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Jürgen Gadau and Jennifer Fewell ... DIVISION OF LABOR is the essence of sociality in insects and its most ... thick glandular epithelium lining her entire body (Hölldobler, Peeters, and .... Workers often do so in ants (e.g., Kikuta and Tsuji 1999; .... Reproductive conflicts in insect societies differ in both proximate and ulti-.