Kerremans, 1912 syn. of plagiatus Ganglbauer, 1889 mequignoni Obenberger, 1930 unavailable syn. of obscuricollis Kiesenwetter, 1857 miao Jendek sp. nov. mikado Obenberger, 1924 syn. of cyaneoniger Saunders, 1873 milletae Baudon, 1968 syn. of coomani Bourgoin, 1925 mio Jendek sp. nov. miwai Obenberger, 1936 mixtulus Obenberger, 1924 unavailable syn. of viridis (Linné, 1758) moerens Saunders, 1873 mokrzeckii Obenberger, 1927 syn. of cuprescens cuprescens (Ménétriés, 1832) mongoliae Obenberger, 1922 syn. of quadrisignatus Marseul, 1866 mongolorum Obenberger, 1936 montanellus Obenberger, 1916 syn. of viridis (Linné, 1758) mopsus Théry, 1930 syn. of baroni Gory & Laporte, 1839 morawitzi Obenberger, 1936 muongoides Jendek sp. nov. muscarinus Baudon unavailable syn. of muscarius Kerremans, 1895 muscarius Kerremans, 1895 mushanus Kurosawa, 1976 syn. of madeci Baudon, 1968 myopic Jendek, 2000 nagaoi Nakane, 1983 syn. of maculifer Saunders, 1873 nakanei Kurosawa, 1963 syn. of yamabusi Miwa & Chûjô, 1940 nalajchanus Cobos, 1968 syn. of friebi Obenberger, 1922 naxi Jendek sp. nov. nelsonodaimio Ohmomo, 2006

notoclavus Jendek, 2000

obtusifera Schaefer, 1949 syn. of cuprescens cuprescens (Ménétriés, 1832) obtusus Abeille de Perrin, 1897 syn. of cuprescens cuprescens (Ménétriés, 1832) obtusus Obenberger unavailable syn. of viridis (Linné, 1758) octostictus Bourgoin, 1922 syn. of decoloratus Kerremans, 1892 odetteae Baudon, 1968 ogatai Ohmomo, 2002 ohmomoi Tôyama, 1985 syn. of yamabusi Miwa & Chûjô, 1940 olivaceidorsis Obenberger, 1917 omissulus Obenberger, 1935 syn. of tyndarus Kerremans, 1913 omissus Kerremans, 1914 syn. of tyndarus Kerremans, 1913 orientalis Harold, 1869 syn. of cyanipennis Gory & Laporte, 1839 ornamentifer Obenberger, 1940 syn. of olivaceidorsis Obenberger, 1917 ornativentris Saunders, 1866 syn. of cyanipennis Gory & Laporte, 1839 ornatoides Jendek sp. nov. panhensis Baudon, 1968 syn. of adelphinus Kerremans, 1895 panphilovi Alexeev, 1959 syn. of asiaticus Kerremans, 1898 paphius Théry, 1942 syn. of moerens Saunders, 1873 paracyaneus Schaefer, 1937 unavailable syn. of pratensis (Ratzeburg, 1837) paralleloides Jendek sp. nov. pecirkai Obenberger, 1916 pekinensis Obenberger, 1924 pekinensis accolus Jendek, 1995 ssp. of pekinensis Obenberger, 1924 peregrinus Kiesenwetter, 1879 perroti Descarpentries & Villiers, 1963

281

CHECKLIST OF SPECIES-GROUP NAMES pidjinus Obenberger, 1924 syn. of auriventris Saunders, 1873

purpureicolor Pic, 1918 unavailable syn. of viridis (Linné, 1758)

pilosovittatus Saunders, 1873

pusillesculptus Obenberger, 1940

piscatorius Obenberger, 1935 syn. of nicolanus Obenberger, 1924

pusillipubis Gebhardt, 1929 syn. of pilosovittatus Saunders, 1873

pistaciophagus Alexeev & Kulinitsh, 1962 syn. of chlorophyllus Abeille de Perrin, 1904

pusztae Obenberger, 1936 syn. of viridis (Linné, 1758)

plagiatus Ganglbauer, 1889

qiang Jendek sp. nov.

planefasciatus Obenberger, 1936 syn. of asiaticus Kerremans, 1898

qinling Jendek, 2000

planipennis Fairmaire, 1888

pyrrhus Kerremans, 1913 syn. of decoloratus Kerremans, 1892

qinlingcola Jendek sp. nov. quadrisignatus Marseul, 1866

plasoni Obenberger, 1917

quadristictulus Obenberger, 1940 syn. of sospes Lewis, 1893

plavilstshikovi Alexeev, 1989

quentini Descarpentries & Villiers, 1963

pleuralis Jendek sp. nov.

quercinus Redtenbacher, 1847 syn. of viridis (Linné, 1758)

pleurostigmus Jendek sp. nov. pliculipennis Obenberger, 1940

raddei Obenberger, 1935 unavailable syn. of suvorovi Obenberger, 1935

politus Weiss unavailable syn. of cuprescens cuprescens (Ménétriés, 1832)

ragusanus Obenberger, 1922 unavailable syn. of viridis (Linné, 1758)

pooli Théry, 1936 syn. of cyanescens cyanescens (Ratzeburg, 1837)

recticollis Jendek sp. nov.

poppiusi Obenberger, 1924 syn. of viridis (Linné, 1758) populneus Schaefer, 1946 syn. of suvorovi Obenberger, 1935

reichardti Obenberger, 1935 reyi Bauduer, 1878 syn. of obscuricollis Kiesenwetter, 1857

praeclarus Rey, 1891 unavailable syn. of viridis (Linné, 1758)

rhodanensis Schaefer, 1934 unavailable syn. of viridis (Linné, 1758)

praeclarus Krogerus, 1925 syn. of pratensis (Ratzeburg, 1837)

ribbei Kiesenwetter, 1879

prasinulus Obenberger, 1924 syn. of smaragdinus smaragdinus Solsky, 1876

ribesi Schaefer, 1946

pratensis (Ratzeburg, 1837)

richteri Alexeev, 1975 syn. of morawitzi Obenberger, 1936 rivalieri Descarpentries & Villiers, 1963

presbys Jendek sp. nov.

robertii Chevrolat, 1840 syn. of pratensis (Ratzeburg, 1837)

priamus Kerremans, 1912

rokuyai Kurosawa, 1976 syn. of madeci Baudon, 1968

prinadai Fisher, 1925 syn. of ribbei Kiesenwetter, 1879) proditor Obenberger, 1917

ronino Obenberger, 1935 syn. of komareki komareki Obenberger, 1926

prolatus Jendek sp. nov.

rosaceus (Scopoli, 1763) syn. of viridis (Linné, 1758)

prosternalis Jendek sp. nov.

rotundicollis Saunders, 1873 syn. of moerens Saunders, 1873

proximus Rey, 1891 syn. of cuprescens cuprescens (Ménétriés, 1832)

rougeoti Descarpentries & Villiers, 1963

przewalskii Obenberger, 1936 pseudalbogularis Alexeev Obenberger, 1936

unavailable

syn.

of

morawitzi

pseudimitans Miwa & Chûjô, 1940 syn. of sinensis sinensis Thomson, 1879 pseudobscuricollis Jendek, 2007 pseudocoeruleus Obenberger, 1930 syn. of pratensis (Ratzeburg, 1837)

rozhkovi Alexeev, 1964 syn. of alashanensis Obenberger, 1936 rubicola Abeille de Perrin, 1897 syn. of cuprescens cuprescens (Ménétriés, 1832) rudicollis Alexeev, 1979 rudis Abeille de Perrin, 1897 syn. of viridis (Linné, 1758) ryukyuensis Tôyama, 1985 syn. of auropictus Kerremans, 1912 sacchari Obenberger, 1940 syn. of alesi Obenberger, 1935 sachalinicola Obenberger, 1940

pseudoconstantini Alexeev, 1981

sagittifer Jendek sp. nov.

pseudofagi Obenberger, 1956 unavailable syn. of suvorovi Obenberger, 1935

saliceti Obenberger, 1924 syn. of betuleti (Ratzeburg, 1837)

pseudograminis Jendek, 2007 syn. of andrusi Baudon, 1968

salicis Frivaldszky, 1877

pseudolimoniastri Cobos, 1968

salicis Csiki unavailable syn. of cuprescens cuprescens (Ménétriés, 1832)

pseudorotundicollis Kurosawa, 1956 syn. of asahinai Kurosawa, 1956

salicicola Roubal, 1936 unavailable syn. of viridis (Linné, 1758)

salicivola Kurosawa, 1963 syn. of viridis (Linné, 1758)

pseudosericans Obenberger, 1936

satoi Kurosawa, 1954

pseudoussuricola Alexeev, 1979 syn. of kaluganus Obenberger, 1940

savalae Stark, 1953 syn. of obscuricollis Kiesenwetter, 1857

pseudroberti Fleischer, 1934 unavailable syn. of cyanescens cyanescens (Ratzeburg, 1837)

schillhammeri Jendek sp. nov.

pterostigma Obenberger, 1927 pumi Jendek sp. nov.

scaberrimus (Ratzeburg, 1837) syn. of laticornis (Illiger, 1803) schilskyi Pic, 1909 unavailable syn. of laticornis (Illiger, 1803) schuleri Schaefer, 1936 unavailable syn. of betuleti (Ratzeburg, 1837)

282

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

seladon Obenberger, 1940 syn. of muscarius Kerremans, 1895

suensoni Obenberger, 1940 syn. of insulicola Kerremans, 1912

semivittatus Tôyama, 1985 syn. of chekiangensis Gebhardt, 1929

suginoi Tôyama, 1985

sentis Jendek sp. nov.

sugiurai Jendek, 2007

sericeus Théry, 1942 syn. of soudeki Obenberger, 1926 serraticornis (Scopoli, 1763) syn. of viridis (Linné, 1758)

sulcaticeps Abeille de Perrin, 1869 syn. of cyanescens cyanescens (Ratzeburg, 1837)

shan Jendek sp. nov.

sulcinotus Jendek, 2007

shaowensis Jendek, 1994

suturicuspidatus Jendek, 2007

shashamboe Kurosawa, 1963

suvorovi Obenberger, 1935

she Jendek sp. nov.

tai Jendek sp. nov.

shibatai Kurosawa, 1964

taigicola Obenberger, 1924 syn. of ribbei Kiesenwetter, 1879

sibiricola Obenberger, 1924 syn. of laticornis (Illiger, 1803)

takahashii Tôyama, 1988

sibiricus Obenberger, 1912

tamanukii Stepanov, 1954 syn. of sachalinicola Obenberger, 1940

sibiricus fukushimenis Jendek, 1994 ssp. of sibiricus Obenberger, 1912)

tazoei Kurosawa, 1985

sichuanus Jendek sp. nov.

teriolensis Obenberger, 1916 syn. of cyanescens cyanescens (Ratzeburg, 1837)

sikkimensis Obenberger, 1928 simoni Schaefer, 1961 unavailable syn. of suvorovi Obenberger, 1935 sinensis Thomson, 1879

tempestivus Lewis, 1893

terracotta Jendek sp. nov. therondi Descarpentries & Villiers, 1963 syn. of tyndarus Kerremans, 1913

sinensis splendidicollis Fairmaire, 1889 ssp. of sinensis Thomson, 1879

thibetanus Obenberger, 1928

sinuatus sachalinensis Obenberger, 1935 ssp. of sinuatus (Olivier, 1790)

tokyoensis Kurosawa, 1985

siskai Jendek sp. nov.

towadensis Miwa & Chûjô, 1940 syn. of viridis (Linné, 1758)

smaragdifrons Ganglbauer, 1889

toyamai Kurosawa, 1985 syn. of ventricosus Fairmaire, 1888

smaragdinus Solsky, 1876

toyoshimai Tôyama, 1988 syn. of esakii Kurosawa, 1964

smaragdinus habros Jendek ssp. nov. of smaragdinus Solsky, 1876

transbaicalensis Obenberger, 1935 syn. of viridis (Linné, 1758)

smetanai Jendek sp. nov.

trinotatus Saunders, 1873

sorocinus Kerremans, 1914 syn. of subrobustus Saunders, 1873

tristis Fügner, 1891 unavailable syn. of betuleti (Ratzeburg, 1837)

sospes Lewis, 1893

truncatus Jendek, 2007

soudeki Obenberger, 1926

tscherepanovi Stepanov, 1954 syn. of fleischeri Obenberger, 1925

souliei Jendek, 1994

tschitscherini Semenov, 1895

spathulifer Jendek sp. nov. spesivcevi Obenberger, 1927 syn. of pilosovittatus Saunders, 1873

tsonglokianus Kerremans unavailable syn. of A. auriventris Saunders, 1873

spiculum Jendek sp. nov.

tsurugianus Iga, 1947 unavailable syn. of fortunatus Lewis, 1893

spinipennis Lewis, 1893

tsushimanus Kurosawa, 1963

stepanovi Alexeev, 1979 syn. of asahinai Kurosawa, 1956

tsutsumiuchii Ohmomo, 2006

steppensis Cobos, 1968 syn. of alashanensis Obenberger, 1936

tuber Jendek sp. nov.

tibialis Lewis, 1893 syn. of ribbei Kiesenwetter, 1879 tonkineus Kerremans, 1895

stigma Jendek sp. nov.

turdus Théry, 1942 syn. of sibiricus sibiricus Obenberger, 1912

strigicollis Rey, 1891 unavailable syn. of pratensis (Ratzeburg, 1837)

tyndarus Kerremans, 1913

subauraticolor Obenberger, 1937 unavailable syn. of obscuricollis Kiesenwetter, 1857

tyrolensis Porta, 1929 unavailable syn. of cyanescens cyanescens (Ratzeburg, 1837) uenoi Kurosawa, 1963

subauratus amurensis Obenberger, 1922 ssp. of subauratus (Gebler, 1833)

ulaangomiensis Cobos, 1972

subcarpathicus Roubal, 1929 unavailable syn. of cyanescens cyanescens (Ratzeburg, 1837)

ulmicola Jendek sp. nov.

subelegans Cobos, 1972 syn. of ulaangomiensis Cobos, 1972 subrobustus Saunders, 1873 subscaber Rey, 1891 unavailable syn. of laticornis (Illiger, 1803) subtilipilis Obenberger, 1940 subviduus Kurosawa, 1957 syn. of viduus Kerremans, 1914 sudai Kurosawa, 1985

ulmi Kurosawa, 1956 syn. of planipennis Fairmaire, 1888 unguiculosus Obenberger, 1936 ussuricola Obenberger, 1924 uzbekistanus Stepanov, 1958 syn. of tschitscherini Semenov, 1895 varius Kerremans, 1895 ventricosus Fairmaire, 1888 vernadskii Obenberger, 1927 syn. of viridis (Linné, 1758)

283

CHECKLIST OF SPECIES-GROUP NAMES viduus Kerremans, 1914 vigil Jendek sp. nov.

1803)

violaceus Nonfried, 1892

wilsoni Baudon, 1968 syn. of quentini Descarpentries & Villiers, 1963

virens Schilsky, 1888 unavailable syn. of cyanescens cyanescens (Ratzeburg, 1837)

wulffiusi Obenberger, 1936 unavailable syn. of ecarinatus Marseul, 1866

virescens Besser unavailable syn. of viridis (Linné, 1758)

yagii Tôyama, 1985 syn. of euonymi Tôyama, 1985

viridicupreus (Geoffroy, 1764) unavailable syn. of viridis (Linné, 1758)

yakushimensis Tôyama, 1985

viridiobscurus Saunders, 1873

yamajii Tôyama, 1985

viridipennis Gory & Laporte, 1839 syn. of viridis (Linné, 1758)

yamawakii Kurosawa, 1957

viridis Weiss unavailable syn. of cuprescens cuprescens (Ménétriés, 1832)

yokoyamai Iga, 1955 syn. of sinuatus sachalinensis Obenberger, 1935

yamabusi Miwa & Chûjô, 1940

viridis (Linné, 1758)

yunnanicola Obenberger, 1936

viridis Seidlitz unavailable syn. of cuprescens cuprescens (Ménétriés, 1832)

yunnanus Obenberger, 1927

vladivostokanus Obenberger, 1922 unavailable syn. of moerens Saunders, 1873

zemani Obenberger, 1925 syn. of pilosovittatus Saunders, 1873

zanthoxylumi Li Meng Lou, 1989

vodaki Obenberger, 1936 syn. of friebi Obenberger, 1922

zhelochovtsevi Alexeev, 1979 syn. of sinuatus sachalinensis Obenberger, 1935

volkovitshi Jendek, 2007

zhongdian Jendek, 2009

voriseki Jendek, 1995

zhuang Jendek sp. nov.

vultaviensis Obenberger, 1917 unavailable syn. of laticornis (Illiger,

zhui Jendek sp. nov.

284

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

adelphinus Kerremans, 1895 aerosus Ganglbauer, 1889 agnatus Kerremans, 1892 alashanensis Obenberger, 1936 alesi Obenberger, 1935 alutaceicollis Obenberger, 1930 andrusi Baudon, 1968 arsenevi Jendek, 2009 asahinai Kurosawa, 1956 ashlocki Baudon, 1968 asiaticus Kerremans, 1898 aureofasciatus sp nov aurigaster sp nov aurinotus sp nov auristernum Obenberger, 1924 aurithorax sp nov auriventris Saunders, 1873 auropictus Kerremans, 1912 bai sp nov baroni Gory & Laporte, 1839 barrati Descarpentries & Villiers, 1963 betuleti (Ratzeburg, 1837) bicuspidatus sp nov bidentulus Ganglbauer, 1889 bituberculatus Jendek, 2007 blatteicollis Bourgoin, 1922 bocaki sp nov bonadonai Descarpentries & Villiers, 1963 bonnottei Bourgoin, 1925 brevinotus sp nov businskorum sp nov caligans Bourgoin, 1925 caudicula sp nov chaetocoxalis sp nov charismaticus Jendek, 2000 chekiangensis Gebhardt, 1929 chujoi Kurosawa, 1985 ciping sp nov compacticornis sp nov

Anhui

x

x

Beijing

Fujian

x x

x x

x

x x x x

x

x

Gansu

x

x

x

Guangdong Guangxi

x

x x x

x

Guizhou

x

x x

x

x

x

x

Hebei

Heilongjiang Henan

x

x

x

x

x x x

x x

x

x

Hubei

Hunan

Hong Kong

x

x x

x x

x x x

x

Jiangxi

x

x

x

x

Liaoning

x

Nei Mongol

x

x

x

Shandong

Shaanxi

x

x x

x x

x

x

x x

x x

Sichuan

Shanxi

x

x

x

x x

x

x

x x

x

x x x

Yunnan

Xizang

x

Zhejiang

x

x x

x

x

x

x

x

x x

x

x

x x

Kyushu

Honshu

Hokkaido

KOREA NORTH

x x

x

KOREA SOUTH

Tsushima

Shikoku

x

x x x x

x x x

x x x x x x

x

x

x x x

MONGOLIA

Buryatiya Respublika

x

Irkutskaya oblast'

Amurskaya oblast'

x x x x

x

x

Khabarovskiy kray

Jilin

Jiangsu

RUSSIA Magadanskaya oblast'

JAPAN Sakhalinskaya oblast'

x

Tyva Respublika

x

x

Zabaykalskiy kray

Yevreyskaya oblast'

x

x x x x x x

x

x x

x

x

Primorskiy kray

CHINA

Sakha Respublika

Xinjiang

Tianjin

Shanghai

Qinghai

Ningxia

REGIONAL DISTRIBUTION OF AGRILUS OF EAST ASIA

285

APPENDIX IV.

Kamchatskiy kray

conspersus sp nov convictor Descarpentries & Villiers, 1963 coomani Bourgoin, 1925 cordicollis sp nov coreanus Obenberger, 1935 cupes Lewis, 1893 cuprescens cuprescens (Ménétriés, 1832) cyaneomicans Nonfried, 1892 cyaneoniger Saunders, 1873 cyaneovirens Bourgoin, 1922 cyanescens cyanescens (Ratzeburg, 1837) cyanipennis Gory & Laporte, 1839 daimio Obenberger, 1936 dali Jendek, 2009 daur sp nov decoloratus Kerremans, 1892 delphinensis Abeille de Perrin, 1897 densetomentosus Obenberger, 1940 dianthus Kerremans, 1892 diao sp nov diaolin Jendek, 2001 discalis Saunders, 1873 diversornatus sp nov dureli sp nov ecarinatus Marseul, 1866 emmerichi Obenberger, 1935 erythrostictus Bourgoin, 1922 esakii Kurosawa, 1964 eunuchus sp nov euonymi Tôyama, 1985 fareastensis Jendek, 1995 fevreae Descarpentries & Villiers, 1963 fissus Obenberger, 1917 fleischeri Obenberger, 1925 foliatus Jendek, 2007 foramenifer sp nov fornicatus sp nov fortunatus Lewis, 1893 friebi Obenberger, 1922

Beijing

Anhui

x x

x x

x

Fujian

x

x

x

Gansu

x

x

Guangxi

x

x

Hebei

Guizhou

x

x

x

x

x

x

x x

x x

x x

Heilongjiang

x

Henan

x

Hunan

Hubei

x x

x

Jiangsu

x

Jilin

x

x

x

x x

x

x x x

x x

x x

Jiangxi

x

Liaoning

x x

Nei Mongol

x

Shaanxi

Shanghai

Qinghai

x

x x x x x

x

x x

x

x x

x x x x x

x x x x

x

x

x

x

x

Shandong

x x

x

Shanxi

x

Sichuan

x

x

x

Yunnan

Zhejiang

x

x

x

x x

x

x x x

Xizang

x

Hokkaido

x

Honshu

x

KOREA NORTH

x

x

KOREA SOUTH

Tsushima

Shikoku

Kyushu

Irkutskaya oblast'

Buryatiya Respublika

Amurskaya oblast'

x x x x x x x

x x

x x x x x x

x x x x

x

x x x x

MONGOLIA

x x x x x x x x x x x x

x x

x x

x

x x x x

x x x x x x

x x x x x x x x x x x x x x x x x x x x

x x

x x

x

x

x

x

Khabarovskiy kray

Hong Kong

Guangdong

RUSSIA Magadanskaya oblast'

JAPAN

x

x x

x x

x

x

x

x

x

x x

Primorskiy kray

CHINA

Sakha Respublika

Xinjiang

Tianjin

Ningxia

REGIONAL DISTRIBUTION OF AGRILUS OF EAST ASIA FRQWLQXDWLRQ

Tyva Respublika

Sakhalinskaya oblast'

x

x x

x

x

x

x

Zabaykalskiy kray

x

x

x

x

286 AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Yevreyskaya oblast'

Kamchatskiy kray

fusciapex sp nov fusiformis sp nov ganglbaueri Semenov, 1891 gaoligong Jendek, 2000 grisator Kerremans, 1893 gussakovskiji Alexeev, 1981 han sp nov hani sp nov hattorii Nakane, 1983 hmong sp nov holzschuhi Jendek, 1994 huashanus Jendek, 2001 hui sp nov hunanus Jendek, 2009 imasakai Tôyama, 1985 imitans Lewis, 1893 improcerus sp nov inamoenus Kerremans, 1892 ineptus Kerremans, 1892 insulicola Kerremans, 1912 jaminae Baudon, 1968 japanocarinatus Ohmomo, 2002 jindrai sp nov kaluganus Obenberger, 1940 kawarai Kurosawa, 1963 komareki komareki Obenberger, 1926 komareki sapporoensis Obenberger, 1935 kostali Jendek, 2007 koyoi Ohmomo, 2002 kuangcenensis Obenberger, 1940 kucerai sp nov kurumi Kurosawa, 1957 lacrima sp nov laetecyanescens Obenberger, 1940 lahu sp nov latipalpis Jendek, 2007 lei sp nov liscapia Jendek, 2003 lisu sp nov

Beijing

x

Fujian

x

Guizhou

x

Hebei

x

Henan

x

x

Hong Kong

x

Hubei

x

Hunan

x

Jilin

x

Nei Mongol

x

x

Shaanxi

x

x x

x

x x

x

Shanxi

x

Sichuan

x

x

x

x

Xinjiang

x

x

Yunnan

x x x x

x

x x x

x x

x

x

x

Zhejiang

x

Tsushima

Shikoku

Kyushu

Honshu

Hokkaido

x

x x

x

x x x x x x

x x x x

x x x x x x

x x x x x

MONGOLIA

Buryatiya Respublika

Amurskaya oblast'

x x x

x

Khabarovskiy kray

Liaoning

Jiangxi

Jiangsu

Heilongjiang

Guangxi

Guangdong

Gansu

Anhui

RUSSIA Magadanskaya oblast'

JAPAN

x

x

x

Primorskiy kray

CHINA

Sakha Respublika

Xizang

Tianjin

Shanghai

Shandong

Qinghai

Ningxia

REGIONAL DISTRIBUTION OF AGRILUS OF EAST ASIA FRQWLQXDWLRQ

Tyva Respublika

Sakhalinskaya oblast'

x x

Zabaykalskiy kray

x

REGIONAL DISTRIBUTION OF AGRILUS OF EAST ASIA 287

Yevreyskaya oblast'

Kamchatskiy kray

Irkutskaya oblast'

KOREA SOUTH

KOREA NORTH

lobatus sp nov loosdregti Baudon, 1965 lubopetri Jendek, 2000 lugu sp nov maculifer Saunders, 1873 madeci Baudon, 1968 magnus sp nov majzlani Jendek, 2007 mali Matsumura, 1924 malloti Théry, 1930 mallotiellus Kurosawa, 1985 manchu sp nov mandala sp nov marginicollis Saunders, 1873 miao sp nov mio sp nov miwai Obenberger, 1936 moerens Saunders, 1873 mongolorum Obenberger, 1936 morawitzi Obenberger, 1936 muongoides sp nov muscarius Kerremans, 1895 myopic Jendek, 2000 naxi sp nov nelsonodaimio Ohmomo, 2006 nicolanus Obenberger, 1924 ningpoensis Obenberger, 1927 nipponigena Obenberger, 1935 niveoguttatus Kerremans, 1892 nivosus Abeille de Perrin, 1900 notoclavus Jendek, 2000 obscuricollis Kiesenwetter, 1857 obscurosericeus Obenberger, 1940 odetteae Baudon, 1968 ogatai Ohmomo, 2002 olivaceidorsis Obenberger, 1917 ornatoides sp nov paralleloides sp nov pekinensis accolus Jendek, 1995

Anhui

x

x

Beijing

x

x

Fujian

x

x x

x

Gansu

x

x

Guangxi

x

x

x

Guizhou

x

Henan

Heilongjiang

Hebei

x

x

x

x x

Hubei

Jiangxi

Hunan

Hong Kong

x

x

x

x

x

x x

x

x

x

x

x x

x

Shandong

x

x

x x

x

x x x

Jiangsu

x

Jilin

x

x

Liaoning

x

Ningxia

x

x

Nei Mongol

x

Qinghai

x

x

x

Shaanxi

x

x x x

x

Sichuan

Shanxi

x

x x

x x

x

x

x

x

Xizang

Xinjiang

x

x x

Yunnan

x x

x

x

x

x x

x

x

x

x

x x x

Honshu Kyushu

x

Shikoku

Hokkaido

Zhejiang

x x

Tsushima

KOREA NORTH

x x

x x x x x x x x x x

x x x

x x x x

Buryatiya Respublika

Amurskaya oblast'

MONGOLIA

x

x x

x

x x x x x

KOREA SOUTH

x x x x x x x

x x x x x

x x x

x

x

Khabarovskiy kray

Guangdong

RUSSIA Magadanskaya oblast'

JAPAN

x

x

x

x

Primorskiy kray

CHINA

Sakha Respublika

Tianjin

Shanghai

REGIONAL DISTRIBUTION OF AGRILUS OF EAST ASIA FRQWLQXDWLRQ

Tyva Respublika

x

x

x

Zabaykalskiy kray

x

x

288 AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Yevreyskaya oblast'

Sakhalinskaya oblast'

Kamchatskiy kray

Irkutskaya oblast'

pekinensis pekinensis Obenberger, 1924 peregrinus Kiesenwetter, 1879 perroti Descarpentries & Villiers, 1963 pilosovittatus Saunders, 1873 plagiatus Ganglbauer, 1889 planipennis Fairmaire, 1888 plasoni plasoni Obenberger, 1917 plavilstshikovi Alexeev, 1989 pleuralis sp nov pleurostigmus sp nov pliculipennis Obenberger, 1940 pratensis (Ratzeburg, 1837) presbys sp nov prolatus sp nov prosternalis sp nov przewalskii Obenberger, 1936 pseudobscuricollis Jendek, 2007 pseudoconstantini Alexeev, 1981 pseudolimoniastri Cobos, 1968 pseudosericans Obenberger, 1936 pterostigma Obenberger, 1927 pumi sp nov pusillesculptus Obenberger, 1940 qiang sp nov qinling Jendek, 2000 qinlingcola sp nov quadrisignatus Marseul, 1866 quentini Descarpentries & Villiers, 1963 recticollis sp nov reichardti Obenberger, 1935 ribbei Kiesenwetter, 1879 ribesi Schaefer, 1946 rivalieri Descarpentries & Villiers, 1963 rougeoti Descarpentries & Villiers, 1963 rudicollis Alexeev, 1979 sachalinicola Obenberger, 1940 sagittifer sp nov satoi Kurosawa, 1954 schillhammeri sp nov

Fujian

Beijing

x

Gansu

x

x

x x

x

x

x

x

Guangdong

x x

x

Guangxi

x

Guizhou

x

x

x x

Hebei

x

Heilongjiang

x

Henan

x

x x x

x

x

x x

x

Hunan

Hubei

x x

x

x x

Jiangxi

Jiangsu

x

x

x

x

x x

Nei Mongol

Liaoning

Jilin

x x x

x

x

x

x x x

x x

Shaanxi

x x

x

x

x x x

x

x x x

x

x

x

Shandong

x

Shanxi

Sichuan

x

x

x

x x x

x

Tianjin

x x x

Xinjiang

x

x

Yunnan

x

Xizang

x

x x x x

x

x

x

x x

x

x

x x

x

Zhejiang

x

KOREA NORTH

Tsushima

Kyushu

Honshu

KOREA SOUTH

Hokkaido

x x

x x

MONGOLIA

Irkutskaya oblast'

Amurskaya oblast'

x

x x

x

x x x

x

x x x x

x

x x

x x x x x x x x

x

x x x x x

Shikoku

x x x x

Buryatiya Respublika

x x x x x x x

x x

x

x x

Khabarovskiy kray

Hong Kong

Anhui

RUSSIA Magadanskaya oblast'

JAPAN

x

Sakhalinskaya oblast'

x x x x

x x

x

x

x

x

x x

Primorskiy kray

CHINA

Sakha Respublika

Shanghai

Qinghai

Ningxia

REGIONAL DISTRIBUTION OF AGRILUS OF EAST ASIA FRQWLQXDWLRQ

Tyva Respublika

x

x x

Zabaykalskiy kray

Yevreyskaya oblast'

x x x

x

x

x

REGIONAL DISTRIBUTION OF AGRILUS OF EAST ASIA 289

Kamchatskiy kray

sentis sp nov shan sp nov shaowensis Jendek, 1994 shashamboe Kurosawa, 1963 she sp nov shibatai Kurosawa, 1964 sibiricus fukushimenis Jendek, 1994 sibiricus sibiricus Obenberger, 1912 sichuanus sp nov sikkimensis Obenberger, 1928 sinensis sinensis Thomson, 1879 sinensis splendidicollis Fairmaire, 1889 sinuatus sachalinensis Obenberger, 1935 siskai sp nov smaragdifrons Ganglbauer, 1889 smaragdinus habros sp nov smaragdinus smaragdinus Solsky, 1876 smetanai sp nov sospes Lewis, 1893 soudeki Obenberger, 1926 souliei Jendek, 1994 spathulifer sp nov spiculum sp nov spinipennis Lewis, 1893 stigma sp nov subauratus amurensis Obenberger, 1922 subrobustus Saunders, 1873 subtilipilis Obenberger, 1940 sudai Kurosawa, 1985 suginoi Tôyama, 1985 sugiurai Jendek, 2007 sulcinotus Jendek, 2007 suturicuspidatus Jendek, 2007 suvorovi Obenberger, 1935 tai sp nov takahashii Tôyama, 1988 tazoei Kurosawa, 1985 tempestivus Lewis, 1893 terracotta sp nov

Anhui

x

x

x x

x x

Beijing

x x

x

Fujian

x

Gansu

x

x

x

Guangdong

x

Guizhou

x

x

x

Heilongjiang

Hebei

x

x

x x

x

x

x

x

Henan

x

x

Hubei

Jiangsu

x

x

x x x

Hunan

x x

x

x

Jiangxi

x

Liaoning

Jilin

x

x

x

x

x x

x

x x

Nei Mongol

x

Shaanxi

Shanghai

x x

x x x

x x

x

x

x

x x

Shandong

x

Shanxi

x x x x

x

x

Sichuan

x x

x

x x

Xinjiang

x

Yunnan

Xizang

x

x

x

x

x x x

x

x x x x x

Zhejiang

x

Shikoku

Kyushu

Honshu

Tsushima

Hokkaido

KOREA NORTH

x x

x x

KOREA SOUTH

x x x x

x x x x x x x x x x

x x x x x x

x

x x x x

MONGOLIA

x

Amurskaya oblast'

x

x x x x x

x

Buryatiya Respublika

x x x x x

x

x

x x x x x x x x

x x x x

x

x x x x x x

x x x

x

x

x

Khabarovskiy kray

Hong Kong

Guangxi

RUSSIA Magadanskaya oblast'

JAPAN

x

x

x

x

x

x

x

Primorskiy kray

CHINA

Sakha Respublika

Tianjin

Qinghai

Ningxia

REGIONAL DISTRIBUTION OF AGRILUS OF EAST ASIA FRQWLQXDWLRQ

Sakhalinskaya oblast'

x

x

Tyva Respublika

x

Zabaykalskiy kray

x

x

x

290 AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Yevreyskaya oblast'

Kamchatskiy kray

Irkutskaya oblast'

thibetanus Obenberger, 1928 tokyoensis Kurosawa, 1985 tonkineus Kerremans, 1895 trinotatus Saunders, 1873 truncatus Jendek, 2007 tschitscherini Semenov, 1895 tsushimanus Kurosawa, 1963 tsutsumiuchii Ohmomo, 2006 tuber sp nov tyndarus Kerremans, 1913 uenoi Kurosawa, 1963 ulaangomiensis Cobos, 1972 ulmicola sp nov unguiculosus Obenberger, 1936 ussuricola Obenberger, 1924 varius Kerremans, 1895 ventricosus Fairmaire, 1888 viduus Kerremans, 1914 vigil sp nov violaceus Nonfried, 1892 viridiobscurus Saunders, 1873 viridis (Linné, 1758) volkovitshi Jendek, 2007 voriseki Jendek, 1995 yakushimensis Tôyama, 1985 yamabusi Miwa & Chûjô, 1940 yamajii Tôyama, 1985 yamawakii Kurosawa, 1957 yunnanicola Obenberger, 1936 yunnanus Obenberger, 1927 zanthoxylumi Li Meng Lou, 1989 zhongdian Jendek, 2009 zhuang sp nov zhui sp nov

Anhui

x

Beijing

x

x x

x

Fujian

x

x

Gansu

Guangdong

x

x x

x

Guangxi

x

Guizhou

x

Hebei

x

x

x x x x

x x

Heilongjiang

x

Henan

x

Hubei

x

x

x

Jiangxi

x

Jilin

Liaoning

x x

x x

x x x

x

Nei Mongol

x

Ningxia

x

Shaanxi

Shandong

x

x x

x x

x x

x x x

x

Shanxi

x

Sichuan

x

x x

x

x

x

Xinjiang

x

Xizang

x

Yunnan

x

Zhejiang

x x x x x

x

x

Kyushu

Honshu

Shikoku

Hokkaido

Tsushima

KOREA NORTH

x

x

x

MONGOLIA

Amurskaya oblast'

x

Buryatiya Respublika

x

x

Kamchatskiy kray

Irkutskaya oblast'

x x x x

x x x x x

x x x x

x x x x

x

x

KOREA SOUTH

x x x x x x x x x x x x x

x x x x x x x

x x x

x

x

x x

x x x

x x

x

x

Khabarovskiy kray

Jiangsu

Hunan

Hong Kong

RUSSIA

x

x x x

x

x

Tyva Respublika

Sakhalinskaya oblast'

x

x x x x x

Magadanskaya oblast'

JAPAN

Primorskiy kray

CHINA

Sakha Respublika

Tianjin

Shanghai

Qinghai

REGIONAL DISTRIBUTION OF AGRILUS OF EAST ASIA FRQWLQXDWLRQ

Zabaykalskiy kray

Yevreyskaya oblast'

x x

x

x

REGIONAL DISTRIBUTION OF AGRILUS OF EAST ASIA 291

292

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Viridis

Viridiobscurus

Viduus

Tempestivus

Sulcicollis

Subrobustus

Spinipennis

Smaragdinus

Sinuatus

Sinensis

Roscidus

Quadrisignatus

Prolatus

Plasoni

Plagiatus

Pekinensis

Occipitalis

Niveoguttatus

Muscarius

Maculifer

Laetecyanescens

Japanocarinatus

Imitans

Hyperici

Densetomentosus

Cyanescens

Cyaneovirens

Cyaneoniger

Biguttatus

Betuleti

Barrati

Angulatus

Albogularis

Acutus

APPENDIX V. MORPHOLOGICAL CHARACTERISTIC OF AGRILUS SPECIES-GROUPS OF EAST ASIA

BODY SIZE

SHAPE

BUILD

COLOR (dorsally)

small ( 5 mm) medium (> 0 5 mm and  10 mm) large (> 10 mm) subparallel (Fig 2A) cuneiform (Fig 2B) fusiform (Fig 2C) slender (Fig 2D) medium robust (Fig 2E) strongly convex unicolored (Fig 2F) bicolored (Fig 2G) tricolored

+ + + + + + + + + + + + + + + + +

+ + + + + + + ± + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + ± ± ± ± + + + + + + + + + + + + + ± + + ±

+ + + + + + + + + + + + + + + + + + + + + + + + + ± + + + + + + + +

+ + + + + + + + + + + + ±

+ + +

+ + + ± + + + + + + + + + + + + + + + ± ± + + + + + +

+ + + + + + + + + ± + + + + + + + + + + + + + ± + +

+ + + + + + + + + + + + + + ± + + + + + + ±

EYES SIZE

small (Fig 2I) moderate (Figs 2N, 2O) large (Figs 2H, 2J, 2P)

+

+ + + + + + + ± + + + ± + ± ± + + ± + ± + + + + + + + + + + + + + + + ± + + + ± + ± + + + + + + + + ± + ± + ± + + + + + + ±

ANTENNAE LENGTH SHAPE ANTENNOMERES (male)

short (Figs 15B, 28B) moderate (Fig 8A) long (Fig 29B) slender (Fig 2DD) stout (Fig 2EE) not modified modified (Fig 2FF)

+ + + + + + + + + + + + + + + + + + ± + + + + + +

± + + + + + + + + +

+ + + + + + + ± + + + + + + + + + + + + + + +

+ + + + + + + + + + + + + + + + + + +

+ + + + + + +

+ + + + + + ± + + + + + + + + + + + + + + + + + + + ± + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + ±

+ + + + + + + + + + + + + ± + + +

+ ± ± + +

PRONOTUM SHAPE

MAX WIDTH

ANTERIOR LOBE

DISK

DISK IMPRESSIONS

PREHUMERUS

PREHUMERUS  shape

PREHUMERUS  extent

PREHUMERUS  anterior end

LATERAL CARINAE  interspace LATERAL CARINAE  convergence LATERAL CARINAE  junction LATERAL CARINAE  narrowest point SCUTELLUM

visually elongate (Figs 10C, 25B) visually square (Fig 12K) transverse (Fig 12g) at anterior margin (Fig 3K) at middle (Fig 3L) at posterior margin (Fig 3M) absent (Fig 3B) moderate (Fig 3D) obvious (Fig 3E) flat (Fig 11E) moderately convex (Fig 12C) strongly convex (Fig 15D) absent medial only medial and lateral (Figs 3Q, 3R)) lateral only posterolateral (Fig 3S) absent (Fig 4A) obsolete (Fig 4B) tubercular (Fig 4C) filamentary (Fig 4D) carinal (Fig 4E) straight (Fig 4B) arcuate (Fig 3A) bisinuate (Figs 3BB, 4E) close to posterior pronotal angle (Fig 3X) to 1/2 of pronotal length (Fig 3Z) beyond 1/2 of pronotal length (Fig 3AA) to anterior pronotal angle (Fig 3BB, 4J) distant from pronotal marginal carina (Fig 4F) close to pronotal marginal carina (Fig 4I) join pronotal marginal carina (Fig 4H) join anterior pronotal angle (Fig 4J) narrow (Fig 4L) moderate (Fig 4N) wide (Fig 4M) convergent (Figs 4L, 4M) subparallel (Fig 4N) absent (Figs 4L, 4N) present (Fig 4M) at posterior angles (Fig 4L) distant from posterior angles (Fig 4M) well developed (Figs 4O, 4Q) rudimentary (Fig 4P)

+ + + ± + + + +

+

+

+ + + + + + + + + + + + + + + + + + + + + + + + +

± + + + + + ± ± + + ± + + + + + + + + ± + + + + + +

± + + + + + + + + + + ± + + + ± + + ± + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + ± + + + + + + + + + + +

± + + + + + + + + + + + + ± + + +

+ + + + + + + + + ± + + ± + + + + + + + + + + + + + + + + + + + + + + + + + + ± +

+ + ± + + + + ± + ± ± + + + + + + + + + + + + +

+ +

+ + + + + + ± +

+ + + + + + + + + + + + + + + + + + + + + ± + + + + + + + + + ± + + + + + + + + + ±

+ + + + + + + + + + ± + +

+ + + + + + + + + + + + + + + + + + ± +

+ + + + + +

+ + + + ± ± + ± + + +

+ + + + + + + + + + + + + + + + + ± + +

+ + + + + + +

+ + + + + + + + ± + + + + + + ± ± + + + + + + + + + + + + + + + + + ± + + + + + ± +

+ + + ± + + + + ± + ± + + + + + + + + + + + + + ± + + + + + + + + + + + + + + + + + + + + + + + + ± + ± ± + + + + ± + + + + + + + + +

+ + + + + + + + + + + +

+ + ± + + + + + + + + +

+ + + + +

+ + + ± + + + + + + + + + + +

+ + + + + ±

+ + + + + + + + ± + + + + + + + +

± + + + + + + + ± + + + ± ± + + + + + + + ± + + + + + + + + + ±

+ +

+ +

+

+ + +

+ + + + + + + + + + + + + + + + + + + + + + +

+ + + + + + + + + + + +

+ + ± + + + + + + + + +

+ + + + + + + + + ± + + ± ± + + + + + + ± + + + + + + +

+ + + + + ± + + + + + + ± + + + + + + + + + + + + + + + + + + ± ± + + + + + + + + ± ± + + + ± + + + ± + + + + + + + + + ± + + + + + + + + + + + ± + + + + + + + + + + + ± + + + + + ± + + + + + + + +

Viridis

Viridiobscurus

Tempestivus

Sulcicollis

Subrobustus

Spinipennis

Smaragdinus

Sinuatus

Sinensis

Roscidus

Quadrisignatus

Prolatus

Plasoni

Plagiatus

Pekinensis

Occipitalis

Niveoguttatus

Muscarius

Maculifer

Laetecyanescens

Japanocarinatus

Imitans

Hyperici

Densetomentosus

Cyanescens

Cyaneovirens

Cyaneoniger

Biguttatus

Betuleti

Barrati

Angulatus

Albogularis

Acutus

Viduus

293

MORPHOLOGICAL CHARACTERISTIC OF AGRILUS SPECIES-GROUPS OF EAST ASIA

ELYTRA COLOR HUMERAL CARINA APICES  arrangement

APICES  shape

PUBESCENCE

PUBESCENCE  color

unicolored (Fig 5N) bicolored (Fig 5P) absent obsolete present (Fig 1C) separate (Fig 5B) conjoint (Fig 5A) arcuate (Fig 5B) subangulate (Fig 5C) subtruncate (Fig 5D) cuspidate (Fig 5E) spinose (Fig 5F absent (Fig 5N) entire or on most of elytra (Fig 5T) sutural wedge or perisutural stripes (Figs 5U, 5V) proximal and/or distal (Figs 5Y, 5Z, 5W, 5X) ornamental (Figs 5AA, 5BB) pairs of pubescent tomentose spots (Fig 5CC) unicolored bicolored (Fig 5BB)

+ + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + ± + + + + + + + + + + + +

flat (Fig 6L) impressed (Fig 6M) with pit (Fig 6N)

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without sexual modifications with sexual modifications arcuate, subangulate or subtruncate (Figs 7A, 7B, 7C) sinuate (Figs 7G, 7H) arcuate (Fig 7I) extended into short protrusion (Fig 7J) extended into long spine (Fig 7K)

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STERNUM METASTERNAL PROJECTION

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ABDOMEN BASAL VENTRITE or METACOXA (male) GROOVE ON LAST VENTRITE PYGIDIUM

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GENITALIA AEDEAGUS

OVIPOSITOR

asymmetrical (Figs 7T, 7U) weakly sclerotised (Fig 7V) markedly dorsoventrally bent (Fig 7W) parameres with obvious membranous lobes (Fig 7X) uritiform (Fig 7R) elongate (Fig 7Q)

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294

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

AGRILUS MORPHOLOGY

Figs. 1A-1D. Agrilus morphology:. 1A) habitus dorsally; 1B) habitus ventrally; 1C) anterior body half laterally; 1D) head and prosternum ventrally.

295

296

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 2A-2FF. Agrilus morphology: Body shape: 2A) subparallel; 2B) cuneiform; 2C) fusiform. Body build: 2D) slender; 2E) robust. Body color: 2F) unicolored; 2G) bicolored. Head shape: 2H) markedly convex; 2I) obviously flat. Medial impression of head: 2J) absent; 2K) present. Frons shape: 2L) flat; 2M) moderately convex; 2N) markedly convex with eyes protruding from head outline. Frons outline: 2O) protruding from head outline; 2P) not protruding from head outline. Vertex outline (postero dorsal view): 2Q) protruding from head outline. Sculpture of frons: 2R) with fine, distinct microsculpture. Predominant sculpture of vertex: 2S) punctures irregular, sparse; 2T) punctures regular; 2U) rugae irregular, dense; 2V) rugae semispherical, sparse; 2W) dense sculpture; 2X) sparse sculpture; 2Y) rugae subparallel, dense; 2Z) subparallel sparse. Lower margin of eye: 2AA) above antennal socket; 2BB) below antennal socket. Shape of epistoma: 2CC) epistoma with raised upper margin. Shape of antennae: 2DD) slender; 2EE) stout; 2FF) sexually modified at male.

AGRILUS MORPHOLOGY

297

Figs. 3A-3JJ. Agrilus morphology: 3A) morphology of pronotum and scutellum. Anterior pronotal lobe: 3B) absent; 3C) vague; 3D) moderate; 3E) obvious. Shape of anterior pronotal lobe: 3F) arcuate; 3G) subangulate. Position of anterior pronotal lobe to anterior angles: 3H) not reaching level of anterior pronotal angles; 3I) at level with anterior pronotal angles; 3J) projecting beyond anterior pronotal angles. Maximal width of pronotum: 3K) at anterior margin; 3L) at middle; 3M) at posterior margin. Anterior margin of pronotum: 3N) wider than posterior; 3O) subequal to posterior; 3P) narrower than posterior. Impressions on pronotal disk: 3Q) medial and lateral; 3R) anteromedial and posteromedial and lateral; 3S) posterolateral. Pronotal sides: 3T) straight; 3U) arcuate; 3V) markedly arcuate; 3W) subangulate. Extent of prehumerus: 3X) close to posterior pronotal angle; 3Y) to third of pronotal length; 3Z) to 1/2 of pronotal length; 3AA) beyond 1/2 of pronotal length; 3BB) to anterior pronotal angle. Apex of posterior pronotal angles: 3CC) sharp; 3DD) blunt. Posterior end of prehumerus: 3EE) joining posterior pronotal angle; 3FF) joining posterior pronotal margin; 3GG) distant from pronotal angle or margin. Shape of posterior pronotal angles: 3HH) acute; 3II) rectangular; 3JJ) obtuse.

298

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 4A-4Q. Agrilus morphology: Prehumerus: 4A) absent; 4B) obsolete; 4C) tubercular; 4D) filamentary; 4E) carinal. Anterior end of prehumerus: 4F) & 4G) distant from pronotal marginal carina; 4H) joining pronotal marginal carina; 4I) close to pronotal marginal carina; 4J) joining anterior pronotal angle. Modification of prehumerus: 4K) obsolete in anterior part. Marginal and submarginal carinae: 4L) convergence moderate, junction absent, interspace narrow, narrowest point at posterior 1/5 1/4 of pronotal length; 4M) convergence obvious, junction present, interspace wide, narrowest point at posterior third of pronotal length; 4N) subparallel, convergence absent, junction absent, interspace moderate. Scutellum: 4O) size moderate, disk not impressed, scutellar carina present; 4P) size rudimentary, disk impressed, scutellar carina obsolete; 4Q) size large, disk not impressed, scutellar carina present.

AGRILUS MORPHOLOGY

299

Figs. 5A-5DD. Agrilus morphology: Elytral apices: 5A) conjointly arcuate; 5B) separately arcuate; 5C) subangulate; 5D) subtruncate; 5E) cuspidate; 5F) spinose; 5G) with spinuliform sutural rim; 5H) transversely truncate; 5I) obliquely truncate with projecting pleural angle; 5J) oblique with projecting sutural angle; 5K) spinose with dominant sutural spine; 5L) spinose with dominant medial spine; 5M) spinose with dominant pleural spine. Color and pubescence of elytra: 5N) unicolored without pubescence; 5O) unicolored with darker hue at apical third; 5P) bicolored with perisutural stripe of alternative color; 5Q bicolored with perisutural drop of alternative color; 5R) bicolored with transverse band of alternative color before apex; 5S) bicolored with apical portion of alternative color; 5T) entirely pubescent; 5U) with sutural wedge of pubescence; 5V) with perisutural stripes of pubescence; 5W) with pubescence in proximal part; 5X) with pubescence in distal part; 5Y) with proximal and distal pubescence; 5Z) bicolored with proximal and distal pubescence; 5AA) bicolored complementary with ornamental pubescence; 5BB) with ornamental bicolored pubescence; 5CC) with tomentose spots; 5DD) with tomentose stripes.

300

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 6A-6AA. Agrilus morphology: Size of prosternal lobe: 6A) narrow; 6B) moderate; 6C) large. Anterior margin of prosternal lobe: 6D) arcuate; 6E) subtruncate; 6F) arcuately emarginate; 6G) angulately emarginate. Emargination of prosternal lobe: 6H) shallow; 6I) moderate; 6J) deep and wide; 6K) deep and narrow. Metasternal projection: 6L) flat; 6M) impressed; 6N) with pit. Sexual modification of prosternum in male: 6O) with medial row of setae; 6P) with longer and denser pubescence. Shape of prosternal process: 6Q) narrowed; 6R) subparallel; 6S) dilated. Tips of angles of prosternal process: 6T) sharp; 6U) blunt. Sides of prosternal process: 6V) straight; 6W) arcuate. Angles of prosternal process: 6X) absent; 6Y) obtuse; 6Z) rectangular; 6AA) acute.

AGRILUS MORPHOLOGY

301

Figs.7A-7AA. Agrilus morphology: Shape of sternal groove on last abdominal ventrite: 7A) arcuate with apical margin laterally emarginate; 7B) subangulate with apical margin subtruncate; 7C) subtruncate; 7D) shallowly arcuately sinuate; 7E) widely arcuately sinuate; 7F) narrowly arcuately sinuate; 7G) deeply arcuately sinuate; 7H) deeply angulately sinuate with apical margin arcuate. Apical margin of pygidium: 7I) arcuate; 7J) extended into short protrusion; 7K) extended into long spine. Size of metafemur: 7L) & 7M) markedly incrassate. Relative size of tarsomere 1: 7N) smaller or subequal to 2 3 combined; 7O) longer than 2 3 combined but shorter than 2 4 combined; 7P) subequal or longer than 2 4 combined. Shape of ovipositor: 7Q) elongate; 7R) uritiform. Shape of aedeagus: 7S) symmetrical; 7T) feebly asymmetrical; 7U) distinctly asymmetrical; 7V) weakly sclerotised; 7W) markedly dorso ventrally bent (lateral view); 7X) parameres with obvious membranous lobes; 7Y) medial lobe distinctly wider than parameres; 7Z) apex of medial lobe markedly obtuse; 7AA) apex of medial lobe sharply pointed.

302

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 8A-8L. Habitus of Agrilus: 8A) cyaneoniger Saunders, 1873 unicolor form, 13 mm, China, Henan; 8B) cyaneoniger Saunders, 1873 bicolored form, 11 mm, Russia, Primorskiy kray; 8C) auristernum Obenberger, 1924 15 mm; 8D) planipennis Fairmaire, 1888 13 mm; 8E) lubopetri Jendek, 2000 ♂, 15 mm; 8F) lubopetri Jendek, 2000 ♀, 13.5 mm; 8G) agnatus Kerremans, 1892 11 mm; 8H) qinling Jendek, 2000 HT, 11.1 mm; 8I) sinensis sinensis Thomson, 1879 9 mm; 8J) sinensis splendidicollis Fairmaire, 1889 14 mm; 8K) coreanus Obenberger, 1935 LT, 11 mm; 8L) magnus sp. nov. HT, 12 mm.

HABITUS OF AGRILUS

303

Figs. 9A-9L. Habitus of Agrilus: 9A) decoloratus Kerremans, 1892 cupreous form, 11.5 mm, China Yunnan; 9B) decoloratus Kerremans, 1892 blue form, 13 mm, China Taiwan; 9C) ventricosus Fairmaire, 1888 11.2 mm; 9D) proditor Obenberger, 1917 LT, 9.6 mm; 9E) sachalinicola Obenberger, 1940 11 mm; 9F) spinipennis Lewis, 1893 9.9 mm; 9G) fleischeri Obenberger, 1925 9.8 mm; 9H) kawarai Kurosawa, 1963 HT, 8.7 mm; 9I) zhui sp. nov. HT, 9.6 mm; 9J) dureli sp. nov. HT, 8.9 mm; 9K) cordicollis sp. nov. HT, 11.3 mm; 9L) insulicola Kerremans, 1912 8.6 mm.

304

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 10A-10L. Habitus of Agrilus: 10A) tokyoensis Kurosawa, 1985 9.7 mm; 10B) ashlocki Baudon, 1968 7.7 mm; 10C) coomani Bourgoin, 1925 9 mm; 10D) fornicatus sp. nov. HT, 10.2 mm; 10E) erythrostictus Bourgoin, 1922 9.7 mm; 10F) hmong sp. nov. HT, 12.8 mm; 10G) cyanipennis Gory & Laporte, 1839 11 mm; 10H) auropictus Kerremans, 1912 9.1 mm; 10I) jaminae Baudon, 1968 6.7 mm; 10J) madeci Baudon, 1968 7.7 mm; 10K) niveoguttatus Kerremans, 1892 8.7 mm; 10L) satoi Kurosawa, 1954 9.5 mm.

HABITUS OF AGRILUS

305

Figs. 11A-11L. Habitus of Agrilus: 11A) fortunatus Lewis, 1893 7.8 mm; 11B) emmerichi Obenberger, 1935 LT, 8.3 mm; 11C) sospes Lewis, 1893 7.8 mm; 11D) prosternalis sp. nov. HT, 6.6 mm; 11E) quadrisignatus Marseul, 1866 5.5 mm; 11F) ornatoides sp. nov. HT, 5.5 mm; 11G) nipponigena Obenberger, 1935 4.8 mm; 11H) maculifer Saunders, 1873 HT, 5.5 mm; 11I) nelsonodaimio Ohmomo, 2006 PT, 6.1 mm; 11J) pleurostigmus sp. nov. HT, 7.1 mm; 11K) daimio Obenberger, 1936 4.1 mm; 11L) imasakai Tôyama, 1985 5.3 mm.

306

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 12A-12L. Habitus of Agrilus: 12A) sudai Kurosawa, 1985 5.4 mm; 12B) shashamboe Kurosawa, 1963 5.8 mm; 12C) dianthus Kerremans, 1892 8 mm; 12D) plasoni plasoni Obenberger, 1917 6.9 mm; 12E) chujoi Kurosawa, 1985 6.7 mm; 12F) huashanus Jendek, 2001 HT, 6.3 mm; 12G) bicuspidatus sp. nov. HT, 7.7 mm; 12H) diaolin Jendek, 2001 HT, 7.3 mm; 12I) imitans Lewis, 1893 7.8 mm; 12J) sichuanus sp. nov. HT, 10.1 mm; 12K) nepalensis Tôyama, 1988 10 mm; 12L) blatteicollis Bourgoin, 1922 10 mm.

HABITUS OF AGRILUS

307

Figs. 13A-13L. Habitus of Agrilus: 13A) gaoligong Jendek, 2000 HT, 10.5 mm; 13B) smaragdifrons Ganglbauer, 1889 6.3 mm; 13C) sikkimensis Obenberger, 1928 6.7 mm; 13D) rivalieri Descarpentries & Villiers, 1963 6.8 mm; 13E) plagiatus Ganglbauer, 1889 6.9 mm; 13F) stigma sp. nov. HT, 7.4 mm; 13G) pumi sp. nov. HT, 6.2 mm; 13H) kostali Jendek, 2007 HT, 7.8 mm; 13I) majzlani Jendek, 2007 HT, 7.9 mm; 13J) miao sp. nov. HT, 6.7 mm; 13K) discalis Saunders, 1873 6.3 mm; 13L) aureofasciatus sp. nov. HT, 8.1 mm.

308

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 14A-14L. Habitus of Agrilus: 14A) spathulifer sp. nov. HT, 5.3 mm; 14B) conspersus sp. nov. HT, 5.9 mm; 14C) sinuatus sachalinensis Obenberger, 1935 8.3 mm; 14D) mali Matsumura, 1924 8.2 mm; 14E) viduus Kerremans, 1914 8.4 mm; 14F) shan sp. nov. HT, 5.4 mm; 14G) compacticornis sp. nov. HT, 5.4 mm; 14H) presbys sp. nov. HT, 7.3 mm; 14I) aurigaster sp. nov. HT, 5.4 mm; 14J) yunnanicola Obenberger, 1936 6.8 mm; 14K) caligans Bourgoin, 1925 4.2 mm; 14L) ineptus Kerremans, 1892 4.2 mm.

HABITUS OF AGRILUS

309

Figs. 15A-15L. Habitus of Agrilus: 15A) subrobustus Saunders, 1873 4.7 mm; 15B) bonadonai Descarpentries & Villiers, 1963 4.2 mm; 15C) ningpoensis Obenberger, 1927 LT, ♀, 6.4 mm; 15D) quentini Descarpentries & Villiers, 1963 4.3 mm; 15E) smaragdinus smaragdinus Solsky, 1876 7.4 mm; 15F) smaragdinus habros ssp. nov. HT, 8.8 mm; 15G) peregrinus Kiesenwetter, 1879 7.0 mm; 15H) priamus Kerremans, 1912 11 mm; 15I) loosdregti Baudon, 1965 11.4 mm; 15J) notoclavus Jendek, 2000 HT, 7.5 mm; 15K) aurinotus sp. nov. HT, 8.8 mm; 15L) odetteae Baudon, 1968 7.6 mm.

310

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 16A-16L. Habitus of Agrilus: 16A) lacrima sp. nov. HT, 7.1 mm; 16B) fusciapex sp. nov. HT, 7.7 mm; 16C) viridis (Linné, 1758) 7.3 mm, Japan, Fukushima, 16D) viridis (Linné, 1758) 7.1 mm, Japan, Tochigi; 16E) viridis (Linné, 1758) 6.8 mm, Mongolia; 16F) viridis (Linné, 1758) 6.3 mm, Mongolia; 16G) viridis (Linné, 1758) 7 mm, Russia, Tyva; 16H) viridis (Linné, 1758) 6.7 mm, Mongolia; 16I) viridis (Linné, 1758) 6.1 mm, Mongolia; 16J) suvorovi Obenberger, 1935 8.6 mm; 16K) cuprescens cuprescens (Ménétriés, 1832) 5.6 mm, Russia, Primorskiy kray; 16L) cuprescens cuprescens (Ménétriés, 1832) 6.2 mm, Russia, Tyva.

HABITUS OF AGRILUS

311

Figs. 17A-17L. Habitus of Agrilus: 17A) ribesi Schaefer, 1946 7.1 mm; 17B) tschitscherini Semenov, 1895 7.3 mm; 17C) pecirkai Obenberger, 1916 5.8 mm; 17D) salicis Frivaldszky, 1877 5.1 mm; 17E) betuleti (Ratzeburg, 1837) 5.7 mm; 17F) fareastensis Jendek, 1995 HT, 5.8 mm; 17G) voriseki Jendek, 1995 HT, 6.0 mm; 17H) delphinensis Abeille de Perrin, 1897 4.7 mm; 17I) pratensis (Ratzeburg, 1837) 4.9 mm, China, Shaanxi; 17J) pratensis (Ratzeburg, 1837) 5.0 mm, Mongolia; 17K) nicolanus Obenberger, 1924 4.6 mm; 17L) soudeki Obenberger, 1926 4.6 mm.

312

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 18A-18L. Habitus of Agrilus: 18A) alutaceicollis Obenberger, 1930 3.8 mm; 18B) cupes Lewis, 1893 6.5 mm; 18C) jindrai sp. nov. HT, 5.7 mm; 18D) mio sp. nov. HT, 8.2 mm; 18E) ogatai Ohmomo, 2002 PT, 5.9 mm; 18F) sibiricus sibiricus Obenberger, 1912 6.5 mm; 18G) sibiricus fukushimenis Jendek, 1994 HT, 5.5 mm; 18H) smetanai sp. nov. HT, 5.9 mm; 18I) laetecyanescens Obenberger, 1940 6.5 mm; 18J) yunnanus Obenberger, 1927 7.8 mm; 18K) vigil sp. nov. HT, 8.1 mm; 18L) aurithorax sp. nov. HT, 6.6 mm.

HABITUS OF AGRILUS

313

Figs. 19A-19L. Habitus of Agrilus: 19A) paralleloides sp. nov. HT, 7.8 mm; 19B) cyanescens cyanescens (Ratzeburg, 1837) 6.3 mm; 19C) bidentulus Ganglbauer, 1889 6.5 mm; 19D) thibetanus Obenberger, 1928 6.5 mm; 19E) arsenevi Jendek, 2009 HT, 6.9 mm; 19F) asahinai Kurosawa, 1956 6.2 mm; 19G) dali Jendek, 2009 HT, 5.8 mm; 19H) zhongdian Jendek, 2009 HT, 7 mm; 19I) alesi Obenberger, 1935 7.4 mm; 19J) auriventris Saunders, 1873 6.6 mm; 19K) tonkineus Kerremans, 1895 8.1 mm; 19L) inamoenus Kerremans, 1892 8.5 mm.

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AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 20A-20L. Habitus of Agrilus: 20A) yamawakii Kurosawa, 1957 10.2 mm; 20B) perroti Descarpentries & Villiers, 1963 HT, 12 mm; 20C) zanthoxylumi Li Meng Lou, 1989 8.5 mm; 20D) grisator Kerremans, 1893 7.4 mm; 20E) olivaceidorsis Obenberger, 1917 6.5 mm; 20F) muscarius Kerremans, 1895 4.1 mm, India, Assam; 20G) muscarius Kerremans, 1895 3.8 mm, Vietnam; 20H) muscarius Kerremans, 1895 4.2 mm, China, Yunnan; 20I) mallotiellus Kurosawa, 1985 5.1 mm; 20J) hunanus Jendek, 2009 HT, 4.1 mm; 20K) liscapia Jendek, 2003 5 mm; 20L) kurumi Kurosawa, 1957 4.8 mm.

HABITUS OF AGRILUS

315

Figs. 21A-21L. Habitus of Agrilus: 21A) ecarinatus Marseul, 1866 6.2 mm; 21B) macrotatopus Obenberger, 1930 LT, 4.8 mm; 21C) nivosus Abeille de Perrin, 1900 6 mm; 21D) mongolorum Obenberger, 1936 LT, 6.5 mm; 21E) pleuralis sp. nov. HT, 6.2 mm; 21F) morawitzi Obenberger, 1936 LT, 6 mm; 21G) aerosus Ganglbauer, 1889 LT, 4.5 mm; 21H) daur sp. nov. HT, 5.5 mm; 21I) przewalskii Obenberger, 1936 LT, 5.2 mm; 21J) pseudosericans Obenberger, 1936 LT, 5.1 mm; 21K) plavilstshikovi Alexeev, 1989 PT, 3.9 mm; 21L) pseudolimoniastri Cobos, 1968 5.9 mm.

316

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 22A-22L. Habitus of Agrilus: 22A) unguiculosus Obenberger, 1936 5.3 mm; 22B) lugu sp. nov. HT, 6.2 mm; 22C) myopic Jendek, 2000 HT, 5.8 mm; 22D) pekinensis pekinensis Obenberger, 1924 6.1 mm; 22E) pekinensis accolus Jendek, 1995 PT, 4.5 mm; 22F) mandala sp. nov. HT, 7.3 mm; 22G) alashanensis Obenberger, 1936 7.2 mm; 22H) souliei Jendek, 1994 HT, 8.0 mm; 22I) adelphinus Kerremans, 1895 8.2 mm; 22J) chaetocoxalis sp. nov. HT, 8.1 mm; 22K) ribbei Kiesenwetter, 1879 6.1 mm, blue form, Japan, Gunma; 22L) ribbei Kiesenwetter, 1879 6 mm, brown green form, Russia, Primorskiy kray.

HABITUS OF AGRILUS

317

Figs. 23A-23L. Habitus of Agrilus: 23A) suginoi Tôyama, 1985 6.6 mm; 23B) sulcinotus Jendek, 2007 HT, 6.6 mm; 23C) reichardti Obenberger, 1935 6.1 mm; 23D) andrusi Baudon, 1968 HT, 5.4 mm; 23E) miwai Obenberger, 1936 5.7 mm; 23F) volkovitshi Jendek, 2007 HT, 5.4 mm; 23G) trinotatus Saunders, 1873 6 mm; 23H) foliatus Jendek, 2007 HT, 5.5 mm; 23I) suturicuspidatus Jendek, 2007 HT, 5.8 mm; 23J) obscuricollis Kiesenwetter, 1857 4.2 mm; 23K) varius Kerremans, 1895 5 mm; 23L) marginicollis Saunders, 1873 5.4 mm.

318

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 24A-24L. Habitus of Agrilus: 24A) fissus Obenberger, 1917 5 mm; 24B) ussuricola Obenberger, 1924 4.8 mm; 24C) truncatus Jendek, 2007 HT, 4.8 mm; 24D) pseudobscuricollis Jendek, 2007 HT, 4.5 mm; 24E) bituberculatus Jendek, 2007 HT, 3.7 mm; 24F) laticornis (Illiger, 1803) 5.8 mm; 24G) latipalpis Jendek, 2007 HT, 3.8 mm; 24H) kaluganus Obenberger, 1940 5 mm; 24I) friebi Obenberger, 1922 4.3 mm; 24J) obscurosericeus Obenberger, 1940 LT, 5 mm; 24K) takahashii Tôyama, 1988 HT, 5.5 mm; 24L) yakushimensis Tôyama, 1985 PT, 4.9 mm.

HABITUS OF AGRILUS

319

Figs. 25A-25L. Habitus of Agrilus: 25A) sentis sp. nov. HT, 9.2 mm; 25B) asiaticus Kerremans, 1898 8.7 mm; 25C) tempestivus Lewis, 1893 6.2 mm; 25D) malloti Théry, 1930 5.6 mm; 25E) fevreae Descarpentries & Villiers, 1963 4.8 mm; 25F) chekiangensis Gebhardt, 1929 4.4 mm; 25G) han sp. nov. HT, 4.1 mm; 25H) esakii Kurosawa, 1964 6.2 mm; 25I) shibatai Kurosawa, 1964 6.6 mm; 25J) tsutsumiuchii Ohmomo, 2006 PT, 7 mm; 25K) diao sp. nov. HT, 5.9 mm; 25L) ciping sp. nov. HT, 6.7 mm.

320

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 26A-26L. Habitus of Agrilus: 26A) she sp. nov. HT, 6.3 mm; 26B) zhuang sp. nov. HT, 6.9 mm; 26C) bonnottei Bourgoin, 1925 10.1 mm; 26D) cyaneovirens Bourgoin, 1922 7.3 mm; 26E) euonymi Tôyama, 1985 5.5 mm; 26F) pterostigma Obenberger, 1927 LT, 6.2 mm; 26G) subtilipilis Obenberger, 1940 LT, 5.0 mm; 26H) rougeoti Descarpentries & Villiers, 1963 4.7 mm; 26I) schillhammeri sp. nov. HT, 6.3 mm; 26J) terracotta sp. nov. HT, 7.2 mm; 26K) yamabusi Miwa & Chûjô, 1940 5.5 mm; 26L) kucerai sp. nov. HT, 5.5 mm.

HABITUS OF AGRILUS

321

Figs. 27A-27L. Habitus of Agrilus: 27A) gussakovskiji Alexeev, 1981 4.7 mm; 27B) sagittifer sp. nov. 5.2 mm; 27C) tsushimanus Kurosawa, 1963 5.3 mm; 27D) hui sp. nov. HT, 3.8 mm; 27E) businskorum sp. nov. HT, 4.0 mm; 27F) manchu sp. nov. HT, 4.6 mm; 27G) brevinotus sp. nov. HT, 4.4 mm; 27H) hani sp. nov. HT, 5.1 mm; 27I) lahu sp. nov. HT, 4.9 mm; 27J) bai sp. nov. HT, 4.6 mm; 27K) naxi sp. nov. HT, 4.6 mm; 27L) qinlingcola sp. nov. HT, 4.5 mm.

322

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 28A-28L. Habitus of Agrilus: 28A) barrati Descarpentries & Villiers, 1963 5 mm; 28B) holzschuhi Jendek, 1994 HT, 7.4 mm; 28C) densetomentosus Obenberger, 1940 4.4 mm; 28D) uenoi Kurosawa, 1963 4.5 mm; 28E) fusiformis sp. nov. HT, 4.1 mm; 28F) viridiobscurus Saunders, 1873 4.4 mm; 28G) caudicula sp. nov. HT, 4.5 mm; 28H) foramenifer sp. nov. HT, 4.4 mm; 28I) lobatus sp. nov. HT, 4.3 mm; 28J) pusillesculptus Obenberger, 1940 4.2 mm; 28K) pliculipennis Obenberger, 1940 4.5 mm; 28L) tai sp. nov. HT, 5.4 mm.

HABITUS OF AGRILUS

323

Figs. 29A-29L. Habitus of Agrilus: 29A) tuber sp. nov. HT, 4.2 mm; 29B) japanocarinatus Ohmomo, 2002 PT, 4.5 mm; 29C) bocaki sp. nov. HT, 4.6 mm; 29D) lei sp. nov. HT, 5.6 mm; 29E) chlorophyllus Abeille de Perrin, 1904 4.3 mm; 29F) eunuchus sp. nov. HT, 4.2 mm; 29G) ulmicola sp. nov. HT, 4.5 mm; 29H) spiculum sp. nov. HT, 5.2 mm; 29I) moerens Saunders, 1873 4.4 mm; 29J) prolatus sp. nov. HT, 4.1 mm; 29K) qiang sp. nov. HT, 4.0 mm; 29L) recticollis sp. nov. HT, 4.2 mm.

324

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 30A-30L. Habitus of Agrilus: 30A) adspersus (Kerremans, 1890) 7.7 mm; 30B) baroni Gory & Laporte, 1839 16 mm; 30C) charismaticus Jendek, 2000 HT, 8.4 mm; 30D) convictor Descarpentries & Villiers, 1963 3.6 mm; 30E) diversornatus sp. nov. HT, 7.1 mm; 30F) ganglbaueri Semenov, 1891 3.6 mm; 30G) hattorii Nakane, 1983 5.2 mm; 30H) improcerus sp. nov. HT, 3.3 mm; 30I) komareki komareki Obenberger, 1926 4.7 mm; 30J) komareki sapporoensis Obenberger, 1935 4.8 mm; 30K) koyoi Ohmomo, 2002 PT, 4.8 mm; 30L) lisu sp. nov. HT, 4.6 mm.

HABITUS OF AGRILUS

325

Figs. 31A-31L. Habitus of Agrilus: 31A) muongoides sp. nov. HT, 4.3 mm; 31B) pilosovittatus Saunders, 1873 4.6 mm; 31C) rudicollis Alexeev, 1979 4.7 mm; 31D) shaowensis Jendek, 1994 HT, 6.3 mm; 31E) siskai sp. nov. HT, 5.8 mm; 31F) subauratus amurensis Obenberger, 1922 golden form, 6.2 mm, Russia, Primorskiy kray; 31G) subauratus amurensis Obenberger, 1922 blue form, 6.4 mm, China, Hebei; 31H) sugiurai Jendek, 2007 PT, 6.2 mm; 31I) tazoei Kurosawa, 1985 8.0 mm; 31J) tyndarus Kerremans, 1913 5.3 mm; 31K) ulaangomiensis Cobos, 1972 5.5 mm; 31L) yamajii Tôyama, 1985 4.5 mm.

326

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 32A-32AA. Aedeagi of Agrilus: 32A) cyaneoniger Saunders, 1873; 32B) planipennis Fairmaire, 1888; 32C) lubopetri Jendek, 2000; 32D) agnatus Kerremans, 1892; 32E) qinling Jendek, 2000; 32F) sinensis splendidicollis Fairmaire, 1889; 32G) coreanus Obenberger, 1935; 32H) magnus sp. nov.; 32I) decoloratus Kerremans, 1892; 32J) ventricosus Fairmaire, 1888; 32K) sachalinicola Obenberger, 1940; 32L) spinipennis Lewis, 1893; 32M) fleischeri Obenberger, 1925; 32N) dureli sp. nov.; 32O) insulicola Kerremans, 1912; 32P) tokyoensis Kurosawa, 1985; 32Q) ashlocki Baudon, 1968; 32R) coomani Bourgoin, 1925; 32S) fornicatus sp. nov.; 32T) erythrostictus Bourgoin, 1922; 32U) hmong sp. nov.; 32V) cyanipennis Gory & Laporte, 1839; 32W) auropictus Kerremans, 1912; 32X) jaminae Baudon, 1968; 32Y) madeci Baudon, 1968; 32Z) niveoguttatus Kerremans, 1892; 32AA) satoi Kurosawa, 1954.

AEDEAGI OF AGRILUS

327

Figs. 33A-33Z. Aedeagi of Agrilus: 33A) fortunatus Lewis, 1893; 33B) emmerichi Obenberger, 1935; 33C) sospes Lewis, 1893; 33D) prosternalis sp. nov.; 33E) quadrisignatus Marseul, 1866; 33F) ornatoides sp. nov.; 33G) nipponigena Obenberger, 1935; 33H) maculifer Saunders, 1873; 33I) nelsonodaimio Ohmomo, 2006; 33J) daimio Obenberger, 1936; 33K) sudai Kurosawa, 1985; 33L) shashamboe Kurosawa, 1963; 33M) dianthus Kerremans, 1892; 33N) plasoni plasoni Obenberger, 1917; 33O) chujoi Kurosawa, 1985; 33P) huashanus Jendek, 2001; 33Q) diaolin Jendek, 2001; 33R) imitans Lewis, 1893; 33S) sichuanus sp. nov.; 33T) nepalensis Tôyama, 1988; 33U) blatteicollis Bourgoin, 1922; 33V) gaoligong Jendek, 2000; 33W) smaragdifrons Ganglbauer, 1889; 33X) sikkimensis Obenberger, 1928; 33Y) rivalieri Descarpentries & Villiers, 1963; 33Z) plagiatus Ganglbauer, 1889.

328

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 34A-34Y. Aedeagi of Agrilus: 34A) stigma sp. nov.; 34B) pumi sp. nov.; 34C) kostali Jendek, 2007; 34D) miao sp. nov.; 34E) discalis Saunders, 1873; 34F) aureofasciatus sp. nov.; 34G) conspersus sp. nov.; 34H) sinuatus sachalinensis Obenberger, 1935; 34I) mali Matsumura, 1924; 34J) viduus Kerremans, 1914; 34K) shan sp. nov.; 34L) compacticornis sp. nov.; 34M) aurigaster sp. nov.; 34N) yunnanicola Obenberger, 1936; 34O) caligans Bourgoin, 1925; 34P) ineptus Kerremans, 1892; 34Q) subrobustus Saunders, 1873; 34R) bonadonai Descarpentries & Villiers, 1963; 34S) quentini Descarpentries & Villiers, 1963; 34T) smaragdinus smaragdinus Solsky, 1876; 34U) peregrinus Kiesenwetter, 1879; 34V) priamus Kerremans, 1912; 34W) loosdregti Baudon, 1965; 34X) notoclavus Jendek, 2000; 34Y) aurinotus sp. nov.

AEDEAGI OF AGRILUS

329

Figs. 35A-35Z. Aedeagi of Agrilus: 35A) odetteae Baudon, 1968; 35B) fusciapex sp. nov.; 35C) viridis (Linné, 1758); 35D) suvorovi Obenberger, 1935; 35E) ribesi Schaefer, 1946; 35F) cuprescens cuprescens (Ménétriés, 1832); 35G) tschitscherini Semenov, 1895; 35H) pecirkai Obenberger, 1916; 35I) salicis Frivaldszky, 1877; 35J) betuleti (Ratzeburg, 1837); 35K) fareastensis Jendek, 1995; 35L) voriseki Jendek, 1995; 35M) delphinensis Abeille de Perrin, 1897; 35N) pratensis (Ratzeburg, 1837); 35O) nicolanus Obenberger, 1924; 35P) soudeki Obenberger, 1926; 35Q) alutaceicollis Obenberger, 1930; 35R) cupes Lewis, 1893; 35S) jindrai sp. nov.; 35T) mio sp. nov.; 35U) ogatai Ohmomo, 2002; 35V) sibiricus sibiricus Obenberger, 1912; 35W) sibiricus fukushimenis Jendek, 1994; 35X) smetanai sp. nov.; 35Y) laetecyanescens Obenberger, 1940; 35Z) yunnanus Obenberger, 1927.

330

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 36A-36Y. Aedeagi of Agrilus: 36A) aurithorax sp. nov.; 36B) paralleloides sp. nov.; 36C) cyanescens cyanescens (Ratzeburg, 1837) aedeagus (left), exposed medial lobe (right); 36D) bidentulus Ganglbauer, 1889 aedeagus (left), exposed medial lobe (right); 36E) thibetanus Obenberger, 1928 aedeagus (left), exposed medial lobe (right); 36F) arsenevi Jendek, 2009; 36G) asahinai Kurosawa, 1956 aedeagus (left), exposed medial lobe (right); 36H) dali Jendek, 2009 aedeagus (left), exposed medial lobe (right); 36I) zhongdian Jendek, 2009 aedeagus (left), exposed medial lobe (right); 36J) alesi Obenberger, 1935; 36K) auriventris Saunders, 1873; 36L) tonkineus Kerremans, 1895; 36M) inamoenus Kerremans, 1892; 36N) yamawakii Kurosawa, 1957; 36O) perroti Descarpentries & Villiers, 1963; 36P) zanthoxylumi Li Meng Lou, 1989; 36Q) grisator Kerremans, 1893; 36R) olivaceidorsis Obenberger, 1917; 36S) muscarius Kerremans, 1895; 36T) mallotiellus Kurosawa, 1985; 36U) hunanus Jendek, 2009; 36V) kurumi Kurosawa, 1957; 36W) ecarinatus Marseul, 1866; 36X) macrotatopus Obenberger, 1930; 36Y) nivosus Abeille de Perrin, 1900.

AEDEAGI OF AGRILUS

331

Figs. 37A-37Y. Aedeagi of Agrilus: 37A) mongolorum Obenberger, 1936; 37B) pleuralis sp. nov.; 37C) morawitzi Obenberger, 1936; 37D) aerosus Ganglbauer, 1889; 37E) daur sp. nov.; 37F) przewalskii Obenberger, 1936; 37G) pseudosericans Obenberger, 1936; 37H) plavilstshikovi Alexeev, 1989; 37I) pseudolimoniastri Cobos, 1968; 37J) unguiculosus Obenberger, 1936; 37K) lugu sp. nov.; 37L) myopic Jendek, 2000; 37M) pekinensis pekinensis Obenberger, 1924; 37N) pekinensis accolus Jendek, 1995; 37O) mandala sp. nov.; 37P) alashanensis Obenberger, 1936; 37Q) souliei Jendek, 1994; 37R) adelphinus Kerremans, 1895; 37S) chaetocoxalis sp. nov.; 37T) ribbei Kiesenwetter, 1879; 37U) suginoi Tôyama, 1985; 37V) sulcinotus Jendek, 2007 aedeagus (left), exposed medial lobe (right); 37W) reichardti Obenberger, 1935; 37X) andrusi Baudon, 1968; 37Y) miwai Obenberger, 1936.

332

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 38A-38Z. Aedeagi of Agrilus: 38A) volkovitshi Jendek, 2007; 38B) trinotatus Saunders, 1873; 38C) foliatus Jendek, 2007; 38D) suturicuspidatus Jendek, 2007; 38E) obscuricollis Kiesenwetter, 1857; 38F) varius Kerremans, 1895; 38G) marginicollis Saunders, 1873; 38H) fissus Obenberger, 1917; 38I) ussuricola Obenberger, 1924; 38J) truncatus Jendek, 2007; 38K) pseudobscuricollis Jendek, 2007; 38L) bituberculatus Jendek, 2007; 38M) laticornis (Illiger, 1803); 38N) latipalpis Jendek, 2007; 38O) kaluganus Obenberger, 1940; 38P) friebi Obenberger, 1922; 38Q) obscurosericeus Obenberger, 1940; 38R) takahashii Tôyama, 1988; 38S) yakushimensis Tôyama, 1985; 38T) sentis sp. nov.; 38U) asiaticus Kerremans, 1898; 38V) tempestivus Lewis, 1893; 38W) malloti Théry, 1930; 38X) fevreae Descarpentries & Villiers, 1963; 38Y) chekiangensis Gebhardt, 1929; 38Z) han sp. nov.

AEDEAGI OF AGRILUS

333

Figs. 39A-39T. Aedeagi of Agrilus: 39A) esakii Kurosawa, 1964; 39B) shibatai Kurosawa, 1964; 39C) tsutsumiuchii Ohmomo, 2006; 39D) ciping sp. nov.; 39E) bonnottei Bourgoin, 1925 aedeagus (left), exposed medial lobe (right); 39F) cyaneovirens Bourgoin, 1922 aedeagus (left), exposed medial lobe (right); 39G) euonymi Tôyama, 1985 aedeagus (left), exposed medial lobe (right); 39H) pterostigma Obenberger, 1927 aedeagus (left), exposed medial lobe (right); 39I) rougeoti Descarpentries & Villiers, 1963 aedeagus (left), exposed medial lobe (right); 39J) schillhammeri sp. nov.; 39K) terracotta sp. nov. aedeagus (left), exposed medial lobe (right); 39L) yamabusi Miwa & Chûjô, 1940; 39M) kucerai sp. nov.; 39N) gussakovskiji Alexeev, 1981 aedeagus (left), exposed medial lobe (right); 39O) sagittifer sp. nov.; 39P) tsushimanus Kurosawa, 1963 aedeagus (left), exposed medial lobe (right); 39Q) hui sp. nov.; 39R) businskorum sp. nov.; 39S) manchu sp. nov. aedeagus (left), exposed medial lobe (right); 39T) brevinotus sp. nov. aedeagus (left), exposed medial lobe (right).

334

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 40A-40X. Aedeagi of Agrilus: 40A) hani sp. nov. aedeagus (left), exposed medial lobe (right); 40B) lahu sp. nov. aedeagus (left), exposed medial lobe (right); 40C) bai sp. nov.; 40D) naxi sp. nov.; 40E) qinlingcola sp. nov.; 40F) barrati Descarpentries & Villiers, 1963; 40G) densetomentosus Obenberger, 1940; 40H) uenoi Kurosawa, 1963; 40I) fusiformis sp. nov.; 40J) viridiobscurus Saunders, 1873; 40K) caudicula sp. nov.; 40L) foramenifer sp. nov.; 40M) lobatus sp. nov.; 40N) pusillesculptus Obenberger, 1940; 40O) pliculipennis Obenberger, 1940; 40P) tai sp. nov.; 40Q) tuber sp. nov.; 40R) japanocarinatus Ohmomo, 2002; 40S) bocaki sp. nov.; 40T) chlorophyllus Abeille de Perrin, 1904; 40U) eunuchus sp. nov.; 40V) ulmicola sp. nov.; 40W) spiculum sp. nov.; 40X) moerens Saunders, 1873.

AEDEAGI OF AGRILUS

335

Figs. 41A-41X. Aedeagi of Agrilus: 41A) prolatus sp. nov.; 41B) qiang sp. nov.; 41C) recticollis sp. nov.; 41D) adspersus (Kerremans, 1890); 41E) baroni Gory & Laporte, 1839; 41F) charismaticus Jendek, 2000; 41G) convictor Descarpentries & Villiers, 1963; 41H) ganglbaueri Semenov, 1891; 41I) hattorii Nakane, 1983; 41J) improcerus sp. nov.; 41K) komareki komareki Obenberger, 1926, parameres artificially expanded; 41L) komareki sapporoensis Obenberger, 1935, parameres artificially expanded; 41M) koyoi Ohmomo, 2002; 41N) lisu sp. nov.; 41O) muongoides sp. nov.; 41P) pilosovittatus Saunders, 1873; 41Q) rudicollis Alexeev, 1979; 41R) shaowensis Jendek, 1994; 41S) subauratus amurensis Obenberger, 1922; 41T) sugiurai Jendek, 2007; 41U) tazoei Kurosawa, 1985; 41V) tyndarus Kerremans, 1913; 41W) ulaangomiensis Cobos, 1972; 41X) yamajii Tôyama, 1985.

336

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 42A-42AA. Agrilus type specimens: 42A) accolus Jendek, 1995 (HT); 42B) acuticornis Abeille de Perrin, 1897 (LT); 42C) adelphinus Kerremans, 1895 (LT); 42D) adspersus (Kerremans, 1890) (LT); 42E) afghanistanicus Alexeev, 1992 (HT); 42F) agilis Kerremans, 1894 (LT); 42G) agnatus Kerremans, 1892 (LT); 42H) ajax Kerremans, 1912 (LT); 42I) alashanensis Obenberger, 1936 (LT); 42J) alazon Lewis, 1893 (LT); 42K) albifrons Reitter, 1897 (ST from MNHN); 42L) alesi Obenberger, 1935 (LT); 42M) altaicola Obenberger, 1935 (LT); 42N) alutaceicollis Obenberger, 1930 (LT); 42O) alutaceus Obenberger, 1916 (LT); 42P) amabilis Gory & Laporte, 1839 (LT); 42Q) amabilis Kerremans, 1893 (HTM); 42R) amurensis Obenberger, 1922 (LT); 42S) angulicollis Cobos, 1968 (HT); 42T) apicalis (Bourgoin, 1923) (LT); 42U) araxicola Abeille de Perrin, 1897 (LT); 42V) aridosus Obenberger, 1936 (LT); 42W) aritai Tôyama, 1985 (HT); 42X) asahinai Kurosawa, 1956 (HT); 42Y) ashlocki Baudon, 1968 (HT); 42Z) asiaticus Kerremans, 1898 (LT); 42AA) asperrimus Marseul, 1866 (LT).

AGRILUS TYPE SPECIMENS

337

Figs. 43A-43Z. Agrilus type specimens: 43A) assamensis Obenberger, 1936 (LT); 43B) ataman Obenberger, 1924 (LT); 43C) attila Obenberger, 1930 (LT); 43D) aubei Gory & Laporte, 1839 (LT); 43E) auristernum Obenberger, 1924 (LT); 43F) auriventris Saunders, 1873 (LT); 43G) auropictus Kerremans, 1912 (LT); 43H) bakinensis Obenberger, 1935 (LT); 43I) barmensis Obenberger, 1936 (LT); 43J) baroni Gory & Laporte, 1839 (ST); 43K) barrati Descarpentries & Villiers, 1963 (HT); 43L) bayeri Obenberger, 1935 (LT); 43M) beppuensis Obenberger, 1936 (LT); 43N) bidentulus Ganglbauer, 1889 (HTM); 43O) blatteiceps Bourgoin, 1925 (HTM); 43P) blatteicollis Bourgoin, 1922 (LT); 43Q) boisreymondi Alexeev & Bílý, 1980 (HT); 43R) bonadonai Descarpentries & Villiers, 1963 (HT); 43S) bonnottei Bourgoin, 1925 (HTM); 43T) boreooccidentalis Obenberger, 1936 (LT); 43U) brevitarsis Lewis, 1893 (LT); 43V) brodjaga Obenberger, 1914 (LT); 43W) brussae Obenberger, 1956 (LT); 43X) brydli Obenberger, 1924 (LT); 43Y) caenus Obenberger, 1924 (LT); 43Z) calcicola Obenberger, 1916 (LT).

338

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 44A-44X. Agrilus type specimens: 44A) caligans Bourgoin, 1925 (LT); 44B) candidiventris Obenberger, 1936 (LT); 44C) capreae (Chevrolat, 1838) (LT); 44D) cardaces Obenberger, 1936 (LT); 44E) cernyi Obenberger, 1925 (LT); 44F) cerskii Obenberger, 1936 (LT); 44G) chankae Obenberger, 1935 (LT); 44H) charbinensis Théry, 1942 (ST from MNHN); 44I) chekiangensis Gebhardt, 1929 (LT); 44J) chinganicus Obenberger, 1922 (LT); 44K) chlorophyllus Abeille de Perrin, 1904 (LT); 44L) chopardi Descarpentries & Villiers, 1963 (HT); 44M) chujoi Kurosawa, 1985 (HT); 44N) cockerelli Fisher, 1925 (HTM); 44O) coelicus Bourgoin, 1922 (LT); 44P) cojbalsanensis Cobos, 1968 (HT); 44Q) communis Obenberger, 1924 (LT); 44R) convictor Descarpentries & Villiers, 1963 (HT); 44S) coomani Bourgoin, 1925 (LT); 44T) corax Obenberger, 1917 (LT); 44U) cupes Lewis, 1893 (LT); 44V) cuprescens (Ménétriés, 1832) (LT); 44W) cyaneoniger Saunders, 1873 (LT); 44X) cyaneoniger Thomson, 1879 (LT).

AGRILUS TYPE SPECIMENS

339

Figs. 45A-45X. Agrilus type specimens: 45A) cyaneovirens Bourgoin, 1922 (LT); 45B) cyanipennis Gory & Laporte, 1839 (LT); 45C) daimio Obenberger, 1936 (LT); 45D) dalbergiae Théry, 1930 (LT); 45E) decoloratus Kerremans, 1892 (HTM); 45F) delphinensis Abeille de Perrin, 1897 (LT); 45G) demuthi Théry, 1942 (HTM); 45H) densetomentosus Obenberger, 1940 (LT); 45I) dentiapex Obenberger, 1940 (LT); 45J) dianthus Kerremans, 1892 (LT); 45K) diolaus Obenberger, 1958 (LT); 45L) discalis Saunders, 1873 (LT); 45M) distinguendus Gory & Laporte, 1839 (LT); 45N) djukini Obenberger, 1935 (LT); 45O) dorri Bourgoin, 1925 (LT); 45P) duporti Bourgoin, 1922 (LT); 45Q) ecarinatus Marseul, 1866 (LT); 45R) egenus Abeille de Perrin, 1895 (LT); 45S) egorovi Alexeev, 1989 (HT); 45T) emeljanovi Alexeev, 1975 (HT); 45U) emisi Ohmomo, 2005 (PT from EJCB); 45V) eogenes Obenberger, 1930 (LT); 45W) epistomalis Abeille de Perrin, 1897 (HTM); 45X) erythrostictus Bourgoin, 1922 (LT).

340

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 46A-46X. Agrilus type specimens: 46A) esakii Kurosawa, 1964 (PT from NSMT); 46B) esfandiarinus Davatchi, 1958 (LT); 46C) euonymi Tôyama, 1985 (HT); 46D) feretrius Obenberger, 1936 (LT); 46E) fevreae Descarpentries & Villiers, 1963 (HT); 46F) filigranus Obenberger, 1916 (HTM); 46G) fissifrons Abeille de Perrin, 1897 (HTM); 46H) fissus Obenberger, 1917 (LT); 46I) fleischeri Obenberger, 1925 (LT); 46J) fleutiauxi Bourgoin, 1922 (LT); 46K) fortunatus Lewis, 1893 (LT); 46L) foveolatus Abeille de Perrin, 1897 (HTM); 46M) freyi Théry, 1939 (ST from MNHN); 46N) friebi Obenberger, 1922 (LT); 46O) ganglbaueri Semenov, 1891 (LT); 46P) gebleri Obenberger, 1924 (LT); 46Q) gerschuni Stepanov, 1958 (HTM); 46R) gobicola Cobos, 1968 (HT); 46S) gracilicornis Ganglbauer, 1889 (HTM); 46T) gracilipes Lewis, 1893 (LT); 46U) graptelytrus Obenberger, 1914 (LT); 46V) grisator Kerremans, 1893 (LT); 46W) gussakovskiji Alexeev, 1981 (HT); 46X) hastuliferoides Gebhardt, 1929 (LT).

AGRILUS TYPE SPECIMENS

341

Figs. 47A-47Z. Agrilus type specimens: 47A) hattorii Nakane, 1983 (HT); 47B) hauserellus Obenberger, 1935 (LT); 47C) hokkaidoensis Tôyama, 1985 (HT); 47D) hoscheki Obenberger, 1916 (LT); 47E) ignoratus Obenberger, 1924 (LT); 47F) imbellis Kerremans, 1892 (LT); 47G) imitans Lewis, 1893 (LT); 47H) impressicollis Marseul, 1866 (LT); 47I) impressifrons Kiesenwetter, 1879 (LT); 47J) inamoenus Kerremans, 1892 (LT); 47K) ineptus Kerremans, 1892 (HTM); 47L) insuspectus Obenberger, 1924 (LT); 47M) italicus Obenberger, 1920 (LT); 47N) jaminae Baudon, 1968 (HT); 47O) japonicus Kerremans, 1898 (LT); 47P) jeanvoinei Descarpentries & Villiers, 1963 (HT); 47Q) jenissejensis Obenberger, 1924 (LT); 47R) kalganicus Obenberger, 1935 (LT); 47S) kaluganus Obenberger, 1940 (LT); 47T) kanssuanus Obenberger, 1930 (LT); 47U) kiangsuanus Obenberger, 1935 (LT); 47V) kinoshitae Obenberger, 1936 (LT); 47W) kiricenkoi Obenberger, 1935 (PLT from NMPC); 47X) klapperichi Obenberger, 1940 (LT); 47Y) komareki Obenberger, 1926 (LT); 47Z) komiyai Tôyama, 1987 (HT).

342

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 48A-48X. Agrilus type specimens: 48A) krasai Obenberger, 1924 (LT); 48B) krogerusi Obenberger, 1924 (LT); 48C) kuldjensis Obenberger, 1936 (LT); 48D) kumamotoensis Obenberger, 1935 (LT); 48E) kumamotoi Tôyama, 1987 (HT); 48F) kurosawai Obenberger, 1940 (LT); 48G) kurumi Kurosawa, 1957 (HT); 48H) kusakievici Obenberger, 1936 (LT); 48I) kuznecovi Obenberger, 1935 (LT); 48J) laetecyanescens Obenberger, 1940 (LT); 48K) lama Obenberger, 1935 (LT); 48L) lameyi Obenberger, 1924 (LT); 48M) lasiolus Obenberger, 1935 (LT); 48N) loosdregti Baudon, 1965 (HT); 48O) lubischevi Stepanov, 1958 (HTM); 48P) lubopetri Jendek, 2000 (HT); 48Q) lugovoi Stepanov, 1958 (LT); 48R) lunlingensis Obenberger, 1958 (LT); 48S) machnovskii Stepanov, 1958 (LT); 48T) madeci Baudon, 1968 (HT); 48U) malloti Kurosawa, 1957 (HT); 48V) malloti Théry, 1930 (LT); 48W) malyi Obenberger, 1936 (LT); 48X) mandjuricus Obenberger, 1922 (LT).

AGRILUS TYPE SPECIMENS

343

Figs. 49A-49Y. Agrilus type specimens: 49A) marcopoli Obenberger, 1930 (LT); 49B) marginicollis Saunders, 1873 (LT); 49C) marinus Obenberger, 1922 (LT); 49D) marquardti Obenberger, 1914 (LT); 49E) melanolucidus Ohmomo, 2005 (PT from EJCB); 49F) melanopterus Solsky, 1876 (LT); 49G) melantatus Obenberger, 1924 (HTM); 49H) mentor Kerremans, 1912 (LT); 49I) mikado Obenberger, 1924 (LT); 49J) milletae Baudon, 1968 (HT); 49K) miwai Obenberger, 1936 (LT); 49L) moerens Saunders, 1873 (LT); 49M) mokrzeckii Obenberger, 1927 (LT); 49N) mongoliae Obenberger, 1922 (LT); 49O) montanellus Obenberger, 1916 (HTM); 49P) mopsus Théry, 1930 (HTM); 49Q) muscarius Kerremans, 1895 (LT); 49R) mushanus Kurosawa, 1976 (HT); 49S) nagaoi Nakane, 1983 (HT); 49T) nalajchanus Cobos, 1968 (HT); 49U) nepalensis Tôyama, 1988 (HT); 49V) nicolanus Obenberger, 1924 (LT); 49W) nigrocoerulans Obenberger, 1924 (LT); 49X) nipponigena Obenberger, 1935 (LT); 49Y) niveoguttatus Kerremans, 1892 (HTM).

344

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 50A-50X. Agrilus type specimens: 50A) nivosus Abeille de Perrin, 1900 (LT); 50B) nodosus Kerremans, 1894 (LT); 50C) nonfriedi Obenberger, 1914 (LT); 50D) obscuricollis Kiesenwetter, 1857 (LT); 50E) obtusus Abeille de Perrin, 1897 (LT); 50F) octostictus Bourgoin, 1922 (LT); 50G) odetteae Baudon, 1968 (HT); 50H) ohmomoi Tôyama, 1985 (PT); 50I) olivaceidorsis Obenberger, 1917 (LT); 50J) omissus Kerremans, 1914 (LT); 50K) ornamentifer Obenberger, 1940 (LT); 50L) ornativentris Saunders, 1866 (LT); 50M) panhensis Baudon, 1968 (HT); 50N) paphius Théry, 1942 (HTM); 50O) pecirkai Obenberger, 1916 (HTM); 50P) pekinensis Obenberger, 1924 (LT); 50Q) peregrinus Kiesenwetter, 1879 (LT); 50R) pidjinus Obenberger, 1924 (LT); 50S) pilosovittatus Saunders, 1873 (LT); 50T) piscatorius Obenberger, 1935 (LT); 50U) pistaciophagus Alexeev & Kulinitsh, 1962 (PT from ZIN); 50V) plagiatus Ganglbauer, 1889 (HTM); 50W) planefasciatus Obenberger, 1936 (LT); 50X) planipennis Fairmaire, 1888 (LT).

AGRILUS TYPE SPECIMENS

345

Figs. 51A-51X. Agrilus type specimens: 51A) plasoni Obenberger, 1917 (HTM); 51B) pliculipennis Obenberger, 1940 (LT); 51C) pooli Théry, 1936 (LT); 51D) poppiusi Obenberger, 1924 (LT); 51E) populneus Schaefer, 1946 (LT); 51F) prasinulus Obenberger, 1924 (LT); 51G) priamus Kerremans, 1912 (LT); 51H) prinadai Fisher, 1925 (HT); 51I) pseudocoeruleus Obenberger, 1930 (LT); 51J) pseudolimoniastri Cobos, 1968 (HT); 51K) pseudorotundicollis Kurosawa, 1956 (HT); 51L) pseudoussuricola Alexeev, 1979 (HT); 51M) pusillesculptus Obenberger, 1940 (LT); 51N) pusillipubis Gebhardt, 1929 (LT); 51O) pusztae Obenberger, 1936 (LT); 51P) pyrrhus Kerremans, 1913 (LT); 51Q) quadrisignatus Marseul, 1866 (LT); 51R) quadristictulus Obenberger, 1940 (LT); 51S) quentini Descarpentries & Villiers, 1963 (HT); 51T) reichardti Obenberger, 1935 (LT); 51U) reyi Bauduer, 1878 (ST from BMNH); 51V) ribbei Kiesenwetter, 1879 (LT); 51W) ribesi Schaefer, 1946 (LT); 51X) rivalieri Descarpentries & Villiers, 1963 (HT).

346

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 52A-52AA. Agrilus type specimens: 52A) robertii Chevrolat, 1840 (LT); 52B) rokuyai Kurosawa, 1976 (HT); 52C) ronino Obenberger, 1935 (LT); 52D) rotundicollis Saunders, 1873 (LT); 52E) rougeoti Descarpentries & Villiers, 1963 (HT); 52F) rubicola Abeille de Perrin, 1897 (LT); 52G) rudicollis Alexeev, 1979 (HT ); 52H) rudis Abeille de Perrin, 1897 (LT); 52I) ryukyuensis Tôyama, 1985 (HT); 52J) sacchari Obenberger, 1940 (LT); 52K) sachalinensis Obenberger, 1935 (LT); 52L) saliceti Obenberger, 1924 (LT); 52M) salicis Frivaldszky, 1877 (LT); 52N) salicivola Kurosawa, 1963 (HT); 52O) sapporoensis Obenberger, 1935 (LT); 52P) satoi Kurosawa, 1954 (HT); 52Q) seladon Obenberger, 1940 (LT); 52R) semivittatus Tôyama, 1985 (HT); 52S) sericeus Théry, 1942 (HTM); 52T) shashamboe Kurosawa, 1963 (PT from NSMT); 52U) shibatai Kurosawa, 1964 (HT); 52V) sibiricola Obenberger, 1924 (LT); 52W) sibiricus Obenberger, 1912 (HTM); 52X) sikkimensis Obenberger, 1928 (LT); 52Y) sinensis Thomson, 1879 (LT); 52Z) smaragdifrons Ganglbauer, 1889 (LT); 52AA) smaragdinus Solsky, 1876 (HTM).

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Figs. 53A-53Z. Agrilus type specimens: 53A) sorocinus Kerremans, 1914 (ST from MNHN); 53B) sospes Lewis, 1893 (LT); 53C) soudeki Obenberger, 1926 (LT); 53D) spesivcevi Obenberger, 1927 (LT); 53E) spinipennis Lewis, 1893 (LT); 53F) splendidicollis Fairmaire, 1889 (LT); 53G) stepanovi Alexeev, 1979 (HT); 53H) steppensis Cobos, 1968 (HT); 53I) subrobustus Saunders, 1873 (LT); 53J) sudai Kurosawa, 1985 (HT); 53K) suensoni Obenberger, 1940 (LT); 53L) suginoi Tôyama, 1985 (HT); 53M) sugiurai Jendek, 2007 (HT); 53N) suvorovi Obenberger, 1935 (LT); 53O) taigicola Obenberger, 1924 (LT); 53P) tazoei Kurosawa, 1985 (HT); 53Q) tempestivus Lewis, 1893 (LT); 53R) teriolensis Obenberger, 1916 (HTM); 53S) therondi Descarpentries & Villiers, 1963 (HT); 53T) thibetanus Obenberger, 1928 (LT); 53U) tibialis Lewis, 1893 (LT); 53V) tokyoensis Kurosawa, 1985 (HT); 53W) tonkineus Kerremans, 1895 (LT); 53X) toyamai Kurosawa, 1985 (HT); 53Y) toyoshimai Tôyama, 1988 (HT); 53Z) transbaicalensis Obenberger, 1935 (LT).

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AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 54A-54X. Agrilus type specimens: 54A) trinotatus Saunders, 1873 (LT); 54B) tscherepanovi Stepanov, 1954 (PT in SZMN); 54C) tschitscherini Semenov, 1895 (LT); 54D) tyndarus Kerremans, 1913 (LT); 54E) uenoi Kurosawa, 1963 (HT); 54F) ulaangomiensis Cobos, 1972 (HT); 54G) ulmi Kurosawa, 1956 (PT from MNMS); 54H) unguiculosus Obenberger, 1936 (LT); 54I) ussuricola Obenberger, 1924 (LT); 54J) uzbekistanus Stepanov, 1958 (LT); 54K) varius Kerremans, 1895 (LT); 54L) ventricosus Fairmaire, 1888 (LT); 54M) vernadskii Obenberger, 1927 (HTM); 54N) viduus Kerremans, 1914 (LT); 54O) viridiobscurus Saunders, 1873 (LT); 54P) viridipennis Gory & Laporte, 1839 (LT); 54Q) vodaki Obenberger, 1936 (LT); 54R) wilsoni Baudon, 1968 (HT); 54S) yamajii Tôyama, 1985 (HT); 54T) yamawakii Kurosawa, 1957 (HT); 54U) yunnanicola Obenberger, 1936 (LT); 54V) yunnanus Obenberger, 1927 (LT); 54W) zemani Obenberger, 1925 (LT); 54X) zhelochovtsevi Alexeev, 1979 (HT).

DATA ANALYSIS

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Figs. 55A-55F. Data analysis of Agrilus of East Asia: 55A) Agrilus taxa authorship 25 most productive authors; 55B) Agrilus synonymy 25 taxa with highest number of synonyms; 55C) Agrilus taxa descriptions number of descriptions in 15 year intervals; 55D) Agrilus taxa description number of descriptions in 15 year intervals and country of origin of primary type; 55E) Examined Agrilus material number of faunal records and specimens by country; 55F) Examined Agrilus material number of faunal records and specimens from China by province displaying 90% most numerous records.

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AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 56A-56F. Data analysis of Agrilus of East Asia: 56A) Examined Agrilus material number of faunal records and specimens from Russia by region displaying 90% most numerous records; 56B) Examined Agrilus material number of faunal records and specimens in 10 year intervals; 56C) Examined Agrilus material number of records by 350 m altitude increment and country. Only records with given altitudes are cited; 56D) Agrilus larval host plants number of host plants by number of Agrilus taxa; 56E) Agrilus larval host plants number of host plants by Agrilus taxa with more than one host plant; 56F) Agrilus larval host plants number of Agrilus taxa by host plant.

DATA ANALYSIS AND BIOGEOGRAPHY OF AGRILUS

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Figs. 57A-57C. Data analysis and biogeography of Agrilus: 57A) Agrilus distribution number of distributional regions for the 50 most widely distributed Agrilus taxa from East Asia. A region is represented by a single country except for those of Canada, China, India, Russia and United States, which are represented by a first level administration unit (e.g. province). Invasive taxa are blue; 57B) Agrilus distribution number of distributional regions by continent for 50 most widely distributed Agrilus taxa from East Asia. See 57A for definition of distributional region; 57C) Target region of East Asia administrative division.

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AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 58A-58B. Biogeography of Agrilus: 58A) Target region of East Asia terrestrial biomes. Source: vector digital map: “Terrestrial Ecoregions of the World”, Version 2.0, 2004, provided by the World Wildlife Fund; 58B) Target region of East Asia relief map with Agrilus collecting sites.

BIOGEOGRAPHY OF AGRILUS

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Figs. 59A-59B. Biogeography of Agrilus: 59A) Examined Agrilus material collecting sites and number of faunal records by region; 59B) Agrilus diversity number of taxa by region.

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AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 60A-60B. Biogeography of Agrilus: 60A) Agrilus diversity and faunal records by region; 60B) Regional endemism of Agrilus number of regionally endemic taxa and their percentage among all regional taxa.

BIOGEOGRAPHY OF AGRILUS

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Figs. 61A-61B. Biogeography of Agrilus: 61A) Regional endemism of Agrilus number of regionally endemic taxa and their percentage among all regional taxa; 61B) Biogeographic origin of Agrilus proportion by region.

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AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 62A-62B. Biogeography of Agrilus: 62A) Biogeographic origin of Agrilus proportion of Palaearctic fauna (exluding East Asian) by region; 62B) Biogeographic origin of Agrilus proportion of Palaearctic (Eurosiberian) fauna by region.

BIOGEOGRAPHY OF AGRILUS

Figs. 63A-63B. Biogeography of Agrilus: 63A) Biogeographic origin of Agrilus origin of Agrilus proportion of Central Asian fauna by region.

357

proportion of East Palaearctic fauna by region; 63B) Biogeographic

358

AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 64A-64B. Biogeography of Agrilus: 64A) Biogeographic origin of Agrilus proportion of Palaearctic (East Asian) fauna by region; 64B) Biogeographic origin of Agrilus proportion of Indo Malayan fauna by region.

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Figs. 65A-65B. Biogeography of Agrilus: 65A) Biogeographic origin of Agrilus taxa extending to Indo Malayan fauna by region; 65B) Regional similarity of Agrilus fauna proportion of shared Mongolian taxa by region.

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AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 66A-66B. Biogeography of Agrilus: 66A) Regional similarity of Agrilus proportion of shared taxa of Primorskyi kray by region; 66B) Regional similarity of Agrilus fauna proportion of shared Japanese taxa by region.

BIOGEOGRAPHY OF AGRILUS

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Figs. 67A-67B. Biogeography of Agrilus: 67A) Regional similarity of Agrilus proportion of shared Yunnan taxa by region; 67B) Estimated biodiversity of Agrilus of Southeast Asia. The numbers display estimated rate between known and undiscovered species.

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AGRILUS (COLEOPTERA, BUPRESTIDAE) OF EAST ASIA

Figs. 68A-68H. Biogeography of Agrilus: Distribution of: 68A) A. viridis (Linné, 1758); 68B) A. cuprescens (Ménétriés, 1832), North American distribution is not displayed; 68C) A. obscuricollis Kiesenwetter, 1857; 68D) A. voriseki Jendek, 1995; 68E) A. ganglbaueri Semenov, 1891; 68F) A. asiaticus Kerremans, 1898; 68G) A. adelphinus Kerremans, 1895; 68H) A. dianthus Kerremans, 1892.

Vasily (= Vasia) Grebennikov was born in 1972 in Volgograd, in the former Soviet Union. After a string of postdocs on various beetle families he become focussed on plant-eating beetles from the Asia-Pacific Region posing a risk to Canadian plant resources. Both authors are entomologists at the Canadian Food Inspection Agency in Ottawa, Canada.

ISBN 978-80-903590-6-2

Eduard (= Edo) Jendek was born in 1959 in Bratislava, in the former Czechoslovakia. Edo researches the worldwide jewel-beetle genus Agrilus with particular emphasis on the Asia-Pacific Region, which he explored through twenty expeditions. Edo provided the first species-level identifications of the infamous Agrilus planipennis Fairmaire, the notorious tree-killer colloquially known as “Emerald Ash Borer”, when the insect had been found established in North America.